294 lines
35 KiB
XML
294 lines
35 KiB
XML
<document id="4259EDC2ED92B28C5CB2A1784E4C3744" ID-CLB-Dataset="9409" ID-DOI="10.11646/zootaxa.4890.3.1" ID-GBIF-Dataset="2565fffa-2704-4101-8fdf-2b863ef19be5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4306124" ID-ZooBank="4D7E2CD3-5350-479F-905A-C2671C414CBC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607097313496" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2020" docId="AE6287F45036FFFBA7FFFA3C7C52FA94" docLanguage="en" docName="zootaxa.4890.3.1.pdf" docOrigin="Zootaxa 4890 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba cooteri Shavrin 2020, sp.n." docType="treatment" docVersion="10" lastPageNumber="310" masterDocId="525BFF8C5031FFF2A768FFAA7D46FFB5" masterDocTitle="New species and records of Paraphloeostiba Steel, 1960 from China and Laos and descriptions of four new species of related genera (Coleoptera: Staphylinidae Omaliinae: Omaliini)" masterLastPageNumber="329" masterPageNumber="301" pageNumber="308" updateTime="1698847096712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DA1E38193BBC0DA05671B239907A57EB">New species and records of Paraphloeostiba Steel, 1960 from China and Laos and descriptions of four new species of related genera (Coleoptera: Staphylinidae Omaliinae: Omaliini)</mods:title>
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<mods:namePart id="CD5669CCFA5C126F271A56C67F776085">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="9D34FF721EB23C01B59867F17D213DDB">Zootaxa</mods:title>
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<mods:date id="99B3C85EB9724F9D98460098C8D56D1F">2020</mods:date>
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<mods:number id="4214BEAED23B32C3535D847EF320E4AB">4890</mods:number>
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<treatment id="AE6287F45036FFFBA7FFFA3C7C52FA94" ID-DOI="http://doi.org/10.5281/zenodo.4332513" ID-GBIF-Taxon="170712162" ID-Zenodo-Dep="4332513" LSID="urn:lsid:plazi:treatment:AE6287F45036FFFBA7FFFA3C7C52FA94" httpUri="http://treatment.plazi.org/id/AE6287F45036FFFBA7FFFA3C7C52FA94" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308">
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<subSubSection id="6ED165695036FFF5A7FFFA3C7CBFFA04" box="[151,505,1430,1457]" pageId="7" pageNumber="308" type="nomenclature">
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<paragraph id="267436E25036FFF5A7FFFA3C7CBFFA04" blockId="7.[151,505,1430,1493]" box="[151,505,1430,1457]" pageId="7" pageNumber="308">
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<heading id="7D3C818E5036FFF5A7FFFA3C7CBFFA04" bold="true" box="[151,505,1430,1457]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
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<emphasis id="14BFEAF05036FFF5A7FFFA3C7CBFFA04" bold="true" box="[151,505,1430,1457]" pageId="7" pageNumber="308">
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<taxonomicName id="E1CB4D615036FFF5A7FFFA3C7CF1FA04" ID-CoL="8Y7TL" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[151,439,1430,1457]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="cooteri" status="sp. nov.">
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<emphasis id="14BFEAF05036FFF5A7FFFA3C7CF1FA04" bold="true" box="[151,439,1430,1457]" italics="true" pageId="7" pageNumber="308">Paraphloeostiba cooteri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0F8C578B5036FFF5A6D6FA3D7CBFFA04" box="[446,505,1431,1457]" pageId="7" pageNumber="308" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6ED165695036FFF5A7FFFA117CEAFA60" box="[151,428,1467,1493]" pageId="7" pageNumber="308" type="description">
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<paragraph id="267436E25036FFF5A7FFFA117CEAFA60" blockId="7.[151,505,1430,1493]" box="[151,428,1467,1493]" pageId="7" pageNumber="308">
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(
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<figureCitation id="BEF02A675036FFF5A7F7FA117DBBFA60" box="[159,253,1467,1493]" captionStart="FIGURE 14" captionStartId="5.[151,250,1346,1370]" captionTargetBox="[151,1437,381,1323]" captionTargetId="figure-121@5.[151,1437,381,1323]" captionTargetPageId="5" captionText="FIGURE 14. Distribution of Paraphloeostiba in south-eastern Asia: P. ampliata (black circle), P. brancuccii (white circles), P. cooteri (black triangle)." figureDoi="http://doi.org/10.5281/zenodo.4306130" httpUri="https://zenodo.org/record/4306130/files/figure.png" pageId="7" pageNumber="308">Figs. 14</figureCitation>
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,
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<figureCitation id="BEF02A675036FFF5A661FA117C17FA60" box="[265,337,1467,1493]" captionStart="FIGURES 6–13" captionStartId="3.[151,264,1932,1956]" captionTargetBox="[156,1429,963,1900]" captionTargetId="figure-363@3.[151,1436,957,1909]" captionTargetPageId="3" captionText="FIGURES 6–13. Abdominal segments of Paraphloeostiba (6–11: P. ampliata; 12–13: P. cooteri): 6, 12—male tergite VIII (dorsal view), 7, 13—male sternite VIII (ventral view), 8—female tergite VIII (dorsal view), 9—female sternite VIII (ventral view), 10—female accessory sclerite, 11—spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306128" httpUri="https://zenodo.org/record/4306128/files/figure.png" pageId="7" pageNumber="308">12–13</figureCitation>
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,
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<figureCitation id="BEF02A675036FFF5A635FA117CE3FA60" box="[349,421,1467,1493]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1959,1983]" captionTargetBox="[391,1191,191,1927]" captionTargetId="figure-27@8.[381,1206,181,1935]" captionTargetPageId="8" captionText="FIGURES 23–26. Aedeagus of Paraphloeostiba (23–24: P. cooteri (paratype), 25–26: P. gayndahensis (Hong Kong)): 23, 25—parameral view, 24, 26—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306136" httpUri="https://zenodo.org/record/4306136/files/figure.png" pageId="7" pageNumber="308">23–24</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6ED165695036FFF5A7FFF9A97F30F8AC" pageId="7" pageNumber="308" type="materials_examined">
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<paragraph id="267436E25036FFF5A7FFF9A97957F93C" blockId="7.[151,1437,1539,2034]" pageId="7" pageNumber="308">
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<emphasis id="14BFEAF05036FFF5A7FFF9A97C84F9A8" bold="true" box="[151,450,1539,1565]" pageId="7" pageNumber="308">Type material examined:</emphasis>
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<materialsCitation id="96A33CBF5036FFF5A6A1F9A97957F93C" ID-GBIF-Occurrence="2989989320" collectingDate="2009-06-22" collectionCode="V, OUMNH" collectorName="W. Tianmu Shan & J. Cooter" country="China" location="Rougemont" municipality="Sieved litter under Ginkgo" pageId="7" pageNumber="308" specimenCount="502" specimenCount-male="500" stateProvince="Zhejiang Prov." typeStatus="holotype">
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<typeStatus id="F97088405036FFF5A6A1F9A97F77F9A8" box="[457,561,1539,1565]" pageId="7" pageNumber="308" type="holotype">Holotype</typeStatus>
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Ƌ (dissected): ‘
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<collectingCountry id="5EDC76725036FFF5A589F9A97E05F9A8" box="[737,835,1539,1565]" name="China" pageId="7" pageNumber="308">CHINA</collectingCountry>
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:
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<collectingRegion id="E40FF8005036FFF5A425F9A97EB3F9AB" box="[845,1013,1539,1566]" country="China" name="Zhejiang" pageId="7" pageNumber="308">Zhejiang Prov.</collectingRegion>
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, |
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<collectorName id="8B3E53345036FFF5A364F9AE798CF9A8" box="[1036,1226,1539,1566]" pageId="7" pageNumber="308">W. Tianmu Shan</collectorName>
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, ca
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<specimenCount id="30CDFD6B5036FFF5A39DF9A9787BF9AB" box="[1269,1341,1539,1566]" pageId="7" pageNumber="308" type="male">
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<quantity id="E1339B075036FFF5A39DF9A9787EF9AB" box="[1269,1336,1539,1566]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="308" unit="m" value="500.0">500m</quantity>
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.
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</specimenCount>
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<collectingMunicipality id="C610AC985036FFF5A238F9A97C37F9F4" pageId="7" pageNumber="308">
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Sieved litter under
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<taxonomicName id="E1CB4D615036FFF5A677F98D7C37F9F4" box="[287,369,1575,1601]" class="Ginkgoopsida" family="Ginkgoaceae" genus="Ginkgo" kingdom="Plantae" order="Ginkgoales" pageId="7" pageNumber="308" phylum="Tracheophyta" rank="genus">
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<emphasis id="14BFEAF05036FFF5A677F98D7C37F9F4" box="[287,369,1575,1601]" italics="true" pageId="7" pageNumber="308">Ginkgo</emphasis>
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</taxonomicName>
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</collectingMunicipality>
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| trees # 225–230. |
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<date id="527510225036FFF5A50AF98D7F9AF9F4" box="[610,732,1575,1601]" pageId="7" pageNumber="308" value="2009-06-22">
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<collectingDate id="4231E9CA5036FFF5A50AF98D7F9AF9F4" box="[610,732,1575,1601]" pageId="7" pageNumber="308" value="2009-06-22">22.vi.2009</collectingDate>
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</date>
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. Leg.,
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<collectorName id="8B3E53345036FFF5A445F9827EDFF9F4" box="[813,921,1575,1602]" pageId="7" pageNumber="308">J. Cooter</collectorName>
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.’ <printed>, ‘
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<location id="231460395036FFF5A346F98D79F0F9F4" LSID="urn:lsid:plazi:treatment:AE6287F45036FFFBA7FFFA3C7C52FA94:231460395036FFF5A346F98D79F0F9F4" box="[1070,1206,1575,1601]" country="China" municipality="Sieved litter under Ginkgo" name="Rougemont" pageId="7" pageNumber="308" stateProvince="Zhejiang Prov.">Rougemont</location>
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| collection’ <printed>, ‘
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<taxonomicName id="E1CB4D615036FFF5A7BCF9E17CCAF9D0" authorityName="Steel" authorityYear="1960" box="[212,396,1611,1637]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
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<location id="231460395036FFF5A7BCF9E17CCAF9D0" LSID="urn:lsid:plazi:treatment:AE6287F45036FFFBA7FFFA3C7C52FA94:231460395036FFF5A7BCF9E17CCAF9D0" box="[212,396,1611,1637]" country="China" municipality="Sieved litter under Ginkgo" name="Paraphloeostiba" pageId="7" pageNumber="308" stateProvince="Zhejiang Prov.">Paraphloeostiba</location>
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</taxonomicName>
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[handwritten] |
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<specimenCount id="30CDFD6B5036FFF5A52AF9E67F24F9D3" box="[578,610,1612,1638]" pageId="7" pageNumber="308" type="generic">sp.</specimenCount>
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[handwritten] | Det. 2009 | G. de Rougemont’ <printed>, ‘
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<typeStatus id="F97088405036FFF5A39EF9E178C9F9D0" box="[1270,1423,1611,1637]" pageId="7" pageNumber="308" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="E1CB4D615036FFF5A7FFF9C57C14F93C" authorityName="Steel" authorityYear="1960" box="[151,338,1647,1673]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
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<emphasis id="14BFEAF05036FFF5A7FFF9C57C14F93C" box="[151,338,1647,1673]" italics="true" pageId="7" pageNumber="308">
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<location id="231460395036FFF5A7FFF9C57C14F93C" LSID="urn:lsid:plazi:treatment:AE6287F45036FFFBA7FFFA3C7C52FA94:231460395036FFF5A7FFF9C57C14F93C" box="[151,338,1647,1673]" country="China" municipality="Sieved litter under Ginkgo" name="Paraphloeostiba" pageId="7" pageNumber="308" stateProvince="Zhejiang Prov.">Paraphloeostiba</location>
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</emphasis>
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</taxonomicName>
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<taxonomicName id="E1CB4D615036FFF5A60DF9DA7CF2F93C" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[357,436,1648,1673]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="cooteri" status="sp. nov.">
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<emphasis id="14BFEAF05036FFF5A60DF9DA7CF2F93C" box="[357,436,1648,1673]" italics="true" pageId="7" pageNumber="308">cooteri</emphasis>
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</taxonomicName>
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<emphasis id="14BFEAF05036FFF5A6D4F9C57CB5F93C" bold="true" box="[444,499,1647,1673]" pageId="7" pageNumber="308">
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<taxonomicNameLabel id="0F8C578B5036FFF5A6D4F9C57CB5F93C" box="[444,499,1647,1673]" pageId="7" pageNumber="308" rank="species">
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||
<specimenCount id="30CDFD6B5036FFF5A6D4F9C57C9BF93C" box="[444,477,1647,1673]" pageId="7" pageNumber="308" type="generic">sp.</specimenCount>
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n.
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</taxonomicNameLabel>
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</emphasis>
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|
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<location id="231460395036FFF5A56FF9C57F38F93C" LSID="urn:lsid:plazi:treatment:AE6287F45036FFFBA7FFFA3C7C52FA94:231460395036FFF5A56FF9C57F38F93C" box="[519,638,1647,1673]" country="China" municipality="Sieved litter under Ginkgo" name="Shavrin A." pageId="7" pageNumber="308" stateProvince="Zhejiang Prov.">Shavrin A.</location>
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||
<collectionCode id="40DAAE275036FFF5A516F9C57FD2F93C" box="[638,660,1647,1673]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="308" type="Museum">V</collectionCode>
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. des. 2020’ <printed> (
|
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<collectionCode id="40DAAE275036FFF5A4FDF9C57940F93C" box="[917,1030,1647,1673]" collectionName="United Kingdom, Oxford, University Museum of Natural History" pageId="7" pageNumber="308">OUMNH</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="267436E25036FFF5A7AFF9397F30F8AC" blockId="7.[151,1437,1539,2034]" pageId="7" pageNumber="308">
|
||
<emphasis id="14BFEAF05036FFF5A7AFF9397C72F918" bold="true" box="[199,308,1683,1709]" pageId="7" pageNumber="308">
|
||
<typeStatus id="F97088405036FFF5A7AFF9397C72F918" box="[199,308,1683,1709]" pageId="7" pageNumber="308" type="paratype">Paratype</typeStatus>
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</emphasis>
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||
1 Ƌ (dissected): ‘
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<collectingCountry id="5EDC76725036FFF5A569F9397F25F918" box="[513,611,1683,1709]" name="China" pageId="7" pageNumber="308">CHINA</collectingCountry>
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:
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<collectingRegion id="E40FF8005036FFF5A518F9397E5CF91B" box="[624,794,1683,1710]" country="China" name="Zhejiang" pageId="7" pageNumber="308">Zhejiang Prov.</collectingRegion>
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, | Lin`an county, m. | W. Tianmu Shan
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<taxonomicNameLabel id="0F8C578B5036FFF5A389F939785EF918" box="[1249,1304,1683,1709]" pageId="7" pageNumber="308" rank="subSpecies">
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N.
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<collectionCode id="40DAAE275036FFF5A396F9397853F918" box="[1278,1301,1683,1709]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="308">R</collectionCode>
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||
.
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||
</taxonomicNameLabel>
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’ <printed>, ‘Flight | interception | trap’ <printed>, ‘Rougemont | collection’ <printed>, ‘
|
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<taxonomicName id="E1CB4D615036FFF5A49CF91D79EAF964" authorityName="Steel" authorityYear="1960" box="[1012,1196,1719,1745]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">Paraphloeostiba</taxonomicName>
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[handwritten] |
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||
<emphasis id="14BFEAF05036FFF5A20CF91D78DAF964" bold="true" box="[1380,1436,1719,1745]" pageId="7" pageNumber="308">n.sp.</emphasis>
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[handwritten] | det. 2002 | G. de Rougemont’ <printed>, ‘
|
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<typeStatus id="F97088405036FFF5A44CF9717EF2F940" box="[804,948,1755,1781]" pageId="7" pageNumber="308" type="paratype">PARATYPE</typeStatus>
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|
|
||
<taxonomicName id="E1CB4D615036FFF5A4A4F97179C1F940" authorityName="Steel" authorityYear="1960" box="[972,1159,1755,1781]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
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<emphasis id="14BFEAF05036FFF5A4A4F97179C1F940" box="[972,1159,1755,1781]" italics="true" pageId="7" pageNumber="308">Paraphloeostiba</emphasis>
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</taxonomicName>
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|
|
||
<taxonomicName id="E1CB4D615036FFF5A3F6F97679ABF940" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[1182,1261,1756,1781]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="cooteri" status="sp. nov.">
|
||
<emphasis id="14BFEAF05036FFF5A3F6F97679ABF940" box="[1182,1261,1756,1781]" italics="true" pageId="7" pageNumber="308">cooteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14BFEAF05036FFF5A39EF9717868F940" bold="true" box="[1270,1326,1755,1781]" pageId="7" pageNumber="308">
|
||
<taxonomicNameLabel id="0F8C578B5036FFF5A39EF9717868F940" box="[1270,1326,1755,1781]" pageId="7" pageNumber="308" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
| Shavrin A.
|
||
<collectionCode id="40DAAE275036FFF5A7D9F9557D81F8AC" box="[177,199,1791,1817]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="308" type="Museum">V</collectionCode>
|
||
. des. 2020’ <red, printed> (
|
||
<collectionCode id="40DAAE275036FFF5A692F9557F2DF8AC" box="[506,619,1791,1817]" pageId="7" pageNumber="308">OUMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6ED165695036FFFBA7AFF8897CC2FB27" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308" type="description">
|
||
<paragraph id="267436E25036FFF5A7AFF8897F04F833" blockId="7.[151,1437,1539,2034]" pageId="7" pageNumber="308">
|
||
<emphasis id="14BFEAF05036FFF5A7AFF8897C17F888" bold="true" box="[199,337,1827,1853]" pageId="7" pageNumber="308">Description</emphasis>
|
||
. Measurements (n=2): HW: 0.36–0.37; HL: 0.20–0.21; AL (
|
||
<typeStatus id="F97088405036FFF5A368F889792FF88B" box="[1024,1129,1827,1854]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
|
||
): 0.44; OL: 0.11; PL: 0.25; PW: 0.43–0.45; ESL: 0.43–0.45; EW: 0.58; AW: 0.60; MTbL: 0.25; MTrL: 0.10 (MTrL 1–4: 0.03; MTrL 5: 0.07); AedL: 0.28; TL: 1.90(
|
||
<typeStatus id="F97088405036FFF5A6E7F8C17CB0F833" box="[399,502,1899,1926]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
|
||
)–1.94.
|
||
</paragraph>
|
||
<paragraph id="267436E25036FFFBA7AFF8257CE8FF4C" blockId="7.[151,1437,1539,2034]" lastBlockId="9.[151,1437,151,1313]" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308">
|
||
Body and antennomeres 6–11 brown; maxillary palpomeres, laterobasal portions of pronotum, median part of elytra (
|
||
<typeStatus id="F97088405036FFF5A662F8197C34F87B" box="[266,370,1971,1998]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
|
||
), paratergites, and apical tergites of abdomen yellow-brown; mouthparts, antennomeres 1–5, and legs yellow. Head with regular, dense, and moderately coarse isodiametric microsculpture; microsculpture of pronotum slightly finer than that of head, without punctation; punctation of elytra irregular, fine and very dense, more distinct in scutellar area and in median portion along suture, microsculpture dense, similar to that on pronotum, but slightly coarser.
|
||
</paragraph>
|
||
<caption id="72B4666A5039FFFAA7FFF80D7E4AF857" ID-DOI="http://doi.org/10.5281/zenodo.4306136" ID-Zenodo-Dep="4306136" httpUri="https://zenodo.org/record/4306136/files/figure.png" pageId="8" pageNumber="309" startId="8.[151,264,1959,1983]" targetBox="[391,1191,191,1927]" targetPageId="8">
|
||
<paragraph id="267436E25039FFFAA7FFF80D7E4AF857" blockId="8.[151,1436,1959,2019]" pageId="8" pageNumber="309">
|
||
<emphasis id="14BFEAF05039FFFAA7FFF80D7C11F80A" bold="true" box="[151,343,1959,1983]" pageId="8" pageNumber="309">FIGURES 23–26.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="E1CB4D615039FFFAA682F80D7FD5F80A" authorityName="Steel" authorityYear="1960" box="[490,659,1959,1983]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="14BFEAF05039FFFAA682F80D7FD5F80A" box="[490,659,1959,1983]" italics="true" pageId="8" pageNumber="309">Paraphloeostiba</emphasis>
|
||
</taxonomicName>
|
||
(23–24:
|
||
<taxonomicName id="E1CB4D615039FFFAA59CF8027E11F80A" authorityName="Shavrin" authorityYear="2020" box="[756,855,1960,1983]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="14BFEAF05039FFFAA59CF8027E11F80A" box="[756,855,1960,1983]" italics="true" pageId="8" pageNumber="309">P. cooteri</emphasis>
|
||
</taxonomicName>
|
||
(paratype), 25–26:
|
||
<taxonomicName id="E1CB4D615039FFFAA34FF8027996F80A" baseAuthorityName="MacLeay" baseAuthorityYear="1873" box="[1063,1232,1959,1983]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="gayndahensis">
|
||
<emphasis id="14BFEAF05039FFFAA34FF8027970F80A" box="[1063,1078,1960,1983]" italics="true" pageId="8" pageNumber="309">P</emphasis>
|
||
.
|
||
<emphasis id="14BFEAF05039FFFAA32EF80D7996F80A" box="[1094,1232,1959,1983]" italics="true" pageId="8" pageNumber="309">gayndahensis</emphasis>
|
||
</taxonomicName>
|
||
(Hong Kong)): 23, 25—parameral view, 24, 26—lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="267436E25038FFFBA7AFFEAB792AFE78" blockId="9.[151,1437,151,1313]" pageId="9" pageNumber="310">Head slightly convex in posterior portions between ocelli and eyes, 1.7–1.8 times as broad as long, with deep and moderately long anteocellar foveae almost reaching level of mid-length of eyes; postocular carina indistinct; distance between ocelli about as long distance between ocellus and posterior margin of eye. Apical palpomere of maxillary palpi about 1.4 times as long as penultimate palpomere. Basal antennomere 1.6 times as long as broad, antennomere 2 indistinctly narrower than basal antennomere, 6–9 about twice as broad as long, apical antennomere about twice as long as 10, from middle slightly narrowing to the moderately wide apex.</paragraph>
|
||
<paragraph id="267436E25038FFFBA7AFFE7D7C8EFDEE" blockId="9.[151,1437,151,1313]" pageId="9" pageNumber="310">Pronotum 1.7–1.8 times as broad as long, 1.1–1.2 times as broad as head, widest approximately in middle, sharply narrowed towards the obtuse posterior angles; anterior angles widely rounded; paramedian longitudinal impressions distinct, wide and long, from medioapical portion extending along almost the whole disc; lateral impressions moderately deep.</paragraph>
|
||
<paragraph id="267436E25038FFFBA7AFFDCE7C41FD17" blockId="9.[151,1437,151,1313]" pageId="9" pageNumber="310">Abdomen slightly wider than elytra, with two indistinct, small and round wing-folding patches on abdominal tergite IV.</paragraph>
|
||
<paragraph id="267436E25038FFFBA7AFFD01791AFCE6" blockId="9.[151,1437,151,1313]" pageId="9" pageNumber="310">
|
||
Male. Posterior margin of abdominal tergite VIII rounded (
|
||
<figureCitation id="BEF02A675038FFFBA43DFD017EE1FD70" box="[853,935,683,709]" captionStart="FIGURES 6–13" captionStartId="3.[151,264,1932,1956]" captionTargetBox="[156,1429,963,1900]" captionTargetId="figure-363@3.[151,1436,957,1909]" captionTargetPageId="3" captionText="FIGURES 6–13. Abdominal segments of Paraphloeostiba (6–11: P. ampliata; 12–13: P. cooteri): 6, 12—male tergite VIII (dorsal view), 7, 13—male sternite VIII (ventral view), 8—female tergite VIII (dorsal view), 9—female sternite VIII (ventral view), 10—female accessory sclerite, 11—spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306128" httpUri="https://zenodo.org/record/4306128/files/figure.png" pageId="9" pageNumber="310">Fig. 12</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII concave (
|
||
<figureCitation id="BEF02A675038FFFBA669FD657C12FD5C" box="[257,340,719,745]" captionStart="FIGURES 6–13" captionStartId="3.[151,264,1932,1956]" captionTargetBox="[156,1429,963,1900]" captionTargetId="figure-363@3.[151,1436,957,1909]" captionTargetPageId="3" captionText="FIGURES 6–13. Abdominal segments of Paraphloeostiba (6–11: P. ampliata; 12–13: P. cooteri): 6, 12—male tergite VIII (dorsal view), 7, 13—male sternite VIII (ventral view), 8—female tergite VIII (dorsal view), 9—female sternite VIII (ventral view), 10—female accessory sclerite, 11—spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306128" httpUri="https://zenodo.org/record/4306128/files/figure.png" pageId="9" pageNumber="310">Fig. 13</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="BEF02A675038FFFBA68AFD657F75FD5C" box="[482,563,719,745]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1959,1983]" captionTargetBox="[391,1191,191,1927]" captionTargetId="figure-27@8.[381,1206,181,1935]" captionTargetPageId="8" captionText="FIGURES 23–26. Aedeagus of Paraphloeostiba (23–24: P. cooteri (paratype), 25–26: P. gayndahensis (Hong Kong)): 23, 25—parameral view, 24, 26—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306136" httpUri="https://zenodo.org/record/4306136/files/figure.png" pageId="9" pageNumber="310">Fig. 23</figureCitation>
|
||
) with narrow basal bulb, gradually narrowed towards middle, with moderately wide, rounded apex; parameres not reaching apex of median lobe, with two pairs of moderately short apical and preapical setae, and three setae on inner edge of apical third of each paramere; internal sac moderately long, with three sclerotized, elongate structures in basal portion. Aedeagus laterally as in
|
||
<figureCitation id="BEF02A675038FFFBA36DFC937911FCE6" box="[1029,1111,825,851]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1959,1983]" captionTargetBox="[391,1191,191,1927]" captionTargetId="figure-27@8.[381,1206,181,1935]" captionTargetPageId="8" captionText="FIGURES 23–26. Aedeagus of Paraphloeostiba (23–24: P. cooteri (paratype), 25–26: P. gayndahensis (Hong Kong)): 23, 25—parameral view, 24, 26—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306136" httpUri="https://zenodo.org/record/4306136/files/figure.png" pageId="9" pageNumber="310">Fig. 24</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="267436E25038FFFBA7AFFCF67CD7FCC3" blockId="9.[151,1437,151,1313]" box="[199,401,860,886]" pageId="9" pageNumber="310">Female unknown.</paragraph>
|
||
<paragraph id="267436E25038FFFBA7AFFC2A7CC2FB27" blockId="9.[151,1437,151,1313]" pageId="9" pageNumber="310">
|
||
<emphasis id="14BFEAF05038FFFBA7AFFC2A7CF6FC2F" bold="true" box="[199,432,896,922]" pageId="9" pageNumber="310">Comparative notes.</emphasis>
|
||
Based on shapes of the posterior angles and the paramedian impressions of the pronotum, on the proportions of elytra, and on the setation of the short parameres,
|
||
<taxonomicName id="E1CB4D615038FFFBA4AEFC0E7973FC08" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[966,1077,932,958]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="cooteri" status="sp. nov.">
|
||
<emphasis id="14BFEAF05038FFFBA4AEFC0E7E91FC08" box="[966,983,932,957]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="14BFEAF05038FFFBA48EFC0E7973FC08" box="[998,1077,932,957]" italics="true" pageId="9" pageNumber="310">cooteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14BFEAF05038FFFBA356FC0E7933FC0B" bold="true" box="[1086,1141,932,958]" pageId="9" pageNumber="310">
|
||
<taxonomicNameLabel id="0F8C578B5038FFFBA356FC0E7933FC0B" box="[1086,1141,932,958]" pageId="9" pageNumber="310" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="E1CB4D615038FFFBA264FC0E7C6AFC57" authority="Kraatz, 1859" authorityName="Kraatz" authorityYear="1859" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="singularis">
|
||
<emphasis id="14BFEAF05038FFFBA264FC0E785BFC08" box="[1292,1309,932,957]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="14BFEAF05038FFFBA244FC0978DAFC08" box="[1324,1436,931,957]" italics="true" pageId="9" pageNumber="310">singularis</emphasis>
|
||
Kraatz, 1859
|
||
</taxonomicName>
|
||
, a species known from
|
||
<collectingCountry id="5EDC76725038FFFBA551FC6D7FEEFC54" box="[569,680,967,993]" name="Sri Lanka" pageId="9" pageNumber="310">Sri Lanka</collectingCountry>
|
||
and the southern
|
||
<collectingCountry id="5EDC76725038FFFBA41BFC6D7EEBFC54" box="[883,941,967,993]" name="India" pageId="9" pageNumber="310">India</collectingCountry>
|
||
(
|
||
<bibRefCitation id="425A4B135038FFFBA4D5FC6D797CFC57" author="Steel, W. O." box="[957,1082,967,994]" pageId="9" pageNumber="310" pagination="141 - 172" refId="ref17997" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960</bibRefCitation>
|
||
), but differs by a smaller body, darker elytra, a wider apical part of the pronotum, a narrower median lobe of the aedeagus, and slightly wider parameres. Regarding, the general shape of the aedeagus,
|
||
<taxonomicName id="E1CB4D615038FFFBA59BFBA57E24FB9D" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[755,866,1038,1064]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="cooteri" status="sp. nov.">
|
||
<emphasis id="14BFEAF05038FFFBA59BFBA57E42FB9D" box="[755,772,1039,1064]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="14BFEAF05038FFFBA47BFBA57E24FB9D" box="[787,866,1039,1064]" italics="true" pageId="9" pageNumber="310">cooteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14BFEAF05038FFFBA401FBA47EE7FB9D" bold="true" box="[873,929,1038,1064]" pageId="9" pageNumber="310">
|
||
<taxonomicNameLabel id="0F8C578B5038FFFBA401FBA47EE7FB9D" box="[873,929,1038,1064]" pageId="9" pageNumber="310" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="E1CB4D615038FFFBA35DFBA578DEFB9D" authority="Watanabe, 1962" authorityName="Watanabe" authorityYear="1962" box="[1077,1432,1038,1064]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="kyushuensis">
|
||
<emphasis id="14BFEAF05038FFFBA35DFBA57900FB9D" box="[1077,1094,1039,1064]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="14BFEAF05038FFFBA33DFBA4799DFB9D" box="[1109,1243,1038,1064]" italics="true" pageId="9" pageNumber="310">kyushuensis</emphasis>
|
||
Watanabe, 1962
|
||
</taxonomicName>
|
||
, described from Kyushu,
|
||
<collectingCountry id="5EDC76725038FFFBA6C6FB987CA8FBF9" box="[430,494,1074,1100]" name="Japan" pageId="9" pageNumber="310">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="425A4B135038FFFBA695FB9B7FFAFBF9" author="Watanabe, Y." box="[509,700,1073,1100]" pageId="9" pageNumber="310" pagination="17 - 18" refId="ref18329" refString="Watanabe, Y. (1962 a) Description of a new species of the genus Paraphloeostiba from Kyushu, Japan (Col., Staphylinidae). The Entomological Review of Japan, 15, 17 - 18." type="journal article" year="1962">Watanabe 1962a</bibRefCitation>
|
||
), from which it differs by a smaller and paler body, the shapes of the paramedian impressions, which are not connected to each other in mediobasal third, and by a narrower median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6ED165695038FFFBA7AFFB31780EFB6F" pageId="9" pageNumber="310" type="distribution">
|
||
<paragraph id="267436E25038FFFBA7AFFB3179D0FB03" blockId="9.[151,1437,151,1313]" box="[199,1174,1179,1206]" pageId="9" pageNumber="310">
|
||
<emphasis id="14BFEAF05038FFFBA7AFFB317C27FB00" bold="true" box="[199,353,1179,1205]" pageId="9" pageNumber="310">Distribution.</emphasis>
|
||
The species is known from the Tianmu
|
||
<collectingRegion id="E40FF8005038FFFBA449FB367E1FFB03" box="[801,857,1180,1206]" country="Myanmar" name="Shan" pageId="9" pageNumber="310">Shan</collectingRegion>
|
||
range in
|
||
<collectingRegion id="E40FF8005038FFFBA4AAFB367960FB03" box="[962,1062,1180,1206]" country="China" name="Zhejiang" pageId="9" pageNumber="310">Zhejiang</collectingRegion>
|
||
(
|
||
<figureCitation id="BEF02A675038FFFBA35DFB3679CFFB03" box="[1077,1161,1180,1206]" captionStart="FIGURE 14" captionStartId="5.[151,250,1346,1370]" captionTargetBox="[151,1437,381,1323]" captionTargetId="figure-121@5.[151,1437,381,1323]" captionTargetPageId="5" captionText="FIGURE 14. Distribution of Paraphloeostiba in south-eastern Asia: P. ampliata (black circle), P. brancuccii (white circles), P. cooteri (black triangle)." figureDoi="http://doi.org/10.5281/zenodo.4306130" httpUri="https://zenodo.org/record/4306130/files/figure.png" pageId="9" pageNumber="310">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="267436E25038FFFBA7AFFB15780EFB6F" blockId="9.[151,1437,151,1313]" box="[199,1352,1215,1242]" pageId="9" pageNumber="310">
|
||
<emphasis id="14BFEAF05038FFFBA7AFFB157C0DFB6C" bold="true" box="[199,331,1215,1241]" pageId="9" pageNumber="310">Bionomics.</emphasis>
|
||
The specimens were collected at an elevation of
|
||
<quantity id="E1339B075038FFFBA41AFB157EFFFB6F" box="[882,953,1215,1242]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="310" unit="m" value="500.0">500 m</quantity>
|
||
a.s.l. by sifting litter under
|
||
<taxonomicName id="E1CB4D615038FFFBA387FB157807FB6C" box="[1263,1345,1215,1241]" class="Ginkgoopsida" family="Ginkgoaceae" genus="Ginkgo" kingdom="Plantae" order="Ginkgoales" pageId="9" pageNumber="310" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="14BFEAF05038FFFBA387FB157807FB6C" box="[1263,1345,1215,1241]" italics="true" pageId="9" pageNumber="310">Ginkgo</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6ED165695038FFFBA7AFFB497C52FA94" pageId="9" pageNumber="310" type="etymology">
|
||
<paragraph id="267436E25038FFFBA7AFFB497C52FA94" blockId="9.[151,1437,151,1313]" pageId="9" pageNumber="310">
|
||
<emphasis id="14BFEAF05038FFFBA7AFFB497C08FB48" bold="true" box="[199,334,1251,1277]" pageId="9" pageNumber="310">Etymology.</emphasis>
|
||
Eponymic, the species is named to honour Jonathan Cooter (Oxford), the collector of the
|
||
<typeStatus id="F97088405038FFFBA204FB4978DAFB48" box="[1388,1436,1251,1277]" pageId="9" pageNumber="310">type</typeStatus>
|
||
specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |