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<document id="F4325F35E46E6CED29173D8F6AEC73AA" ID-CLB-Dataset="21692" ID-DOI="10.5252/geodiversitas2020v42a13" ID-GBIF-Dataset="d64d995c-06c4-4541-aae0-737c86d030a9" ID-ISSN="1638-9395" ID-Zenodo-Dep="3894810" ID-ZooBank="urn:lsid:zoobank.org:pub:52D33BB7-2713-4F9D-AACF-3FAA90FDB31F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592234070810" checkinUser="valdenar" docAuthor="Solé, Floréal, Marandat, Bernard &amp; Lihoreau, Fabrice" docDate="2020" docId="03C08782FFDCFFE9FEF23F19FD4FF85E" docLanguage="en" docName="GEODIVERSITAS.42.13.185-214.pdf.imf" docOrigin="Geodiversitas 42 (13)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Oxyaenoides aumelasiensis Solé &amp; Marandat &amp; Lihoreau 2020, n. sp" docType="treatment" docUuid="AFFF7C27-FB65-49C0-A788-96352B214851" docUuidSource="ZooBank" docVersion="9" lastPageNumber="192" masterDocId="FFF9FFFAFFDBFFE0FFE73A2CFF8FFFB3" masterDocTitle="The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian" masterLastPageNumber="214" masterPageNumber="185" pageNumber="190" updateTime="1701999769407" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="91C917DB10B36FFC622F02A720FD64CA">The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian</mods:title>
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<mods:namePart id="570B7224078F06468F6E67C7603D912B">Solé, Floréal</mods:namePart>
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<mods:namePart id="A62483E3E1CBFA7BC72F86E7F59D85B9">Marandat, Bernard</mods:namePart>
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<mods:namePart id="568AB6EF4C83BCC6B9B6C71F1EFDDE5C">Lihoreau, Fabrice</mods:namePart>
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<mods:title id="5D098C080627EE81FF76010D87E7831C">Geodiversitas</mods:title>
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<mods:date id="B3467935A4518F18FF5AFD7F127E331A">2020</mods:date>
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<treatment id="03C08782FFDCFFE9FEF23F19FD4FF85E" ID-GBIF-Taxon="164597322" LSID="urn:lsid:zoobank.org:act:AFFF7C27-FB65-49C0-A788-96352B214851" httpUri="http://treatment.plazi.org/id/03C08782FFDCFFE9FEF23F19FD4FF85E" lastPageId="9" lastPageNumber="192" pageId="7" pageNumber="190">
<subSubSection id="C373651FFFDCFFE7FEF23F19FE62FAC3" pageId="7" pageNumber="190" type="nomenclature">
<paragraph id="8BD63694FFDCFFE7FEF23F19FE62FAC3" blockId="7.[277,630,1333,1392]" pageId="7" pageNumber="190">
<heading id="D09E81F8FFDCFFE7FEF23F19FDFDFAE3" box="[277,626,1333,1360]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="190" reason="2">
<taxonomicName id="4C694D17FFDCFFE7FEF23F19FDBBFAE3" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[277,564,1333,1360]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="190" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFDCFFE7FEF23F19FDBBFAE3" bold="true" box="[277,564,1333,1360]" italics="true" pageId="7" pageNumber="190">Oxyaenoides aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDCFFE7FDDB3F1AFDFDFAE3" box="[572,626,1334,1360]" pageId="7" pageNumber="190" rank="species">n. sp</taxonomicNameLabel>
</heading>
. (Fig. 2)
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE7FF633FB9FC88FA15" box="[132,775,1429,1446]" pageId="7" pageNumber="190" type="description">
<paragraph id="8BD63694FFDCFFE7FF633FB9FC88FA15" blockId="7.[132,775,1429,1446]" box="[132,775,1429,1446]" pageId="7" pageNumber="190">urn:lsid:zoobank.org:act:AFFF7C27-FB65-49C0-A788-96352B214851</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE7FF633FE2FED4F9AF" pageId="7" pageNumber="190" type="materials_examined">
<paragraph id="8BD63694FFDCFFE7FF633FE2FED4F9AF" blockId="7.[132,775,1486,1609]" pageId="7" pageNumber="190">
<materialsCitation id="3B013CC9FFDCFFE7FF633FE2FED4F9AF" ID-GBIF-Occurrence="2635845302" collectionCode="UM-AUM" pageId="7" pageNumber="190" specimenCode="UM-AUM693" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D28836FFDCFFE7FF633FE2FF60FA57" box="[132,239,1486,1509]" pageId="7" pageNumber="190" type="holotype">HOLOTYPE</typeStatus>
. —
<specimenCode id="DBCF9EEFFFDCFFE7FEFB3FE3FE3FFA54" box="[284,432,1487,1511]" collectionCode="UM-AUM" pageId="7" pageNumber="190">UM-AUM693</specimenCode>
, left mandible bearing m3, the anterior root and talonid of the m1, the posterior alveolus of the p4, and the alveoli of m2.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE7FF633C1CFE5FF89B" pageId="7" pageNumber="190" type="etymology">
<paragraph id="8BD63694FFDCFFE7FF633C1CFD07F9FA" blockId="7.[132,775,1486,1609]" box="[132,648,1584,1609]" pageId="7" pageNumber="190">
ETYMOLOGY. — Latin, from Aumelas (
<typeStatus id="54D28836FFDCFFE7FDE63C1DFDA3F9FA" box="[513,556,1585,1609]" pageId="7" pageNumber="190">type</typeStatus>
locality).
</paragraph>
<paragraph id="8BD63694FFDCFFE7FF633C72FE5FF89B" blockId="7.[132,775,1630,1877]" pageId="7" pageNumber="190">
<materialsCitation id="3B013CC9FFDCFFE7FF633C72FC8AF975" ID-GBIF-Occurrence="2635845307" pageId="7" pageNumber="190">REFERRED SPECIMENS. — UM-AUM686, right M2; UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2; UM-AUM695, labial part of a M1.</materialsCitation>
HORIZON AND AGE. — Lacustrine limestone of Montpellier, proposed as Lutetian in age in previous study but here considered possibly late Ypresian/early Lutetian in age
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE7FF633D10FDCCF8E6" box="[132,579,1852,1877]" pageId="7" pageNumber="190" type="distribution">
<paragraph id="8BD63694FFDCFFE7FF633D10FDCCF8E6" blockId="7.[132,775,1630,1877]" box="[132,579,1852,1877]" pageId="7" pageNumber="190">
TYPE LOCALITY. — Aumelas, Hérault,
<collectingCountry id="F37E7604FFDCFFE7FE1C3D12FDB0F8E6" box="[507,575,1854,1877]" name="France" pageId="7" pageNumber="190">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE7FF633D46FEE9F85F" pageId="7" pageNumber="190" type="diagnosis">
<paragraph id="8BD63694FFDCFFE7FF633D46FEE9F85F" blockId="7.[132,775,1898,2028]" pageId="7" pageNumber="190">
DIAGNOSIS. — The species is 15% smaller than
<emphasis id="B91DEA86FFDCFFE7FDB23D47FD53F831" box="[597,732,1898,1922]" italics="true" pageId="7" pageNumber="190">O. bicuspidens</emphasis>
and 25% smaller than
<emphasis id="B91DEA86FFDCFFE7FEDB3DA9FE49F82E" box="[316,454,1925,1949]" italics="true" pageId="7" pageNumber="190">O. bicuspidens</emphasis>
and
<emphasis id="B91DEA86FFDCFFE7FE1C3DA9FDE3F82E" box="[507,620,1925,1949]" italics="true" pageId="7" pageNumber="190">O. schlosseri</emphasis>
. It differs from
<emphasis id="B91DEA86FFDCFFE7FF633D8CFF7AF804" box="[132,245,1951,1975]" italics="true" pageId="7" pageNumber="190">O. lindgreni</emphasis>
and
<emphasis id="B91DEA86FFDCFFE7FEC53D8CFE27F804" box="[290,424,1951,1975]" italics="true" pageId="7" pageNumber="190">O. bicuspidens</emphasis>
by a more buccally aligned metastyle on molars. It differs from
<emphasis id="B91DEA86FFDCFFE7FE633D96FD83F861" box="[388,524,1978,2002]" italics="true" pageId="7" pageNumber="190">O. bicuspidens</emphasis>
and
<emphasis id="B91DEA86FFDCFFE7FDD83D96FD20F861" box="[575,687,1978,2002]" italics="true" pageId="7" pageNumber="190">O. schlosseri</emphasis>
by a less retroflexed protoconid.
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE7FCCA3AF5FBC9FA1C" pageId="7" pageNumber="190" type="description">
<paragraph id="8BD63694FFDCFFE7FCCA3AF5FBCBFF42" blockId="7.[813,1092,217,241]" box="[813,1092,217,241]" pageId="7" pageNumber="190">
MEASUREMENTS. —
<tableCitation id="C6EB032FFFDCFFE7FC113AF5FBCBFF42" box="[1014,1092,217,241]" captionStart="TABLE" captionStartId="7.[132,143,219,236]" captionText="TABLE 3. — Measurements (in mm) of Oxyaenoides aumelasiensis n. sp. from Aumelas (MP10-MP11?). *, based on roots and/or alveoli;Body mass (c. 4.24 kg) estimated after Morlo (1999)." pageId="7" pageNumber="190">Table 3.</tableCitation>
</paragraph>
<paragraph id="8BD63694FFDCFFE7FCCA3B34FC32FE83" blockId="7.[813,1456,280,1455]" box="[813,957,280,306]" pageId="7" pageNumber="190">DESCRIPTION</paragraph>
<paragraph id="8BD63694FFDCFFE7FCCA3B15FB12FDE1" blockId="7.[813,1456,280,1455]" pageId="7" pageNumber="190">
The P2 has a very small lingual fold that corresponds to an incipient parastyle. The metastyle is short and low (
<figureCitation id="13522A11FFDCFFE7FAB23B75FCCEFE20" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@8.[132,1455,215,1878]" captionTargetPageId="8" captionText="FIG. 2. — Oxyaenoides aumelasiensis n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM693 (Holotype), left mandible bearing m3, the anterior root and talonid of the m1, and the posterior alveolus of the p4, and the alveoli of m2; A, occlusal view; B, lingual view; C, labial view; D-G, UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2; D, occlusal view of left maxillary; E, labial view of the left maxillary; F, occlusal view of the right maxillary; G, labial view of the right maxillary (reversed view);H-I, UM-AUM686,right M2; H, occlusal view; I, labial view. Scale bars: A-G, 1 cm; H, I, 50 mm." figureDoi="http://doi.org/10.5281/zenodo.3894814" httpUri="https://zenodo.org/record/3894814/files/figure.png" pageId="7" pageNumber="190">Fig. 2D, E</figureCitation>
). There is a small diastema between the P2 and P3 (
<figureCitation id="13522A11FFDCFFE7FA863B55FCCBFE00" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@8.[132,1455,215,1878]" captionTargetPageId="8" captionText="FIG. 2. — Oxyaenoides aumelasiensis n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM693 (Holotype), left mandible bearing m3, the anterior root and talonid of the m1, and the posterior alveolus of the p4, and the alveoli of m2; A, occlusal view; B, lingual view; C, labial view; D-G, UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2; D, occlusal view of left maxillary; E, labial view of the left maxillary; F, occlusal view of the right maxillary; G, labial view of the right maxillary (reversed view);H-I, UM-AUM686,right M2; H, occlusal view; I, labial view. Scale bars: A-G, 1 cm; H, I, 50 mm." figureDoi="http://doi.org/10.5281/zenodo.3894814" httpUri="https://zenodo.org/record/3894814/files/figure.png" pageId="7" pageNumber="190">Fig. 2F, G</figureCitation>
). The P3 and P4 are mesiodistally elongated. The P3 is characterized by the presence of a distinct parastyle and a strong metastyle. There is a large lingual bulge but no protocone. The P4 also displays a very large parastyle and metastyle; the latter is very tall. The protocone projects lingually and is mesiodistally narrow (
<figureCitation id="13522A11FFDCFFE7FBFC3814FBD6FDE1" box="[1051,1113,568,594]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@8.[132,1455,215,1878]" captionTargetPageId="8" captionText="FIG. 2. — Oxyaenoides aumelasiensis n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM693 (Holotype), left mandible bearing m3, the anterior root and talonid of the m1, and the posterior alveolus of the p4, and the alveoli of m2; A, occlusal view; B, lingual view; C, labial view; D-G, UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2; D, occlusal view of left maxillary; E, labial view of the left maxillary; F, occlusal view of the right maxillary; G, labial view of the right maxillary (reversed view);H-I, UM-AUM686,right M2; H, occlusal view; I, labial view. Scale bars: A-G, 1 cm; H, I, 50 mm." figureDoi="http://doi.org/10.5281/zenodo.3894814" httpUri="https://zenodo.org/record/3894814/files/figure.png" pageId="7" pageNumber="190">Fig. 2</figureCitation>
D-G).
</paragraph>
<paragraph id="8BD63694FFDCFFE7FCA43874FC52FC22" blockId="7.[813,1456,280,1455]" pageId="7" pageNumber="190">
The two known molars have a very similar morphology (
<figureCitation id="13522A11FFDCFFE7FCD13854FC79FD21" box="[822,1014,632,658]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@8.[132,1455,215,1878]" captionTargetPageId="8" captionText="FIG. 2. — Oxyaenoides aumelasiensis n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM693 (Holotype), left mandible bearing m3, the anterior root and talonid of the m1, and the posterior alveolus of the p4, and the alveoli of m2; A, occlusal view; B, lingual view; C, labial view; D-G, UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2; D, occlusal view of left maxillary; E, labial view of the left maxillary; F, occlusal view of the right maxillary; G, labial view of the right maxillary (reversed view);H-I, UM-AUM686,right M2; H, occlusal view; I, labial view. Scale bars: A-G, 1 cm; H, I, 50 mm." figureDoi="http://doi.org/10.5281/zenodo.3894814" httpUri="https://zenodo.org/record/3894814/files/figure.png" pageId="7" pageNumber="190">Fig. 2D, E, H, I</figureCitation>
). On M1, the parastyle is present and bulbous but is mesiodistally short and narrow. The metastyle is mesiodistally elongated. The paracone and metacone are partially fused; only the apexes are fully separated. The protocone is low. The protofossa is very mesiodistally short and narrow. The paraconule is present. The metaconule is absent. The M1 is morphologically identical to the M2 except that the parastyle is wider and the metastyle is less distally located.
</paragraph>
<paragraph id="8BD63694FFDCFFE7FCA439BBFC51FBE3" blockId="7.[813,1456,280,1455]" pageId="7" pageNumber="190">
The mandible is narrow. The masseteric fossa is deep. As in
<taxonomicName id="4C694D17FFDCFFE7FC94399BFC62FC62" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[883,1005,951,977]" class="Mammalia" family="Hyaenodontidae" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDCFFE7FC94399BFC62FC62" box="[883,1005,951,977]" italics="true" pageId="7" pageNumber="190">Matthodon</emphasis>
</taxonomicName>
, a deep fossa for insertion of the temporal muscle is present along the anterior margin of the coronoid crest. The latter is distally inclined and the angle between the coronoid crest and the body of the dentary is close to 120°.
</paragraph>
<paragraph id="8BD63694FFDCFFE7FCA43E7AFBC9FA1C" blockId="7.[813,1456,280,1455]" pageId="7" pageNumber="190">
UM/AUM693 is a fragment of a left mandible that bears the talonid of m1 and a complete m3 (
<figureCitation id="13522A11FFDCFFE7FB103E5AFAB4FB23" box="[1271,1339,1142,1168]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@8.[132,1455,215,1878]" captionTargetPageId="8" captionText="FIG. 2. — Oxyaenoides aumelasiensis n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM693 (Holotype), left mandible bearing m3, the anterior root and talonid of the m1, and the posterior alveolus of the p4, and the alveoli of m2; A, occlusal view; B, lingual view; C, labial view; D-G, UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2; D, occlusal view of left maxillary; E, labial view of the left maxillary; F, occlusal view of the right maxillary; G, labial view of the right maxillary (reversed view);H-I, UM-AUM686,right M2; H, occlusal view; I, labial view. Scale bars: A-G, 1 cm; H, I, 50 mm." figureDoi="http://doi.org/10.5281/zenodo.3894814" httpUri="https://zenodo.org/record/3894814/files/figure.png" pageId="7" pageNumber="190">Fig. 2</figureCitation>
A-C). The talonid of the m1 bears three distinct cusps (entoconid, hypoconid, and hypoconulid). The entoconid is lower than the hypoconulid, which is distally located, and lower than the hypoconid. Contrary to the m1, the m3 is complete. The paraconid is tall, long and well mesially located. There is no metaconid. The m3 is thus characterized by a cutting trigonid. The talonid is very mesiodistally short, low, and narrow with only one cusp (hypoconulid?) present. The precingulid is developed.
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDCFFE9FCCA3FF8FD4FF85E" lastPageId="9" lastPageNumber="192" pageId="7" pageNumber="190" type="discussion">
<paragraph id="8BD63694FFDCFFE7FCCA3FF8FC3BFA5F" blockId="7.[813,1456,1492,2030]" box="[813,948,1492,1518]" pageId="7" pageNumber="190">DISCUSSION</paragraph>
<paragraph id="8BD63694FFDCFFE7FCCA3FD9FBF5F83D" blockId="7.[813,1456,1492,2030]" pageId="7" pageNumber="190">
The fossils from Aumelas are characterized by a very cutting morphology (e.g., long metastyle on P4 and molars, absence of metaconid, long paracristid on m3). Only one hyaenodont genus known from the Ypresian and Lutetian of Europe has molars that are characterized by the absence of a metaconid, a hypercarnivorous feature:
<taxonomicName id="4C694D17FFDCFFE7FACE3CB8FA3FF91D" authorityName="Matthes" authorityYear="1967" box="[1321,1456,1684,1710]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDCFFE7FACE3CB8FA3FF91D" box="[1321,1456,1684,1710]" italics="true" pageId="7" pageNumber="190">Oxyaenoides</emphasis>
</taxonomicName>
(MP10-?MP13) (
<bibRefCitation id="EFF84B65FFDCFFE7FC173C98FB03F97D" author="MATTHES H. W." box="[1008,1164,1716,1742]" journalOrPublisher="Geologie Jahrbuch" pageId="7" pageNumber="190" pagination="452 - 456" part="16" refId="ref15146" refString="MATTHES H. W. 1967. - Erstmaliger Nachweis eines Vertreters der Oxyaeninae Trouessart 1885 (Creodonta) in Europa. Geologie Jahrbuch 16: 452 - 456." title="Erstmaliger Nachweis eines Vertreters der Oxyaeninae Trouessart 1885 (Creodonta) in Europa" type="journal article" year="1967">Matthes 1967</bibRefCitation>
; Lange-Badré &amp; Haubold 1990;
<bibRefCitation id="EFF84B65FFDCFFE7FC933CF8FBA7F95D" author="Sole" box="[884,1064,1748,1774]" etAl="et al." firstAuthor="Sole" pageId="7" pageNumber="190" pagination="878 - 917" refId="ref15999" refString="SOLE F., FALCONNET J. &amp; LAURENT Y. 2014 a. - New proviverrines (Hyaenodontida) from the early Eocene of Europe; phylogeny and ecological evolution of the Proviverrinae. Zoological Journal of the Linnean Society 171: 878 - 917. https: // doi. org / 10.1111 / zoj. 12155" type="journal article" year="2014" yearSuffix="a">
Solé
<emphasis id="B91DEA86FFDCFFE7FC4E3CF9FC52F95D" box="[937,989,1748,1774]" italics="true" pageId="7" pageNumber="190">et al.</emphasis>
2014a
</bibRefCitation>
). The teeth found at Aumelas share with
<taxonomicName id="4C694D17FFDCFFE7FC813CD8FC63F8BD" authorityName="Matthes" authorityYear="1967" box="[870,1004,1780,1806]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDCFFE7FC813CD8FC63F8BD" box="[870,1004,1780,1806]" italics="true" pageId="7" pageNumber="190">Oxyaenoides</emphasis>
</taxonomicName>
mesiodistally short and narrow talonids bearing crestiform entoconids on molars. This genus is known both in northwestern (
<bibRefCitation id="EFF84B65FFDCFFE7FB8C3D1FFA8FF8FE" author="MATTHES H. W." box="[1131,1280,1843,1869]" journalOrPublisher="Geologie Jahrbuch" pageId="7" pageNumber="190" pagination="452 - 456" part="16" refId="ref15146" refString="MATTHES H. W. 1967. - Erstmaliger Nachweis eines Vertreters der Oxyaeninae Trouessart 1885 (Creodonta) in Europa. Geologie Jahrbuch 16: 452 - 456." title="Erstmaliger Nachweis eines Vertreters der Oxyaeninae Trouessart 1885 (Creodonta) in Europa" type="journal article" year="1967">Matthes 1967</bibRefCitation>
; Lange-Badré &amp; Haubold 1990;
<bibRefCitation id="EFF84B65FFDCFFE7FC3B3D7FFB01F8DD" author="Sole" box="[988,1166,1875,1902]" etAl="et al." firstAuthor="Sole" pageId="7" pageNumber="190" pagination="878 - 917" refId="ref15999" refString="SOLE F., FALCONNET J. &amp; LAURENT Y. 2014 a. - New proviverrines (Hyaenodontida) from the early Eocene of Europe; phylogeny and ecological evolution of the Proviverrinae. Zoological Journal of the Linnean Society 171: 878 - 917. https: // doi. org / 10.1111 / zoj. 12155" type="journal article" year="2014" yearSuffix="a">
Solé
<emphasis id="B91DEA86FFDCFFE7FBF73D78FBCCF8DE" box="[1040,1091,1875,1901]" italics="true" pageId="7" pageNumber="190">et al.</emphasis>
2014a
</bibRefCitation>
,
<bibRefCitation id="EFF84B65FFDCFFE7FB7D3D7FFB50F8DD" author="Sole" box="[1178,1247,1875,1902]" firstAuthor="Sole" pageId="7" pageNumber="190" pagination="1049 - 1072" refId="ref16133" refString="SOLE F., FALCONNET J. &amp; VIDALENC D. 2015 a. - New fossil Hyaenodonta (Mammalia, Placentalia) from the Ypresian and Lutetian of France and the evolution of the Proviverrinae in southern Europe. Palaeontology 58: 1049 - 1072. https: // doi. org / 10.1111 / pala. 12198" type="journal article" year="2015" yearSuffix="a">2015a</bibRefCitation>
) and southwestern Europe (
<bibRefCitation id="EFF84B65FFDCFFE7FC683D5FFBE6F83E" author="CROCHET B. &amp; CROUZEL F. &amp; LANGE- BADRE B." box="[911,1129,1907,1933]" journalOrPublisher="Comptes rendus hebdomadaires de l'Academie des sciences, S e rie D" pageId="7" pageNumber="190" pagination="1597 - 1600" part="282" publicationUrl="https://gallica.bnf.fr/ark:/12148/bpt6k54844424/f27.item" refId="ref13167" refString="CROCHET B., CROUZEL F. &amp; LANGE- BADRE B. 1976. - Consequences de la decouverte du genre Oxyaenoides Matthes, sur la datation du Poudingue de Palassou. Comptes rendus hebdomadaires de l'Academie des sciences, S e rie D 282: 1597 - 1600. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 54844424 / f 27. item" title="Consequences de la decouverte du genre Oxyaenoides Matthes, sur la datation du Poudingue de Palassou" type="journal article" year="1976">
Crochet
<emphasis id="B91DEA86FFDCFFE7FC153D58FBA7F83E" box="[1010,1064,1907,1933]" italics="true" pageId="7" pageNumber="190">et al.</emphasis>
1976
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD63694FFDCFFE9FCA43DBFFE95FB03" blockId="7.[813,1456,1492,2030]" lastBlockId="9.[132,776,951,2030]" lastPageId="9" lastPageNumber="192" pageId="7" pageNumber="190">
Three species of
<taxonomicName id="4C694D17FFDCFFE7FC1D3DBFFB0DF81E" authorityName="Matthes" authorityYear="1967" box="[1018,1154,1939,1965]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDCFFE7FC1D3DBFFB0DF81E" box="[1018,1154,1939,1965]" italics="true" pageId="7" pageNumber="190">Oxyaenoides</emphasis>
</taxonomicName>
are presently known:
<emphasis id="B91DEA86FFDCFFE7FA9E3DB8FC72F87E" italics="true" pageId="7" pageNumber="190">
<taxonomicName id="4C694D17FFDCFFE7FA9E3DB8FC03F87E" authorityName="Matthes" authorityYear="1967" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">Oxyaenoides</taxonomicName>
lindgreni
</emphasis>
from MP10 (
<bibRefCitation id="EFF84B65FFDCFFE7FB4E3D9FFAA8F87E" author="Rich" box="[1193,1319,1971,1997]" firstAuthor="Rich" pageId="7" pageNumber="190" pagination="1 - 72" refId="ref15539" refString="RICH T. H. 1971. - Deltatheridia, Carnivora, and Condylarthra (Mammalia) of the early Eocene, Paris Basin, France. University of California Publications in Geological Sciences 88: 1 - 72." type="journal article" year="1971">Rich 1971</bibRefCitation>
;
<bibRefCitation id="EFF84B65FFDCFFE7FADE3D9FFCF9F85E" author="Sole" etAl="et al." firstAuthor="Sole" pageId="7" pageNumber="190" pagination="878 - 917" refId="ref15999" refString="SOLE F., FALCONNET J. &amp; LAURENT Y. 2014 a. - New proviverrines (Hyaenodontida) from the early Eocene of Europe; phylogeny and ecological evolution of the Proviverrinae. Zoological Journal of the Linnean Society 171: 878 - 917. https: // doi. org / 10.1111 / zoj. 12155" type="journal article" year="2014" yearSuffix="a">
Solé
<emphasis id="B91DEA86FFDCFFE7FA923D98FA3FF87E" box="[1397,1456,1971,1997]" italics="true" pageId="7" pageNumber="190">et al.</emphasis>
2014a
</bibRefCitation>
),
<emphasis id="B91DEA86FFDCFFE7FC763DFFFB29F85E" box="[913,1190,2003,2029]" italics="true" pageId="7" pageNumber="190">
<taxonomicName id="4C694D17FFDCFFE7FC763DFFFB92F85E" authorityName="Matthes" authorityYear="1967" box="[913,1053,2003,2029]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">Oxyaenoides</taxonomicName>
bicuspidens
</emphasis>
from MP11 (
<bibRefCitation id="EFF84B65FFDCFFE9FAB73DFFFF31FC62" author="MATTHES H. W." journalOrPublisher="Geologie Jahrbuch" lastPageId="9" lastPageNumber="192" pageId="7" pageNumber="190" pagination="452 - 456" part="16" refId="ref15146" refString="MATTHES H. W. 1967. - Erstmaliger Nachweis eines Vertreters der Oxyaeninae Trouessart 1885 (Creodonta) in Europa. Geologie Jahrbuch 16: 452 - 456." title="Erstmaliger Nachweis eines Vertreters der Oxyaeninae Trouessart 1885 (Creodonta) in Europa" type="journal article" year="1967">Matthes 1967</bibRefCitation>
;
<bibRefCitation id="EFF84B65FFD2FFE9FF2C399BFE11FC62" author="CROCHET B. &amp; CROUZEL F. &amp; LANGE- BADRE B." box="[203,414,951,977]" journalOrPublisher="Comptes rendus hebdomadaires de l'Academie des sciences, S e rie D" pageId="9" pageNumber="192" pagination="1597 - 1600" part="282" publicationUrl="https://gallica.bnf.fr/ark:/12148/bpt6k54844424/f27.item" refId="ref13167" refString="CROCHET B., CROUZEL F. &amp; LANGE- BADRE B. 1976. - Consequences de la decouverte du genre Oxyaenoides Matthes, sur la datation du Poudingue de Palassou. Comptes rendus hebdomadaires de l'Academie des sciences, S e rie D 282: 1597 - 1600. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 54844424 / f 27. item" title="Consequences de la decouverte du genre Oxyaenoides Matthes, sur la datation du Poudingue de Palassou" type="journal article" year="1976">
Crochet
<emphasis id="B91DEA86FFD2FFE9FECC399BFED0FC62" box="[299,351,951,977]" italics="true" pageId="9" pageNumber="192">et al.</emphasis>
1976
</bibRefCitation>
; Lange-Badré &amp; Haubold 1990) and
<emphasis id="B91DEA86FFD2FFE9FF5239FAFE2DFC43" box="[181,418,982,1008]" italics="true" pageId="9" pageNumber="192">
<taxonomicName id="4C694D17FFD2FFE9FF5239FAFEB2FC43" authorityName="Matthes" authorityYear="1967" box="[181,317,982,1008]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">Oxyaenoides</taxonomicName>
schlosseri
</emphasis>
from?MP13 (
<bibRefCitation id="EFF84B65FFD2FFE9FDA039FAFC8FFC42" author="Rutimeyer" box="[583,768,982,1009]" firstAuthor="Rutimeyer" pageId="9" pageNumber="192" pagination="1 - 153" refId="ref15704" refString="RUTIMEYER L. 1891. - Die Eocane Saugethier-Welt von Egerkingen. Abhandlungen der Schweizerischen Palaeontologischen Gesellschaft 18: 1 - 153. https: // doi. org / 10.5962 / bhl. title. 158813" type="journal article" year="1891">Rütimeyer 1891</bibRefCitation>
; Lange-Badré 1972;
<bibRefCitation id="EFF84B65FFD2FFE9FE8939DAFDBBFBA2" author="Sole" box="[366,564,1014,1041]" etAl="et al." firstAuthor="Sole" pageId="9" pageNumber="192" pagination="1049 - 1072" refId="ref16133" refString="SOLE F., FALCONNET J. &amp; VIDALENC D. 2015 a. - New fossil Hyaenodonta (Mammalia, Placentalia) from the Ypresian and Lutetian of France and the evolution of the Proviverrinae in southern Europe. Palaeontology 58: 1049 - 1072. https: // doi. org / 10.1111 / pala. 12198" type="journal article" year="2015" yearSuffix="a">
Solé
<emphasis id="B91DEA86FFD2FFE9FE4F39DBFE6DFBA3" box="[424,482,1014,1040]" italics="true" pageId="9" pageNumber="192">et al.</emphasis>
2015a
</bibRefCitation>
). The
<typeStatus id="54D28836FFD2FFE9FD6339DBFD69FBA2" box="[644,742,1015,1041]" pageId="9" pageNumber="192" type="syntype">syntypes</typeStatus>
of
<emphasis id="B91DEA86FFD2FFE9FF633E3BFE86FB83" box="[132,265,1046,1073]" italics="true" pageId="9" pageNumber="192">O. schlosseri</emphasis>
are from Egerkingen
<emphasis id="B91DEA86FFD2FFE9FDE23E3AFD9EFB82" bold="true" box="[517,529,1046,1073]" pageId="9" pageNumber="192">γ</emphasis>
(Lange-Badré 1972), but the MP reference-level of this site (possibly MP13) is uncertain (
<bibRefCitation id="EFF84B65FFD2FFE9FEC73E7AFD87FBC3" author="BIOCHROM' 97 (collective)" box="[288,520,1110,1136]" editor="AGUILAR J. - P. &amp; LEGENDRE S. &amp; MICHAUX J." journalOrPublisher="Memoires et Travaux de l'EPHE, Institut de Montpellier" pageId="9" pageNumber="192" pagination="769 - 805" refId="ref12622" refString="BIOCHROM' 97 (collective) 1997. - Syntheses et tableaux de correlations, in AGUILAR J. - P., LEGENDRE S. &amp; MICHAUX J. (eds), Actes du Congres BiochroM' 97. Memoires et Travaux de l'EPHE, Institut de Montpellier: 769 - 805." title="Syntheses et tableaux de correlations" type="book chapter" volumeTitle="Actes du Congres BiochroM' 97" year="1997">BiochroM97 1997</bibRefCitation>
), as is Aigues-Vives 2 where a mandible of
<emphasis id="B91DEA86FFD2FFE9FE653E5AFD83FB23" box="[386,524,1142,1168]" italics="true" pageId="9" pageNumber="192">O. schlosseri</emphasis>
was discovered (
<bibRefCitation id="EFF84B65FFD2FFE9FD303E5AFE86FB02" author="Sole" etAl="et al." firstAuthor="Sole" pageId="9" pageNumber="192" pagination="1049 - 1072" refId="ref16133" refString="SOLE F., FALCONNET J. &amp; VIDALENC D. 2015 a. - New fossil Hyaenodonta (Mammalia, Placentalia) from the Ypresian and Lutetian of France and the evolution of the Proviverrinae in southern Europe. Palaeontology 58: 1049 - 1072. https: // doi. org / 10.1111 / pala. 12198" type="journal article" year="2015" yearSuffix="a">
Solé
<emphasis id="B91DEA86FFD2FFE9FF633EBBFF34FB03" box="[132,187,1174,1200]" italics="true" pageId="9" pageNumber="192">et al.</emphasis>
2015a
</bibRefCitation>
).
</paragraph>
<caption id="DF16661CFFD3FFE8FF633D51FD3EF85A" ID-DOI="http://doi.org/10.5281/zenodo.3894814" ID-Zenodo-Dep="3894814" httpUri="https://zenodo.org/record/3894814/files/figure.png" pageId="8" pageNumber="191" startId="8.[132,143,1917,1934]" targetBox="[132,1455,215,1878]" targetPageId="8">
<paragraph id="8BD63694FFD3FFE8FF633D51FD3EF85A" blockId="8.[132,1456,1917,2025]" pageId="8" pageNumber="191">
<subScript id="17ED34D1FFD3FFE8FF633D51FF12F83D" attach="left" box="[132,157,1917,1934]" fontSize="5" pageId="8" pageNumber="191">FIG</subScript>
. 2. —
<taxonomicName id="4C694D17FFD3FFE8FF333D51FE38F83D" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[212,439,1917,1934]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="8" pageNumber="191" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFD3FFE8FF333D51FE38F83D" box="[212,439,1917,1934]" italics="true" pageId="8" pageNumber="191">Oxyaenoides aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD3FFE8FE5A3D51FE62F83D" box="[445,493,1917,1934]" pageId="8" pageNumber="191" rank="species">n. sp.</taxonomicNameLabel>
from Aumelas (MP10-MP11?):
<emphasis id="B91DEA86FFD3FFE8FD173D51FC9DF83D" bold="true" box="[752,786,1917,1934]" pageId="8" pageNumber="191">A -C</emphasis>
, UM-AUM693 (Holotype), left mandible bearing m3, the anterior root and talonid of the m1, and the posterior alveolus of the p4, and the alveoli of m2;
<emphasis id="B91DEA86FFD3FFE8FD2A3DB8FD55F816" bold="true" box="[717,730,1940,1957]" pageId="8" pageNumber="191">A</emphasis>
, occlusal view;
<emphasis id="B91DEA86FFD3FFE8FCB83DB8FCE3F816" bold="true" box="[863,876,1940,1957]" pageId="8" pageNumber="191">B</emphasis>
, lingual view;
<emphasis id="B91DEA86FFD3FFE8FC043DB8FC7EF816" bold="true" box="[995,1009,1940,1957]" pageId="8" pageNumber="191">C</emphasis>
, labial view;
<emphasis id="B91DEA86FFD3FFE8FBBA3DB8FB0FF816" bold="true" box="[1117,1152,1940,1957]" pageId="8" pageNumber="191">D -G</emphasis>
, UM-AUM692, combination of a left maxillary fragment bearing the P2-M1, a right maxillary fragment bearing the P3-P4, the protocone of the M1 and the alveoli of the P2;
<emphasis id="B91DEA86FFD3FFE8FB163D86FB70F808" bold="true" box="[1265,1279,1962,1979]" pageId="8" pageNumber="191">D</emphasis>
, occlusal view of left maxillary;
<emphasis id="B91DEA86FFD3FFE8FF3F3DEDFF6BF861" bold="true" box="[216,228,1985,2002]" pageId="8" pageNumber="191">E</emphasis>
, labial view of the left maxillary;
<emphasis id="B91DEA86FFD3FFE8FE0A3DEDFE77F861" bold="true" box="[493,504,1985,2002]" pageId="8" pageNumber="191">F</emphasis>
, occlusal view of the right maxillary;
<emphasis id="B91DEA86FFD3FFE8FCC13DEDFCBBF861" bold="true" box="[806,820,1985,2002]" pageId="8" pageNumber="191">G</emphasis>
, labial view of the right maxillary (reversed view);
<emphasis id="B91DEA86FFD3FFE8FB2E3DEDFB6AF861" bold="true" box="[1225,1253,1985,2002]" pageId="8" pageNumber="191">H -I</emphasis>
, UM-AUM686,right M2;
<emphasis id="B91DEA86FFD3FFE8FF633DF4FF1DF85A" bold="true" box="[132,146,2008,2025]" pageId="8" pageNumber="191">H</emphasis>
, occlusal view;
<emphasis id="B91DEA86FFD3FFE8FEF13DF4FE93F85A" bold="true" box="[278,284,2008,2025]" pageId="8" pageNumber="191">I</emphasis>
, labial view. Scale bars: A-G, 1 cm; H, I, 50 mm.
</paragraph>
</caption>
<caption id="DF16661CFFD2FFE9FF633AF7FDAEFEA9" pageId="9" pageNumber="192">
<paragraph id="8BD63694FFD2FFE9FF633AF7FDAEFEA9" blockId="9.[132,775,219,282]" pageId="9" pageNumber="192">
<subScript id="17ED34D1FFD2FFE9FF633AF7FF3DFF5F" attach="left" box="[132,178,219,236]" fontSize="5" pageId="9" pageNumber="192">TABLE</subScript>
4. — Measurements (in mm) of
<taxonomicName id="4C694D17FFD2FFE9FE513AF7FD06FF5F" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[438,649,219,236]" class="Mammalia" family="Hyaenodontidae" genus="Leonhardtina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="meridianum" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FE513AF7FD06FF5F" box="[438,649,219,236]" italics="true" pageId="9" pageNumber="192">Leonhardtina meridianum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FD6A3AF7FD33FF5F" box="[653,700,219,236]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
from Aumelas and Rouzilhac (MP10-MP11?).Body mass (
<emphasis id="B91DEA86FFD2FFE9FDFC3ADEFDA4FEB0" box="[539,555,242,259]" italics="true" pageId="9" pageNumber="192">c.</emphasis>
843.29 g) of
<taxonomicName id="4C694D17FFD2FFE9FD753ADEFC88FEB0" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[658,775,242,259]" class="Mammalia" family="Hyaenodontidae" genus="Leonhardtina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="meridianum" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FD753ADEFC88FEB0" box="[658,775,242,259]" italics="true" pageId="9" pageNumber="192">L. meridianum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FF633B25FF3BFEA9" box="[132,180,265,282]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
from Aumelas estimated after
<bibRefCitation id="EFF84B65FFD2FFE9FE543B25FD92FEA9" author="MORLO M." box="[435,541,265,282]" journalOrPublisher="Geobios" pageId="9" pageNumber="192" pagination="297 - 305" part="32" refId="ref15217" refString="MORLO M. 1999. - Niche structure and evolution in creodont (Mammalia) faunas of the European and North American Eocene. Geobios 32: 297 - 305. https: // doi. org / 10.1016 / S 0016 - 6995 (99) 80043 - 6" title="Niche structure and evolution in creodont (Mammalia) faunas of the European and North American Eocene" type="journal article" year="1999">Morlo (1999)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BD63694FFD2FFE9FE973B7FFD52FCEC" pageId="9" pageNumber="192">
<table id="F969C434FFD2001FFF613B7FFD52FCEC" box="[134,733,339,863]" gridcols="6" gridrows="20" pageId="9" pageNumber="192">
<tr id="355934D6FFD2001FFF613B7FFD52FED4" box="[134,733,339,359]" gridrow="0" pageId="9" pageNumber="192" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<th id="76885DAAFFD2001FFE973B7FFD52FED4" box="[368,733,339,359]" colspan="3" colspanRight="2" gridcol="3" gridrow="0" pageId="9" pageNumber="192">
<taxonomicName id="4C694D17FFD2FFE9FE973B7FFDECFED4" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[368,611,339,359]" class="Mammalia" family="Hyaenodontidae" genus="Leonhardtina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="meridianum" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FE973B7FFDECFED4" box="[368,611,339,359]" italics="true" pageId="9" pageNumber="192">Leonhardtina meridianum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FD8E3B7FFD10FED4" box="[617,671,339,359]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr id="355934D6FFD2001FFF613B5BFD52FE38" box="[134,733,375,395]" gridrow="1" pageId="9" pageNumber="192" rowspan-0="1" rowspan-1="1">
<td id="76885DAAFFD2001FFEA73B5BFE51FE38" box="[320,478,375,395]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="9" pageNumber="192">from Aumelas</td>
<td id="76885DAAFFD2001FFDA73B5BFD52FE38" box="[576,733,375,395]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="9" pageNumber="192">from Rouzilhac</td>
</tr>
<tr id="355934D6FFD2001FFF613BB7FD52FE1C" box="[134,733,411,431]" gridrow="2" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613BB7FF65FE1C" box="[134,234,411,431]" colspan="2" colspanRight="1" gridcol="0" gridrow="2" pageId="9" pageNumber="192">
<emphasis id="B91DEA86FFD2FFE9FF613BB7FF4BFE1C" bold="true" box="[134,196,411,431]" pageId="9" pageNumber="192">Locus</emphasis>
</th>
<td id="76885DAAFFD2001FFEA73BB7FEC2FE1C" box="[320,333,411,431]" gridcol="2" gridrow="2" pageId="9" pageNumber="192">
<emphasis id="B91DEA86FFD2FFE9FEA73BB7FEC2FE1C" bold="true" box="[320,333,411,431]" pageId="9" pageNumber="192">n</emphasis>
</td>
<td id="76885DAAFFD2001FFE973BB7FE51FE1C" box="[368,478,411,431]" gridcol="3" gridrow="2" pageId="9" pageNumber="192">
<emphasis id="B91DEA86FFD2FFE9FE503BB7FE58FE1C" bold="true" box="[439,471,411,431]" pageId="9" pageNumber="192">OR</emphasis>
</td>
<td id="76885DAAFFD2001FFDA73BB7FDC1FE1C" box="[576,590,411,431]" gridcol="4" gridrow="2" pageId="9" pageNumber="192">
<emphasis id="B91DEA86FFD2FFE9FDA63BB7FDC1FE1C" bold="true" box="[577,590,411,431]" pageId="9" pageNumber="192">n</emphasis>
</td>
<td id="76885DAAFFD2001FFD4F3BB7FD52FE1C" box="[680,733,411,431]" gridcol="5" gridrow="2" pageId="9" pageNumber="192">
<emphasis id="B91DEA86FFD2FFE9FD503BB7FD58FE1C" bold="true" box="[695,727,411,431]" pageId="9" pageNumber="192">OR</emphasis>
</td>
</tr>
<tr id="355934D6FFD2001FFF613B92FD52FE61" box="[134,733,446,466]" gridrow="3" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613B92FF27FE61" box="[134,168,446,466]" gridcol="0" gridrow="3" pageId="9" pageNumber="192">M2</th>
<td id="76885DAAFFD2001FFF313B92FF65FE61" box="[214,234,446,466]" gridcol="1" gridrow="3" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73B92FEC2FE61" box="[320,333,446,466]" gridcol="2" gridrow="3" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973B92FE51FE61" box="[368,478,446,466]" gridcol="3" gridrow="3" pageId="9" pageNumber="192">4.96</td>
<td id="76885DAAFFD2001FFDA73B92FDC1FE61" box="[576,590,446,466]" gridcol="4" gridrow="3" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F3B92FD52FE61" box="[680,733,446,466]" gridcol="5" gridrow="3" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF613BFAFD52FE59" box="[134,733,470,490]" gridrow="4" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF313BFAFF65FE59" box="[214,234,470,490]" gridcol="1" gridrow="4" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA73BFAFEC2FE59" box="[320,333,470,490]" gridcol="2" gridrow="4" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973BFAFE51FE59" box="[368,478,470,490]" gridcol="3" gridrow="4" pageId="9" pageNumber="192">6.07</td>
<td id="76885DAAFFD2001FFDA73BFAFDC1FE59" box="[576,590,470,490]" gridcol="4" gridrow="4" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F3BFAFD52FE59" box="[680,733,470,490]" gridcol="5" gridrow="4" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF613BC2FD52FDB1" box="[134,733,494,514]" gridrow="5" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613BC2FF27FDB1" box="[134,168,494,514]" gridcol="0" gridrow="5" pageId="9" pageNumber="192">M3</th>
<td id="76885DAAFFD2001FFF313BC2FF65FDB1" box="[214,234,494,514]" gridcol="1" gridrow="5" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73BC2FEC2FDB1" box="[320,333,494,514]" gridcol="2" gridrow="5" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973BC2FE51FDB1" box="[368,478,494,514]" gridcol="3" gridrow="5" pageId="9" pageNumber="192">2.91</td>
<td id="76885DAAFFD2001FFDA73BC2FDC1FDB1" box="[576,590,494,514]" gridcol="4" gridrow="5" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F3BC2FD52FDB1" box="[680,733,494,514]" gridcol="5" gridrow="5" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF61382AFD52FDA9" box="[134,733,518,538]" gridrow="6" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF31382AFF65FDA9" box="[214,234,518,538]" gridcol="1" gridrow="6" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA7382AFEC2FDA9" box="[320,333,518,538]" gridcol="2" gridrow="6" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE97382AFE51FDA9" box="[368,478,518,538]" gridcol="3" gridrow="6" pageId="9" pageNumber="192">5.49</td>
<td id="76885DAAFFD2001FFDA7382AFDC1FDA9" box="[576,590,518,538]" gridcol="4" gridrow="6" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F382AFD52FDA9" box="[680,733,518,538]" gridcol="5" gridrow="6" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF613833FD52FD80" box="[134,733,543,563]" gridrow="7" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613833FF27FD80" box="[134,168,543,563]" gridcol="0" gridrow="7" pageId="9" pageNumber="192">p2</th>
<td id="76885DAAFFD2001FFF313833FF65FD80" box="[214,234,543,563]" gridcol="1" gridrow="7" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73833FEC2FD80" box="[320,333,543,563]" gridcol="2" gridrow="7" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFE973833FE51FD80" box="[368,478,543,563]" gridcol="3" gridrow="7" pageId="9" pageNumber="192"></td>
<td id="76885DAAFFD2001FFDA73833FDC1FD80" box="[576,590,543,563]" gridcol="4" gridrow="7" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F3833FD52FD80" box="[680,733,543,563]" gridcol="5" gridrow="7" pageId="9" pageNumber="192">6</td>
</tr>
<tr id="355934D6FFD2001FFF61381BFD52FDF8" box="[134,733,567,587]" gridrow="8" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF31381BFF65FDF8" box="[214,234,567,587]" gridcol="1" gridrow="8" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA7381BFEC2FDF8" box="[320,333,567,587]" gridcol="2" gridrow="8" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFE97381BFE51FDF8" box="[368,478,567,587]" gridcol="3" gridrow="8" pageId="9" pageNumber="192"></td>
<td id="76885DAAFFD2001FFDA7381BFDC1FDF8" box="[576,590,567,587]" gridcol="4" gridrow="8" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F381BFD52FDF8" box="[680,733,567,587]" gridcol="5" gridrow="8" pageId="9" pageNumber="192">2.7</td>
</tr>
<tr id="355934D6FFD2001FFF613863FD52FDD0" box="[134,733,591,611]" gridrow="9" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613863FF27FDD0" box="[134,168,591,611]" gridcol="0" gridrow="9" pageId="9" pageNumber="192">p3</th>
<td id="76885DAAFFD2001FFF313863FF65FDD0" box="[214,234,591,611]" gridcol="1" gridrow="9" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73863FEC2FDD0" box="[320,333,591,611]" gridcol="2" gridrow="9" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973863FE51FDD0" box="[368,478,591,611]" gridcol="3" gridrow="9" pageId="9" pageNumber="192">6.5</td>
<td id="76885DAAFFD2001FFDA73863FDC1FDD0" box="[576,590,591,611]" gridcol="4" gridrow="9" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F3863FD52FDD0" box="[680,733,591,611]" gridcol="5" gridrow="9" pageId="9" pageNumber="192">7</td>
</tr>
<tr id="355934D6FFD2001FFF61384BFD52FDC8" box="[134,733,615,635]" gridrow="10" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF31384BFF65FDC8" box="[214,234,615,635]" gridcol="1" gridrow="10" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA7384BFEC2FDC8" box="[320,333,615,635]" gridcol="2" gridrow="10" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE97384BFE51FDC8" box="[368,478,615,635]" gridcol="3" gridrow="10" pageId="9" pageNumber="192">3.19</td>
<td id="76885DAAFFD2001FFDA7384BFDC1FDC8" box="[576,590,615,635]" gridcol="4" gridrow="10" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F384BFD52FDC8" box="[680,733,615,635]" gridcol="5" gridrow="10" pageId="9" pageNumber="192">3.2</td>
</tr>
<tr id="355934D6FFD2001FFF613853FD52FD20" box="[134,733,639,659]" gridrow="11" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613853FF27FD20" box="[134,168,639,659]" gridcol="0" gridrow="11" pageId="9" pageNumber="192">p4</th>
<td id="76885DAAFFD2001FFF313853FF65FD20" box="[214,234,639,659]" gridcol="1" gridrow="11" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73853FEC2FD20" box="[320,333,639,659]" gridcol="2" gridrow="11" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973853FE51FD20" box="[368,478,639,659]" gridcol="3" gridrow="11" pageId="9" pageNumber="192">5.39</td>
<td id="76885DAAFFD2001FFDA73853FDC1FD20" box="[576,590,639,659]" gridcol="4" gridrow="11" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F3853FD52FD20" box="[680,733,639,659]" gridcol="5" gridrow="11" pageId="9" pageNumber="192">5.4</td>
</tr>
<tr id="355934D6FFD2001FFF6138BBFD52FD18" box="[134,733,663,683]" gridrow="12" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF3138BBFF65FD18" box="[214,234,663,683]" gridcol="1" gridrow="12" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA738BBFEC2FD18" box="[320,333,663,683]" gridcol="2" gridrow="12" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE9738BBFE51FD18" box="[368,478,663,683]" gridcol="3" gridrow="12" pageId="9" pageNumber="192">2.51</td>
<td id="76885DAAFFD2001FFDA738BBFDC1FD18" box="[576,590,663,683]" gridcol="4" gridrow="12" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F38BBFD52FD18" box="[680,733,663,683]" gridcol="5" gridrow="12" pageId="9" pageNumber="192">2.7</td>
</tr>
<tr id="355934D6FFD2001FFF613883FD52FD70" box="[134,733,687,707]" gridrow="13" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613883FF27FD70" box="[134,168,687,707]" gridcol="0" gridrow="13" pageId="9" pageNumber="192">m1</th>
<td id="76885DAAFFD2001FFF313883FF65FD70" box="[214,234,687,707]" gridcol="1" gridrow="13" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73883FEC2FD70" box="[320,333,687,707]" gridcol="2" gridrow="13" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973883FE51FD70" box="[368,478,687,707]" gridcol="3" gridrow="13" pageId="9" pageNumber="192">4.95</td>
<td id="76885DAAFFD2001FFDA73883FDC1FD70" box="[576,590,687,707]" gridcol="4" gridrow="13" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F3883FD52FD70" box="[680,733,687,707]" gridcol="5" gridrow="13" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF6138EBFD52FD68" box="[134,733,711,731]" gridrow="14" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF3138EBFF65FD68" box="[214,234,711,731]" gridcol="1" gridrow="14" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA738EBFEC2FD68" box="[320,333,711,731]" gridcol="2" gridrow="14" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE9738EBFE51FD68" box="[368,478,711,731]" gridcol="3" gridrow="14" pageId="9" pageNumber="192">2.94</td>
<td id="76885DAAFFD2001FFDA738EBFDC1FD68" box="[576,590,711,731]" gridcol="4" gridrow="14" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F38EBFD52FD68" box="[680,733,711,731]" gridcol="5" gridrow="14" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF6138F3FD52FD40" box="[134,733,735,755]" gridrow="15" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF6138F3FF27FD40" box="[134,168,735,755]" gridcol="0" gridrow="15" pageId="9" pageNumber="192">m2</th>
<td id="76885DAAFFD2001FFF3138F3FF65FD40" box="[214,234,735,755]" gridcol="1" gridrow="15" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA738F3FEC2FD40" box="[320,333,735,755]" gridcol="2" gridrow="15" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE9738F3FE51FD40" box="[368,478,735,755]" gridcol="3" gridrow="15" pageId="9" pageNumber="192">5.49</td>
<td id="76885DAAFFD2001FFDA738F3FDC1FD40" box="[576,590,735,755]" gridcol="4" gridrow="15" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F38F3FD52FD40" box="[680,733,735,755]" gridcol="5" gridrow="15" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF6138DBFD52FCB8" box="[134,733,759,779]" gridrow="16" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF3138DBFF65FCB8" box="[214,234,759,779]" gridcol="1" gridrow="16" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA738DBFEC2FCB8" box="[320,333,759,779]" gridcol="2" gridrow="16" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE9738DBFE51FCB8" box="[368,478,759,779]" gridcol="3" gridrow="16" pageId="9" pageNumber="192">3.21</td>
<td id="76885DAAFFD2001FFDA738DBFDC1FCB8" box="[576,590,759,779]" gridcol="4" gridrow="16" pageId="9" pageNumber="192">0</td>
<td id="76885DAAFFD2001FFD4F38DBFD52FCB8" box="[680,733,759,779]" gridcol="5" gridrow="16" pageId="9" pageNumber="192"></td>
</tr>
<tr id="355934D6FFD2001FFF613923FD52FC90" box="[134,733,783,803]" gridrow="17" pageId="9" pageNumber="192">
<th id="76885DAAFFD2001FFF613923FF27FC90" box="[134,168,783,803]" gridcol="0" gridrow="17" pageId="9" pageNumber="192">m3</th>
<td id="76885DAAFFD2001FFF313923FF65FC90" box="[214,234,783,803]" gridcol="1" gridrow="17" pageId="9" pageNumber="192">L</td>
<td id="76885DAAFFD2001FFEA73923FEC2FC90" box="[320,333,783,803]" gridcol="2" gridrow="17" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973923FE51FC90" box="[368,478,783,803]" gridcol="3" gridrow="17" pageId="9" pageNumber="192">5.78</td>
<td id="76885DAAFFD2001FFDA73923FDC1FC90" box="[576,590,783,803]" gridcol="4" gridrow="17" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F3923FD52FC90" box="[680,733,783,803]" gridcol="5" gridrow="17" pageId="9" pageNumber="192">5.9</td>
</tr>
<tr id="355934D6FFD2001FFF613904FD52FC8F" box="[134,733,808,828]" gridrow="18" pageId="9" pageNumber="192" rowspan-0="1">
<td id="76885DAAFFD2001FFF313904FF65FC8F" box="[214,234,808,828]" gridcol="1" gridrow="18" pageId="9" pageNumber="192">W</td>
<td id="76885DAAFFD2001FFEA73904FEC2FC8F" box="[320,333,808,828]" gridcol="2" gridrow="18" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973904FE51FC8F" box="[368,478,808,828]" gridcol="3" gridrow="18" pageId="9" pageNumber="192">3.67</td>
<td id="76885DAAFFD2001FFDA73904FDC1FC8F" box="[576,590,808,828]" gridcol="4" gridrow="18" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F3904FD52FC8F" box="[680,733,808,828]" gridcol="5" gridrow="18" pageId="9" pageNumber="192">4</td>
</tr>
<tr id="355934D6FFD2001FFF613967FD52FCEC" box="[134,733,843,863]" gridrow="19" pageId="9" pageNumber="192" rowspan-1="1">
<th id="76885DAAFFD2001FFF613967FF27FCEC" box="[134,168,843,863]" gridcol="0" gridrow="19" pageId="9" pageNumber="192">MD</th>
<td id="76885DAAFFD2001FFEA73967FEC2FCEC" box="[320,333,843,863]" gridcol="2" gridrow="19" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFE973967FE51FCEC" box="[368,478,843,863]" gridcol="3" gridrow="19" pageId="9" pageNumber="192">10.48</td>
<td id="76885DAAFFD2001FFDA73967FDC1FCEC" box="[576,590,843,863]" gridcol="4" gridrow="19" pageId="9" pageNumber="192">1</td>
<td id="76885DAAFFD2001FFD4F3967FD52FCEC" box="[680,733,843,863]" gridcol="5" gridrow="19" pageId="9" pageNumber="192">10.68</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD63694FFD2FFE9FF7C3E9AFDACFADC" blockId="9.[132,776,951,2030]" pageId="9" pageNumber="192">
The most striking feature of the fossils from Aumelas is their small size compared to other
<taxonomicName id="4C694D17FFD2FFE9FE5F3EFAFDB6FB43" authorityName="Matthes" authorityYear="1967" box="[440,569,1238,1264]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFD2FFE9FE5F3EFAFDB6FB43" box="[440,569,1238,1264]" italics="true" pageId="9" pageNumber="192">Oxyaenoides</emphasis>
</taxonomicName>
material. The teeth are 15% smaller than those of
<emphasis id="B91DEA86FFD2FFE9FE2C3EDAFDC5FABC" box="[459,586,1269,1296]" italics="true" pageId="9" pageNumber="192">O. lindgreni</emphasis>
and 25% smaller than those of
<emphasis id="B91DEA86FFD2FFE9FEF03F3AFE22FA9C" box="[279,429,1301,1328]" italics="true" pageId="9" pageNumber="192">O. bicuspidens</emphasis>
and
<emphasis id="B91DEA86FFD2FFE9FE043F3AFDD0FA9C" box="[483,607,1301,1328]" italics="true" pageId="9" pageNumber="192">O. schlosseri</emphasis>
. The size of the new taxon is actually almost equivalent to that of
<emphasis id="B91DEA86FFD2FFE9FD733F19FF70FADC" italics="true" pageId="9" pageNumber="192">Preregidens langebadrae</emphasis>
(Saint-Papoul;
<emphasis id="B91DEA86FFD2FFE9FE4F3F7AFE36FADC" box="[424,441,1366,1391]" italics="true" pageId="9" pageNumber="192">c.</emphasis>
MP8+9).
</paragraph>
<paragraph id="8BD63694FFD2FFE9FF7C3F59FEF8F93D" blockId="9.[132,776,951,2030]" pageId="9" pageNumber="192">
<taxonomicName id="4C694D17FFD2FFE9FF7C3F59FE2DFA3C" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[155,418,1397,1423]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FF7C3F59FE2DFA3C" box="[155,418,1397,1423]" italics="true" pageId="9" pageNumber="192">Oxyaenoides aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FE413F5AFE52FA23" box="[422,477,1398,1424]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
differs from those of
<emphasis id="B91DEA86FFD2FFE9FD513F59FF38FA1C" italics="true" pageId="9" pageNumber="192">O. lindgreni</emphasis>
and
<emphasis id="B91DEA86FFD2FFE9FF0D3FB9FEF0FA1C" box="[234,383,1429,1455]" italics="true" pageId="9" pageNumber="192">O. bicuspidens</emphasis>
by a more buccally aligned metastyle on upper molars. It also differs from
<emphasis id="B91DEA86FFD2FFE9FDD43F99FD5BFA7C" box="[563,724,1461,1487]" italics="true" pageId="9" pageNumber="192">O. bicuspidens</emphasis>
and
<emphasis id="B91DEA86FFD2FFE9FF633FF9FE8EFA5C" box="[132,257,1493,1519]" italics="true" pageId="9" pageNumber="192">O. schlosseri</emphasis>
by a mesially oriented protoconid this feature accommodates an elongate preprotocristid, a structure found on cutting molars.
<taxonomicName id="4C694D17FFD2FFE9FEB03C38FDE6F99D" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[343,617,1556,1582]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FEB03C38FDE6F99D" box="[343,617,1556,1582]" italics="true" pageId="9" pageNumber="192">Oxyaenoides aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FD953C39FD21F99C" box="[626,686,1557,1583]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
appears more primitive than northwestern species by having a buccally aligned metastyle on upper molars and the less retroflexed protoconid on molars.
</paragraph>
<paragraph id="8BD63694FFD2FFE9FF7C3CB9FE0BF8FE" blockId="9.[132,776,951,2030]" pageId="9" pageNumber="192">
To conclude, the fossils of
<taxonomicName id="4C694D17FFD2FFE9FE503CB8FD48F91D" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[439,711,1684,1710]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FE503CB8FD48F91D" box="[439,711,1684,1710]" italics="true" pageId="9" pageNumber="192">Oxyaenoides aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FD2A3CB9FC88F91C" box="[717,775,1685,1711]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
share with the other
<taxonomicName id="4C694D17FFD2FFE9FEB43C98FE40F97D" authorityName="Matthes" authorityYear="1967" box="[339,463,1716,1742]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFD2FFE9FEB43C98FE40F97D" box="[339,463,1716,1742]" italics="true" pageId="9" pageNumber="192">Oxyaenoides</emphasis>
</taxonomicName>
species several features including: a reduced metaconid and reduced talonid morphology. However, based on the small size and distinctive features, we distinguish
<taxonomicName id="4C694D17FFD2FFE9FEE63D38FE23F89D" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[257,428,1812,1838]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="192" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFD2FFE9FEE63D38FE23F89D" box="[257,428,1812,1838]" italics="true" pageId="9" pageNumber="192">O. aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFD2FFE9FE553D38FE63F89D" box="[434,492,1812,1838]" pageId="9" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
from previously described species of
<taxonomicName id="4C694D17FFD2FFE9FF163D1FFE0FF8FE" authorityName="Matthes" authorityYear="1967" box="[241,384,1843,1869]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">Oxyaenoides</taxonomicName>
.
</paragraph>
<paragraph id="8BD63694FFD2FFE9FF7C3D7FFD4FF85E" blockId="9.[132,776,951,2030]" pageId="9" pageNumber="192">
The fossils from Aumelas are important because they are the first to provide so much information on the morphology of the upper premolars of the genus
<taxonomicName id="4C694D17FFD2FFE9FDED3DBFFD04F81E" authorityName="Matthes" authorityYear="1967" box="[522,651,1939,1965]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFD2FFE9FDED3DBFFD04F81E" box="[522,651,1939,1965]" italics="true" pageId="9" pageNumber="192">Oxyaenoides</emphasis>
</taxonomicName>
. The fossils from Aumelas represent the smallest hypercarnivorous hyaenodont ever discovered in the Ypresian and Lutetian.
</paragraph>
</subSubSection>
</treatment>
</document>