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<document id="C40DCBB70AB8EAD18D4F462F18ADC053" ID-DOI="10.5281/zenodo.201036" ID-GBIF-Dataset="e32d8614-7fee-4095-921d-fcd6c8dfe476" ID-ISSN="1175-5326" ID-Zenodo-Dep="201036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460272492680" checkinUser="plazi" docAuthor="Mendoza, Jose Christopher E. &amp; Ng, Peter K. L." docDate="2011" docId="E3508794B30DB92BAEC841DCFA84FDD0" docLanguage="en" docName="zt02809p066.pdf" docOrigin="Zootaxa 2809" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leelumnus Mendoza &amp; Ng, 2011, gen. nov." docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="1F69FFECB30CB929AE5F414BFFD4FF8F" masterDocTitle="Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae)" masterLastPageNumber="66" masterPageNumber="58" pageNumber="59" updateTime="1698247077057" updateUser="plazi">
<mods:mods id="564AF10E0A6C193BA4A4D3EA7820362B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="00569C787AD7E76AE02E4864AF053B55">Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae)</mods:title>
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<mods:name id="1A3A7291D7CFCBBEE2B0DCFAA7C0FD5C" type="personal">
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<mods:namePart id="94C2F7311B2E95C5C3408C0424F80C64">Mendoza, Jose Christopher E.</mods:namePart>
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<mods:name id="796D1F4C62BFBC09B9C64D303790D4D5" type="personal">
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<mods:namePart id="FC6E0083957EED24833E47621C67302F">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="2AF6EE1A688D0022994A35AE1720662D">text</mods:typeOfResource>
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<mods:title id="8485C1336ECBEF6530942AE60BAD9DB5">Zootaxa</mods:title>
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<mods:part id="9EBA8891B225236D6C99493EEC91C1CD">
<mods:date id="C3137B20472C949BE5BE11572D1A6C32">2011</mods:date>
<mods:detail id="1115E2D691437406F9B14A7A4B4F3B9C" type="volume">
<mods:number id="4774DB6B47530A9AF638D1C43D9004C4">2809</mods:number>
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<mods:classification id="297F744B79C099EB05E772AEF9103B02">journal article</mods:classification>
<mods:identifier id="477D2FA01D5BDDB2933AB9094E6A24ED" type="DOI">10.5281/zenodo.201036</mods:identifier>
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<mods:identifier id="47EBCA5C91ACCB711EF9E9EDF7353AC9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2A5A4E82974D7F78295CD184562D51E9" type="Zenodo-Dep">201036</mods:identifier>
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<treatment id="E3508794B30DB92BAEC841DCFA84FDD0" ID-DOI="http://doi.org/10.5281/zenodo.6184233" ID-GBIF-Taxon="119395388" ID-Zenodo-Dep="6184233" LSID="urn:lsid:plazi:treatment:E3508794B30DB92BAEC841DCFA84FDD0" httpUri="http://treatment.plazi.org/id/E3508794B30DB92BAEC841DCFA84FDD0" lastPageId="2" lastPageNumber="60" pageId="1" pageNumber="59">
<subSubSection id="23E36509B30DB928AEC841DCFE47FF3E" box="[151,403,151,177]" pageId="1" pageNumber="59" type="nomenclature">
<paragraph id="6B463682B30DB928AEC841DCFE47FF3E" blockId="1.[151,403,151,177]" box="[151,403,151,177]" pageId="1" pageNumber="59">
<heading id="300E81EEB30DB928AEC841DCFE47FF3E" bold="true" box="[151,403,151,177]" fontSize="11" level="1" pageId="1" pageNumber="59" reason="1">
<emphasis id="598DEA90B30DB928AEC841DCFE47FF3E" bold="true" box="[151,403,151,177]" pageId="1" pageNumber="59">
<taxonomicName id="ACF94D01B30DB928AEC841DCFECAFF3E" ID-CoL="7PMJ8" box="[151,286,151,177]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30DB928AEC841DCFECAFF3E" bold="true" box="[151,286,151,177]" italics="true" pageId="1" pageNumber="59">Leelumnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42BE57EBB30DB928AF7A41DCFE47FF3E" box="[293,403,151,177]" pageId="1" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="23E36509B30DB928AEC84191FCE2FF60" box="[151,822,217,240]" pageId="1" pageNumber="59" type="materials_examined">
<paragraph id="6B463682B30DB928AEC84191FCE2FF60" blockId="1.[151,822,217,240]" box="[151,822,217,240]" pageId="1" pageNumber="59">
<emphasis id="598DEA90B30DB928AEC84191FEF6FF7F" bold="true" box="[151,290,217,240]" pageId="1" pageNumber="59">
<typeStatus id="B4428820B30DB928AEC84191FF18FF60" box="[151,204,218,239]" pageId="1" pageNumber="59">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="ACF94D01B30DB928AF764192FE32FF60" box="[297,486,217,239]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="species" species="radium" status="sp. nov.">
<emphasis id="598DEA90B30DB928AF764192FE32FF60" box="[297,486,217,239]" italics="true" pageId="1" pageNumber="59">Leelumnus radium</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30DB928AFB24191FDE9FF60" bold="true" box="[493,573,218,239]" pageId="1" pageNumber="59">
<taxonomicNameLabel id="42BE57EBB30DB928AFB24191FDE9FF60" box="[493,573,218,239]" pageId="1" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="23E36509B30DB928AEC8405DFE61FD19" pageId="1" pageNumber="59" type="diagnosis">
<paragraph id="6B463682B30DB928AEC8405DFE61FD19" blockId="1.[151,1437,278,2031]" pageId="1" pageNumber="59">
<emphasis id="598DEA90B30DB928AEC8405DFEC5FEA0" bold="true" box="[151,273,278,303]" pageId="1" pageNumber="59">Diagnosis.</emphasis>
Carapace transversely subhexagonal, broader than long; regions poorly defined; dorsal surface convex, smooth, sparsely setose. Front moderately projecting, bilobed; anterior margins gently concave, granulate, mesial angles more advanced than lateral, no lateral lobule visible. Fronto-orbital margin, epistome and buccal field relatively narrow transversely; fronto-orbital width about 0.4 times maximum carapace width. Orbits relatively short, rounded; eyes with short ocular peduncle. Anterolateral margin granulate; divided into 4 teeth; first tooth smallest, connected to exorbital angle by broad, horizontal, granulate crest; subsequent teeth projecting laterally, curving anteriorly, second tooth broadly triangular, third and fourth teeth narrower, acute. Chelipeds distinctly unequal, external surface of male major chela smooth, glabrous. Ambulatory legs smooth, without granules on margins of all articles; densely setose on carpi, propodi and dactyli. Male thoracic sternum and abdomen relatively broad; male telson triangular. G1 slender, slightly sinuous; distal tip flared/widened, flattened, covered with several simple, stiff, spiniform setae.
</paragraph>
</subSubSection>
<subSubSection id="23E36509B30DB928AE9A43E8FE4AFCA9" pageId="1" pageNumber="59" type="etymology">
<paragraph id="6B463682B30DB928AE9A43E8FE4AFCA9" blockId="1.[151,1437,278,2031]" pageId="1" pageNumber="59">
<emphasis id="598DEA90B30DB928AE9A43E8FE98FD34" bold="true" box="[197,332,675,699]" pageId="1" pageNumber="59">Etymology.</emphasis>
The new genus is named in honor of Dr Lee Seng Tee. Through the Lee Foundation, he has been instrumental in helping the Raffles Museum and the National University of
<collectingCountry id="13EE7612B30DB928ADAD438DFBB0FD51" box="[1010,1124,710,734]" name="Singapore" pageId="1" pageNumber="59">Singapore</collectingCountry>
build a new natural history museum for the country. The genus name is formed by an arbitrary combination of his family name Lee and
<taxonomicName id="ACF94D01B30DB928AB0F43A0FF6AFCA8" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30DB928AB0F43A0FF6AFCA8" italics="true" pageId="1" pageNumber="59">Pilumnus</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="23E36509B30DB928AE9A4278FD16FB3D" pageId="1" pageNumber="59" type="discussion">
<paragraph id="6B463682B30DB928AE9A4278FD16FB3D" blockId="1.[151,1437,278,2031]" pageId="1" pageNumber="59">
<emphasis id="598DEA90B30DB928AE9A4278FEEDFCC4" bold="true" box="[197,313,819,843]" pageId="1" pageNumber="59">Remarks.</emphasis>
Although there is general consensus that the Pilumnoidea is a monophyletic entity (Ng
<emphasis id="598DEA90B30DB928AB4B427FFA99FCC4" box="[1300,1357,819,843]" italics="true" pageId="1" pageNumber="59">et al.</emphasis>
2008), the classification of the various families is not satisfactory. On the basis of the carapace and general features,
<taxonomicName id="ACF94D01B30DB928AB354213FF3CFC1C" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30DB928AB354213FF3CFC1C" italics="true" pageId="1" pageNumber="59">Leelumnus</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30DB928AEB04230FE8DFC1C" bold="true" box="[239,345,891,915]" pageId="1" pageNumber="59">
<taxonomicNameLabel id="42BE57EBB30DB928AEB04230FE8DFC1C" box="[239,345,891,915]" pageId="1" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is best classified in the
<taxonomicName id="ACF94D01B30DB928AC354231FC65FC1D" authority="Samouelle, 1819" authorityName="Samouelle" authorityYear="1819" box="[618,945,890,914]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="family">
Pilumnidae
<bibRefCitation id="0F684B73B30DB928ACAE4231FC65FC1D" author="Samouelle" box="[753,945,890,914]" pageId="1" pageNumber="59" refString="Samouelle, G. (1819) The Entomologist's Useful Compendium, or an Introduction to the Knowledge of the British Insects. London: Thomas Boys, 496 pp." type="book" year="1819">Samouelle, 1819</bibRefCitation>
</taxonomicName>
, primarily due to the presence of complete, well-defined longitudinal ridges on the endostome (endostomial crests), a relatively broader male abdomen with a semicircular telson, the G1 being long and sinuous, and the G2 is very short and sigmoid in shape (Ng 1998;
<bibRefCitation id="0F684B73B30DB928AB064289FF07FC71" author="Davie" pageId="1" pageNumber="59" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda - Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. W. (Eds.), Zoological Catalogue of Australia, Vol. 19.3 B. Melbourne: CSIRO Publishing. xiv + 641 pp." type="book" year="2002">Davie 2002</bibRefCitation>
; Ng
<emphasis id="598DEA90B30DB928AF5942A3FEEAFC71" box="[262,318,998,1023]" italics="true" pageId="1" pageNumber="59">et al.</emphasis>
2008).
<taxonomicName id="ACF94D01B30DB928AFCB42ADFDD9FC71" box="[404,525,998,1022]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30DB928AFCB42ADFDD9FC71" box="[404,525,998,1022]" italics="true" pageId="1" pageNumber="59">Leelumnus</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30DB928AC4A42ACFDAAFC70" bold="true" box="[533,638,999,1023]" pageId="1" pageNumber="59">
<taxonomicNameLabel id="42BE57EBB30DB928AC4A42ACFDAAFC70" box="[533,638,999,1023]" pageId="1" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is superficially similar to
<taxonomicName id="ACF94D01B30DB928ADFD42ADFA4CFC71" authority="Turkay &amp; Schuhmacher, 1985" authorityName="Turkay &amp; Schuhmacher" authorityYear="1985" box="[930,1432,998,1022]" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30DB928ADFD42ADFBE8FC71" box="[930,1084,998,1022]" italics="true" pageId="1" pageNumber="59">Latopilumnus</emphasis>
<bibRefCitation id="0F684B73B30DB928AA1C42ADFA4CFC71" author="Turkay" box="[1091,1432,998,1022]" pageId="1" pageNumber="59" refString="Turkay, M. &amp; Schuhmacher, H. (1985) Latopilumnus tubicolus n. gen. n. sp., eine neue korallenassoziierte Krabbe, die die Bildung einer Wohnhohle induzierte. (Crustacea: Decapoda: Pilumnidae). Senckenbergiana maritima, 17 (1 / 3), 55 - 63." type="journal article" year="1985">Türkay &amp; Schuhmacher, 1985</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="ACF94D01B30DB928AE994540FE69FBAD" authority="Ng, 2002" authorityName="Ng" authorityYear="2002" box="[198,445,1034,1059]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30DB928AE994540FE98FBAC" box="[198,332,1035,1059]" italics="true" pageId="1" pageNumber="59">Aniptumnus</emphasis>
Ng, 2002
</taxonomicName>
, in the following features: 1) the general form of the carapace, which is transversely subhexagonal, dorsally convex, relatively smooth and sparsely setose; 2) the absence of a lateral lobe on the frontal margin; 3) the quadridentate carapace anterolateral margin, where the two anterior teeth are relatively low and broad while the posterior two are more developed and acute; 4) the distinctly unequal chelipeds; and 5) the relatively broad male thoracic sternum and abdomen.
</paragraph>
</subSubSection>
<subSubSection id="23E36509B30DB92BAE9A45F5FBC1FEB0" lastPageId="2" lastPageNumber="60" pageId="1" pageNumber="59" type="reference_group">
<paragraph id="6B463682B30DB928AE9A45F5FC03F925" blockId="1.[151,1437,278,2031]" pageId="1" pageNumber="59">
<taxonomicName id="ACF94D01B30DB928AE9A45F5FEEAFB59" ID-CoL="7PMJ8" box="[197,318,1214,1238]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30DB928AE9A45F5FEEAFB59" box="[197,318,1214,1238]" italics="true" pageId="1" pageNumber="59">Leelumnus</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30DB928AF1945F4FE64FB58" bold="true" box="[326,432,1215,1239]" pageId="1" pageNumber="59">
<taxonomicNameLabel id="42BE57EBB30DB928AF1945F4FE64FB58" box="[326,432,1215,1239]" pageId="1" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="ACF94D01B30DB928AC1945F5FD18FB59" ID-CoL="7NPWK" box="[582,716,1214,1238]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30DB928AC1945F5FD18FB59" box="[582,716,1214,1238]" italics="true" pageId="1" pageNumber="59">Aniptumnus</emphasis>
</taxonomicName>
by 1) the absence of prominent granules on the distal border of the basis-ischium and the ventral border of the merus of P5 (
<figureCitation id="F3C22A07B30DB928AD1945A9FC5CFB75" box="[838,904,1250,1274]" captionStart="FIGURE 5" captionStartId="6.[151,250,1884,1907]" captionTargetBox="[195,1380,194,1825]" captionTargetId="figure@6.[185,1398,193,1867]" captionTargetPageId="6" captionText="FIGURE 5. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, left third maxilliped, external view (setae denuded); B, anterior thoracic sternum; C, posterior thoracic sternum and penis; D, abdomen with telson; E, coxa, basis-ischium and merus, and, F, propodus and dactylus of right P 5, dorsal view; G, left G 1, external view; distal tip of left G 1, I, external view, J, internal view; H, left G 2, external view. Scale bars: A H = 1.0 mm" httpUri="https://zenodo.org/record/201041/files/figure.png" pageId="1" pageNumber="59">Fig. 5</figureCitation>
E) (vs. present); 2) having the anterior margins of the frontal lobes concave (
<figureCitation id="F3C22A07B30DB928AFB9444DFDFEFA91" box="[486,554,1286,1310]" captionStart="FIGURE 1" captionStartId="3.[151,250,1885,1908]" captionTargetBox="[151,1436,195,1860]" captionTargetId="figure@3.[151,1436,195,1861]" captionTargetPageId="3" captionText="FIGURE 1. Leelumnus radium gen. et sp. nov. A, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore; habitus, recently ethanol-preserved coloration; B, paratype male, 14.6 × 9.8 mm (ZRC 2011.0039), colour in life, in-situ on encrusting community." httpUri="https://zenodo.org/record/201037/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
,
<figureCitation id="F3C22A07B30DB928AC65444DFD9CFA91" box="[570,584,1286,1310]" captionStart="FIGURE 2" captionStartId="4.[151,250,1978,2001]" captionTargetBox="[286,1301,193,1956]" captionTargetId="figure@4.[286,1301,193,1957]" captionTargetPageId="4" captionText="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="1" pageNumber="59">2</figureCitation>
A) (vs. straight or slightly convex); 3) having a relatively longer and more horizontal granulate crest between the first anterolateral tooth and exorbital angle (
<figureCitation id="F3C22A07B30DB928AA704461FBA5FACD" box="[1071,1137,1322,1346]" captionStart="FIGURE 1" captionStartId="3.[151,250,1885,1908]" captionTargetBox="[151,1436,195,1860]" captionTargetId="figure@3.[151,1436,195,1861]" captionTargetPageId="3" captionText="FIGURE 1. Leelumnus radium gen. et sp. nov. A, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore; habitus, recently ethanol-preserved coloration; B, paratype male, 14.6 × 9.8 mm (ZRC 2011.0039), colour in life, in-situ on encrusting community." httpUri="https://zenodo.org/record/201037/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
,
<figureCitation id="F3C22A07B30DB928AA214461FB58FACD" box="[1150,1164,1322,1346]" captionStart="FIGURE 2" captionStartId="4.[151,250,1978,2001]" captionTargetBox="[286,1301,193,1956]" captionTargetId="figure@4.[286,1301,193,1957]" captionTargetPageId="4" captionText="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="1" pageNumber="59">2</figureCitation>
A) (vs. shorter and more oblique); 4) the proportionately narrower fronto-orbital margin, epistome and buccal fields, short and rounded orbits with proportionately smaller eyes, the fronto-orbital width being about 0.4 times the maximum carapace width (
<figureCitation id="F3C22A07B30DB928AEB844DDFEFEFA21" box="[231,298,1430,1454]" captionStart="FIGURE 2" captionStartId="4.[151,250,1978,2001]" captionTargetBox="[286,1301,193,1956]" captionTargetId="figure@4.[286,1301,193,1957]" captionTargetPageId="4" captionText="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="1" pageNumber="59">Fig. 2</figureCitation>
A, B) (vs. wider, with relatively larger eyes, fronto-orbital width about 0.5 times maximum carapace width); 5) the relatively sharper, more angular internal infra-orbital tooth (
<figureCitation id="F3C22A07B30DB928AD8844F1FBCEFA5D" box="[983,1050,1466,1490]" captionStart="FIGURE 2" captionStartId="4.[151,250,1978,2001]" captionTargetBox="[286,1301,193,1956]" captionTargetId="figure@4.[286,1301,193,1957]" captionTargetPageId="4" captionText="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="1" pageNumber="59">Fig. 2</figureCitation>
B) (vs. more rounded); 6) the relatively low orbital margins (
<figureCitation id="F3C22A07B30DB928AF8A4495FDCEFA79" box="[469,538,1502,1526]" captionStart="FIGURE 2" captionStartId="4.[151,250,1978,2001]" captionTargetBox="[286,1301,193,1956]" captionTargetId="figure@4.[286,1301,193,1957]" captionTargetPageId="4" captionText="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="1" pageNumber="59">Fig. 2</figureCitation>
A, B) (vs. raised, appearing cristate); 6) the more triangular male telson with almost straight lateral margins (
<figureCitation id="F3C22A07B30DB928AC574749FD8EF995" box="[520,602,1538,1562]" captionStart="FIGURE 2" captionStartId="4.[151,250,1978,2001]" captionTargetBox="[286,1301,193,1956]" captionTargetId="figure@4.[286,1301,193,1957]" captionTargetPageId="4" captionText="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="1" pageNumber="59">Figs. 2</figureCitation>
C, 5D) (vs. semicircular with lateral margins rounded); 7) the distinctly flared/widened and flattened distal tip of the G1 (
<figureCitation id="F3C22A07B30DB928ACE4476DFD28F9B1" box="[699,764,1574,1598]" captionStart="FIGURE 5" captionStartId="6.[151,250,1884,1907]" captionTargetBox="[195,1380,194,1825]" captionTargetId="figure@6.[185,1398,193,1867]" captionTargetPageId="6" captionText="FIGURE 5. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, left third maxilliped, external view (setae denuded); B, anterior thoracic sternum; C, posterior thoracic sternum and penis; D, abdomen with telson; E, coxa, basis-ischium and merus, and, F, propodus and dactylus of right P 5, dorsal view; G, left G 1, external view; distal tip of left G 1, I, external view, J, internal view; H, left G 2, external view. Scale bars: A H = 1.0 mm" httpUri="https://zenodo.org/record/201041/files/figure.png" pageId="1" pageNumber="59">Fig. 5</figureCitation>
G, IJ) (vs. blunt, but not flared or flattened); and 8) the proportionately smaller vulvae which do not reach sternal suture 5/6 and only have a small sternal cover (
<figureCitation id="F3C22A07B30DB928AB534701FA9AF9ED" box="[1292,1358,1610,1634]" captionStart="FIGURE 4" captionStartId="5.[151,250,1938,1961]" captionTargetBox="[151,1436,1354,1917]" captionTargetId="figure@5.[151,1436,1354,1918]" captionTargetPageId="5" captionText="FIGURE 4. Female thoracic sternum and vulvae. A, Leelumnus radium gen. et sp. nov., paratype, 12.7 × 8.0 mm (ZRC 2011.0037), Singapore; B, Aniptumnus quadridentatus (De Man, 1895), 11.0 × 7.1 mm (ZRC 1989.3649 3660), Singapore." httpUri="https://zenodo.org/record/201040/files/figure.png" pageId="1" pageNumber="59">Fig. 4</figureCitation>
A) (vs. relatively larger vulvae, touching sternite 5 with a large sternal cover;
<figureCitation id="F3C22A07B30DB928ADE84725FC2FF909" box="[951,1019,1646,1670]" captionStart="FIGURE 4" captionStartId="5.[151,250,1938,1961]" captionTargetBox="[151,1436,1354,1917]" captionTargetId="figure@5.[151,1436,1354,1918]" captionTargetPageId="5" captionText="FIGURE 4. Female thoracic sternum and vulvae. A, Leelumnus radium gen. et sp. nov., paratype, 12.7 × 8.0 mm (ZRC 2011.0037), Singapore; B, Aniptumnus quadridentatus (De Man, 1895), 11.0 × 7.1 mm (ZRC 1989.3649 3660), Singapore." httpUri="https://zenodo.org/record/201040/files/figure.png" pageId="1" pageNumber="59">Fig. 4</figureCitation>
B) (see
<bibRefCitation id="0F684B73B30DB928AA0B4725FAE4F909" author="Garth" box="[1108,1328,1646,1670]" pageId="1" pageNumber="59" refString="Garth, J. S. &amp; Kim, H. S. (1983) Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908 - 1909. Journal of Natural History, 17, 663 - 729." type="journal article" year="1983">Garth &amp; Kim 1983</bibRefCitation>
: 706, fig. 11; Ng 2002: 213, fig. 1; Ng &amp;
<bibRefCitation id="0F684B73B30DB928AFA747D9FDADF925" author="Clark" box="[504,633,1682,1706]" pageId="1" pageNumber="59" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilummnidae) with larval comparisons. Journal of Natural History, 42 (9 - 12), 885 - 912." type="journal article" year="2008">Clark 2008</bibRefCitation>
: 901, 904, figs. 13, 14, 1921).
</paragraph>
<paragraph id="6B463682B30DB928AE9A47FDFDEAF861" blockId="1.[151,1437,278,2031]" pageId="1" pageNumber="59">
<taxonomicName id="ACF94D01B30DB928AE9A47FDFEEAF941" ID-CoL="7PMJ8" box="[197,318,1718,1742]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30DB928AE9A47FDFEEAF941" box="[197,318,1718,1742]" italics="true" pageId="1" pageNumber="59">Leelumnus</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30DB928AF1947FCFE65F940" bold="true" box="[326,433,1719,1743]" pageId="1" pageNumber="59">
<taxonomicNameLabel id="42BE57EBB30DB928AF1947FCFE65F940" box="[326,433,1719,1743]" pageId="1" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="ACF94D01B30DB928AC1647FDFD37F941" ID-CoL="7PMH7" box="[585,739,1718,1742]" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30DB928AC1647FDFD37F941" box="[585,739,1718,1742]" italics="true" pageId="1" pageNumber="59">Latopilumnus</emphasis>
</taxonomicName>
in the absence of prominent granules on the distal margin of the basis-ischium and the ventral margin of the merus of the last ambulatory leg (
<figureCitation id="F3C22A07B30DB928AA7B4791FBB2F97D" box="[1060,1126,1754,1778]" captionStart="FIGURE 5" captionStartId="6.[151,250,1884,1907]" captionTargetBox="[195,1380,194,1825]" captionTargetId="figure@6.[185,1398,193,1867]" captionTargetPageId="6" captionText="FIGURE 5. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, left third maxilliped, external view (setae denuded); B, anterior thoracic sternum; C, posterior thoracic sternum and penis; D, abdomen with telson; E, coxa, basis-ischium and merus, and, F, propodus and dactylus of right P 5, dorsal view; G, left G 1, external view; distal tip of left G 1, I, external view, J, internal view; H, left G 2, external view. Scale bars: A H = 1.0 mm" httpUri="https://zenodo.org/record/201041/files/figure.png" pageId="1" pageNumber="59">Fig. 5</figureCitation>
E). In addition to the differences in the condition of the front, anterolateral margin, orbits, male telson and vulvae noted above for
<taxonomicName id="ACF94D01B30DB928AB6B47B4FE92F8B4" ID-CoL="7NPWK" authority="Leelumnus" authorityName="Leelumnus" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30DB928AB6B47B4FF6AF8B4" italics="true" pageId="1" pageNumber="59">Aniptumnus</emphasis>
,
<emphasis id="598DEA90B30DB928AE924668FE92F8B4" box="[205,326,1827,1851]" italics="true" pageId="1" pageNumber="59">Leelumnus</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30DB928AF104668FE6EF8B4" bold="true" box="[335,442,1827,1851]" pageId="1" pageNumber="59">
<taxonomicNameLabel id="42BE57EBB30DB928AF104668FE6EF8B4" box="[335,442,1827,1851]" pageId="1" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="ACF94D01B30DB928AC0C4668FD39F8B4" ID-CoL="7PMH7" box="[595,749,1827,1851]" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30DB928AC0C4668FD39F8B4" box="[595,749,1827,1851]" italics="true" pageId="1" pageNumber="59">Latopilumnus</emphasis>
</taxonomicName>
in having 1) the external surface of the major chela smooth (
<figureCitation id="F3C22A07B30DB928AEC0460DFF34F8D1" box="[159,224,1862,1886]" captionStart="FIGURE 3" captionStartId="5.[151,250,1220,1243]" captionTargetBox="[283,1304,193,1198]" captionTargetId="figure@5.[283,1304,193,1199]" captionTargetPageId="5" captionText="FIGURE 3. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, major chela, external view; B, minor chela, external view." httpUri="https://zenodo.org/record/201039/files/figure.png" pageId="1" pageNumber="59">Fig. 3</figureCitation>
A) (vs. prominently granular); 2) the dorsal margin of the ambulatory meri smooth (
<figureCitation id="F3C22A07B30DB928AAD2460DFB1AF8D1" box="[1165,1230,1862,1886]" captionStart="FIGURE 5" captionStartId="6.[151,250,1884,1907]" captionTargetBox="[195,1380,194,1825]" captionTargetId="figure@6.[185,1398,193,1867]" captionTargetPageId="6" captionText="FIGURE 5. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, left third maxilliped, external view (setae denuded); B, anterior thoracic sternum; C, posterior thoracic sternum and penis; D, abdomen with telson; E, coxa, basis-ischium and merus, and, F, propodus and dactylus of right P 5, dorsal view; G, left G 1, external view; distal tip of left G 1, I, external view, J, internal view; H, left G 2, external view. Scale bars: A H = 1.0 mm" httpUri="https://zenodo.org/record/201041/files/figure.png" pageId="1" pageNumber="59">Fig. 5</figureCitation>
E) (vs. granular or denticulate); 3) the ambulatory carpi without a distinct granulate, submedian, longitudinal ridge (vs. present); and 4) the distal tip of the G1 distinctly flared/widened and flattened (
<figureCitation id="F3C22A07B30DB928AD2946C5FC63F829" box="[886,951,1934,1958]" captionStart="FIGURE 5" captionStartId="6.[151,250,1884,1907]" captionTargetBox="[195,1380,194,1825]" captionTargetId="figure@6.[185,1398,193,1867]" captionTargetPageId="6" captionText="FIGURE 5. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, left third maxilliped, external view (setae denuded); B, anterior thoracic sternum; C, posterior thoracic sternum and penis; D, abdomen with telson; E, coxa, basis-ischium and merus, and, F, propodus and dactylus of right P 5, dorsal view; G, left G 1, external view; distal tip of left G 1, I, external view, J, internal view; H, left G 2, external view. Scale bars: A H = 1.0 mm" httpUri="https://zenodo.org/record/201041/files/figure.png" pageId="1" pageNumber="59">Fig. 5</figureCitation>
G, IJ) (vs. tapering to a sharp point, sometimes recurved) (see
<bibRefCitation id="0F684B73B30DB928AF2046F9FD82F845" author="Garth" box="[383,598,1970,1994]" pageId="1" pageNumber="59" refString="Garth, J. S. &amp; Kim, H. S. (1983) Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908 - 1909. Journal of Natural History, 17, 663 - 729." type="journal article" year="1983">Garth &amp; Kim 1983</bibRefCitation>
: 709, fig. 12;
<bibRefCitation id="0F684B73B30DB928ACB046F9FBE9F845" author="Turkay" box="[751,1085,1970,1994]" pageId="1" pageNumber="59" refString="Turkay, M. &amp; Schuhmacher, H. (1985) Latopilumnus tubicolus n. gen. n. sp., eine neue korallenassoziierte Krabbe, die die Bildung einer Wohnhohle induzierte. (Crustacea: Decapoda: Pilumnidae). Senckenbergiana maritima, 17 (1 / 3), 55 - 63." type="journal article" year="1985">Türkay &amp; Schuhmacher 1985</bibRefCitation>
: 56, 58, figs. 1, 2, pl. 1, figs. 1 4; Ng &amp;
<bibRefCitation id="0F684B73B30DB928AEA5469DFEAFF861" author="Clark" box="[250,379,2006,2030]" pageId="1" pageNumber="59" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilummnidae) with larval comparisons. Journal of Natural History, 42 (9 - 12), 885 - 912." type="journal article" year="2008">Clark 2008</bibRefCitation>
: 886, figs. 112).
</paragraph>
<paragraph id="6B463682B30EB92BAE9A41DCFBC1FEB0" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="60">
The relatively narrow front and general carapace features of
<taxonomicName id="ACF94D01B30EB92BAD2841D3FC24FF3F" ID-CoL="7PMJ8" box="[887,1008,152,176]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="598DEA90B30EB92BAD2841D3FC24FF3F" box="[887,1008,152,176]" italics="true" pageId="2" pageNumber="60">Leelumnus</emphasis>
</taxonomicName>
<emphasis id="598DEA90B30EB92BADA641D3FBB7FF3F" bold="true" box="[1017,1123,152,176]" pageId="2" pageNumber="60">
<taxonomicNameLabel id="42BE57EBB30EB92BADA641D3FBB7FF3F" box="[1017,1123,152,176]" pageId="2" pageNumber="60" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
superficially resemble species of the Indo-Pacific pilumnoid genus
<taxonomicName id="ACF94D01B30EB92BAC3B41F6FC4CFF5B" ID-CoL="7PS4H" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[612,920,188,212]" class="Malacostraca" family="Galenidae" genus="Parapanope" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30EB92BAC3B41F6FD3BFF5B" box="[612,751,189,212]" italics="true" pageId="2" pageNumber="60">Parapanope</emphasis>
De
<bibRefCitation id="0F684B73B30EB92BAD4341F6FC4CFF5B" author="Man" box="[796,920,188,212]" pageId="2" pageNumber="60" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere. 8 (4), 485 - 609, pls. 12 - 14 [vol. 9, 1897]." type="journal article" year="1895">Man, 1895</bibRefCitation>
</taxonomicName>
. However, members of this genus do not have endostomial crests, the anterolateral teeth are broader and more lobiform, the male abdomen is relatively more slender with the telson more elongate, and the G1 is more sinuous (
<bibRefCitation id="0F684B73B30EB92BADC7404FFBFFFE93" author="Guinot" box="[920,1067,260,284]" pageId="2" pageNumber="60" refString="Guinot, D. (1985) Revision du genre Parapanope De Man, 1895 (Crustacea Decapoda Brachyura), avec description de trois especes nouvelles. Bulletin du Museum national d'Histoire naturelle, Paris, (4) 7 (A 3), 677 - 707, pls. 1 - 4." type="journal article" year="1985">Guinot 1985</bibRefCitation>
;
<bibRefCitation id="0F684B73B30EB92BAA67404FFAC7FE93" author="Guinot" box="[1080,1299,260,285]" pageId="2" pageNumber="60" refString="Guinot, D. &amp; Ng, P. K. L. (1988) Observations nouvelles sur Parapanope singaporensis Ng et Guinot, 1985, synonyme de P." type="book chapter" year="1988">Guinot &amp; Ng 1988</bibRefCitation>
). Moreover,
<taxonomicName id="ACF94D01B30EB92BAEC84062FEF6FECF" ID-CoL="7PS4H" box="[151,290,297,320]" class="Malacostraca" family="Galenidae" genus="Parapanope" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="598DEA90B30EB92BAEC84062FEF6FECF" box="[151,290,297,320]" italics="true" pageId="2" pageNumber="60">Parapanope</emphasis>
</taxonomicName>
is currently placed in the
<taxonomicName id="ACF94D01B30EB92BAC1B406CFBC5FEB0" ID-CoL="7NJGM" authority="Alcock, 1898 (Ng et al. 2008)" authorityName="Alcock, 1898 (Ng et al." authorityYear="2008" box="[580,1041,295,320]" class="Malacostraca" family="Galenidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="family">
Galenidae
<bibRefCitation id="0F684B73B30EB92BACE3406CFC87FEB0" author="Alcock" box="[700,851,295,319]" pageId="2" pageNumber="60" refString="Alcock, A. (1898) Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The family Xanthidae. Journal of the Asiatic Society of Bengal, Calcutta, 67 (1), 67 - 233." type="journal article" year="1898">Alcock, 1898</bibRefCitation>
(Ng
<emphasis id="598DEA90B30EB92BADD24062FC11FECF" box="[909,965,296,320]" italics="true" pageId="2" pageNumber="60">et al.</emphasis>
2008)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="23E36509B30EB92BAE9A4000FA84FDD0" pageId="2" pageNumber="60" type="materials_examined">
<paragraph id="6B463682B30EB92BAE9A4000FA84FDD0" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="60">
<emphasis id="598DEA90B30EB92BAE9A4000FE0CFEEB" bold="true" box="[197,472,331,356]" pageId="2" pageNumber="60">Comparative material.</emphasis>
<taxonomicName id="ACF94D01B30EB92BAFBE4007FC34FEEB" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[481,992,332,356]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="598DEA90B30EB92BAFBE4007FCCEFEEB" box="[481,794,332,356]" italics="true" pageId="2" pageNumber="60">Aniptumnus quadridentatus</emphasis>
(De
<bibRefCitation id="0F684B73B30EB92BAD094006FC03FEEB" author="Man" box="[854,983,332,356]" pageId="2" pageNumber="60" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere. 8 (4), 485 - 609, pls. 12 - 14 [vol. 9, 1897]." type="journal article" year="1895">Man, 1895</bibRefCitation>
)
</taxonomicName>
:
<specimenCount id="7DFFFD0BB30EB92BADB24007FB91FEEB" box="[1005,1093,332,356]" pageId="2" pageNumber="60" type="male">6 males</specimenCount>
(6.7 ×
<quantity id="AC019B67B30EB92BAAC14007FB23FEEB" box="[1182,1271,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="60" unit="mm" value="4.4">4.4 mm</quantity>
14.6 × 10.0 mm),
<specimenCount id="7DFFFD0BB30EB92BAE844024FE9DFE08" box="[219,329,367,391]" pageId="2" pageNumber="60" type="female">6 females</specimenCount>
(7.9 ×
<quantity id="AC019B67B30EB92BAFC04024FE22FE08" box="[415,502,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="60" unit="mm" value="5.3">5.3 mm</quantity>
11.0 ×
<quantity id="AC019B67B30EB92BAC3A4024FD6AFE08" box="[613,702,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="60" unit="mm" value="7.1">7.1 mm</quantity>
) (
<collectionCode id="0DE8AE47B30EB92BAC894024FCDAFE08" box="[726,782,367,391]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
1989.36493660), from barnacle clumps on kelong pole, with mussels, Lim Chu Kang mangroves,
<collectingCountry id="13EE7612B30EB92BAC2740DFFD39FE23" box="[632,749,404,428]" name="Singapore" pageId="2" pageNumber="60">Singapore</collectingCountry>
.
<taxonomicName id="ACF94D01B30EB92BACA640DFFB3CFE23" authority="Ng &amp; Clark, 2008" authorityName="Ng &amp; Clark" authorityYear="2008" box="[761,1256,404,429]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="598DEA90B30EB92BACA640DFFBD9FE23" box="[761,1037,404,428]" italics="true" pageId="2" pageNumber="60">Aniptumnus vietnamicus</emphasis>
Ng &amp;
<bibRefCitation id="0F684B73B30EB92BAA0140DFFB3CFE23" author="Clark" box="[1118,1256,404,428]" pageId="2" pageNumber="60" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilummnidae) with larval comparisons. Journal of Natural History, 42 (9 - 12), 885 - 912." type="journal article" year="2008">Clark, 2008</bibRefCitation>
</taxonomicName>
:
<typeStatus id="B4428820B30EB92BAAAA40DFFA89FE23" box="[1269,1373,404,428]" pageId="2" pageNumber="60" type="holotype">Holotype</typeStatus>
male (7.9 ×
<quantity id="AC019B67B30EB92BAEBB40FCFEE8FE40" box="[228,316,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="60" unit="mm" value="5.7">5.7 mm</quantity>
) (
<collectionCode id="0DE8AE47B30EB92BAF0B40FCFE58FE40" box="[340,396,439,463]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
1970.1.10.1),
<typeStatus id="B4428820B30EB92BAC6C40F3FD47FE5F" box="[563,659,440,464]" pageId="2" pageNumber="60" type="paratype">paratype</typeStatus>
female (8.8 ×
<quantity id="AC019B67B30EB92BAD6240FCFC42FE40" box="[829,918,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="60" unit="mm" value="6.5">6.5 mm</quantity>
) (
<collectionCode id="0DE8AE47B30EB92BADF240FCFC31FE40" box="[941,997,439,463]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
1970.1.10.2), Nhatrang Bay,
<collectingCountry id="13EE7612B30EB92BAB6840FCFA4DFE40" box="[1335,1433,439,463]" name="Vietnam" pageId="2" pageNumber="60">Vietnam</collectingCountry>
, coll. R. Serène,
<date id="1F471042B30EB92BAF154097FE35FE7B" box="[330,481,476,500]" pageId="2" pageNumber="60" value="1966-04-10">10 Apr. 1966</date>
.
<taxonomicName id="ACF94D01B30EB92BAFB34097FC06FE7B" authority="Ng &amp; Clark, 2008: 2" authorityName="Ng &amp; Clark" authorityPageNumber="2" authorityYear="2008" box="[492,978,476,501]" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="conicus">
<emphasis id="598DEA90B30EB92BAFB34097FD36FE7B" box="[492,738,476,500]" italics="true" pageId="2" pageNumber="60">Latopilumnus conicus</emphasis>
Ng &amp;
<bibRefCitation id="0F684B73B30EB92BAD6F4097FC6DFE7B" author="Clark" box="[816,953,476,500]" pageId="2" pageNumber="60" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilummnidae) with larval comparisons. Journal of Natural History, 42 (9 - 12), 885 - 912." type="journal article" year="2008">Clark, 2008</bibRefCitation>
:
<specimenCount id="7DFFFD0BB30EB92BAD9B4097FC06FE7B" box="[964,978,476,500]" pageId="2" pageNumber="60" type="male">2</specimenCount>
</taxonomicName>
male
<typeStatus id="B4428820B30EB92BAA484096FB56FE7A" box="[1047,1154,477,501]" pageId="2" pageNumber="60" type="paratype">paratypes</typeStatus>
(3.3 ×
<quantity id="AC019B67B30EB92BAA8B4097FAF8FE7B" box="[1236,1324,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="60" unit="mm" value="2.3">2.3 mm</quantity>
, 3.9 ×
<quantity id="AC019B67B30EB92BAB264097FF13FD98" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="60" unit="mm" value="2.8">2.8 mm</quantity>
),
<specimenCount id="7DFFFD0BB30EB92BAE8340B4FE9EFD98" box="[220,330,511,535]" pageId="2" pageNumber="60" type="female">2 females</specimenCount>
(4.8 ×
<quantity id="AC019B67B30EB92BAFFD40B4FE2FFD98" box="[418,507,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="60" unit="mm" value="3.3">3.3 mm</quantity>
, 5.0 ×
<quantity id="AC019B67B30EB92BAC1140B4FD73FD98" box="[590,679,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="60" unit="mm" value="3.4">3.4 mm</quantity>
) (
<collectionCode id="0DE8AE47B30EB92BACE040B4FD23FD98" box="[703,759,511,535]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
2003.0343), Conic Island Cave, 22º21´54˝N, 114º23´22˝E,
<collectingCountry id="13EE7612B30EB92BAEC8436EFEF5FDB2" box="[151,289,549,573]" name="Hong Kong" pageId="2" pageNumber="60">Hong Kong</collectingCountry>
, coll.
<date id="1F471042B30EB92BAF33436FFDDEFDB3" box="[364,522,548,572]" pageId="2" pageNumber="60" value="2002-10-25">25 Oct. 2002</date>
.
<taxonomicName id="ACF94D01B30EB92BAC47436FFC0BFDB3" authority="De Man, 1914" authorityName="De Man" authorityYear="1914" box="[536,991,548,572]" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="malardi">
<emphasis id="598DEA90B30EB92BAC47436FFCC0FDB3" box="[536,788,548,572]" italics="true" pageId="2" pageNumber="60">Latopilumnus malardi</emphasis>
(De
<bibRefCitation id="0F684B73B30EB92BAD0B436EFC02FDB3" author="Man" box="[852,982,548,572]" pageId="2" pageNumber="60" refString="Man, J. G. De (1914) Description de deux especes nouvelles du Pilumnus Leach et d'une jeune femelle du Pil. longicornis Hilgd., decouvertes dans des coquilles vides de balanes. Bulletin du Societe zoologie Francais, 39, 330 - 343." type="journal article" year="1914">Man, 1914</bibRefCitation>
)
</taxonomicName>
:
<typeStatus id="B4428820B30EB92BADB2436FFB50FDB3" box="[1005,1156,548,572]" pageId="2" pageNumber="60" type="paralectotype">paralectotype</typeStatus>
female (7.4 ×
<quantity id="AC019B67B30EB92BAB66436FFA40FDB3" box="[1337,1428,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="60" unit="mm" value="5.2">5.2 mm</quantity>
) (RMNH-D2187), in clumps of
<taxonomicName id="ACF94D01B30EB92BAFAB4303FC57FDD0" authority="Cirripedia" authorityName="Cirripedia" box="[500,899,583,608]" class="Maxillopoda" family="Balanidae" genus="Balanus" kingdom="Animalia" order="Sessilia" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="tintinnabulum">
<emphasis id="598DEA90B30EB92BAFAB4303FD20FDEF" box="[500,756,584,608]" italics="true" pageId="2" pageNumber="60">Balanus tintinnabulum</emphasis>
(Cirripedia)
</taxonomicName>
,
<collectingCountry id="13EE7612B30EB92BADD2430CFBC2FDD0" box="[909,1046,583,607]" name="Madagascar" pageId="2" pageNumber="60">Madagascar</collectingCountry>
, coll. L.M. Malard, no date.
</paragraph>
</subSubSection>
</treatment>
</document>