357 lines
41 KiB
XML
357 lines
41 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="ECDAF5B877D124C1802B904D5509173E" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole maizina Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="0E6A56CB-6415-4402-A3C9-80FFD4F86BF7" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:title id="A6551AC6AA33EABBAEA00C7DA5858E4D">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart id="51174999B5522CD160BA3DD2D7D219F6">Salata, Sebastian</mods:namePart>
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<mods:namePart id="98AD6557200E5A1A3853FE1E3E7D02C8">Fisher, Brian L.</mods:namePart>
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<mods:title id="D53413313D9239CCED8F0F663DB2AD3D">ZooKeys</mods:title>
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<mods:part id="D3DE303594F1513BAB576CD268B11612">
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<mods:date id="EA95F7FD5CB3DE8E05F2C86AB2CE3E68">2020</mods:date>
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<mods:detail id="19A7F650929EA79B4B6F93A77E4213FC" type="volume">
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<mods:number id="BFF14EBA2EA41ED0F6390C9204B9E5E6">905</mods:number>
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<mods:url id="E60A44E1885D6067288B3703B5B53859">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="5BDDC7F457D55CB4AE8CC8E25083D025">journal article</mods:classification>
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<mods:identifier id="8B5FEEE6C7F8DF6D598686C01A4BA2F8" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<treatment id="ECDAF5B877D124C1802B904D5509173E" ID-GBIF-Taxon="161777038" LSID="urn:lsid:zoobank.org:act:0E6A56CB-6415-4402-A3C9-80FFD4F86BF7" httpUri="http://treatment.plazi.org/id/ECDAF5B877D124C1802B904D5509173E" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="6C0973F3CE9EBAD20E17FDECB4FE76D9" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="F5587B5469A0C1A348C5BAD992E22A4F" pageId="0" pageNumber="1">
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<taxonomicName id="1558B67E0464B96EFD130A3A2AB8D40F" ID-CoL="CMLFT" LSID="http://zoobank.org/0E6A56CB-6415-4402-A3C9-80FFD4F86BF7" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina" status="sp. nov.">Pheidole maizina</taxonomicName>
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<taxonomicNameLabel id="A7C2B12DCB6F1E2729B45B9714CF5830" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="1FED6A2ACB7A2534A0B65422E651A303" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">Figs 27A-F</figureCitation>
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<figureCitation id="A4F90C67C6B79D240B95ABC6DF6EF47D" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85C</figureCitation>
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<figureCitation id="BF927FD7BBE60476B10E811F36B29710" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87G</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="D58D0C409CFDF72E766DD1E105059F43" pageId="0" pageNumber="1" type="type material">
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<paragraph id="C3ECB16F68C45FFEC4867C31B2ACB761" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="23DC03030A6C5254360CD8D36A1B52B3" pageId="0" pageNumber="1">
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<emphasis id="5C7C9A2812373ED53C2B7D4A73B55748" bold="true" pageId="0" pageNumber="1">
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<emphasis id="4B7B5309DCA428FED2A6CC26C41E56C5" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toamasina;
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<normalizedToken id="52DF1ED5BFCF35323E3A1A31D9BD9103" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="F6AC8B2FF3C7EF79E6329963FEC689DE" originalValue="Spéciale">Speciale</normalizedToken>
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||
Ambatovaky, Sandrangato River;
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<geoCoordinate id="394064CD6C8149A6099C8468F341C850" degrees="16.80561" direction="south" orientation="latitude" precision="1" value="-16.80561">-16.80561</geoCoordinate>
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,
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<geoCoordinate id="4C68E98506D4FDC1CC10BAD4B415F55B" degrees="49.29507" direction="east" orientation="longitude" precision="1" value="49.29507">49.29507</geoCoordinate>
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; alt. 480 m; 27 Feb 2010; B.L. Fisher et al. leg.; BLF25039, CASENT0162231 (CASC).
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<emphasis id="32CD660FA7F21109E1688D5F54B54A5D" bold="true" pageId="0" pageNumber="1">
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<emphasis id="452C74304A4902BBEE6E8938BE0824F1" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •2w., 1q.; same data as for holotype; CASENT0162230, CASENT0923223, CASENT0872222 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="1BE4647D6F69C5FBDC88F491623A0CCF" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="8754A8CB5B0524B142FE928DCE695318" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="059D91A5C0D663476BE57119FE5844AC" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="EA3D5CD994C48391CB13BDAB73814E86" bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •3w., 1s.; 6.5 km SSW Befingotra,
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<normalizedToken id="971B433D56B598C955E65027E8301007" originalValue="Rés">Res</normalizedToken>
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||
. Anjanaharibe-Sud; -14.75, 49.5; alt. 875 m; 19 Oct 1994; B.L. Fisher leg.; CASENT0198389, CASENT0198390 (CASC). •7w., 4s., 1q.; 9.2 km WSW Befingotra,
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<normalizedToken id="6B51EE98236854FCB2385C6B2A21D9E5" originalValue="Rés">Res</normalizedToken>
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. Anjanaharibe-Sud; -14.75, 49.46667; alt. 1200 m; 8 Nov 1994; B.L. Fisher leg.; CASENT0198391, CASENT0198393, CASENT0198394, CASENT0235113, CASENT0235114 (CASC). -
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<emphasis id="36D3A5E76D831BE52CB5EE1B1AB96CAF" bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
|
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: •1w.; Parc National de Ranomafana, Vatoharanana River, 4.1 km 231°SW Ranomafana; -21.29, 47.43333; alt. 1100 m; 27 Mar 2003; B.L. Fisher et al. leg.; CASENT0497619 (CASC). -
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<emphasis id="79ACEF4D54150C3E139BEF087B98F12D" bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
|
||
: •3w., 2s.;
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|
||
<normalizedToken id="251988074DB79B6325F61466775B350C" originalValue="Spéciale">Speciale</normalizedToken>
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||
Marotandrano, Marotandrano 48.3 km S Mandritsara;
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<geoCoordinate id="D3609B27C9BD465185B72A4152886B7B" degrees="16.28322" direction="south" orientation="latitude" precision="1" value="-16.28322">-16.28322</geoCoordinate>
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,
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<geoCoordinate id="EF097638061F20CCBDB42DF6E9035D24" degrees="48.81443" direction="east" orientation="longitude" precision="1" value="48.81443">48.81443</geoCoordinate>
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; alt. 865 m; 7 Dec 2007; B.L. Fisher et al. leg.; CASENT0134242, CASENT0134279, CASENT0235121 (CASC). -
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||
<emphasis id="43685CE54CE7ED141F607EE48A576A33" bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •1w.; Ambanizana, Parc National Masoala;
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<geoCoordinate id="7DF4F29AB26A5D66A30D5DFCEFC4BA07" degrees="15.57222" direction="south" orientation="latitude" precision="1" value="-15.57222">-15.57222</geoCoordinate>
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,
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<geoCoordinate id="CCD272FEFA3D77DD6DC3E076C9EFD144" degrees="50.00694" direction="east" orientation="longitude" precision="1" value="50.00694">50.00694</geoCoordinate>
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; alt. 930-1110 m; 2 Mar 2003; Andriamalala et al. leg.; CASENT0073482 (CASC). •1w.; F.C. Andriantantely;
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<geoCoordinate id="26612C1046214A86020ED74888BECDB8" degrees="18.695" direction="south" orientation="latitude" precision="55" value="-18.695">-18.695</geoCoordinate>
|
||
,
|
