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<document ID-CLB-Dataset="3848" ID-DOI="http://dx.doi.org/10.3897/zookeys.1062.66845" ID-GBIF-Dataset="e73f66c3-4c75-4c11-9ad1-be2def71e368" ID-PMC="PMC8530997" ID-Pensoft-Pub="1313-2970-1062-73" ID-Pensoft-UUID="2BCBF5DF56B450B1A9A88480A9F8B9CE" ID-PubMed="34720618" ID-ZooBank="310272E8F5DD4DAB84603AFCCD2E9C45" ModsDocID="1313-2970-1062-73" checkinTime="1634290158865" checkinUser="pensoft" docAuthor="Zhang, Jianshuang, Yu, Hao &amp; Li, Shuqiang" docDate="2021" docId="6B51C26A88B759F397FBC556977F3155" docLanguage="en" docName="ZooKeys 1062: 73-122" docOrigin="ZooKeys 1062" docPubDate="2021-10-14" docSource="http://dx.doi.org/10.3897/zookeys.1062.66845" docTitle="Matidia xieqian Yu &amp; Li 2021, sp. nov." docType="treatment" docUuid="C948F60A-43D6-4898-BF0D-CD89EFF124A6" docUuidSource="ZooBank" docVersion="6" id="2BCBF5DF56B450B1A9A88480A9F8B9CE" lastPageNumber="73" masterDocId="2BCBF5DF56B450B1A9A88480A9F8B9CE" masterDocTitle="On the clubionid spiders (Araneae, Clubionidae) from Xishuangbanna, China, with descriptions of two new genera and seven new species" masterLastPageNumber="122" masterPageNumber="73" pageNumber="73" updateTime="1732931016866" updateUser="ExternalLinkService">
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<mods:title id="059BF845158B78F9E91C95AE23BA8049">On the clubionid spiders (Araneae, Clubionidae) from Xishuangbanna, China, with descriptions of two new genera and seven new species</mods:title>
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<mods:namePart id="1E705D9CF1A57D6AA09362A1929453B6">Zhang, Jianshuang</mods:namePart>
<mods:affiliation id="5D788CFC7921D823267F06A1C8C4E96D">School of Life Sciences, Guizhou Normal University, Guiyang, Guizhou, China</mods:affiliation>
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<mods:namePart id="193CC4772A93A297C61EFC224060A692">Yu, Hao</mods:namePart>
<mods:affiliation id="E6BC0DF3F57C1249EB225C2ABCB9E7A9">School of Biological Sciences, Guizhou Education University, Guiyang, Guizhou, China</mods:affiliation>
<mods:nameIdentifier id="2AC5D64EDAA27A4F160C49AD7B973202" type="email">insect1986@126.com</mods:nameIdentifier>
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<mods:namePart id="ED54137937A2DFE6FF851007FFDBB6AA">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="F18E0937B23C829C94FFD12B33B0DEA5" type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="79F3FADFEC60CF76A5BEFC7C7860C18B">Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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<mods:date id="F86231745AEA0CA751571AD5DC443EAD">2021</mods:date>
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<mods:number id="5A9680B3AE8B3B9C4B4044293F882108">2021-10-14</mods:number>
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<treatment id="6B51C26A88B759F397FBC556977F3155" ID-GBIF-Taxon="189468733" LSID="urn:lsid:zoobank.org:act:C948F60A-43D6-4898-BF0D-CD89EFF124A6" httpUri="http://treatment.plazi.org/id/6B51C26A88B759F397FBC556977F3155" lastPageNumber="73" pageId="0" pageNumber="73" scope_family="Clubionidae" scope_order="Araneae">
<subSubSection id="3BC66FC153DA2BEEC727155B59BD9F84" pageId="0" pageNumber="73" type="nomenclature">
<paragraph id="E6E8D251FB8F5877B7A5C2F54CCD1C27" pageId="0" pageNumber="73">
<taxonomicName id="1D3A141467F34CB6E5D38A6AA4882F65" ID-CoL="98YMF" LSID="http://zoobank.org/C948F60A-43D6-4898-BF0D-CD89EFF124A6" authority="Yu &amp; Li" authorityName="Yu &amp; Li" authorityYear="2021" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia xieqian" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="xieqian" status="sp. nov.">Matidia xieqian Yu &amp; Li</taxonomicName>
<taxonomicNameLabel id="BE86BC9A88512368C6D55E64BA83461B" pageId="0" pageNumber="73">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6E286B60422DC1729B3BF3B4BA16E982" pageId="0" pageNumber="73" type="description">
<paragraph id="0F1D5A64C491651AF00AD8B00B4EC30D" pageId="0" pageNumber="73">
<figureCitation id="945C3A26137C077FBD8336706C1421ED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of the holotype of Matidia xieqian sp. nov., left palp and bulb (A, B, E, F) and flipped right bulb (C, D) A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, ventrolateral view F retrolateral view. Arrows (E) point at three processes of tegular apophysis. Abbreviations: C = conductor; E = embolus; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.20 mm (equal for A, B, equal for C-F)." figureDoi="10.3897/zookeys.1062.66845.figure5" httpUri="https://binary.pensoft.net/fig/599509" pageId="0" pageNumber="73">Figs 5</figureCitation>
<figureCitation id="CD785124BDBBA4B026052E3742103CE1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="721DF52AAFB2C6780E282C6AEDEAEA44" pageId="0" pageNumber="73" type="type material">
<paragraph id="03F2E86F0BB34C26E920CC2213D23A85" pageId="0" pageNumber="73">Type material.</paragraph>
<paragraph id="B878F033AE504396A5FF46842D964EB1" pageId="0" pageNumber="73">
<emphasis id="EDDCF086F166F40AF73B427A82B47BCC" bold="true" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
♂ (IZCAS-Ar34728), China:
<emphasis id="3CBCBDCA450BCCD705F8ED29A391F9CF" bold="true" pageId="0" pageNumber="73">Yunnan</emphasis>
: Xishuangbanna: Mengla County: Menglun Town: Menglun Nature Reserve: secondary tropical montane evergreen broad-leaved forest (
<geoCoordinate id="CB2D6761DD518D0DE2DE27D3BB1C356F" degrees="21" direction="north" minutes="57.784" orientation="latitude" precision="1" value="21.963066">21°57.784'N</geoCoordinate>
,
<geoCoordinate id="B9EA99C85E599E746AAAD3EE37097B7B" degrees="101" direction="east" minutes="11.947" orientation="longitude" precision="1" value="101.19912">101°11.947'E</geoCoordinate>
, 895 m), 6 August 2007, G. Zheng leg.
<emphasis id="F29FCB9DB008B5B693335D7DBFA834E8" bold="true" italics="true" pageId="0" pageNumber="73">Paratype</emphasis>
: 1♀ (IZCAS-Ar34729), secondary tropical montane evergreen broad-leaved forest (
<geoCoordinate id="2008D010BDCD1C43381FC14ED935BD5F" degrees="21" direction="north" minutes="57.534" orientation="latitude" precision="1" value="21.9589">21°57.534'N</geoCoordinate>
,
<geoCoordinate id="BB45169F89C99E4DA1976C763860010E" degrees="101" direction="east" minutes="12.300" orientation="longitude" precision="1" value="101.205">101°12.300'E</geoCoordinate>
, 860 m), 4 August 2007, G. Zheng leg.
</paragraph>
</subSubSection>
<subSubSection id="1AB2A1006CDAAECFD1CA58E4ED099D16" pageId="0" pageNumber="73" type="materials_examined">
<paragraph id="710EC5F1D12679B1F86B7327AD357140" pageId="0" pageNumber="73">Other material examined.</paragraph>
<paragraph id="595E3591F8A548FADCF8FCB849544A58" pageId="0" pageNumber="73">
<materialsCitation id="252477819DEBEAE44BBE0D8DB7F563C7" collectingDate="2011-08-12" collectorName="G. Zheng" country="China" county="Mengla County" elevation="1088" latitude="21.9784" location="Menglun Town" longLatPrecision="1" longitude="101.32913" municipality="Xishuangbanna" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<collectingCountry id="825DBC5C0E0C6637E128A46BC44230B9" name="China">China</collectingCountry>
:
<emphasis id="C5C27E338D5C8787320086AAD7A269EC" bold="true" pageId="0" pageNumber="73">
<collectingRegion id="7B0AE9BBC2ADEAE034CF8A950C44A077" country="China" name="Yunnan">Yunnan</collectingRegion>
</emphasis>
:
<collectingMunicipality id="098195E194E9C20AA7E252B308BF8B7E">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="C40993D67124CDCBEC1371E3C7D68D51">Mengla County</collectingCounty>
:
<location id="906723F9A58F36EE9B31EC609336DBD6" LSID="urn:lsid:plazi:treatment:6B51C26A88B759F397FBC556977F3155:906723F9A58F36EE9B31EC609336DBD6" country="China" county="Mengla County" latitude="21.9784" longLatPrecision="1" longitude="101.32913" municipality="Xishuangbanna" name="Menglun Town" stateProvince="Yunnan">Menglun Town</location>
:
<location id="B1CC92B9D859758C0CA2776C6C681195" LSID="urn:lsid:plazi:treatment:6B51C26A88B759F397FBC556977F3155:B1CC92B9D859758C0CA2776C6C681195" country="China" county="Mengla County" latitude="21.9784" longLatPrecision="1" longitude="101.32913" municipality="Xishuangbanna" name="Menglun Nature Reserve" stateProvince="Yunnan">Menglun Nature Reserve</location>
:
<specimenCount id="FBD3DD73F562B551B627816D8EA96705" type="male">1♂</specimenCount>
(YHCLU0126),
<quantity id="9BC5ED5CFFE8AA1690E5AB01A31ED4BD" metricMagnitude="4" metricUnit="m" metricValue="4.8" unit="km" value="48.0">48 km</quantity>
landmark, seasonal rainforest (
<geoCoordinate id="ABBFA36BCBE1A1E341DD0962BB32A3B7" degrees="21" direction="north" minutes="58.704" orientation="latitude" precision="1" value="21.9784">21°58.704'N</geoCoordinate>
,
<geoCoordinate id="F3BAF872463D19CD119ED00B41F270CC" degrees="101" direction="east" minutes="19.748" orientation="longitude" precision="1" value="101.32913">101°19.748'E</geoCoordinate>
,
<quantity id="388A30171AED4661EE1DEC08BCFD3866" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" unit="m" value="1088.0">
<elevation id="DB28D5D3508198453EF5A0D050BE490A" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" unit="m" value="1088.0">1088 m</elevation>
</quantity>
),
<collectingDate id="69EC14DA76FE9B8612DAE01F88237E56" value="2011-08-12">12 August 2011</collectingDate>
,
<collectorName id="382898A388EDBAE1590413951FBE5FAB">G. Zheng</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="F24EFB75F3350ED8C438856D65D62298" collectingDate="2012-06-30" collectorName="Q. Zhao" country="China" elevation="1137" latitude="22.083366" location="Wengnan Village" longLatPrecision="1" longitude="100.36681" municipality="Meng'a Town" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<collectingMunicipality id="9A1B5AFC09360CDAB73F5FB2270F50DB">
<normalizedToken id="0D7B058D7B96628233BF3378415F0DDE" originalValue="Menga">Meng'a</normalizedToken>
Town
</collectingMunicipality>
:
<specimenCount id="DF50BF48D469A0463150B78964340AF3" type="female">1♀</specimenCount>
(YHCLU0127),
<location id="08704761484FA396B1D1B8FE90E7478F" LSID="urn:lsid:plazi:treatment:6B51C26A88B759F397FBC556977F3155:08704761484FA396B1D1B8FE90E7478F" country="China" latitude="22.083366" longLatPrecision="1" longitude="100.36681" municipality="Meng'a Town" name="Wengnan Village" stateProvince="Yunnan">Wengnan Village</location>
, secondary seasonal rain forest (
<geoCoordinate id="03E69A5AD71DE357CABFBF98E738B8BF" degrees="22" direction="north" minutes="05.002" orientation="latitude" precision="1" value="22.083366">22°05.002'N</geoCoordinate>
,
<geoCoordinate id="8D9EA384ACFB5178A5650C1995187CFC" degrees="100" direction="east" minutes="22.009" orientation="longitude" precision="1" value="100.36681">100°22.009'E</geoCoordinate>
,
<quantity id="CA29DB591142BA5F9EEEC8CD85365F60" metricMagnitude="3" metricUnit="m" metricValue="1.137" unit="m" value="1137.0">
<elevation id="51D57FAD671A087407F65B969ED02299" metricMagnitude="3" metricUnit="m" metricValue="1.137" unit="m" value="1137.0">1137 m</elevation>
</quantity>
),
<collectingDate id="524F1849C1329814BA1C6E8626E6C2B7" value="2012-06-30">30 June 2012</collectingDate>
,
<collectorName id="1E884B32860E83A7DD6EE62130412B9F">Q. Zhao</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="77006E6F9BD3787362615D5B0881D632" pageId="0" pageNumber="73" type="etymology">
<paragraph id="9EB3D610CCA1BD18FAF9CA6A52A6DF95" pageId="0" pageNumber="73">Etymology.</paragraph>
<paragraph id="3F3FC87D6F0612A082B15980FEC69697" pageId="0" pageNumber="73">
The specific name is derived from the Chinese pinyin
<emphasis id="1E95014B5E9719B37C88684DBA3A8390" italics="true" pageId="0" pageNumber="73">
<normalizedToken id="1D0099890442D3F3258B935D95F78667" originalValue="xiè">xie</normalizedToken>
<normalizedToken id="7F01B56998CCC71E14DCCD114C70C7E4" originalValue="qián">qian</normalizedToken>
</emphasis>
, which means crab claw, referring to the concave retrolateral tibial apophysis with two branches resembling a crab claw; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="37A567F7F840B91DD86EDCCD4788D02C" pageId="0" pageNumber="73" type="diagnosis">
<paragraph id="8CEAFDC89514504C3D5BA68B99C00412" pageId="0" pageNumber="73">Diagnosis.</paragraph>
<paragraph id="E815062DB5533C89BFA63D48360A2388" pageId="0" pageNumber="73">
Males of
<taxonomicName id="AD5DE1C2824C86E3F0F4EA2F4DC08AD6" lsidName="M. xieqian" pageId="0" pageNumber="73" rank="species" species="xieqian">
<emphasis id="250970041495202B3AC73A0F7FB4A8E4" italics="true" pageId="0" pageNumber="73">M. xieqian</emphasis>
</taxonomicName>
sp. nov. can be distinguished from other
<taxonomicName id="FAACD906B085C9FD3B4D99D6AEF710B3" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="E4DB21E4FB01CD2D5CCCB971F084277B" italics="true" pageId="0" pageNumber="73">Matidia</emphasis>
</taxonomicName>
species by the branched retrolateral tibial apophysis (Fig.
<figureCitation id="19EB0FDF08B35368B18563D94A7C1F99" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of the holotype of Matidia xieqian sp. nov., left palp and bulb (A, B, E, F) and flipped right bulb (C, D) A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, ventrolateral view F retrolateral view. Arrows (E) point at three processes of tegular apophysis. Abbreviations: C = conductor; E = embolus; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.20 mm (equal for A, B, equal for C-F)." figureDoi="10.3897/zookeys.1062.66845.figure5" httpUri="https://binary.pensoft.net/fig/599509" pageId="0" pageNumber="73">5B</figureCitation>
) and a tegular apophysis with three processes (Fig.
<figureCitation id="8FE0393444D39267B39574F9F15B8292" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of the holotype of Matidia xieqian sp. nov., left palp and bulb (A, B, E, F) and flipped right bulb (C, D) A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, ventrolateral view F retrolateral view. Arrows (E) point at three processes of tegular apophysis. Abbreviations: C = conductor; E = embolus; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.20 mm (equal for A, B, equal for C-F)." figureDoi="10.3897/zookeys.1062.66845.figure5" httpUri="https://binary.pensoft.net/fig/599509" pageId="0" pageNumber="73">5E</figureCitation>
) (vs. RTA unbranched, tegular apophysis absent or present but with only process in other species). Females of this species resemble those of
<taxonomicName id="23A5B95C12E3407AB580CE61E8B00C07" lsidName="M. simia" pageId="0" pageNumber="73" rank="species" species="simia">
<emphasis id="7F92692BBCA135D8FED293360F8DDD57" italics="true" pageId="0" pageNumber="73">M. simia</emphasis>
</taxonomicName>
Deeleman-Reinhold, 2001 in having an epigynal atrium, lacking in all other
<taxonomicName id="9D6ACF5B0585C9705D5C6483BB20E084" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B6049E27C076463B1D2C31DA23B403EB" italics="true" pageId="0" pageNumber="73">Matidia</emphasis>
</taxonomicName>
species, but differ by the: (1) copulatory ducts 3
<normalizedToken id="BF411FC5EBF01D5138FEF002855FE9FB" originalValue="×">x</normalizedToken>
longer than spermathecae (Fig.
<figureCitation id="BC0A8355BD1242D271DA160548445720" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6C, D</figureCitation>
) (vs. &lt;2
<normalizedToken id="5F71FEF3DCE63D7CDDC3F72F0950454A" originalValue="×">x</normalizedToken>
longer than spermathecae); (2) spermathecae situated posteriorly to bursae (Fig.
<figureCitation id="DA54DA8A67CCACD7D8A1DB9182D6A593" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6C, D</figureCitation>
) (vs. anteriorly); (3) spermathecae not coiled (Fig.
<figureCitation id="CBD80D2E74730784A95C0463AD3B37C4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6C, D</figureCitation>
) (vs. coiled).
</paragraph>
<caption id="CB9651098AEA77AFE4B4A5DFCDA33B8B" doi="10.3897/zookeys.1062.66845.figure5" httpUri="https://binary.pensoft.net/fig/599509" pageId="0" pageNumber="73" start="Figure 5" startId="F5">
<paragraph id="6EF05C4653EB9381F22231F01662CEB5" pageId="0" pageNumber="73">
<emphasis id="5396208408559C72A6100C07A70ABDFB" bold="true" pageId="0" pageNumber="73">Figure 5.</emphasis>
Male palp of the holotype of
<taxonomicName id="344342824B0E7241850C84F8D7282C17" authorityName="Yu &amp; Li" authorityYear="2021" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia xieqian" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="xieqian">
<emphasis id="57AAAE34831739E5E948DD516B9E8693" italics="true" pageId="0" pageNumber="73">Matidia xieqian</emphasis>
</taxonomicName>
sp. nov., left palp and bulb (
<emphasis id="4ACBBAAF4F86FCA90104601C54D29123" bold="true" pageId="0" pageNumber="73">A, B, E, F</emphasis>
) and flipped right bulb (
<emphasis id="932C94B64CA0B0305D57683CE69B8ADA" bold="true" pageId="0" pageNumber="73">C, D</emphasis>
)
<emphasis id="56B4544181B990B0BDB07DB777CC9ECD" bold="true" pageId="0" pageNumber="73">A</emphasis>
prolateral view
<emphasis id="E6E74070C571C232B931FEC33CBFD30C" bold="true" pageId="0" pageNumber="73">B</emphasis>
retrolateral view
<emphasis id="2E88826A4EF2ED53A1A526C574651777" bold="true" pageId="0" pageNumber="73">C</emphasis>
bulb, prolateral view
<emphasis id="C97EB08B62D7ADBA2FD1A1396C4B46F2" bold="true" pageId="0" pageNumber="73">D</emphasis>
bulb, ventral view
<emphasis id="3DA90E8665515152D9FE74D16C2822AE" bold="true" pageId="0" pageNumber="73">E</emphasis>
bulb, ventrolateral view
<emphasis id="21BF98387B4A62CB2FD4A03C3E05BEA4" bold="true" pageId="0" pageNumber="73">F</emphasis>
retrolateral view. Arrows (
<emphasis id="DD650E6D8CA99CFC18F2B748F40FED78" bold="true" pageId="0" pageNumber="73">E</emphasis>
) point at three processes of tegular apophysis. Abbreviations: C = conductor; E = embolus; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.20 mm (equal for
<emphasis id="140D45CAEA0530155DE12C6DC1AAF967" bold="true" pageId="0" pageNumber="73">A, B</emphasis>
, equal for
<emphasis id="BA58E9941053B4CA7544B7DD39545EEB" bold="true" pageId="0" pageNumber="73">C-F</emphasis>
).
</paragraph>
</caption>
<caption id="966F27FE904FF3E74C9A11F43DC567FE" doi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73" start="Figure 6" startId="F6">
<paragraph id="4BF9BD1E0565C4E729697AB605634D42" pageId="0" pageNumber="73">
<emphasis id="12F5825D2100CD915CD4CA27D136A447" bold="true" pageId="0" pageNumber="73">Figure 6.</emphasis>
<taxonomicName id="BB7B7EAB5B31DBCDF4AB57A4666EA08E" authorityName="Yu &amp; Li" authorityYear="2021" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia xieqian" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="xieqian">
<emphasis id="D895280792A293AAA9BD8A2E149F2387" italics="true" pageId="0" pageNumber="73">Matidia xieqian</emphasis>
</taxonomicName>
sp. nov., female paratype and male holotype, epigyne (
<emphasis id="C534D20B1490AF3C4A0BD8646F8C9C8D" bold="true" pageId="0" pageNumber="73">A-D</emphasis>
), male habitus (
<emphasis id="9B0018046DFE2D0CEFAF930148EE3B77" bold="true" pageId="0" pageNumber="73">E, F</emphasis>
) and female habitus (
<emphasis id="75A7437280942834746A91A9CBA5DFE4" bold="true" pageId="0" pageNumber="73">G, H</emphasis>
)
<emphasis id="2924D0AE66A4B908CAB66B62D6694841" bold="true" pageId="0" pageNumber="73">A</emphasis>
intact, ventral view
<emphasis id="D225125AC9BAD9A9D91709646F250834" bold="true" pageId="0" pageNumber="73">B</emphasis>
cleared, ventral view
<emphasis id="1E476D69169AB4FBA842848D5DDB1F77" bold="true" pageId="0" pageNumber="73">C</emphasis>
cleared, dorsal view
<emphasis id="98AE8D170411FFE136B8085F2796A59C" bold="true" pageId="0" pageNumber="73">D</emphasis>
cleared, dorsal view; path of copulatory duct marked
<emphasis id="95BCBF617A124FEC36B41BD67E0E46A0" bold="true" pageId="0" pageNumber="73">E</emphasis>
dorsal view
<emphasis id="2EDD63EF46FCDDE1F592BD7F5CF73B39" bold="true" pageId="0" pageNumber="73">F</emphasis>
lateral view
<emphasis id="1DC2F8703924C5CEAE0F35957A34FF3D" bold="true" pageId="0" pageNumber="73">G</emphasis>
dorsal view
<emphasis id="2C72D6D022E3A18591085E19624947A7" bold="true" pageId="0" pageNumber="73">H</emphasis>
ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for
<emphasis id="5E062AE207C74DB7E28E662396DCA100" bold="true" pageId="0" pageNumber="73">A-D</emphasis>
); 1 mm (equal for
<emphasis id="EA0BFE5731B5C512A384C2DD4922AE38" bold="true" pageId="0" pageNumber="73">E, F</emphasis>
, equal for
<emphasis id="A2C87B0D350B400A621715EED669373F" bold="true" pageId="0" pageNumber="73">G, H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C95321089E20AEEAE793F5E3A2E47FE2" pageId="0" pageNumber="73" type="description">
<paragraph id="64F8AD34FB1F333BF5BC4E4A014815DB" pageId="0" pageNumber="73">Description.</paragraph>
<paragraph id="BDA51FB4C29A59911D972D9AFE069695" pageId="0" pageNumber="73">
<emphasis id="093BA6D9A595B2943262A9C5CD4A12B7" bold="true" pageId="0" pageNumber="73">Male</emphasis>
(holotype) (Fig.
<figureCitation id="ED068E459D7D40925F4357410C295FB5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6E, F</figureCitation>
). Total length 5.30; carapace 1.99 long, 1.50 wide; opisthosoma 3.32 long, 0.98 wide. Carapace yellowish brown posteriorly and centrally, dark anteriorly and marginally, without distinct pattern; cervical groove and radial groove distinct, fovea indistinct. Eyes: AER slightly recurved, PER wider than AER, almost straight in dorsal view. AME dark, other eyes light, with black rings. Eye sizes and interdistances: AME 0.08, ALE 0.10, PME 0.10, PLE 0.10, AME-AME 0.05, AME-ALE 0.05, PME-PME 0.22, PME-PLE 0.06, MOQL 0.28, MOQA 0.24, MOQP 0.44. Chelicerae coloured as ocular region, with 3 promarginal and 2 retromarginal teeth. Labium and endites light brown. Sternum yellowish white. Legs uniformly yellowish orange. Leg measurements: I 12.71(3.35, 5.08, 2.93, 1.35), II 10.53 (3.02, 4.26, 2.27, 0.98), III 6.02 (1.75, 2.11, 1.67, 0.50), IV 11.02 (3.11, 3.64, 3.46, 0.81). Abdomen lanceolate, white, dorsum with a lengthwise, white heart mark reaching posterior half of abdomen; pair of muscle depressions on both sides of heart-shaped mark; venter, spinnerets yellowish white.
</paragraph>
<paragraph id="32A528862D155D801F31263A55202F7C" pageId="0" pageNumber="73">
Palp (Fig.
<figureCitation id="BB316FF0D127B0D00EBA41BA8992A47B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of the holotype of Matidia xieqian sp. nov., left palp and bulb (A, B, E, F) and flipped right bulb (C, D) A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, ventrolateral view F retrolateral view. Arrows (E) point at three processes of tegular apophysis. Abbreviations: C = conductor; E = embolus; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.20 mm (equal for A, B, equal for C-F)." figureDoi="10.3897/zookeys.1062.66845.figure5" httpUri="https://binary.pensoft.net/fig/599509" pageId="0" pageNumber="73">5A-F</figureCitation>
): Tibia short, ~ 1/4-1/3 of cymbium length, with two apophyses: PTA subtriangular in prolateral view, with a bulky base and blunt tip, ~ 1/2 of tibia length; RTA approximately as long as tibia, heavily sclerotised, strongly expanded, directed antero-dorsally, concave with two branches, shaped like a crab claw, ventral branch thumb-shaped, dorsal branch finger-like. Bulb elongated, with distinct, sinuate sperm duct. Tegular apophysis well developed,&gt; 2/3 of tegulum length, with three processes: apico-prolateral process sharp, tooth-shaped, meso-prolateral process blunt, trapezoidal, baso-prolateral process represented by a blunt flange. Embolus wide, approximately as long as tegulum, originating at ~ 8
<normalizedToken id="EE3271F0785626D8DEE7F9512094CAB0" originalValue="oclock">o'clock</normalizedToken>
, beak-shaped tip terminated at ~ 1
<normalizedToken id="71B4F8E00ED5F9DA91C9BF56B08C59A5" originalValue="oclock">o'clock</normalizedToken>
position. Conductor thick, membranous, ~ 1.1-1.2
<normalizedToken id="ADEF62123DC6A8DBE38CA9CA364333B7" originalValue="×">x</normalizedToken>
longer than embolus, originating baso-retrolaterally from tegulum, spoon-shaped distally.
</paragraph>
<paragraph id="761B019B18B32B8F7C68CD53DFD02B6A" pageId="0" pageNumber="73">
<emphasis id="06013C84CDBBF389840C93025DD12422" bold="true" pageId="0" pageNumber="73">Female</emphasis>
(paratype IZCAS-Ar34729): Total length 5.46; carapace 2.10 long, 1.47 wide; opisthosoma 3.41 long, 0.98 wide. General characters as in male (Fig.
<figureCitation id="0400B84FE7E7D153EB9F70477D130203" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6G, H</figureCitation>
). Eye sizes and interdistances: AME 0.07, ALE 0.10, PME 0.11, PLE 0.09, AME-AME 0.06, AME-ALE 0.05, PME-PME 0.22, PME-PLE 0.05, MOQL 0.53, MOQA 0.21, MOQP 0.42. Legs white, without distinct markings. Leg measurements: I 10.10 (2.82, 4.15, 2.26, 0.87), II 6.59 (1.87, 2.37, 1.60, 0.75), III 5.62 (1.75, 1.89, 1.54, 0.45), IV 9.92 (3.05, 3.18, 2.98, 0.70).
</paragraph>
<paragraph id="A03F982A1AA688616AA276AC104F53BE" pageId="0" pageNumber="73">
Epigyne (Fig.
<figureCitation id="B4BD77B0B09D7BA57F04413528019180" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6A-D</figureCitation>
): Plate fan-shaped, length subequal to width, margins distinctly delimited, spermathecae and bursae indistinct through integument. Atrium large, ~ 1/4 of epigyne length and 1/3 of epigyne width, located at posterior portion of epigynal plate. Copulatory openings indistinct, located at basolateral atrial borders. Hyaline copulatory ducts ascending anteriorly, following atrial borders, curved at half their length to form a loop, and ascending to spermathecae. Spermathecae bean-shaped, ~ 1.5
<normalizedToken id="0CABF197CFB1851B5411A2D582AFA272" originalValue="×">x</normalizedToken>
longer than wide, at lateral portion of vulva, separated by three diameters. Fertilisation ducts as long as spermathecae, blade-shaped. Bursae hyaline, oval, much larger than spermathecae, ~ 1.5
<normalizedToken id="B3EA62C72958880C605D17CEB0EA843B" originalValue="×">x</normalizedToken>
longer than wide, situated anteriorly, separated by ~ 0.8 diameters, surface semi-transparent.
</paragraph>
</subSubSection>
<subSubSection id="A7F658A2640DD9CE8F6E76145A797B7B" pageId="0" pageNumber="73" type="distribution">
<paragraph id="3D9DD0F40E6A24437514F56F9EDDA7DC" pageId="0" pageNumber="73">Distribution.</paragraph>
<paragraph id="9074C3F9A7132B63D1FEC41D2BC3F4E8" pageId="0" pageNumber="73">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</document>