treatments-xml/data/D7/32/40/D7324026FFF7FFC692ACFB99FB1E7769.xml

<document id="7C871CBAB4AC68C803681C4F8DE51B2F" ID-CLB-Dataset="263449" ID-DOI="10.11646/phytotaxa.607.1.7" ID-GBIF-Dataset="0c8c007a-9892-47b6-9740-7811adbe9909" ID-ISSN="1179-3163" ID-Zenodo-Dep="8212241" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1691078460740" checkinUser="plazi" docAuthor="Lu, Ngan Thi, Zhang, Liang, Thanh, Nguyen Trung, Toan, Le Chi &amp; Zhang, Li-Bing" docDate="2023" docId="D7324026FFF7FFC692ACFB99FB1E7769" docLanguage="en" docName="phytotaxa.607.1.7.pdf" docOrigin="Phytotaxa 607 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.607.1.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Polystichum xuansonense Li Bing Zhang, N. T. Lu &amp; Liang Zhang 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="2B0B385EFFF3FFC39224FFDFFFB7700E" masterDocTitle="Three new species and one new record of the fern genus Polystichum (Dryopteridaceae) from Vietnam" masterLastPageNumber="94" masterPageNumber="85" pageNumber="89" updateTime="1699550717687" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2C92BDEF00F3493632C0056E4C4D332C">Three new species and one new record of the fern genus Polystichum (Dryopteridaceae) from Vietnam</mods:title>
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<mods:namePart id="E911D5831AA5C7B92E79874854C41DDA">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="C9AF56A4B41484EE9B9C30C961895EA9">vnmngan @ gmail. com; https: // orcid. org / 0000 - 0001 - 9823 - 2018</mods:affiliation>
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<mods:namePart id="F514629A3E182E73DB33868ACD17BD57">Zhang, Liang</mods:namePart>
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<mods:affiliation id="32402CC0683E8602448CB7C70AC9F7C7">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China &amp; zhangliang @ mail. kib. ac. cn; https: // orcid. org / 0000 - 0003 - 3784 - 3135</mods:affiliation>
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<mods:namePart id="86C5BA86A0641E9BB7F173EA9917A961">Thanh, Nguyen Trung</mods:namePart>
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<mods:affiliation id="6CA8BA89EE8FF36D5F8A100E637F9DDB">Faculty of Biology, VNU University of Science, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam &amp; thanhntsh @ gmail. com; https: // orcid. org / 0000 - 0003 - 4734 - 3775</mods:affiliation>
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<mods:namePart id="E11BD349A7C556DC6C233970AAAEEDCB">Toan, Le Chi</mods:namePart>
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<mods:affiliation id="D410715A5C01CF4FCD4A4129106D7125">Department of Botany, Hanoi Pedagogical University 2, 32 Nguyen Van Linh, Xuan Hoa, Phuc Yen, Vietnam &amp; lechitoan @ hpu 2. edu. vn; https: // orcid. org / 0000 - 0002 - 5339 - 4440</mods:affiliation>
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<mods:namePart id="E23152002D70415B91614246A749ACD2">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="798A31B9D83035B942532A804E766235">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Sichuan 610041, China &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<taxonomicName id="989B8AB3FFF7FFC792ACFB99FCC2746E" ID-CoL="BSTY4" authority="Li Bing Zhang, N.T.Lu &amp; Liang Zhang" authorityName="Li Bing Zhang, N. T. Lu &amp; Liang Zhang" authorityYear="2023" box="[136,885,1094,1121]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="xuansonense" status="sp. nov.">
<emphasis id="6DEF2D22FFF7FFC792ACFB99FE02746F" bold="true" box="[136,437,1094,1121]" italics="true" pageId="4" pageNumber="89">Polystichum xuansonense</emphasis>
Li Bing Zhang, N.T.Lu &amp; Liang Zhang
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, 
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</emphasis>
(
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&amp; 
<figureCitation id="C7A0EDB5FFF7FFC79672FB99FBD0746E" box="[1110,1127,1094,1120]" captionStart="FIGURE 4" captionStartId="6.[136,234,1465,1487]" captionTargetBox="[280,1308,190,1440]" captionTargetId="figure-27@6.[280,1308,190,1441]" captionTargetPageId="6" captionText="FIGURE 4. Polystichum xuansonense Li Bing Zhang, N.T.Lu &amp; Liang Zhang.—A. Lower portion of plant.—B. Habit." figureDoi="http://doi.org/10.5281/zenodo.8212253" httpUri="https://zenodo.org/record/8212253/files/figure.png" pageId="4" pageNumber="89">4</figureCitation>
).
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</paragraph>
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<emphasis id="6DEF2D22FFF7FFC792ACFB4DFF6D74A9" bold="true" box="[136,218,1170,1191]" pageId="4" pageNumber="89">Type:—</emphasis>
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. 
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: 
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, 
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, 
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, limestone mountain, 
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, 
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, elev. ca. 
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, 
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, 
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<collectorName id="F26E94E6FFF7FFC79144FB69FC4574C5" box="[864,1010,1206,1227]" pageId="4" pageNumber="89">Li Bing Zhang</collectorName>
, 
<collectorName id="F26E94E6FFF7FFC791DAFB69FBCC74C5" box="[1022,1147,1206,1227]" pageId="4" pageNumber="89">Liang Zhang</collectorName>
&amp; 
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. Lu
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6565 (
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!;
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!, 
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)
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.
</paragraph>
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<emphasis id="6DEF2D22FFF7FFC792ACFB21FEBC751D" bold="true" box="[136,267,1278,1299]" italics="true" pageId="4" pageNumber="89">Diagnosis:—</emphasis>
This species is somewhat similar to 
<taxonomicName id="989B8AB3FFF7FFC79049FB21FC49751D" authority="Ching (1958: 234)" authorityName="Ching" authorityPageNumber="234" authorityYear="1958" box="[621,1022,1278,1300]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="excellens">
<emphasis id="6DEF2D22FFF7FFC79049FB21FCF4751D" box="[621,835,1278,1299]" italics="true" pageId="4" pageNumber="89">Polystichum excellens</emphasis>
<bibRefCitation id="3B0A8CC1FFF7FFC7916EFB21FC49751D" author="Ching, R. C." box="[842,1022,1278,1300]" pageId="4" pageNumber="89" refId="ref4455" refString="Ching, R. C. (1958) Icones Filicarum Sinicarum, vol. 5. The Metropolitan Museum of Natural History, Academia Sinica, Nanking (Nanjing) and the Fan Memorial Institute of Biology, Peiping (Beijing), 102 pp." type="book" year="1958">Ching (1958: 234)</bibRefCitation>
</taxonomicName>
in having oblong-lanceolate pinnae, but the former has middle pinnae with undulate to bluntly dentate acroscopic margins, proximal margins of pinnae parallel to or overlapping the rachis, and rounded pinna apex, whereas the latter has middle pinnae with low obtuse teeth or nearly repand acroscopic margins, proximal margins of pinnae forming an angle of 15–30° with the rachis, and acute pinna apex.
</paragraph>
</subSubSection>
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Plants medium-sized, evergreen, (13–) 
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tall. Rhizomes erect, ca. 
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long, ca. 
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in diam.; roots brown blackish when dried, up to 
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long, ca. 
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in diam. Fronds 9–14 per rhizome, 
<quantity id="98635CD5FFF7FFC7966BFA0DFB7575E3" box="[1103,1218,1490,1517]" metricMagnitude="-1" metricUnit="m" metricValue="2.65" metricValueMax="4.3" metricValueMin="1.0" pageId="4" pageNumber="89" unit="cm" value="26.5" valueMax="43.0" valueMin="10.0">10–43 cm</quantity>
; stipe brown, 
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long, ca. 1.0 mm in diam., basal stipe scales brown, membranous, lanceolate-ovate, 0.2–1.1 × 
<quantity id="98635CD5FFF7FFC7969BFA29FAF0761F" box="[1215,1351,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="4.1" metricValueMin="2.2" pageId="4" pageNumber="89" unit="mm" value="3.15" valueMax="4.1" valueMin="2.2">2.2–4.1 mm</quantity>
, margins entire, apex short acuminate; distal petiole scales lanceolate, subulate, 0.2–0.5 × 1.0– 
<quantity id="98635CD5FFF7FFC79664F9C5FB10763B" box="[1088,1191,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="2.12" pageId="4" pageNumber="89" unit="mm" value="2.12">2.12 mm</quantity>
, membranous, margins subentire to irregular dentate, apex long-acuminate or caudate. Laminae 1-pinnate, linear-lanceolate, 
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long, 
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wide, apex acuminate; rachises ca. 
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in diam., stramineous, adaxially sulcate; abaxially and adaxially with subulate scales, membranous, light brown, margins long outward, apex long-caudate. Pinnae 17–32 pairs, alternate, attached at 70° (–90°) angles to rachis, separate from each other, 
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distant (between two adjacent points), acroscopic auriculate; basalmost pinnae rhombic-ovate, 0.5–1.2 × 2.0–6.0 mm, shortly stalked, 2 basal pairs ca. 
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apart, apices acute or obtuse, bases obviously asymmetrical, acroscopic margins blunt dentate, basiscopic margins entire; middle pinnae largest, oblong-lanceolate, 0.4–0.7 × 
<quantity id="98635CD5FFF7FFC7914AF8C9FC5B773F" box="[878,1004,1814,1841]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="4" pageNumber="89" unit="cm" value="1.9" valueMax="2.5" valueMin="1.3">1.3–2.5 cm</quantity>
, shortly stalked, bases cuneate, proximal margins parallel to or overlapping the rachis, acroscopic margins undulate to bluntly dentate, basiscopic margins entire or slightly dentate upward, apices rounded; adaxially glabrous, abaxially sparsely covered with linear light brown microscales; frond texture papery; venation pinnate, distinct abaxially, slightly distinct adaxially, lateral veins nearly reaching pinna margin. Sori small, terminal on lateral veins of fertile pinnae, 
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in diam., closer to margins than to midrib, 1–12 on acroscopic side and 1–5 on basiscopic side, centers ca. 
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apart from one another; indusia present, light brown, entire.
</paragraph>
</subSubSection>
<caption id="0BE4A1B8FFF6FFC692ACFAA2FB4775D5" ID-DOI="http://doi.org/10.5281/zenodo.8212251" ID-Zenodo-Dep="8212251" httpUri="https://zenodo.org/record/8212251/files/figure.png" pageId="5" pageNumber="90" startId="5.[136,234,1405,1427]" targetBox="[363,1225,190,1380]" targetPageId="5" targetType="figure">
<paragraph id="5F24F130FFF6FFC692ACFAA2FB4775D5" blockId="5.[136,1452,1405,1499]" pageId="5" pageNumber="90">
<emphasis id="6DEF2D22FFF6FFC692ACFAA2FEB4759C" bold="true" box="[136,259,1405,1427]" pageId="5" pageNumber="90">FIGURE 3.</emphasis>
<taxonomicName id="989B8AB3FFF6FFC6932CFAA2FC34759D" authority="Li Bing Zhang, N. T. Lu &amp; Liang Zhang." authorityName="Li Bing Zhang, N. T. Lu &amp; Liang Zhang." box="[264,899,1405,1427]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="xuansonense">
<emphasis id="6DEF2D22FFF6FFC6932CFAA2FDB7759D" box="[264,512,1405,1427]" italics="true" pageId="5" pageNumber="90">Polystichum xuansonense</emphasis>
Li Bing Zhang, N.T.Lu &amp; Liang Zhang.
</taxonomicName>
—A &amp; D. Abaxial views of upper portions of lamina.—B &amp; C. Adaxial views of upper portions of lamina.—E. Adaxial view of lower portion of lamina.—F. Indusia and portion of pinna showing veins and sori.—G. Scales from proximal and distal stipes.—H. Portion of lamina showing auricles and rachis scales.
</paragraph>
</caption>
<subSubSection id="1781A2BBFFF6FFC69299F9D6FE457649" pageId="5" pageNumber="90" type="etymology">
<paragraph id="5F24F130FFF6FFC69299F9D6FE457649" blockId="5.[136,1452,1545,1896]" pageId="5" pageNumber="90">
<emphasis id="6DEF2D22FFF6FFC69299F9D6FED3762D" bold="true" box="[189,356,1545,1571]" pageId="5" pageNumber="90">Etymology:—</emphasis>
The species epithet is from Xuan Son, the name of the national park in northern 
<collectingCountry id="278CB1A0FFF6FFC696DDF9D6FAEC762D" box="[1273,1371,1545,1571]" name="Vietnam" pageId="5" pageNumber="90">Vietnam</collectingCountry>
, where the new species was discovered.
</paragraph>
</subSubSection>
<subSubSection id="1781A2BBFFF6FFC69299F98EFBD57665" box="[189,1122,1617,1644]" pageId="5" pageNumber="90" type="distribution">
<paragraph id="5F24F130FFF6FFC69299F98EFBD57665" blockId="5.[136,1452,1545,1896]" box="[189,1122,1617,1644]" pageId="5" pageNumber="90">
<emphasis id="6DEF2D22FFF6FFC69299F98EFEC27665" bold="true" box="[189,373,1617,1643]" pageId="5" pageNumber="90">Distribution:—</emphasis>
This species is currently known to be endemic to the 
<typeStatus id="80204F92FFF6FFC691EDF98DFC4E7662" box="[969,1017,1618,1644]" pageId="5" pageNumber="90">type</typeStatus>
location.
</paragraph>
</subSubSection>
<subSubSection id="1781A2BBFFF6FFC69299F9AAFB1E7769" pageId="5" pageNumber="90" type="discussion">
<paragraph id="5F24F130FFF6FFC69299F9AAFA8176F5" blockId="5.[136,1452,1545,1896]" pageId="5" pageNumber="90">
<emphasis id="6DEF2D22FFF6FFC69299F9AAFEB77681" bold="true" box="[189,256,1653,1679]" pageId="5" pageNumber="90">Notes</emphasis>
:—Both 
<taxonomicName id="989B8AB3FFF6FFC69346F9AAFD347681" box="[354,643,1653,1679]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="xuansonense">
<emphasis id="6DEF2D22FFF6FFC69346F9AAFD347681" box="[354,643,1653,1679]" italics="true" pageId="5" pageNumber="90">Polystichum xuansonense</emphasis>
</taxonomicName>
and 
<taxonomicName id="989B8AB3FFF6FFC6909FF9A9FC887681" authorityName="Ching" authorityYear="1958" box="[699,831,1653,1679]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="excellens">
<emphasis id="6DEF2D22FFF6FFC6909FF9A9FC887681" box="[699,831,1653,1679]" italics="true" pageId="5" pageNumber="90">P. excellens</emphasis>
</taxonomicName>
occur in northern 
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and the latter is common in limestone areas in 
<collectingRegion id="9D5F3FD2FFF6FFC6937DF946FE7C76BA" box="[345,459,1689,1716]" country="Vietnam" name="Cao Bang" pageId="5" pageNumber="90">Cao Bang</collectingRegion>
, 
<collectingRegion id="9D5F3FD2FFF6FFC693F2F945FDF376BD" box="[470,580,1689,1716]" country="Vietnam" name="Ha Giang" pageId="5" pageNumber="90">Ha Giang</collectingRegion>
, and 
<collectingRegion id="9D5F3FD2FFF6FFC6905BF945FD5D76BD" box="[639,746,1689,1716]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="90">Hoa Binh</collectingRegion>
provinces (
<bibRefCitation id="3B0A8CC1FFF6FFC69149F946FBAE76BD" author="Lu, T. N. &amp; Zhang, L. &amp; Zhang, L. - B." box="[877,1049,1689,1715]" pageId="5" pageNumber="90" pagination="115 - 123" refId="ref4903" refString="Lu, T. N., Zhang, L. &amp; Zhang, L. - B. (2014 a) Ten new records of Polystichum (Dryopteridaceae) for Vietnam and lectotypification of Polystichum atroviridissimum and P. fimbriatum. Phytotaxa 164 (2): 115 - 123. https: // doi. org / 10.11646 / phytotaxa. 164.2.5" type="journal article" year="2014">
Lu 
<emphasis id="6DEF2D22FFF6FFC691B1F945FC7A76BD" box="[917,973,1689,1715]" italics="true" pageId="5" pageNumber="90">et al.</emphasis>
2014a
</bibRefCitation>
) but not found in 
<collectingRegion id="9D5F3FD2FFF6FFC696C6F946FA1B76BD" box="[1250,1452,1689,1715]" country="Vietnam" name="Phu Tho" pageId="5" pageNumber="90">Phu Tho Province</collectingRegion>
where the former was discovered. 
<taxonomicName id="989B8AB3FFF6FFC6902FF962FC9B76D9" box="[523,812,1725,1751]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="xuansonense">
<emphasis id="6DEF2D22FFF6FFC6902FF962FC9B76D9" box="[523,812,1725,1751]" italics="true" pageId="5" pageNumber="90">Polystichum xuansonense</emphasis>
</taxonomicName>
is quite rare and was found only in one small location in relatively dry forest with about 30 individuals. It might occur elsewhere and more explorations are needed.
</paragraph>
<paragraph id="5F24F130FFF6FFC69299F8DAFB1E7769" blockId="5.[136,1452,1545,1896]" pageId="5" pageNumber="90">
Our unpublished DNA data shows that 
<taxonomicName id="989B8AB3FFF6FFC6904BF8D9FCAE7711" box="[623,793,1798,1823]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="xuansonense">
<emphasis id="6DEF2D22FFF6FFC6904BF8D9FCAE7711" box="[623,793,1798,1823]" italics="true" pageId="5" pageNumber="90">P. xuansonense</emphasis>
</taxonomicName>
might be closely related to 
<taxonomicName id="989B8AB3FFF6FFC6966DF8D9FACA7711" authority="Christ (1906: 238)" authorityName="Christ" authorityPageNumber="238" authorityYear="1906" box="[1097,1405,1797,1823]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="dielsii">
<emphasis id="6DEF2D22FFF6FFC6966DF8D9FB1F7711" box="[1097,1192,1797,1823]" italics="true" pageId="5" pageNumber="90">P. dielsii</emphasis>
Christ (1906: 238)
</taxonomicName>
, but the two are morphologically quite different from each other: 
<taxonomicName id="989B8AB3FFF6FFC6910BF8F5FC6D774D" box="[815,986,1834,1859]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="xuansonense">
<emphasis id="6DEF2D22FFF6FFC6910BF8F5FC6D774D" box="[815,986,1834,1859]" italics="true" pageId="5" pageNumber="90">P. xuansonense</emphasis>
</taxonomicName>
has oblong-lanceolate pinnae, whereas 
<taxonomicName id="989B8AB3FFF6FFC697B3F8F5FF7B7769" authorityName="Christ" authorityYear="1906" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="dielsii">
<emphasis id="6DEF2D22FFF6FFC697B3F8F5FF7B7769" italics="true" pageId="5" pageNumber="90">P. dielsii</emphasis>
</taxonomicName>
has oblong pinnae. The latter occurs in 
<collectingCountry id="278CB1A0FFF6FFC690A9F892FD677769" box="[653,720,1869,1895]" name="China" pageId="5" pageNumber="90">China</collectingCountry>
and 
<collectingCountry id="278CB1A0FFF6FFC69121F892FCD37769" box="[773,868,1869,1895]" name="Vietnam" pageId="5" pageNumber="90">Vietnam</collectingCountry>
(
<bibRefCitation id="3B0A8CC1FFF6FFC69157F892FB2A7769" author="Zhang, L. - B. &amp; Barrington, D. S." box="[883,1181,1869,1895]" pageId="5" pageNumber="90" pagination="629 - 713" refId="ref6175" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum. In: Wu, Z. - Y., Raven, P. H. &amp; Hong, D. - Y. (Eds.) Flora of China, vols. 2 - 3. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 629 - 713." type="book chapter" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>