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<geoCoordinate id="F12D685F6F01A399D71AF39DF09359F2" degrees="48.81333" direction="east" orientation="longitude" precision="1" value="48.81333">48.81333</geoCoordinate>
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; alt. 530 m; 4 Dec 1998; Ratsirarson leg.; CASENT0198237 (CASC). •19w., 1s.; Montagne
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<normalizedToken id="55A77DBBB9DC2BB1B92DE35A9AF3490C" originalValue="d’Akirindro">d'Akirindro</normalizedToken>
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||
7.6 km 341°NNW Ambinanitelo;
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<geoCoordinate id="C1A50111E33809F11F142013C1F412B9" degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
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,
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<geoCoordinate id="4769FEE79D6BABBE7259C194C6AFB03C" degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
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; alt. 600 m; 17 Mar 2003; B.L. Fisher et al. leg.; CASENT0039054, CASENT0039064, CASENT0039103, CASENT0039188, CASENT0039200, CASENT0039209, CASENT0039219, CASENT0235138, CASENT0496323, CASENT0496345, CASENT0496346, CASENT0496347 (CASC). •15w., 4s.; Montagne
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||
<normalizedToken id="B7761BA99ABD55450D1A201F2E05234C" originalValue="d’Anjanaharibe">d'Anjanaharibe</normalizedToken>
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||
, 18.0 km 21°NNE Ambinanitelo;
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||
<geoCoordinate id="44D89A28EF7686DECC18463329A9F735" degrees="15.18833" direction="south" orientation="latitude" precision="1" value="-15.18833">-15.18833</geoCoordinate>
|
||
,
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||
<geoCoordinate id="D0313B664E2B57DF7ABC98862E1794B3" degrees="49.615" direction="east" orientation="longitude" precision="55" value="49.615">49.615</geoCoordinate>
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||
; alt. 470 m; 8 Mar 2003; B.L. Fisher et al. leg.; CASENT0037623, CASENT0037626, CASENT0048951, CASENT0495382, CASENT0495383, CASENT0495384, CASENT0495480, CASENT0495481, CASENT0495482 (CASC). •1w.;
|
||
<normalizedToken id="C54A256C4EC52F9D60D0CD68B500056F" originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken id="111C61933EF043634F078F398CD3F1FD" originalValue="Spéciale">Speciale</normalizedToken>
|
||
Ambatovaky, Sandrangato River;
|
||
<geoCoordinate id="F5E5B2447CADC1DDC2EB954F7127004B" degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="73D392A7B0946EB457F5D0827F5E61E3" degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
|
||
; alt. 520 m; 22 Feb 2010; B.L. Fisher et al. leg.; CASENT0160480 (CASC). •2w.;
|
||
<normalizedToken id="9AF4CCBC7397989A22DDF149294D5ABA" originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken id="AA8CE0F510757C51EF98B5B1D24921CB" originalValue="Spéciale">Speciale</normalizedToken>
|
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Ambatovaky, Sandrangato River;
|
||
<geoCoordinate id="AD998C038A3D9C5178F4CDEEAD1F4157" degrees="16.7702" direction="south" orientation="latitude" precision="5" value="-16.7702">-16.7702</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2A42999D06E6A8FD760B510BD02D77A8" degrees="49.26638" direction="east" orientation="longitude" precision="1" value="49.26638">49.26638</geoCoordinate>
|
||
; alt. 470 m; 23 Feb 2010; B.L. Fisher et al. leg.; CASENT0161455 (CASC). •1w.;
|
||
<normalizedToken id="ED1EED3A367874C2AFC4BC7166449A42" originalValue="Réserve">Reserve</normalizedToken>
|
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<normalizedToken id="B0E54C4D3C4F00260C6CDB8DFF826DF4" originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato River;
|
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<geoCoordinate id="E3E11E01E474B6009D728EC8AE277288" degrees="16.8162" direction="south" orientation="latitude" precision="5" value="-16.8162">-16.8162</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="71E85703490A64F81442DB57B2325C51" degrees="49.29202" direction="east" orientation="longitude" precision="1" value="49.29202">49.29202</geoCoordinate>
|
||
; alt. 425 m; 25 Feb 2010; B.L. Fisher et al. leg.; CASENT0161854 (CASC). •2w.;
|
||
<normalizedToken id="F82D4544A8C747A911606EDE36EA6FA1" originalValue="Réserve">Reserve</normalizedToken>
|
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<normalizedToken id="C5F78383DC6DE549FFB17DC6262669E1" originalValue="Spéciale">Speciale</normalizedToken>
|
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Ambatovaky, Sandrangato River;
|
||
<geoCoordinate id="9F5C1B07E2D509B3EB40E53FC26923D7" degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
|
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,
|
||
<geoCoordinate id="4E2B7B6B1EA76772D3FDAE4DDBC0B098" degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
|
||
; alt. 400 m; 26 Feb 2010; B.L. Fisher et al. leg.; CASENT0162160 (CASC). •4w., 2s., 1q.;
|
||
<normalizedToken id="768F5EFFE1FD388CA3E2147438C7EC57" originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken id="2E9EA1026F9E8B4E1175D55E5275F212" originalValue="Spéciale">Speciale</normalizedToken>
|
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Ambatovaky, Sandrangato River;
|
||
<geoCoordinate id="A12EBB41864635EB371F3A16716CC93F" degrees="16.77274" direction="south" orientation="latitude" precision="1" value="-16.77274">-16.77274</geoCoordinate>
|
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,
|
||
<geoCoordinate id="46C804CCD19E376210AB592F6B0A5371" degrees="49.26551" direction="east" orientation="longitude" precision="1" value="49.26551">49.26551</geoCoordinate>
|
||
; alt. 450 m; 20 Feb 2010; B.L. Fisher et al. leg.; CASENT0162675, CASENT0162686, CASENT0162688, CASENT0162689 (CASC). •4w., 4s., 3q.;
|
||
<normalizedToken id="6B910CCABCEF751E69CAD715CE63FD3C" originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken id="885EB6C0746580E7608299B2628AED5A" originalValue="Spéciale">Speciale</normalizedToken>
|
||
Ambatovaky, Sandrangato River;
|
||
<geoCoordinate id="660D6B59B81B9EAFE3880F5203049D5F" degrees="16.80561" direction="south" orientation="latitude" precision="1" value="-16.80561">-16.80561</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="20E5127619EFE11D49B9D1FCB50552A7" degrees="49.29507" direction="east" orientation="longitude" precision="1" value="49.29507">49.29507</geoCoordinate>
|
||
; alt. 480 m; 27 Feb 2010; B.L. Fisher et al. leg.; CASENT0163009, CASENT0163019, CASENT0163021, CASENT0163027, CASENT0163032, CASENT0163040, CASENT0163073 (CASC). •2w.;
|
||
<normalizedToken id="9A02201DB07B15DE83DA3E66FF7866CF" originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken id="5F42203BA7B809EBEBAD32FB5BAD1B72" originalValue="Spéciale">Speciale</normalizedToken>
|
||
Ambatovaky, Sandrangato River;
|
||
<geoCoordinate id="D5A1C690AB0BD56322F4445AE267975C" degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4AEEEE88E3FC2E2327842481D0F18557" degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
|
||
; alt. 520 m; 22 Feb 2010; B.L. Fisher et al. leg.; CASENT0163856, CASENT0163945 (CASC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23C67CEDD78330CEDC97B3D91AF4BC3A" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="BA6675D90A5026970490B291A88BEAD4" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="524DDE41E71D7B56BEF8E83B2416249A" pageId="0" pageNumber="1">
|
||
<emphasis id="782CABCFE36462EC97B9134F0A3D3C3D" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="C920D11442E111902FD4BC1FFA5605C5" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
Large species: HL: 2.38-2.71 (2.55), HW: 2.17-2.54 (2.4), WL: 1.6-1.95 (1.76); propodeal spines very long (PSL: 0.48-0.59 (0.54)); head in full-face view trapezoid, widened posteriorly; sides of the head with sparse, relatively long, erect pilosity; frons with longitudinal, sparse, and directed outward rugae, interspaces with dense and fine rugulae, sculpture weakening posteriorly; first gastral tergite shagreened, at least on its basal part; body dark brown to black; inner hypostomal teeth distinct, triangular and thick, with rounded apex, closely spaced; outer hypostomal teeth distinct, low, lobe-like, with wide base and tops directed outward, smaller and thinner than inner hypostomal teeth.
|
||
<emphasis id="2214DF7AFB0077601705F4A9BCF2A129" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="2704D541811397B17E5CE59A75BEF0FF" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
Large species: HL: 0.85-1.03 (0.93), HW: 0.8-1.0 (0.89), WL: 1.21-1.47 (1.36); propodeal spines very long (PSL: 0.34-0.44 (0.39)); scape, when laid back, surpassing posterior head margin by one-third of its length; head sculpture rugoreticulate; mesosoma with dense rugoreticulation, sometimes dorsum with weaker sculpture, but never smooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75763D74E3E02D2BBA5682E58966C3E9" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="2DB6DB034384B0DB271DC9D3568AB099" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="9EB6536174AFA7C9E5DC9984FF0D0576" pageId="0" pageNumber="1">
|
||
<emphasis id="C88BB0645EB54FB316658E392524765A" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="19F21C386DFF6100A49346E9AE0B63DE" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 2.38-2.71 (2.55); HW: 2.17-2.54 (2.4); SL: 0.95-1.17 (1.08); EL: 0.24-0.3 (0.27); WL: 1.6-1.95 (1.76); PSL: 0.48-0.59 (0.54); MTL: 1.16-1.28 (1.23); PNW: 0.8-0.95 (0.9); PTW: 0.24-0.31 (0.28); PPW: 0.7-0.92 (0.83); CI: 91.3-98.5 (94.0); SI: 43.6-46.3 (45.2); PSLI: 19.8-22.6 (21.2); PPI: 31.0-36.0 (34.1); PNI: 35.5-38.8 (37.4); MTI: 49.5-53.8 (51.2).
|
||
<emphasis id="455DEDA546D7A1117D1D23EC5247B136" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="4A67F604BE986D7601C8DCC601C51E8C" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
In full-face view trapezoid, widened posteriorly (Fig.
|
||
<figureCitation id="D8B4059403B53DC0E1324260E84F075F" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27B</figureCitation>
|
||
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with sparse, relatively long, erect pilosity; whole head with moderately sparse, long, erect setae. Antennal scrobes absent. Occipital lobes shiny, and smooth at posterior part, anterior part with fine and dense rugulae; genae shiny, with fine and dense rugulae, sometimes posterior part smooth or with reduced sculpture; malar area and frons with longitudinal, sparse, and directed outward rugae, interspaces with dense and fine rugulae; lateral sides of head shiny, with dense and fine rugulae and thin, indistinct, longitudinal rugae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, shallow, and wide; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching mid-length of head; pilosity suberect to erect (Fig.
|
||
<figureCitation id="01A1C615C73456DE0A9F22302D20355D" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27B, D</figureCitation>
|
||
). Inner hypostomal teeth distinct, triangular, and thick, with rounded apex, closely spaced; outer hypostomal teeth distinct, low, lobe-like, with wide base and tops directed outward, smaller and thinner than inner hypostomal teeth (Fig.
|
||
<figureCitation id="C63A5ED92C7C9055528086285E9B31A7" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85C</figureCitation>
|
||
).
|
||
<emphasis id="6F6FD0AA3E9C18B680A07C68D95A1F91" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="C66E9B3CAC6C9C1AED3929046025B538" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum short, low, and evenly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines very long, massive basally, with acute apex; humeral area laterally weakly produced (Fig.
|
||
<figureCitation id="339A090C97280C9C86462365888F578B" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D</figureCitation>
|
||
). Surface shiny, with dense rugoreticulation, mesosomal dorsum and propodeum with weaker sculpture, but never smooth. Pilosity relatively dense, long, and erect (Fig.
|
||
<figureCitation id="829A71D7516A920812E108169632B89D" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
|
||
).
|
||
<emphasis id="8A5BDF1A06A4D337CE91B6FDDCDE9863" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="4014C13C6578E0DD02B9ED52C82A7123" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shagreened; peduncle relatively long and thin; node triangular with rounded apex, in rear view node dorsoventrally slightly depressed; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="CCA0CF51919A86B118A1C9D73535EA51" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
|
||
).
|
||
<emphasis id="13BEBE9BD410AD3FBAC15122E197E589" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="BD0A3A3FBAFFC3C7004E370050BDDAEA" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shagreened; in dorsal view sides with acute, and short to moderately short angular projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="36C566CA9C62DEF74A2E0938AF410309" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
|
||
).
|
||
<emphasis id="5AB44873B945566D6CCB573FE9EC7E30" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="55B1965191783040FE74999E647641B2" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
First gastral tergite shagreened, at least on its basal part; pilosity sparse, long and erect (Fig.
|
||
<figureCitation id="32BFDFD851CC970826466B1B7A3B1578" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
|
||
).
|
||
<emphasis id="F2AD6D60A93EEC2413C31CEE396F5518" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="69A2EF8ABBC0C5345B93CE8C97847920" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, dark brown to black (Fig.
|
||
<figureCitation id="341AE4BE5B6B48FAD218F8CBB3DDA655" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B861B8A8D1508BE63FC3D3816A9C9460" doi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
|
||
<paragraph id="F537971E2B53ED2C20D3CB729BF9A21F" pageId="0" pageNumber="1">
|
||
<emphasis id="C099EE5CE66024B6302A1FAC89CCF065" bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
|
||
<taxonomicName id="B2B8726F036D2CB0756D8BF15B720B0E" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
|
||
<emphasis id="480D0EA8E78D627FD2807FE55582B7BA" italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., full-face view (
|
||
<emphasis id="C6DB4D44F784C7820F2FA22C07FDF552" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis id="F5216DF83BE43A17A4DA3B8F0F8443E9" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis id="3853414C88C81AECEA7841DA309A73D4" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of paratype minor worker (CASENT0923223) and full-face view (
|
||
<emphasis id="D4ABE1B6F4630A8CD6B6F1D7179E63AA" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis id="41E9069B3A1B09CBB54269EC5412DECE" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis id="4A02AB61FFD494A08264D09C77ED9822" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of holotype major worker (CASENT0162231).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="278D3618FA4B3EE1746744BC4B8287E8" pageId="0" pageNumber="1">
|
||
<emphasis id="212825B68B1935CF7349F3D578C40566" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="B77CA2FA655DAEB41B67FA8CE6F0AE39" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.85-1.03 (0.93); HW: 0.8-1.0 (0.89); SL: 1.07-1.2 (1.16); EL: 0.16-0.21 (0.18); WL: 1.21-1.47 (1.36); PSL: 0.34-0.44 (0.39); MTL: 0.91-1.1 (1.01); PNW: 0.55-0.66 (0.61); PTW: 0.1-0.16 (0.13); PPW: 0.19-0.28 (0.23); CI: 93.3-101.0 (95.5); SI: 122.0-135.6 (130.7); PSLI: 40.0-44.9 (42.3); PPI: 50.5-60.2 (55.3); PNI: 66.3-70.4 (68.7); MTI: 111.2-117.0 (114.1).
|
||
<emphasis id="DBA9CA73C01158030C08A20C5DCFCE58" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="CC2D29499F7A57BEAF1524CBEE1C853A" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin convex; occipital carina narrow, weakly developed (Fig.
|
||
<figureCitation id="2694B1B5C95C3F1E5B4E64220AD855A8" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27A</figureCitation>
|
||
). Pilosity sparse, long, and erect. Sculpture variable, finely to strongly rugoreticulate on the whole surface; only centre of frons sometimes with sculpture reduced or smooth; antennal sockets with sparse carinae curved outward. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae present. Scape, when laid back, surpassing posterior head margin by one-third of its length; pilosity suberect to erect (Fig.
|
||
<figureCitation id="74905F5509C1A0A5AECBDF9335722715" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27A, C</figureCitation>
|
||
).
|
||
<emphasis id="FB35DEC58882A32A43A9CDBE9D2F00AA" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="DB87CF2011FC018E6D1D6E521824EF41" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, long, and slightly convex; promesonotal groove absent; metanotal groove very indistinct; propodeal spines very long, massive basally, with acute apex (Fig.
|
||
<figureCitation id="A3909F5F5F7764117F5D8B7D6AB29901" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C</figureCitation>
|
||
). Surface shiny, with dense rugoreticulation, sometimes dorsum with weaker sculpture, but never smooth. Pilosity sparse, long, and erect (Fig.
|
||
<figureCitation id="47A79CB89E41E5884111A8B3E863FA73" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
|
||
).
|
||
<emphasis id="1BA98266AE8BE3ED89418BC8177CAA17" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="CC3FBE8BD56DF0FEB8E4A4AF26C2C8EC" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Finely shagreened; peduncle long and thin; node triangular with rounded apex; with few long, erect setae (Fig.
|
||
<figureCitation id="8886AB0FE7BCD4ECB39E5766B3357772" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
|
||
).
|
||
<emphasis id="849E2CAB45871A4B374152D891BC400C" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="BDFE8584586987C8F614EEC00F0D68D5" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Moderately shagreened; moderately short, low, and convex; with few long, erect setae at the anterior edge (Fig.
|
||
<figureCitation id="729EAC09A9084157A40F01D95E558845" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
|
||
).
|
||
<emphasis id="0C4121F40A6C07D43B7EC0DDBD50093D" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="59791AA1CC06335319B8FE292AEC845C" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With few long, erect setae (Fig.
|
||
<figureCitation id="CE33BB1C5D96919938E291076BA6F5AB" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
|
||
).
|
||
<emphasis id="2DDC619A5243477BB7F4C2ED7D2ADF02" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="717B2D3F3A94310D165E603488FA58C4" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, dark brown to black (Fig.
|
||
<figureCitation id="F43E725E5F2101FA051E7119DC16EBBD" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F162629C002D9458731AAF1367F7D04B" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="267CF3A2607669DA870881B8D74AD826" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="FD71FA498BF260328D44ADD74F3BD962" pageId="0" pageNumber="1">Malagasy for dark, in reference to dark body colouration.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F9BE83647D4CFF087C55E8718C53EA7F" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="44D1AE93FF4892D197082EB6281F20AE" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="1D96B2D969797D70DF571B3F7B31CA97" pageId="0" pageNumber="1">The species was collected at between 400-1200 m in elevation, in rainforest, montane rainforest, and transitional humid forest. Nests were located in in rotten logs and stumps, and in rotten sticks on the ground.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9F753FC221D8C26F81917015BD2D5344" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="CD6D215A9F2CDA2B1B6FB456BAF71205" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="0E02E85B8266445EC70C136EB9304D6D" pageId="0" pageNumber="1">
|
||
This species is most similar to
|
||
<taxonomicName id="C97F9FD5B70889BAFCB0D5E9D7F8C443" lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
|
||
<emphasis id="4D2C5B333F4927BF1DF4A348DF97A0B9" italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="D97EFFC89EA6C5C72367E4E99A30F507" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="F9C06758BE9FF91280C9BEC809DDB5BF" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="9854DDA62FDB844307596311EB148897" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
|
||
<emphasis id="8C3654F6B96EBBD618183469A06213D3" italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName id="0C4F74C2D792F8A1B37A75D9EC51CC79" lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
|
||
<emphasis id="EBBF4068964C058CC42CA512F03DFE1D" italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
|
||
</taxonomicName>
|
||
by having distinct rugulae on at least half of the genae, tips of outer hypostomal teeth directed outward, never smooth katepisternum, densely shagreened surface of first gastral tergite, and darker body colouration.
|
||
<emphasis id="48F7D036A11A35ADC20FB84FA22A5E65" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="52E64333C3CA389468C821025D51775B" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="F9F8051C3E64E14E26236CB6C9F5EEAB" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
|
||
<emphasis id="F68BB74BA6823723FF8FA70072DDE5E2" italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName id="AC5A3A91EC3BA70B92C48E3423927B31" lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
|
||
<emphasis id="E32573B63B414824CEBA7AB2BF797EAC" italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
|
||
</taxonomicName>
|
||
by rugoreticulate head sculpture with never smooth genae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |