153 lines
15 KiB
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153 lines
15 KiB
XML
<document id="13D157AE195F077939A0710BD5D87A1C" ID-CLB-Dataset="32988" ID-DOI="http://dx.doi.org/10.3897/zookeys.675.13252" ID-GBIF-Dataset="724d6de0-f19a-457d-9339-84c836e19c38" ID-PMC="PMC5523889" ID-Pensoft-Pub="1313-2970-675-129" ID-PubMed="28769682" ID-ZooBank="E5C8F163D61547B98418CEE8D71A7DAB" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-675-129" ModsDocOrigin="ZooKeys 675" ModsDocTitle="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species" checkinTime="1495538522124" checkinUser="pensoft" docAuthor="Budha, Prem B., Naggs, Fred & Backeljau, Thierry" docDate="2017" docId="8E7B838BE5016E1A92F06F455F7A6660" docLanguage="en" docName="ZooKeys 675: 129-156" docOrigin="ZooKeys 675" docSource="http://dx.doi.org/10.3897/zookeys.675.13252" docTitle="Rishetia hastula Benson 1860" docType="treatment" docVersion="8" lastPageNumber="134" masterDocId="FF8BE96F4A658939FFA88167FF8E6E18" masterDocTitle="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species" masterLastPageNumber="156" masterPageNumber="129" pageNumber="133" updateTime="1732877711202" updateUser="ExternalLinkService">
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<mods:title id="C73531AD0F527F2C0923EBC310489840">Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species</mods:title>
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<mods:namePart id="943EB2D2B03D1635FC2DCA2A6379C000">Budha, Prem B.</mods:namePart>
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<mods:namePart id="79503DB1605FC5B9DC471BA8E33E0942">Naggs, Fred</mods:namePart>
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<mods:namePart id="6EC7FD14A1B236D46D1D592280455548">Backeljau, Thierry</mods:namePart>
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<mods:date id="BFF55E13D0D816E919B3BBACF9E5C95E">2017</mods:date>
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<mods:detail id="60E0B3B892952B1076AD35F8806B4374" type="volume">
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<mods:number id="45814B75BF5DDC906BB9157884E0EE68">675</mods:number>
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<mods:classification id="0296D17848E3E54945A4529A62E564BB">journal article</mods:classification>
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<mods:identifier id="306E4451E070B7CEE1880868CEF445D0" type="DOI">http://dx.doi.org/10.3897/zookeys.675.13252</mods:identifier>
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<treatment id="8E7B838BE5016E1A92F06F455F7A6660" ID-GBIF-Taxon="130844515" LSID="urn:lsid:plazi:treatment:8E7B838BE5016E1A92F06F455F7A6660" httpUri="http://treatment.plazi.org/id/8E7B838BE5016E1A92F06F455F7A6660" lastPageId="5" lastPageNumber="134" pageId="4" pageNumber="133" scope_class="Gastropoda" scope_order="Stylommatophora" scope_subFamily="Glessulinae">
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<subSubSection id="432432158BBB46FE761C839137E614C1" pageId="4" pageNumber="133" type="nomenclature">
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<paragraph id="D1610A2CF4B74DB57ABC4BC8A3765528" pageId="4" pageNumber="133">
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<taxonomicName id="36E0AEF1A07050F400A2E2194798973E" authority="Benson, 1860" authorityName="Benson" authorityYear="1860" family="Subulinidae" genus="Rishetia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rishetia hastula" order="Gastropoda" pageId="4" pageNumber="133" rank="species" species="hastula">Rishetia hastula (Benson, 1860)</taxonomicName>
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Figs 2D, 6
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</paragraph>
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</subSubSection>
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<subSubSection id="0B95394C0612B023C4B1EDBF2EED325F" pageId="5" pageNumber="134" type="reference_group">
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<paragraph id="3DC60EEA19577961D6976F11E210C4F4" pageId="5" pageNumber="134">
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<taxonomicName id="80B0195DB1737DA2B1BF0D28C73C4F66" family="Subulinidae" genus="Achatina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Achatina hastula" order="Gastropoda" pageId="5" pageNumber="134" rank="species" species="hastula">
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<pageBreakToken id="9D8BAA03AFB07FAC84C0FD45640E4ED0" pageId="5" pageNumber="134" start="start">Achatina</pageBreakToken>
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hastula
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</taxonomicName>
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B.:
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<bibRefCitation id="F7E4C013FB5F000C13029374F47C3BAF" author="Benson, WH" journalOrPublisher="The Annals and Magazine of Natural History" pageId="14" pageNumber="143" pagination="460 - 466" title="On new land shells from Darjiling with a series of new Indian species of Achatina." volume="5" year="1860">Benson 1860</bibRefCitation>
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, p. 461.
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</paragraph>
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<paragraph id="92F33173C69A405798A8015598069D3A" pageId="5" pageNumber="134">
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<taxonomicName id="1ADB14EB2BF4B99109AF4BAB424294AA" family="Subulinidae" genus="Achatina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Achatina (Electra) hastula" order="Gastropoda" pageId="5" pageNumber="134" rank="species" species="hastula" subGenus="Electra">Achatina (Electra) hastula</taxonomicName>
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Benson: Hanley and Theobald 1876, pl. 18, fig. 4.
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</paragraph>
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<paragraph id="9CF269E8D7966BC1F8B1F365390AA0A1" pageId="5" pageNumber="134">
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<taxonomicName id="AEF60AB6AD2302DDD376130197001B1E" family="Subulinidae" genus="Stenogyra" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Stenogyra (Glessula) hastula" order="Gastropoda" pageId="5" pageNumber="134" rank="species" species="hastula" subGenus="Glessula">Stenogyra (Glessula) hastula</taxonomicName>
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, Benson:
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<bibRefCitation id="69D25855663FE4D2C41E27642E0974A1" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="14" pageNumber="143" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda." url="http://dx.doi.org/10.5962/bhl.title.54249" year="1878">Nevill 1878</bibRefCitation>
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, p. 169.
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</paragraph>
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<paragraph id="9CF412177F70D3A3505B3191C13C78FD" pageId="5" pageNumber="134">
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<taxonomicName id="48874CEE7E40A52759FA8A6B00DE1D4A" family="Subulinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. lessula" order="Gastropoda" pageId="5" pageNumber="134" rank="subspecies" species="lessula" subspecies="hastula">G. lessula hastula</taxonomicName>
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(Benson): Pilsbry 1909, p. 93.
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</paragraph>
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<paragraph id="A19D871488259D6EEA9C501734311EF3" pageId="5" pageNumber="134">
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<taxonomicName id="A7471681B79F34941C28B4C53DDCB44B" family="Subulinidae" genus="Glessula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Glessula hastula" order="Gastropoda" pageId="5" pageNumber="134" rank="species" species="hastula">Glessula hastula</taxonomicName>
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Benson:
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<bibRefCitation id="7DAB3525852F7E1D7F923EC7B2C2F129" author="Gude, GK" journalOrPublisher="London" pageId="14" pageNumber="143" title="The fauna of British India including Ceylon and Burma" volumeTitle="Mollusca" year="1914">Gude 1914</bibRefCitation>
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, p. 414.
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</paragraph>
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<paragraph id="E964D37D75FDD3E609DD97B775B30EEF" pageId="5" pageNumber="134">
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<taxonomicName id="73103FFBC7FB5EFC300B3150FD8AEFEC" family="Subulinidae" genus="Glessula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Glessula (Rishetia) hastula" order="Gastropoda" pageId="5" pageNumber="134" rank="species" species="hastula" subGenus="Rishetia">Glessula (Rishetia) hastula</taxonomicName>
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Benson:
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<bibRefCitation id="BB87C840EE4A1F88B62DDE5297DED769" author="Godwin-Austen, AA" journalOrPublisher="Land and freshwater Mollusca of India, Conchologia Indica" pageId="14" pageNumber="143" pagination="1 - 65" title="Genus Glessula." volume="3" year="1920">Godwin-Austen 1920</bibRefCitation>
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, p.16.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4161D926CC726D98A53D7F3C6AF1D4B" pageId="5" pageNumber="134" type="materials_examined">
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<paragraph id="D9397D3AD13A4668DEC161457DEA799C" pageId="5" pageNumber="134">Material examined.</paragraph>
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<paragraph id="3829F02005EA71847919FB98B31BAF00" pageId="5" pageNumber="134">
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CDZMTU059/24 shells and CDZMTU059P/5 specimens, Chitwan National Park, Central Nepal, riverine forest, opposite bank of Rapti River at Sauraha, 142-211 m,
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<geoCoordinate id="B9C02494FA53BC8CE50DEC03BCE9494C" direction="north" orientation="latitude" precision="1" value="27.571774">27.571774N</geoCoordinate>
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,
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<geoCoordinate id="5F66572BE08A7D5CDBEC3C754D3CE58F" direction="east" orientation="longitude" precision="1" value="84.48952">84.489514E</geoCoordinate>
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, 8.XI.2008. CDZMTU060/6 shells, Kumrose Community Forest, 197 m,
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<geoCoordinate id="E7E2E1F32015BB7DF7CA463E7BDE94E8" direction="north" orientation="latitude" precision="1" value="27.556519">27.556519N</geoCoordinate>
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,
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<geoCoordinate id="099E76718F32B1E921EAC84D6104D154" direction="east" orientation="longitude" precision="1" value="84.553024">84.553028E</geoCoordinate>
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, 21.X.2008. CDZMTU061/1 shell, Baghmara Community Forest, 201 m,
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<geoCoordinate id="11EDD20960BE58ADE4F9034B6B790B8B" direction="north" orientation="latitude" precision="1" value="27.5775">27.57750N</geoCoordinate>
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,
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<geoCoordinate id="0BC0D54FF3E4845EDC49A2413EF01EE0" direction="east" orientation="longitude" precision="1" value="84.46602">84.466017E</geoCoordinate>
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, 20.X.2008, leg. P.B. Budha.
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<taxonomicName id="03E957A3D62017D85E724EEA148E1DE4" lsidName="R. hastula" pageId="5" pageNumber="134" rank="species" species="hastula">R. hastula</taxonomicName>
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(Benson, 1860) at NHMUK, Godwin-Austen colln. Reg. No. 3557.03.VII.1.
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</paragraph>
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</subSubSection>
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<subSubSection id="43237D764204D93704CF643E80539687" pageId="5" pageNumber="134" type="type locality">
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<paragraph id="A5CDFC05790C63B035A107F982558D5F" pageId="5" pageNumber="134">Type locality.</paragraph>
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<paragraph id="C4E8A02872B322AC7E379BF741BDA94E" pageId="5" pageNumber="134">West Bengal "Pankabari (= Pankhabari), prope Darjeeling", India.</paragraph>
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</subSubSection>
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<subSubSection id="0C14FEA58B1C4867076C6E63D2341A94" pageId="5" pageNumber="134" type="distribution">
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<paragraph id="820A70FF0141EE4BE0EDE32B82C6D9CD" pageId="5" pageNumber="134">Distribution.</paragraph>
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<paragraph id="2282E1D6808CAE2C5D24272C47A0831E" pageId="5" pageNumber="134">
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NE India, Nepal (
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<bibRefCitation id="BA5FAF268CFA4BB073E151C852D3A20B" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="14" pageNumber="143" pagination="1 - 48" title="Annotated checklist of terrestrial gastropods of Nepal." url="http://dx.doi.org/10.3897/zookeys.492.9175" volume="392" year="2015">Budha et al. 2015</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="5F15E3717AC0BED664C855DFA51FDEF4" pageId="5" pageNumber="134" type="shell">
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<paragraph id="DA6046C9DA7630A11285632E9FBCBF30" pageId="5" pageNumber="134">Shell.</paragraph>
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<paragraph id="A33608C082C3BAE9CFC0990DE029E8D1" pageId="5" pageNumber="134">
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Measurements (n = 8): SH 9.5-13.6 mm, SW 3.5-4.1 mm, HA 3.3-3.6 mm, WA 2.0-2.5 mm, Wh 8.0-10.0; approx. 3
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<normalizedToken id="CB903DB1A06DE375DEE2B94B418A6528" originalValue="×">x</normalizedToken>
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higher than wide, slender, elongate, brownish, with dense radial riblets all over the shell. The first whorl blunt and smooth, fine riblets starting from the second whorl. Suture deep, spire rounded. Aperture ovate elliptical, nearly 2
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<normalizedToken id="AB5FCFBBEEE3CBB8C82E325075D3DC67" originalValue="×">x</normalizedToken>
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higher than wide, peristome thin, columellar margin calloused whitish, obliquely truncate at the base.
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</paragraph>
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</subSubSection>
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<subSubSection id="3CFE043BA08FFC5096DC2B96EF001B0A" pageId="5" pageNumber="134" type="genitalia">
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<paragraph id="160FDE2876BAD158E56F5C2E8ACF4C38" pageId="5" pageNumber="134">Genitalia</paragraph>
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<paragraph id="3E6E801AB652C3A0B48CE67197F49E8F" pageId="5" pageNumber="134">(n = 5) (Fig. 6). Flagellum very short tubular sac; there is a very short knob-like epiphallic caecum. Penis cylindrical, basal end comparatively narrow, swollen at the middle and proximal portions cylindrical. Epiphallus basally swollen and the proximal portion tapering. The penial retractor muscle far apart from the flagellum. Gametolytic sac balloon-like, separated from the gametolytic duct by a narrow neck. A mature dissected specimen contained 4-5 gelatinous eggs in the spermoviduct. Another mature specimen contained a spermatophore in the gametolytic sac. This is the first observation of a glessuline spermatophore (Fig. 6B). Vagina short, nearly half the length of the penis. Albumen gland elongate and yellowish; hermaphroditic duct thinner than the albumen gland in the observed specimens.</paragraph>
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<caption id="8FD7AFC79856004446C4C286FE781C71" pageId="5" pageNumber="134">
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<paragraph id="1FFFC46C21F3E662C4F25DCEFEF86D5E" pageId="5" pageNumber="134">
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Figure 6. Genitalia of
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<taxonomicName id="1628BB143C497EC1E2CCAB04C78AF493" class="Gastropoda" family="Achatinidae" genus="Rishetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rishetia hastula" order="Stylommatophora" pageId="5" pageNumber="134" phylum="Mollusca" rank="species" species="hastula">Rishetia hastula</taxonomicName>
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, CDZMTU059P, A General view B Spermatophore of the same, Chitwan National Park, riverine forest opposite bank of Sauraha, Rapti River.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="1028F3667FFBB2C9BF47CE92440DE4B2" pageId="5" pageNumber="134" type="remarks">
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<paragraph id="A2A52438C0939D022031EEDAD7C22EDE" pageId="5" pageNumber="134">Remarks.</paragraph>
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<paragraph id="8B3E15643F05D01AF31BA58E4ED1CF55" pageId="5" pageNumber="134">
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<taxonomicName id="E3D2107DA244FD786D63335F8E06CC37" lsidName="R. hastula" pageId="5" pageNumber="134" rank="species" species="hastula">R. hastula</taxonomicName>
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is common in subtropical riverine floodplain forest leaf litter at lower altitudes (up to 300 m) in Chitwan National Park and the adjacent bufferzone community forests.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |