186 lines
19 KiB
XML
186 lines
19 KiB
XML
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<mods:title id="C081C0AEB557944F8724457B5692736B">Revision and taxonomic position of genera Afghanorthochirus Lourenço & Vachon, Baloorthochirus Kova ík, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kova ík, Pakistanorthochirus Lourenço, and Asian Orthochirus Karsch, with descriptions of twelve new species (Scorpiones, Buthidae)</mods:title>
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<mods:namePart id="773847FD412D0F862B98F92A4D2FF775">Kovařík, Baloorthochirus becvari</mods:namePart>
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<mods:roleTerm id="13BDF03E30240AC360538E0FFBCE727F">Author</mods:roleTerm>
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<mods:namePart id="7116BD625BDDE6E0CF1DE12DE150DAFA">1996</mods:namePart>
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<mods:title id="8FDC42D5703C830057A4C3610785DC59">Euscorpius</mods:title>
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<mods:part id="27E973544CA2642F2A17EEC1F7205DDC">
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<mods:date id="3799766628A77BF23746FBB8797C82F2">2004</mods:date>
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<mods:detail id="936E5E22A32A6C9F54AB6E476327E58A" type="volume">
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<mods:number id="515EB24E0B2F629EE3BCE06E7C109B7D">16</mods:number>
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<subSubSection id="C338C2DD3C335B29FC6EF8CBFB7DF8C2" box="[995,1276,1799,1826]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8B9D91563C335B29FC6EF8CBFB7DF8C2" blockId="3.[995,1276,1799,1826]" box="[995,1276,1799,1826]" pageId="3" pageNumber="2">
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<heading id="D0D5263A3C335B29FC6EF8CBFB7DF8C2" box="[995,1276,1799,1826]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2" reason="4">
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<taxonomicName id="4C22EAD53C335B29FC6EF8CBFB7DF8C2" ID-CoL="62HVH" authority="Simon, 1882" authorityName=": Simon" authorityYear="1889" box="[995,1276,1799,1826]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B9564D443C335B29FC6EF8CBFBDEF8C2" bold="true" box="[995,1119,1799,1826]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
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Simon, 1882
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C338C2DD3C335B2EFCB3F884FEB2FEDA" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="reference_group">
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<paragraph id="8B9D91563C335B2EFCB3F884FDC9FF1D" blockId="3.[830,1441,1864,1918]" lastBlockId="4.[192,802,199,682]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<taxonomicName id="4C22EAD53C335B29FCB3F884FBF5F8BF" ID-CoL="62HVH" authority="Simon, 1882: 248" authorityName=": Simon" authorityPageNumber="248" authorityYear="1889" box="[830,1140,1864,1887]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B9564D443C335B29FCB3F884FC28F8BF" box="[830,937,1864,1887]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
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Simon, 1882: 248
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</taxonomicName>
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;
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<bibRefCitation id="EFB3ECA73C335B29FB0CF884FABCF8BF" author="KARSCH" box="[1153,1341,1864,1887]" pageId="3" pageNumber="2" pagination="75 - 79" refId="ref27060" refString="KARSCH, F. 1886. Skorpionologische Beitrage. Berliner Entomologische Zeitschrift, 30: 75 - 79." type="journal article" year="1886">Karsch, 1886: 76</bibRefCitation>
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; Werner, 1934: 270; Vachon, 1980: 253; Sissom, 1990: 101;
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<bibRefCitation id="EFB3ECA73C345B2EFF4DFF0BFE1AFF3E" author="KOVARIK" box="[192,411,199,222]" pageId="4" pageNumber="3" pagination="177 - 181" refId="ref27390" refString="KOVARIK, F. 1996. Baloorthochirus becvari gen. et sp. n. from Pakistan, and taxonomic position of Orthochirus luteipes (Scorpiones: Buthidae). Acta Societatis Zoologicae Bohemicae, 60: 177 - 181." type="journal article" year="1996">Kovařík, 1996: 177</bibRefCitation>
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; Kovařík, 1998: 117; Fet & Lowe, 2000: 88; Lourenço, 2001: 176, 179.
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</paragraph>
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<paragraph id="8B9D91563C345B2EFF4DFEC8FEB2FEDA" blockId="4.[192,802,199,682]" pageId="4" pageNumber="3">
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<taxonomicName id="4C22EAD53C345B2EFF4DFEC8FEAAFEFB" ID-CoL="62HVH" authorityName=": Simon" authorityYear="1889" box="[192,299,260,283]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B9564D443C345B2EFF4DFEC8FEAAFEFB" box="[192,299,260,283]" italics="true" pageId="4" pageNumber="3">Butheolus</emphasis>
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</taxonomicName>
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(in part): Kraepelin, 1899: 34; Kraepelin, 1903: 564.
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</paragraph>
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</subSubSection>
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<subSubSection id="C338C2DD3C345B2EFF4DFE8EFD33FDF1" pageId="4" pageNumber="3" type="description">
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<paragraph id="8B9D91563C345B2EFF4DFE8EFE8FFE12" blockId="4.[192,802,199,682]" pageId="4" pageNumber="3">
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=
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<taxonomicName id="4C22EAD53C345B2EFF6CFE8EFDD2FEB9" authority="Pocock, 1895: 314" authorityName="Pocock" authorityPageNumber="314" authorityYear="1895" box="[225,595,322,345]" class="Arachnida" family="Buthidae" genus="Nanobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B9564D443C345B2EFF6CFE8EFEE3FEB9" box="[225,354,322,345]" italics="true" pageId="4" pageNumber="3">Nanobuthus</emphasis>
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Pocock, 1895: 314
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</taxonomicName>
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, TS:
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<taxonomicName id="4C22EAD53C345B2EFD2CFE8EFE42FE97" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" class="Arachnida" family="Buthidae" genus="Nanobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="andersoni">
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<emphasis id="B9564D443C345B2EFD2CFE8EFEAAFE97" italics="true" pageId="4" pageNumber="3">Nanobuthus andersoni</emphasis>
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Pocock, 1895
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</taxonomicName>
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=
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<taxonomicName id="4C22EAD53C345B2EFE6DFEACFE80FE76" authority="(Pocock, 1895)" authorityName="Kovařík & 1996" authorityYear="2004" baseAuthorityName="Pocock" baseAuthorityYear="1895" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="andersoni" status="comb. nov.">
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<emphasis id="B9564D443C345B2EFE6DFEACFD3CFE97" box="[480,701,352,375]" italics="true" pageId="4" pageNumber="3">Butheolus andersoni</emphasis>
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(Pocock, 1895)
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</taxonomicName>
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,
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<emphasis id="B9564D443C345B2EFE83FEB3FEF0FE76" bold="true" box="[270,369,383,406]" pageId="4" pageNumber="3">
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<taxonomicNameLabel id="A265F03F3C345B2EFE83FEB3FEF0FE76" box="[270,369,383,406]" pageId="4" pageNumber="3" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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; Kraepelin, 1899: 38; Kraepelin, 1903: 564; Kraepelin, 1905: 337; Werner, 1934: 270; Perez, 1974: 27; Fet & Lowe, 2000: 186; Lourenço, 2001: 179.
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<emphasis id="B9564D443C345B2EFF4DFE17FE8FFE12" bold="true" box="[192,270,475,498]" pageId="4" pageNumber="3">Syn. n.</emphasis>
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</paragraph>
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<paragraph id="8B9D91563C345B2EFF4DFE36FD33FDF1" blockId="4.[192,802,199,682]" box="[192,690,506,529]" pageId="4" pageNumber="3">
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<taxonomicName id="4C22EAD53C345B2EFF4DFE36FE47FDF1" authority="" box="[192,454,506,529]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Nanobuthus">
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<emphasis id="B9564D443C345B2EFF4DFE36FEAAFDF1" box="[192,299,506,529]" italics="true" pageId="4" pageNumber="3">Butheolus</emphasis>
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(
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<emphasis id="B9564D443C345B2EFEB7FE36FE3AFDF1" box="[314,443,506,529]" italics="true" pageId="4" pageNumber="3">Nanobuthus</emphasis>
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)
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</taxonomicName>
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: Vachon, 1975: 1598.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338C2DD3C345B2EFF4DFDD4FD25FD6B" pageId="4" pageNumber="3" type="reference_group">
|
||
<paragraph id="8B9D91563C345B2EFF4DFDD4FE21FD8D" blockId="4.[192,802,199,682]" pageId="4" pageNumber="3">
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||
<treatmentCitationGroup id="AB32B6783C345B2EFF4DFDD4FE21FD8D" pageId="4" pageNumber="3">
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||
=
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||
<treatmentCitation id="0A83B7473C345B2EFF5BFDD4FD80FDCF" author="HIRST" box="[214,513,536,559]" page="462" pageId="4" pageNumber="3" year="1911">
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||
<taxonomicName id="4C22EAD53C345B2EFF5BFDD4FD80FDCF" ID-CoL="5YVN" authority="Hirst, 1911: 462" authorityName="Hirst" authorityPageNumber="462" authorityYear="1911" box="[214,513,536,559]" class="Arachnida" family="Buthidae" genus="Neobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9564D443C345B2EFF5BFDD4FEC9FDCF" box="[214,328,536,559]" italics="true" pageId="4" pageNumber="3">Neobuthus</emphasis>
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||
<bibRefCitation id="EFB3ECA73C345B2EFEC2FDD4FD80FDCF" author="HIRST" box="[335,513,536,559]" pageId="4" pageNumber="3" pagination="462 - 473" refId="ref26937" refString="HIRST, S. 1911. Descriptions of new Scorpions. Annals and Magazine of Natural History, 8 (8): 462 - 473." type="journal article" year="1911">Hirst, 1911: 462</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
; Werner, 1934: 270; Fet & Lowe, 2000: 186; Lourenço, 2001: 179 (syn. by Kovařík, 2003: 137).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9D91563C345B2EFF4DFDB8FD25FD6B" blockId="4.[192,802,199,682]" box="[192,676,628,651]" pageId="4" pageNumber="3">
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<taxonomicName id="4C22EAD53C345B2EFF4DFDB8FE36FD6B" authority="" box="[192,439,628,651]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Neobuthus">
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<emphasis id="B9564D443C345B2EFF4DFDB8FEAAFD6B" box="[192,299,628,651]" italics="true" pageId="4" pageNumber="3">Butheolus</emphasis>
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(
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<emphasis id="B9564D443C345B2EFEB7FDB8FE2DFD6B" box="[314,428,628,651]" italics="true" pageId="4" pageNumber="3">Neobuthus</emphasis>
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)
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</taxonomicName>
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: Vachon, 1975: 1598.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C338C2DD3C345B2EFF4DFD5FFD5EFD4A" box="[192,735,659,682]" pageId="4" pageNumber="3" type="materials_examined">
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<paragraph id="8B9D91563C345B2EFF4DFD5FFD5EFD4A" blockId="4.[192,802,199,682]" box="[192,735,659,682]" pageId="4" pageNumber="3">
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<typeStatus id="54992FF43C345B2EFF4DFD5FFF79FD49" box="[192,248,659,682]" pageId="4" pageNumber="3">TYPE</typeStatus>
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SPECIES:
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<taxonomicName id="4C22EAD53C345B2EFED2FD5FFD5AFD4A" authority="Simon, 1882" authorityName="Simon" authorityYear="1882" box="[351,731,659,682]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="thalassinus">
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<emphasis id="B9564D443C345B2EFED2FD5FFDC9FD4A" box="[351,584,659,682]" italics="true" pageId="4" pageNumber="3">Butheolus thalassinus</emphasis>
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Simon, 1882
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C338C2DD3C345B2EFF4DFD1CFDB4FBB7" pageId="4" pageNumber="3" type="diagnosis">
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<paragraph id="8B9D91563C345B2EFF4DFD1CFDB4FBB7" blockId="4.[192,802,720,1111]" pageId="4" pageNumber="3">
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DIAGNOSIS: Patella of pedipalp without ventral trichobothria. Dorsal trichobothria of femur arranged in beta-configuration. Trichobothrium
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<emphasis id="B9564D443C345B2EFDCEFCC2FDD1FCC5" box="[579,592,782,805]" italics="true" pageId="4" pageNumber="3">d</emphasis>
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<subScript id="17A693133C345B2EFDDCFCD4FDDBFCC7" attach="left" box="[593,602,792,807]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
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of pedipalp femur present on dorsal surface but often smaller than other trichobothria. Tibial spurs present on third and fourth legs. Pectines with fulcra and densely hirsute. Movable fingers of pedipalps with 6–10 rows of granules and 3–5 distal granules. Carapace, in lateral view, distinctly inclined downward from median eyes to anterior margin. Mesosoma with three carinae, of which two may be less conspicuous. Metasomal segments ventrally smooth or granulated. Telson bulbous, aculeus shorter or as long as vesicle. Total length under
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<quantity id="4CDA3CB33C345B2EFE6EFB8CFDB0FBB7" box="[483,561,1088,1111]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="3" unit="mm" value="60.0">60 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C338C2DD3C345B2EFF4DFBB2FDCEF9E5" pageId="4" pageNumber="3" type="discussion">
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<paragraph id="8B9D91563C345B2EFF4DFBB2FDCEF9E5" blockId="4.[192,802,1150,1541]" pageId="4" pageNumber="3">
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COMMENTS. Although the genus
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<taxonomicName id="4C22EAD53C345B2EFDAEFBB2FD25FB75" authorityName="Pocock" authorityYear="1895" box="[547,676,1150,1173]" class="Arachnida" family="Buthidae" genus="Nanobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9564D443C345B2EFDAEFBB2FD25FB75" box="[547,676,1150,1173]" italics="true" pageId="4" pageNumber="3">Nanobuthus</emphasis>
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||
</taxonomicName>
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||
is accepted in recent papers (Lourenço, 2001: 179; Kovařík, 2003: 142) as valid, after examination of most species of
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||
<taxonomicName id="4C22EAD53C345B2EFF4DFB16FEAAFB11" authorityName=": Simon" authorityYear="1889" box="[192,299,1242,1265]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9564D443C345B2EFF4DFB16FEAAFB11" box="[192,299,1242,1265]" italics="true" pageId="4" pageNumber="3">Butheolus</emphasis>
|
||
</taxonomicName>
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and related genera I come to the conclusion that
|
||
<taxonomicName id="4C22EAD53C345B2EFF7DFB34FDFDFAEF" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" box="[240,636,1272,1295]" class="Arachnida" family="Buthidae" genus="Nanobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="andersoni">
|
||
<emphasis id="B9564D443C345B2EFF7DFB34FE62FAEF" box="[240,483,1272,1295]" italics="true" pageId="4" pageNumber="3">Nanobuthus andersoni</emphasis>
|
||
Pocock, 1895
|
||
</taxonomicName>
|
||
does not differ enough, namely from the similar
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<taxonomicName id="4C22EAD53C345B2EFDA3FADBFEEEFAAD" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="anthracinus">
|
||
<emphasis id="B9564D443C345B2EFDA3FADBFCA0FACE" box="[558,801,1303,1326]" italics="true" pageId="4" pageNumber="3">Butheolus anthracinus</emphasis>
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||
(Pocock, 1895)
|
||
</taxonomicName>
|
||
, to warrant placement in a separate genus. Opinions on what characters should be regarded as generic vary. In the species concerned the primary difference is in the width of the metasoma; however, this character cannot be deemed generic, as is clear from examples of other genera (
|
||
<taxonomicName id="4C22EAD53C345B2EFD95FA03FD0BFA06" authorityName="Karsch" authorityYear="1886" box="[536,650,1487,1510]" class="Arachnida" family="Buthidae" genus="Babycurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9564D443C345B2EFD95FA03FD0BFA06" box="[536,650,1487,1510]" italics="true" pageId="4" pageNumber="3">Babycurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C22EAD53C345B2EFD2BFA03FC9AFA06" authorityName="Peters" authorityYear="1861" box="[678,795,1487,1510]" class="Arachnida" family="Buthidae" genus="Uroplectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9564D443C345B2EFD2BFA03FC9AFA06" box="[678,795,1487,1510]" italics="true" pageId="4" pageNumber="3">Uroplectes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C22EAD53C345B2EFF4DFA22FEBBF9E5" box="[192,314,1518,1541]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9564D443C345B2EFF4DFA22FEBBF9E5" box="[192,314,1518,1541]" italics="true" pageId="4" pageNumber="3">Hottentotta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C22EAD53C345B2EFECAFA22FE42F9E5" authorityName="Pocock" authorityYear="1890" box="[327,451,1518,1541]" class="Arachnida" family="Buthidae" genus="Parabuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9564D443C345B2EFECAFA22FE42F9E5" box="[327,451,1518,1541]" italics="true" pageId="4" pageNumber="3">Parabuthus</emphasis>
|
||
</taxonomicName>
|
||
, and others).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |