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<document id="B81F8306B1822E5A2AD1ABFDBD357747" ID-CLB-Dataset="57168" ID-DOI="10.31610/zsr/2020.29.1.17" ID-GBIF-Dataset="2a1f83d5-8801-4113-97a4-1b396a462e28" ID-ISSN="2410-0226" ID-Zenodo-Dep="10124220" ID-ZooBank="urn:lsid:zoobank.org:pub:F21075F1-3741-4670-B69D-EF178CF5D765" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688480101663" checkinUser="felipe" docAuthor="Kaplin, V. G." docDate="2020" docId="96448788FF8D487346D9F9DDFDF8F9C0" docLanguage="en" docName="ZoosystRoss.29.1.17-22.pdf" docOrigin="Zoosystematica Rossica (Zoosyst. Rossica) 29 (1)" docSource="http://dx.doi.org/10.31610/zsr/2020.29.1.17" docStyle="DocumentStyle:8B7E50B51175314E898B46063364B751.1:ZoosystRoss.2019-.journal_article" docStyleId="8B7E50B51175314E898B46063364B751" docStyleName="ZoosystRoss.2019-.journal_article" docStyleVersion="1" docTitle="Petrobiellus sachalinensis Kaplin 2020, sp. nov." docType="treatment" docUuid="806355BC-A68A-4DC0-BD72-3ECDB81CC636" docUuidSource="ZooBank" docVersion="7" lastPageNumber="22" masterDocId="6A7DFFF0FF8C48764649FFBDFF86FF87" masterDocTitle="A new species of bristletails of the genus Petrobiellus (Microcoryphia: Machilidae) from Sakhalin" masterLastPageNumber="22" masterPageNumber="17" pageNumber="18" updateTime="1700050020898" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E02283BCECF4D8B36531450FEB0ECBE9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="980781BC4A7EF94EC21AC73CB25B26A1">
<mods:title id="37270BBC2D9A00E53A590DFD6731DCD6">A new species of bristletails of the genus Petrobiellus (Microcoryphia: Machilidae) from Sakhalin</mods:title>
</mods:titleInfo>
<mods:name id="14DFCB6B1C6B8DAE3051D61A409BECBF" type="personal">
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<mods:namePart id="D5542E1BD929C3FD36C58F061504D9EA">Kaplin, V. G.</mods:namePart>
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<mods:title id="B9DC08C1650F7E19E4EF12F5E7044131">Zoosystematica Rossica</mods:title>
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<mods:part id="164479682CA3F2DC57F6C18D1E7B228A">
<mods:date id="61C0682F6FC397B353AFAB49FB3D46C9">2020</mods:date>
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<mods:title id="C354F1FA83792AB751DF8F87510E31C9">Zoosyst. Rossica</mods:title>
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<mods:number id="FB51B4755DB2CCCEF43F8FEE15FE7DA5">2020-04-14</mods:number>
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<mods:number id="DAC402443700CBD36DEE27128F74BCA1">29</mods:number>
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<mods:url id="A25DCE487F485738ED45674FFB1107E0">http://dx.doi.org/10.31610/zsr/2020.29.1.17</mods:url>
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<mods:classification id="3B83CB88F275E1F9019163E7F80BCF96">journal article</mods:classification>
<mods:identifier id="6BFD40138ECAF84BE2BD4F11554C58A1" type="CLB-Dataset">57168</mods:identifier>
<mods:identifier id="906CBB889D2FC7E97CB387D54A5CF774" type="DOI">10.31610/zsr/2020.29.1.17</mods:identifier>
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<treatment id="96448788FF8D487346D9F9DDFDF8F9C0" ID-DOI="http://doi.org/10.5281/zenodo.8114970" ID-GBIF-Taxon="210811786" ID-Zenodo-Dep="8114970" LSID="urn:lsid:zoobank.org:act:806355BC-A68A-4DC0-BD72-3ECDB81CC636" httpUri="http://treatment.plazi.org/id/96448788FF8D487346D9F9DDFDF8F9C0" lastPageId="5" lastPageNumber="22" pageId="1" pageNumber="18">
<subSubSection id="56F76515FF8D487746D9F9DDFDDEF9FC" box="[144,600,1632,1659]" pageId="1" pageNumber="18" type="nomenclature">
<paragraph id="1E52369EFF8D487746D9F9DDFDDEF9FC" blockId="1.[144,600,1632,1659]" box="[144,600,1632,1659]" pageId="1" pageNumber="18">
<heading id="451A81F2FF8D487746D9F9DDFDDEF9FC" box="[144,600,1632,1659]" fontSize="11" level="2" pageId="1" pageNumber="18" reason="2">
<emphasis id="2C99EA8CFF8D487746D9F9DDFDDEF9FC" bold="true" box="[144,600,1632,1659]" pageId="1" pageNumber="18">
<taxonomicName id="D9ED4D1DFF8D487746D9F9DDFE6BF9FC" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[144,493,1632,1659]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8D487746D9F9DDFE6BF9FC" bold="true" box="[144,493,1632,1659]" italics="true" pageId="1" pageNumber="18">Petrobiellus sachalinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37AA57F7FF8D487747BDF9DCFDDEF9FC" box="[500,600,1633,1659]" pageId="1" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="56F76515FF8D487746D9F92EFEA4F92A" box="[144,290,1683,1709]" pageId="1" pageNumber="18" type="description">
<paragraph id="1E52369EFF8D487746D9F92EFEA4F92A" blockId="1.[144,290,1683,1709]" box="[144,290,1683,1709]" pageId="1" pageNumber="18">
(
<figureCitation id="86D62A1BFF8D487746D0F92EFE9EF92A" box="[153,280,1683,1709]" captionStart-0="Fig" captionStart-1="Figs 2–9" captionStart-2="Figs 10–13" captionStartId-0="1.[619,661,721,745]" captionStartId-1="2.[159,213,1474,1498]" captionStartId-2="4.[159,215,1565,1589]" captionTargetBox-0="[144,1429,181,745]" captionTargetBox-1="[159,1444,181,1447]" captionTargetBox-2="[159,1000,181,1539]" captionTargetId-0="figure-553@1.[618,1429,181,702]" captionTargetId-1="figure-79@2.[159,1444,181,1447]" captionTargetId-2="figure-367@4.[159,1000,181,1539]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionText-0="Fig. 1. Petrobiellus sachalinensis sp. nov., general view." captionText-1="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." captionText-2="Figs 10–13. Petrobiellus sachalinensis sp. nov., holotype (female). 10, urosternite and urocoxites (part) V; 11, urocoxites VIII (part) and IX, with posterior gonapophyses; 12, distal part of anterior gonapophy- ses; 13, distal part of posterior gonapophyses. Scale bar: 0.1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.10135591" figureDoi-1="http://doi.org/10.5281/zenodo.10135593" figureDoi-2="http://doi.org/10.5281/zenodo.10135595" httpUri-0="https://zenodo.org/record/10135591/files/figure.png" httpUri-1="https://zenodo.org/record/10135593/files/figure.png" httpUri-2="https://zenodo.org/record/10135595/files/figure.png" pageId="1" pageNumber="18">Figs 1–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="56F76515FF8D487746FCF976FEB9F801" pageId="1" pageNumber="18" type="materials_examined">
<paragraph id="1E52369EFF8D487746FCF976FD15F8C2" blockId="1.[144,764,1739,1926]" pageId="1" pageNumber="18">
<materialsCitation id="AE853CC3FF8D487746FCF976FD15F8C2" ID-GBIF-Occurrence="4144867302" collectingDate="2019-08-08" collectorName="V. Kaplin" country="Russia" latitude="50.9" location="Sakhalin I." longLatPrecision="1093" longitude="142.15" municipality="Female" pageId="1" pageNumber="18" specimenCount="1" stateProvince="Sakhalin Prov." typeStatus="holotype">
<emphasis id="2C99EA8CFF8D487746FCF976FEA5F963" box="[181,291,1739,1764]" italics="true" pageId="1" pageNumber="18">
<typeStatus id="C156883CFF8D487746FCF976FE98F963" box="[181,286,1739,1764]" pageId="1" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="08EBFD17FF8D48774763F976FEFBF963" box="[298,381,1739,1764]" pageId="1" pageNumber="18" type="female">Female</specimenCount>
(slide-mounted),
<emphasis id="2C99EA8CFF8D48774403F971FD1AF963" bold="true" box="[586,668,1740,1764]" pageId="1" pageNumber="18">
<collectingCountry id="66FA760EFF8D48774403F971FD1AF963" box="[586,668,1740,1764]" name="Russia" pageId="1" pageNumber="18">Russia</collectingCountry>
</emphasis>
,
<collectingRegion id="DC29F87CFF8D487744E2F976FF69F883" country="Russia" name="Sakhalin" pageId="1" pageNumber="18">
<emphasis id="2C99EA8CFF8D487744E2F976FF6DF883" italics="true" pageId="1" pageNumber="18">Sakhalin Prov</emphasis>
.
</collectingRegion>
, western shore of northern
<location id="1B326045FF8D48774471F951FD3CF883" LSID="urn:lsid:plazi:treatment:96448788FF8D487346D9F9DDFDF8F9C0:1B326045FF8D48774471F951FD3CF883" box="[568,698,1772,1797]" country="Russia" latitude="50.9" longLatPrecision="1093" longitude="142.15" municipality="Female" name="Sakhalin I." pageId="1" pageNumber="18" stateProvince="Sakhalin Prov.">Sakhalin I.</location>
, near
<location id="1B326045FF8D487746D9F8B1FE57F8A2" LSID="urn:lsid:plazi:treatment:96448788FF8D487346D9F9DDFDF8F9C0:1B326045FF8D487746D9F8B1FE57F8A2" box="[144,465,1804,1829]" country="Russia" latitude="50.9" longLatPrecision="1093" longitude="142.15" municipality="Female" name="Aleksandrovsk-Sakhalinsky" pageId="1" pageNumber="18" stateProvince="Sakhalin Prov.">Aleksandrovsk-Sakhalinsky</location>
,
<geoCoordinate id="7BD95059FF8D48774792F8B1FDBCF8A3" box="[475,570,1803,1829]" degrees="50" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="18" precision="925" value="50.9">50°54′N</geoCoordinate>
<geoCoordinate id="7BD95059FF8D48774477F8B0FD2CF8A3" box="[574,682,1803,1829]" degrees="142" direction="east" minutes="09" orientation="longitude" pageId="1" pageNumber="18" precision="925" value="142.15">142°09′E</geoCoordinate>
, shoreline, under stones,
<date id="6A53105EFF8D4877472DF890FE6BF8C2" box="[356,493,1837,1861]" pageId="1" pageNumber="18" value="2019-08-08">
<collectingDate id="7A17E9B6FF8D4877472DF890FE6BF8C2" box="[356,493,1837,1861]" pageId="1" pageNumber="18" value="2019-08-08">8.VIII.2019</collectingDate>
</date>
,
<collectorName id="B3185348FF8D487747B1F893FDE3F8C2" box="[504,613,1836,1861]" pageId="1" pageNumber="18">V. Kaplin</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="1E52369EFF8D487746FCF8F3FEB9F801" blockId="1.[144,764,1739,1926]" pageId="1" pageNumber="18">
<materialsCitation id="AE853CC3FF8D487746FCF8F3FEBDF801" ID-GBIF-Occurrence="4144867301" collectingDate="2019-08-08" collectorName="V. Kaplin" country="Russia" latitude="50.9" location="Aleksandrovsk-Sakhalinsky" longLatPrecision="1093" longitude="142.15" municipality="Female" pageId="1" pageNumber="18" specimenCount="12" specimenCount-female="12" stateProvince="Sakhalin" typeStatus="paratype">
<emphasis id="2C99EA8CFF8D487746FCF8F3FEA8F8E2" box="[181,302,1870,1893]" italics="true" pageId="1" pageNumber="18">
<typeStatus id="C156883CFF8D487746FCF8F3FEACF8E2" box="[181,298,1870,1893]" pageId="1" pageNumber="18" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="08EBFD17FF8D4877477DF8F3FE2CF8E1" box="[308,426,1869,1894]" pageId="1" pageNumber="18" type="female">11 females</specimenCount>
(
<specimenCount id="08EBFD17FF8D487747F2F8F3FD91F8E1" box="[443,535,1869,1894]" pageId="1" pageNumber="18" type="female">1 female</specimenCount>
on slide), same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="56F76515FF8D48734506FC9AFEE7FEBA" lastPageId="5" lastPageNumber="22" pageId="1" pageNumber="18" type="description">
<paragraph id="1E52369EFF8D48774506FC9AFAE0F8B9" blockId="1.[809,1429,806,1927]" pageId="1" pageNumber="18">
<emphasis id="2C99EA8CFF8D48774506FC9AFC60FCC7" box="[847,998,807,832]" italics="true" pageId="1" pageNumber="18">Description.</emphasis>
Female. Body length (not including antennae):
<quantity id="D9159B7BFF8D487745ADFCF7FB0FFCE2" box="[996,1161,842,870]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.16" metricValueMin="0.84" pageId="1" pageNumber="18" unit="mm" value="10.0" valueMax="11.6" valueMin="8.4">8.4–11.6 mm</quantity>
; body width:
<quantity id="D9159B7BFF8D4877437DFCF6FCDDFC0D" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="1" pageNumber="18" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
; cercal length:
<quantity id="D9159B7BFF8D48774263FCD3FB4CFC0D" box="[1066,1226,878,906]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.7" metricValueMin="5.5" pageId="1" pageNumber="18" unit="mm" value="5.6" valueMax="5.7" valueMin="5.5">5.5–5.7 mm</quantity>
; total width of compound eyes: 0.94–1.00 mm; eye length:
<quantity id="D9159B7BFF8D48774302FC2EFC24FC54" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.699999999999999" metricValueMin="5.4" pageId="1" pageNumber="18" unit="mm" value="0.5549999999999999" valueMax="0.57" valueMin="0.54">0.54– 0.57 mm</quantity>
; paired ocelli width:
<quantity id="D9159B7BFF8D48774287FC0AFA16FC54" box="[1230,1424,951,979]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="1" pageNumber="18" unit="mm" value="0.5" valueMax="0.51" valueMin="0.49">0.49–0.51 mm</quantity>
, length:
<quantity id="D9159B7BFF8D487745CFFC60FBBEFC70" box="[902,1080,989,1015]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" pageId="1" pageNumber="18" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18 mm</quantity>
. Middle and hind coxal styli length:
<quantity id="D9159B7BFF8D487745C3FBBDFBCFFB9B" box="[906,1097,1024,1052]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="1" pageNumber="18" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60–0.65 mm</quantity>
. Ovipositor length:
<quantity id="D9159B7BFF8D48774303FBBDFC1BFBC7" metricMagnitude="-3" metricUnit="m" metricValue="3.425" metricValueMax="3.5" metricValueMin="3.35" pageId="1" pageNumber="18" unit="mm" value="3.425" valueMax="3.5" valueMin="3.35">3.35– 3.50 mm</quantity>
. General body colour whitish or slightly yellowish, almost without pigment. Antennal base, frons, lateral parts of clypeus, basal part of first and third articles of maxillary palps, mandibulae, hypopharynx, and first, third and basal half of second tarsomeres with brown or brown-violet pigment of weak or medium intensity. Maxillary and labial palps, flagellum and pedicellus of antennae, legs, coxal and abdominal styli without scales. Scapus with very sparse scales. Colour of scales whitish or light gray. Metanotum also with one pair, urotergites with three pairs of large spots of brown scales (
<figureCitation id="86D62A1BFF8D487745B1FA66FBC6FA71" box="[1016,1088,1499,1526]" captionStart="Fig" captionStartId="1.[619,661,721,745]" captionTargetBox="[144,1429,181,745]" captionTargetId="figure-553@1.[618,1429,181,702]" captionTargetPageId="1" captionText="Fig. 1. Petrobiellus sachalinensis sp. nov., general view." figureDoi="http://doi.org/10.5281/zenodo.10135591" httpUri="https://zenodo.org/record/10135591/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
). Antennae very long, 1.5– 1.9 times as long as body. Distal chains of flagellum divided into 14–23 annuli (
<figureCitation id="86D62A1BFF8D48774236F999FB4BF9B8" box="[1151,1229,1572,1599]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="1" pageNumber="18">Fig. 2</figureCitation>
). Clypeus without long thin bristles. Cercus approximately 0.48– 0.55 times as long as body, with about 21–24 divisions. Apex of cercus with one apical spike (
<figureCitation id="86D62A1BFF8D48774373F92FFA02F92B" box="[1338,1412,1682,1708]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="1" pageNumber="18">Fig. 3</figureCitation>
). All divisions of cerci, except for apical one or two, with 1–3 colorless spines on inner side supporting cerci above substrate. Divisions in middle part of cerci also with one outer lateral spine (
<figureCitation id="86D62A1BFF8D48774340F899FAD1F8B9" box="[1289,1367,1828,1854]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="1" pageNumber="18">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="1E52369EFF8D48744506F8F5FEC6F801" blockId="1.[809,1429,806,1927]" lastBlockId="2.[159,779,1648,1926]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
Compound eyes dark, with bluish tinge (in ethanol). Ratio of length to width of compound eye about 1.13–1.15; ratio of contact line to length of eyes about 0.45. Paired ocelli sole-shaped, brown, with narrow white bordering, 2.8–3.0 times as wide as long. Distance between inner margins of paired ocelli 0.13–0.15 and between their outer margins 0.96–0.98 times total width of compound eyes (
<figureCitation id="86D62A1BFF8E48744770F8F5FE00F8E4" box="[313,390,1863,1891]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="2" pageNumber="19">Fig. 5</figureCitation>
). Frons clearly convex between paired ocelli.
</paragraph>
<caption id="4A926616FF8E487446D6FA7FFD2DF9BD" ID-DOI="http://doi.org/10.5281/zenodo.10135593" ID-Zenodo-Dep="10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="2" pageNumber="19" startId="2.[159,213,1474,1498]" targetBox="[159,1444,181,1447]" targetPageId="2" targetType="figure">
<paragraph id="1E52369EFF8E487446D6FA7FFD2DF9BD" blockId="2.[159,1444,1474,1595]" pageId="2" pageNumber="19">
<emphasis id="2C99EA8CFF8E487446D6FA7FFE94FA5D" bold="true" box="[159,274,1474,1498]" pageId="2" pageNumber="19">Figs 2–9.</emphasis>
<taxonomicName id="D9ED4D1DFF8E48744751FA7FFDC1FA5C" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[280,583,1474,1499]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8E48744751FA7FFDC1FA5C" box="[280,583,1474,1499]" italics="true" pageId="2" pageNumber="19">Petrobiellus sachalinensis</emphasis>
</taxonomicName>
<emphasis id="2C99EA8CFF8E48744404FA7EFD2DFA5C" bold="true" box="[589,683,1475,1499]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="37AA57F7FF8E48744404FA7EFD2DFA5C" box="[589,683,1475,1499]" pageId="2" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (female).
<emphasis id="2C99EA8CFF8E487445DFFA7FFC22FA5D" bold="true" box="[918,932,1474,1498]" pageId="2" pageNumber="19">2</emphasis>
, distal chain of flagellum;
<emphasis id="2C99EA8CFF8E48744294FA7FFB6DFA5D" bold="true" box="[1245,1259,1474,1498]" pageId="2" pageNumber="19">3</emphasis>
, apex of cercus;
<emphasis id="2C99EA8CFF8E487446D6FA5FFF2BFA7D" bold="true" box="[159,173,1506,1530]" pageId="2" pageNumber="19">4</emphasis>
, divisions in middle part of cercus;
<emphasis id="2C99EA8CFF8E48744401FA5EFDD0FA7C" bold="true" box="[584,598,1507,1531]" pageId="2" pageNumber="19">5</emphasis>
, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum);
<emphasis id="2C99EA8CFF8E48744483F9BFFD5EF99D" bold="true" box="[714,728,1538,1562]" pageId="2" pageNumber="19">6</emphasis>
, maxillary palpus;
<emphasis id="2C99EA8CFF8E487445F8F9BEFC38F99C" bold="true" box="[945,958,1539,1563]" pageId="2" pageNumber="19">7</emphasis>
, labial palpus and labium (part);
<emphasis id="2C99EA8CFF8E48744371F9BFFAC0F99D" bold="true" box="[1336,1350,1538,1562]" pageId="2" pageNumber="19">8</emphasis>
, apex of mandible;
<emphasis id="2C99EA8CFF8E4874475CF99FFEA5F9BD" bold="true" box="[277,291,1570,1594]" pageId="2" pageNumber="19">9</emphasis>
, fore leg (part). Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="4A926616FF8E48744571F9CCFB74F924" ID-Table-UUID="4A926616FF8E48744571F9CCFB74F924" httpUri="http://table.plazi.org/id/4A926616FF8E48744571F9CCFB74F924" pageId="2" pageNumber="19" startId="2.[824,884,1649,1670]" targetBox="[835,1412,1740,1913]" targetIsTable="true" targetPageId="2" targetType="table">
<paragraph id="1E52369EFF8E48744571F9CCFB74F924" blockId="2.[824,1444,1648,1700]" pageId="2" pageNumber="19">
<emphasis id="2C99EA8CFF8E48744571F9CCFC08F901" bold="true" box="[824,910,1649,1670]" pageId="2" pageNumber="19">Table 1.</emphasis>
Ratios of the length to width of the leg parts in females of
<taxonomicName id="D9ED4D1DFF8E487445DBF930FB1EF923" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[914,1176,1677,1700]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8E487445DBF930FB1EF923" box="[914,1176,1677,1700]" italics="true" pageId="2" pageNumber="19">Petrobiellus sachalinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37AA57F7FF8E487442D7F933FB74F924" box="[1182,1266,1678,1699]" pageId="2" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="1E52369EFF8E4874450AF962FA02F8FE" pageId="2" pageNumber="19">
<table id="6CEDC43EFF8EB789450AF971FA02F8FE" box="[835,1412,1740,1913]" gridcols="4" gridrows="4" pageId="2" pageNumber="19">
<tr id="A0DD34DCFF8EB789450AF971FA02F88F" box="[835,1412,1740,1800]" gridrow="0" pageId="2" pageNumber="19">
<th id="E30C5DA0FF8EB789450AF971FC26F88F" box="[835,928,1740,1800]" gridcol="0" gridrow="0" pageId="2" pageNumber="19">Segments</th>
<th id="E30C5DA0FF8EB78945ADF971FBC3F88F" box="[996,1093,1740,1800]" gridcol="1" gridrow="0" pageId="2" pageNumber="19">fore</th>
<th id="E30C5DA0FF8EB7894234F971FB6DF88F" box="[1149,1259,1740,1800]" gridcol="2" gridrow="0" pageId="2" pageNumber="19">Pair of legs middle</th>
<th id="E30C5DA0FF8EB789436AF971FA02F88F" box="[1315,1412,1740,1800]" gridcol="3" gridrow="0" pageId="2" pageNumber="19">hind</th>
</tr>
<tr id="A0DD34DCFF8EB789450AF8AAFA02F8AA" box="[835,1412,1815,1837]" gridrow="1" pageId="2" pageNumber="19">
<th id="E30C5DA0FF8EB789450AF8AAFC26F8AA" box="[835,928,1815,1837]" gridcol="0" gridrow="1" pageId="2" pageNumber="19">Femur</th>
<td id="E30C5DA0FF8EB78945ADF8AAFBC3F8AA" box="[996,1093,1815,1837]" gridcol="1" gridrow="1" pageId="2" pageNumber="19">2.64–2.83</td>
<td id="E30C5DA0FF8EB7894234F8AAFB6DF8AA" box="[1149,1259,1815,1837]" gridcol="2" gridrow="1" pageId="2" pageNumber="19">2.68–2.74</td>
<td id="E30C5DA0FF8EB789436AF8AAFA02F8AA" box="[1315,1412,1815,1837]" gridcol="3" gridrow="1" pageId="2" pageNumber="19">2.83–2.91</td>
</tr>
<tr id="A0DD34DCFF8EB789450AF880FA02F8D3" box="[835,1412,1853,1876]" gridrow="2" pageId="2" pageNumber="19">
<th id="E30C5DA0FF8EB789450AF880FC26F8D3" box="[835,928,1853,1876]" gridcol="0" gridrow="2" pageId="2" pageNumber="19">Tibia</th>
<td id="E30C5DA0FF8EB78945ADF880FBC3F8D3" box="[996,1093,1853,1876]" gridcol="1" gridrow="2" pageId="2" pageNumber="19">3.00–3.22</td>
<td id="E30C5DA0FF8EB7894234F880FB6DF8D3" box="[1149,1259,1853,1876]" gridcol="2" gridrow="2" pageId="2" pageNumber="19">2.82–3.12</td>
<td id="E30C5DA0FF8EB789436AF880FA02F8D3" box="[1315,1412,1853,1876]" gridcol="3" gridrow="2" pageId="2" pageNumber="19">3.68–3.74</td>
</tr>
<tr id="A0DD34DCFF8EB789450AF8DEFA02F8FE" box="[835,1412,1891,1913]" gridrow="3" pageId="2" pageNumber="19">
<th id="E30C5DA0FF8EB789450AF8DEFC26F8FE" box="[835,928,1891,1913]" gridcol="0" gridrow="3" pageId="2" pageNumber="19">Tarsus</th>
<td id="E30C5DA0FF8EB78945ADF8DEFBC3F8FE" box="[996,1093,1891,1913]" gridcol="1" gridrow="3" pageId="2" pageNumber="19">6.50–6.67</td>
<td id="E30C5DA0FF8EB7894234F8DEFB6DF8FE" box="[1149,1259,1891,1913]" gridcol="2" gridrow="3" pageId="2" pageNumber="19">6.40-6.65</td>
<td id="E30C5DA0FF8EB789436AF8DEFA02F8FE" box="[1315,1412,1891,1913]" gridcol="3" gridrow="3" pageId="2" pageNumber="19">9.12–9.36</td>
</tr>
</table>
</paragraph>
<caption id="4A926616FF8F487546D9FF05FE0CFF6D" ID-Table-UUID="4A926616FF8F487546D9FF05FE0CFF6D" httpUri="http://table.plazi.org/id/4A926616FF8F487546D9FF05FE0CFF6D" pageId="3" pageNumber="20" startId="3.[144,203,184,205]" targetBox="[154,712,270,558]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="1E52369EFF8F487546D9FF05FE0CFF6D" blockId="3.[144,758,183,235]" pageId="3" pageNumber="20">
<emphasis id="2C99EA8CFF8F487546D9FF05FF62FF4A" bold="true" box="[144,228,184,205]" pageId="3" pageNumber="20">Table 2.</emphasis>
Number of spines at the legs in females of
<taxonomicName id="D9ED4D1DFF8F487544DDFF0AFEB2FF6C" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8F487544DDFF0AFEB2FF6C" italics="true" pageId="3" pageNumber="20">Petrobiellus sachalinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37AA57F7FF8F48754770FF68FE0CFF6D" box="[313,394,213,234]" pageId="3" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="4A926616FF8F48754560FF05FA86FF6D" ID-Table-UUID="4A926616FF8F48754560FF05FA86FF6D" httpUri="http://table.plazi.org/id/4A926616FF8F48754560FF05FA86FF6D" pageId="3" pageNumber="20" startId="3.[809,868,184,205]" targetBox="[820,1418,268,560]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="1E52369EFF8F48754560FF05FA86FF6D" blockId="3.[809,1417,183,235]" pageId="3" pageNumber="20">
<emphasis id="2C99EA8CFF8F48754560FF05FCFBFF4A" bold="true" box="[809,893,184,205]" pageId="3" pageNumber="20">Table 3.</emphasis>
Ratios of the lengths of some abdominal structures in females of
<taxonomicName id="D9ED4D1DFF8F487545E7FF69FB2CFF6C" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[942,1194,212,235]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8F487545E7FF69FB2CFF6C" box="[942,1194,212,235]" italics="true" pageId="3" pageNumber="20">Petrobiellus sachalinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37AA57F7FF8F487542F9FF68FA86FF6D" box="[1200,1280,213,234]" pageId="3" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="1E52369EFF8F487546FFFE98FD31FDA9" pageId="3" pageNumber="20">
<table id="6CEDC43EFF8FB78946D3FEB3FD4EFDA9" box="[154,712,270,558]" gridcols="4" gridrows="5" pageId="3" pageNumber="20">
<tr id="A0DD34DCFF8FB78946D3FEB3FD4EFED6" box="[154,712,270,337]" gridrow="0" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FEB3FE9CFED6" box="[154,282,270,337]" gridcol="0" gridrow="0" pageId="3" pageNumber="20">Segments</th>
<th id="E30C5DA0FF8FB789473FFEB3FE18FED6" box="[374,414,270,337]" gridcol="1" gridrow="0" pageId="3" pageNumber="20">fore</th>
<th id="E30C5DA0FF8FB78947ACFEB3FDD3FED6" box="[485,597,270,337]" gridcol="2" gridrow="0" pageId="3" pageNumber="20">Pair of legs middle</th>
<th id="E30C5DA0FF8FB78944D0FEB3FD4EFED6" box="[665,712,270,337]" gridcol="3" gridrow="0" pageId="3" pageNumber="20">hind</th>
</tr>
<tr id="A0DD34DCFF8FB78946D3FEDAFD4EFE3C" box="[154,712,359,443]" gridrow="1" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FEDAFE9CFE3C" box="[154,282,359,443]" gridcol="0" gridrow="1" pageId="3" pageNumber="20">1st Tarso- meres 2nd</th>
<td id="E30C5DA0FF8FB789473FFEDAFE18FE3C" box="[374,414,359,443]" gridcol="1" gridrow="1" pageId="3" pageNumber="20">2 4</td>
<td id="E30C5DA0FF8FB78947ACFEDAFDD3FE3C" box="[485,597,359,443]" gridcol="2" gridrow="1" pageId="3" pageNumber="20">2 6</td>
<td id="E30C5DA0FF8FB78944D0FEDAFD4EFE3C" box="[665,712,359,443]" gridcol="3" gridrow="1" pageId="3" pageNumber="20">4–5 7</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FE02FD4EFE51" box="[154,712,447,470]" gridrow="2" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FE02FE9CFE51" box="[154,282,447,470]" gridcol="0" gridrow="2" pageId="3" pageNumber="20">3rd</th>
<td id="E30C5DA0FF8FB789473FFE02FE18FE51" box="[374,414,447,470]" gridcol="1" gridrow="2" pageId="3" pageNumber="20">0</td>
<td id="E30C5DA0FF8FB78947ACFE02FDD3FE51" box="[485,597,447,470]" gridcol="2" gridrow="2" pageId="3" pageNumber="20">0</td>
<td id="E30C5DA0FF8FB78944D0FE02FD4EFE51" box="[665,712,447,470]" gridcol="3" gridrow="2" pageId="3" pageNumber="20">0</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FE56FD4EFD85" box="[154,712,491,514]" gridrow="3" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FE56FE9CFD85" box="[154,282,491,514]" gridcol="0" gridrow="3" pageId="3" pageNumber="20">Tibia</th>
<td id="E30C5DA0FF8FB789473FFE56FE18FD85" box="[374,414,491,514]" gridcol="1" gridrow="3" pageId="3" pageNumber="20">0</td>
<td id="E30C5DA0FF8FB78947ACFE56FDD3FD85" box="[485,597,491,514]" gridcol="2" gridrow="3" pageId="3" pageNumber="20">0</td>
<td id="E30C5DA0FF8FB78944D0FE56FD4EFD85" box="[665,712,491,514]" gridcol="3" gridrow="3" pageId="3" pageNumber="20">1–2</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FDA4FD4EFDA9" box="[154,712,537,558]" gridrow="4" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FDA4FE9CFDA9" box="[154,282,537,558]" gridcol="0" gridrow="4" pageId="3" pageNumber="20">Femur</th>
<td id="E30C5DA0FF8FB789473FFDA4FE18FDA9" box="[374,414,537,558]" gridcol="1" gridrow="4" pageId="3" pageNumber="20">0</td>
<td id="E30C5DA0FF8FB78947ACFDA4FDD3FDA9" box="[485,597,537,558]" gridcol="2" gridrow="4" pageId="3" pageNumber="20">0</td>
<td id="E30C5DA0FF8FB78944D0FDA4FD4EFDA9" box="[665,712,537,558]" gridcol="3" gridrow="4" pageId="3" pageNumber="20">0</td>
</tr>
</table>
</paragraph>
<paragraph id="1E52369EFF8F48754508FEA7FAD0FDA8" pageId="3" pageNumber="20">
<table id="6CEDC43EFF8FB789457DFEB1FA0CFDB7" box="[820,1418,268,560]" gridcols="4" gridrows="7" pageId="3" pageNumber="20">
<tr id="A0DD34DCFF8FB789457DFEB1FA0CFEC7" box="[820,1418,268,320]" gridrow="0" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFEB1FC05FEC7" box="[820,899,268,320]" gridcol="0" gridrow="0" pageId="3" pageNumber="20">Urites</th>
<th id="E30C5DA0FF8FB78945ECFEB1FBBEFEC7" box="[933,1080,268,320]" gridcol="1" gridrow="0" pageId="3" pageNumber="20">Urosternite / urocoxite</th>
<th id="E30C5DA0FF8FB789422DFEB1FB4CFEC7" box="[1124,1226,268,320]" gridcol="2" gridrow="0" pageId="3" pageNumber="20">Stylus / urocoxite</th>
<th id="E30C5DA0FF8FB78942BEFEB1FA0CFEC7" box="[1271,1418,268,320]" gridcol="3" gridrow="0" pageId="3" pageNumber="20">Apical spine / stylus</th>
</tr>
<tr id="A0DD34DCFF8FB789457DFEECFA0CFEEF" box="[820,1418,337,360]" gridrow="1" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFEECFC05FEEF" box="[820,899,337,360]" gridcol="0" gridrow="1" pageId="3" pageNumber="20">II</th>
<td id="E30C5DA0FF8FB78945ECFEECFBBEFEEF" box="[933,1080,337,360]" gridcol="1" gridrow="1" pageId="3" pageNumber="20">0.69</td>
<td id="E30C5DA0FF8FB789422DFEECFB4CFEEF" box="[1124,1226,337,360]" gridcol="2" gridrow="1" pageId="3" pageNumber="20">0.85</td>
<td id="E30C5DA0FF8FB78942BEFEECFA0CFEEF" box="[1271,1418,337,360]" gridcol="3" gridrow="1" pageId="3" pageNumber="20">0.34</td>
</tr>
<tr id="A0DD34DCFF8FB789457DFEC7FA0CFE08" box="[820,1418,378,399]" gridrow="2" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFEC7FC05FE08" box="[820,899,378,399]" gridcol="0" gridrow="2" pageId="3" pageNumber="20">III</th>
<td id="E30C5DA0FF8FB78945ECFEC7FBBEFE08" box="[933,1080,378,399]" gridcol="1" gridrow="2" pageId="3" pageNumber="20">0.71</td>
<td id="E30C5DA0FF8FB789422DFEC7FB4CFE08" box="[1124,1226,378,399]" gridcol="2" gridrow="2" pageId="3" pageNumber="20">0.84</td>
<td id="E30C5DA0FF8FB78942BEFEC7FA0CFE08" box="[1271,1418,378,399]" gridcol="3" gridrow="2" pageId="3" pageNumber="20">0.34</td>
</tr>
<tr id="A0DD34DCFF8FB789457DFE1FFA0CFE30" box="[820,1418,418,439]" gridrow="3" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFE1FFC05FE30" box="[820,899,418,439]" gridcol="0" gridrow="3" pageId="3" pageNumber="20">IV</th>
<td id="E30C5DA0FF8FB78945ECFE1FFBBEFE30" box="[933,1080,418,439]" gridcol="1" gridrow="3" pageId="3" pageNumber="20">0.70</td>
<td id="E30C5DA0FF8FB789422DFE1FFB4CFE30" box="[1124,1226,418,439]" gridcol="2" gridrow="3" pageId="3" pageNumber="20">0.81</td>
<td id="E30C5DA0FF8FB78942BEFE1FFA0CFE30" box="[1271,1418,418,439]" gridcol="3" gridrow="3" pageId="3" pageNumber="20">0.34</td>
</tr>
<tr id="A0DD34DCFF8FB789457DFE74FA0CFE67" box="[820,1418,457,480]" gridrow="4" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFE74FC05FE67" box="[820,899,457,480]" gridcol="0" gridrow="4" pageId="3" pageNumber="20">V–VII</th>
<td id="E30C5DA0FF8FB78945ECFE74FBBEFE67" box="[933,1080,457,480]" gridcol="1" gridrow="4" pageId="3" pageNumber="20">0.67–0.68</td>
<td id="E30C5DA0FF8FB789422DFE74FB4CFE67" box="[1124,1226,457,480]" gridcol="2" gridrow="4" pageId="3" pageNumber="20">0.74–0.76</td>
<td id="E30C5DA0FF8FB78942BEFE74FA0CFE67" box="[1271,1418,457,480]" gridcol="3" gridrow="4" pageId="3" pageNumber="20">0.36–0.37</td>
</tr>
<tr id="A0DD34DCFF8FB789457DFE4CFA0CFD8F" box="[820,1418,497,520]" gridrow="5" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFE4CFC05FD8F" box="[820,899,497,520]" gridcol="0" gridrow="5" pageId="3" pageNumber="20">VIII</th>
<td id="E30C5DA0FF8FB78945ECFE4CFBBEFD8F" box="[933,1080,497,520]" gridcol="1" gridrow="5" pageId="3" pageNumber="20">–</td>
<td id="E30C5DA0FF8FB789422DFE4CFB4CFD8F" box="[1124,1226,497,520]" gridcol="2" gridrow="5" pageId="3" pageNumber="20">0.86</td>
<td id="E30C5DA0FF8FB78942BEFE4CFA0CFD8F" box="[1271,1418,497,520]" gridcol="3" gridrow="5" pageId="3" pageNumber="20">0.31</td>
</tr>
<tr id="A0DD34DCFF8FB789457DFDA7FA0CFDB7" box="[820,1418,538,560]" gridrow="6" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB789457DFDA7FC05FDB7" box="[820,899,538,560]" gridcol="0" gridrow="6" pageId="3" pageNumber="20">IX</th>
<td id="E30C5DA0FF8FB78945ECFDA7FBBEFDB7" box="[933,1080,538,560]" gridcol="1" gridrow="6" pageId="3" pageNumber="20">–</td>
<td id="E30C5DA0FF8FB789422DFDA7FB4CFDB7" box="[1124,1226,538,560]" gridcol="2" gridrow="6" pageId="3" pageNumber="20">0.78</td>
<td id="E30C5DA0FF8FB78942BEFDA7FA0CFDB7" box="[1271,1418,538,560]" gridcol="3" gridrow="6" pageId="3" pageNumber="20">0.28</td>
</tr>
</table>
</paragraph>
<paragraph id="1E52369EFF8F48754560FDFAFAEDFDDA" blockId="3.[809,1387,582,605]" box="[809,1387,582,605]" pageId="3" pageNumber="20">
<tableNote id="E30B3710FF8F48754560FDFAFAEDFDDA" box="[809,1387,582,605]" pageId="3" pageNumber="20" targetBox="[820,1418,268,560]" targetPageId="3">
<emphasis id="2C99EA8CFF8F48754560FDFAFCDEFDDB" box="[809,856,583,604]" italics="true" pageId="3" pageNumber="20">Note</emphasis>
. Length of the stylus does not include apical spines.
</tableNote>
</paragraph>
<caption id="4A926616FF8F487546D9FD24FB84FD29" ID-Table-UUID="4A926616FF8F487546D9FD24FB84FD29" box="[144,1026,664,687]" httpUri="http://table.plazi.org/id/4A926616FF8F487546D9FD24FB84FD29" pageId="3" pageNumber="20" startId="3.[144,203,665,686]" targetBox="[154,1412,720,1799]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="1E52369EFF8F487546D9FD24FB84FD29" blockId="3.[144,1026,664,687]" box="[144,1026,664,687]" pageId="3" pageNumber="20">
<emphasis id="2C99EA8CFF8F487546D9FD24FF62FD29" bold="true" box="[144,228,665,686]" pageId="3" pageNumber="20">Table 4.</emphasis>
Main morphological differences between females of the species of
<taxonomicName id="D9ED4D1DFF8F487545C0FD25FC7BFD28" authorityName="Silvestri" authorityYear="1943" box="[905,1021,664,687]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="2C99EA8CFF8F487545C0FD25FC7BFD28" box="[905,1021,664,687]" italics="true" pageId="3" pageNumber="20">Petrobiellus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="1E52369EFF8F4875469AFD6DFAA7F880" pageId="3" pageNumber="20">
<table id="6CEDC43EFF8FB78946D3FD6DFA02F880" box="[154,1412,720,1799]" gridcols="4" gridrows="15" pageId="3" pageNumber="20">
<tr id="A0DD34DCFF8FB78946D3FD6DFA02FD60" box="[154,1412,720,743]" gridrow="0" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FD6DFD96FD60" box="[154,528,720,743]" gridcol="0" gridrow="0" pageId="3" pageNumber="20">Morphological characters</th>
<th id="E30C5DA0FF8FB7894461FD6DFCA9FD60" box="[552,815,720,743]" gridcol="1" gridrow="0" pageId="3" pageNumber="20">
<taxonomicName id="D9ED4D1DFF8F4875447EFD6FFCADFD61" authority="Kaplin, 1980" authorityName="Kaplin" authorityYear="1980" box="[567,811,720,743]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="kusakini">
<emphasis id="2C99EA8CFF8F4875447EFD6FFD27FD60" box="[567,673,720,743]" italics="true" pageId="3" pageNumber="20">P. kusakini</emphasis>
<bibRefCitation id="7A7C4B6FFF8F487544EEFD6DFCADFD61" author="Kaplin V. G." box="[679,811,720,743]" pageId="3" pageNumber="20" pagination="3 - 9" refId="ref2321" refString="Kaplin V. G. 1980. New species of bristletails (Microcoryphia, Machilidae) from the Kuril Islands and Primorsky Krai. In: Lehr P. A. (Ed.). Taksonomiya nasekomykh Dal'nego Vostoka [Taxonomy of insects of the Far East]: 3 - 9. Vladivostok. (In Russian)." type="book chapter" year="1980">Kaplin, 1980</bibRefCitation>
</taxonomicName>
</th>
<th id="E30C5DA0FF8FB7894506FD6DFBE6FD60" box="[847,1120,720,743]" gridcol="2" gridrow="0" pageId="3" pageNumber="20">
<taxonomicName id="D9ED4D1DFF8F4875451AFD6FFBD8FD60" authority="Uchida, 1954" authorityName="Uchida" authorityYear="1954" box="[851,1118,720,743]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="curvistylis">
<emphasis id="2C99EA8CFF8F4875451AFD6FFC49FD60" box="[851,975,720,743]" italics="true" pageId="3" pageNumber="20">P. curvistylis</emphasis>
<bibRefCitation id="7A7C4B6FFF8F4875459CFD6DFBD8FD60" author="Uchida H." box="[981,1118,720,743]" pageId="3" pageNumber="20" pagination="1 - 17" refId="ref2500" refString="Uchida H. 1954. Apterygota of the Hachijo-Jima and its adjacent islands. Science Reports of the Faculty of Literature and Science, Hirosaki University, 1 (1): 1 - 17." type="journal article" year="1954">Uchida, 1954</bibRefCitation>
</taxonomicName>
</th>
<th id="E30C5DA0FF8FB789423EFD6DFA02FD60" box="[1143,1412,720,743]" gridcol="3" gridrow="0" pageId="3" pageNumber="20">
<taxonomicName id="D9ED4D1DFF8F487542CEFD6FFAA4FD60" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[1159,1314,720,743]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8F487542CEFD6FFAA4FD60" box="[1159,1314,720,743]" italics="true" pageId="3" pageNumber="20">P. sachalinensis</emphasis>
</taxonomicName>
<emphasis id="2C99EA8CFF8F4875436EFD6CFAFEFD61" bold="true" box="[1319,1400,721,742]" pageId="3" pageNumber="20">
<taxonomicNameLabel id="37AA57F7FF8F4875436EFD6CFAFEFD61" box="[1319,1400,721,742]" pageId="3" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="A0DD34DCFF8FB78946D3FD45FA02FCAB" box="[154,1412,760,812]" gridrow="1" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FD45FD96FCAB" box="[154,528,760,812]" gridcol="0" gridrow="1" pageId="3" pageNumber="20">General body colour</th>
<td id="E30C5DA0FF8FB7894461FD45FCA9FCAB" box="[552,815,760,812]" gridcol="1" gridrow="1" pageId="3" pageNumber="20">yellowish</td>
<td id="E30C5DA0FF8FB7894506FD45FBE6FCAB" box="[847,1120,760,812]" gridcol="2" gridrow="1" pageId="3" pageNumber="20">rufous</td>
<td id="E30C5DA0FF8FB789423EFD45FA02FCAB" box="[1143,1412,760,812]" gridcol="3" gridrow="1" pageId="3" pageNumber="20">whitish or slightly yel- lowish</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FC80FA02FC09" box="[154,1412,829,910]" gridrow="2" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FC80FD96FC09" box="[154,528,829,910]" gridcol="0" gridrow="2" pageId="3" pageNumber="20">Colour of scales on body surface</th>
<td id="E30C5DA0FF8FB7894461FC80FCA9FC09" box="[552,815,829,910]" gridcol="1" gridrow="2" pageId="3" pageNumber="20">dark brownish</td>
<td id="E30C5DA0FF8FB7894506FC80FBE6FC09" box="[847,1120,829,910]" gridcol="2" gridrow="2" pageId="3" pageNumber="20">brownish</td>
<td id="E30C5DA0FF8FB789423EFC80FA02FC09" box="[1143,1412,829,910]" gridcol="3" gridrow="2" pageId="3" pageNumber="20">whitish or light gray, with several relatively large spots of brown scales</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FC22FA02FB8B" box="[154,1412,927,1036]" gridrow="3" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FC22FD96FB8B" box="[154,528,927,1036]" gridcol="0" gridrow="3" pageId="3" pageNumber="20">Distribution of hypodermal pigment</th>
<td id="E30C5DA0FF8FB7894461FC22FCA9FB8B" box="[552,815,927,1036]" gridcol="1" gridrow="3" pageId="3" pageNumber="20">head, maxillary palps, mandibles, maxillae, thorax and legs highly pigmented</td>
<td id="E30C5DA0FF8FB7894506FC22FBE6FB8B" box="[847,1120,927,1036]" gridcol="2" gridrow="3" pageId="3" pageNumber="20">scapus and pedicellus of antenna, trochanter, femur and tibia of all legs with blackish purple pigment</td>
<td id="E30C5DA0FF8FB789423EFC22FA02FB8B" box="[1143,1412,927,1036]" gridcol="3" gridrow="3" pageId="3" pageNumber="20">head and tarsi with brown or brown-violet hypoder- mal pigment of weak or medium intensity</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FBA0FA02FBD6" box="[154,1412,1053,1105]" gridrow="4" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FBA0FD96FBD6" box="[154,528,1053,1105]" gridcol="0" gridrow="4" pageId="3" pageNumber="20">Colour of paired ocelli</th>
<td id="E30C5DA0FF8FB7894461FBA0FCA9FBD6" box="[552,815,1053,1105]" gridcol="1" gridrow="4" pageId="3" pageNumber="20">dark, without white bordering</td>
<td id="E30C5DA0FF8FB7894506FBA0FBE6FBD6" box="[847,1120,1053,1105]" gridcol="2" gridrow="4" pageId="3" pageNumber="20">dark</td>
<td id="E30C5DA0FF8FB789423EFBA0FA02FBD6" box="[1143,1412,1053,1105]" gridcol="3" gridrow="4" pageId="3" pageNumber="20">brown, with white bor- dering</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FBDFFA02FB34" box="[154,1412,1122,1203]" gridrow="5" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FBDFFD96FB34" box="[154,528,1122,1203]" gridcol="0" gridrow="5" pageId="3" pageNumber="20">Shape of paired ocelli</th>
<td id="E30C5DA0FF8FB7894461FBDFFCA9FB34" box="[552,815,1122,1203]" gridcol="1" gridrow="5" pageId="3" pageNumber="20">pistiliform, strongly nar- rowed in the middle parts</td>
<td id="E30C5DA0FF8FB7894506FBDFFBE6FB34" box="[847,1120,1122,1203]" gridcol="2" gridrow="5" pageId="3" pageNumber="20">pistiliform, either of them almost separated in mid- dle into two parts</td>
<td id="E30C5DA0FF8FB789423EFBDFFA02FB34" box="[1143,1412,1122,1203]" gridcol="3" gridrow="5" pageId="3" pageNumber="20">almost sole-shaped</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FB79FA02FB7F" box="[154,1412,1220,1272]" gridrow="6" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FB79FD96FB7F" box="[154,528,1220,1272]" gridcol="0" gridrow="6" pageId="3" pageNumber="20">Ratio of length to width of com- pound eye</th>
<td id="E30C5DA0FF8FB7894461FB79FCA9FB7F" box="[552,815,1220,1272]" gridcol="1" gridrow="6" pageId="3" pageNumber="20">1.09</td>
<td id="E30C5DA0FF8FB7894506FB79FBE6FB7F" box="[847,1120,1220,1272]" gridcol="2" gridrow="6" pageId="3" pageNumber="20">1.00–1.02</td>
<td id="E30C5DA0FF8FB789423EFB79FA02FB7F" box="[1143,1412,1220,1272]" gridcol="3" gridrow="6" pageId="3" pageNumber="20">1.13–1.15</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FAB4FA02FABB" box="[154,1412,1289,1340]" gridrow="7" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FAB4FD96FABB" box="[154,528,1289,1340]" gridcol="0" gridrow="7" pageId="3" pageNumber="20">Ratios of lengths of apical and pre- ceding articles of maxillary palps</th>
<td id="E30C5DA0FF8FB7894461FAB4FCA9FABB" box="[552,815,1289,1340]" gridcol="1" gridrow="7" pageId="3" pageNumber="20">0.83</td>
<td id="E30C5DA0FF8FB7894506FAB4FBE6FABB" box="[847,1120,1289,1340]" gridcol="2" gridrow="7" pageId="3" pageNumber="20">0.57</td>
<td id="E30C5DA0FF8FB789423EFAB4FA02FABB" box="[1143,1412,1289,1340]" gridcol="3" gridrow="7" pageId="3" pageNumber="20">0.60–0.62</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FAF0FA02FA06" box="[154,1412,1357,1409]" gridrow="8" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FAF0FD96FA06" box="[154,528,1357,1409]" gridcol="0" gridrow="8" pageId="3" pageNumber="20">Ratios of lengths of 5th and 4th articles of maxillary palps</th>
<td id="E30C5DA0FF8FB7894461FAF0FCA9FA06" box="[552,815,1357,1409]" gridcol="1" gridrow="8" pageId="3" pageNumber="20">1.20</td>
<td id="E30C5DA0FF8FB7894506FAF0FBE6FA06" box="[847,1120,1357,1409]" gridcol="2" gridrow="8" pageId="3" pageNumber="20">1.10</td>
<td id="E30C5DA0FF8FB789423EFAF0FA02FA06" box="[1143,1412,1357,1409]" gridcol="3" gridrow="8" pageId="3" pageNumber="20">1.04–1.06</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FA2FFA02FA2E" box="[154,1412,1426,1449]" gridrow="9" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FA2FFD96FA2E" box="[154,528,1426,1449]" gridcol="0" gridrow="9" pageId="3" pageNumber="20">Number of spines on 2nd tarsomere</th>
<td id="E30C5DA0FF8FB7894461FA2FFCA9FA2E" box="[552,815,1426,1449]" gridcol="1" gridrow="9" pageId="3" pageNumber="20">16–26</td>
<td id="E30C5DA0FF8FB7894506FA2FFBE6FA2E" box="[847,1120,1426,1449]" gridcol="2" gridrow="9" pageId="3" pageNumber="20">?</td>
<td id="E30C5DA0FF8FB789423EFA2FFA02FA2E" box="[1143,1412,1426,1449]" gridcol="3" gridrow="9" pageId="3" pageNumber="20">4–7</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FA07FA02FA6A" box="[154,1412,1466,1517]" gridrow="10" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FA07FD96FA6A" box="[154,528,1466,1517]" gridcol="0" gridrow="10" pageId="3" pageNumber="20">Ratio of length of styli to width of coxae</th>
<td id="E30C5DA0FF8FB7894461FA07FCA9FA6A" box="[552,815,1466,1517]" gridcol="1" gridrow="10" pageId="3" pageNumber="20">1.61</td>
<td id="E30C5DA0FF8FB7894506FA07FBE6FA6A" box="[847,1120,1466,1517]" gridcol="2" gridrow="10" pageId="3" pageNumber="20">1.14</td>
<td id="E30C5DA0FF8FB789423EFA07FA02FA6A" box="[1143,1412,1466,1517]" gridcol="3" gridrow="10" pageId="3" pageNumber="20">1.05–1.09</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3FA42FA02F9E1" box="[154,1412,1535,1638]" gridrow="11" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3FA42FD96F9E1" box="[154,528,1535,1638]" gridcol="0" gridrow="11" pageId="3" pageNumber="20">I Ratios of lengths of urosterni- II te and urocoxite VII</th>
<td id="E30C5DA0FF8FB7894461FA42FCA9F9E1" box="[552,815,1535,1638]" gridcol="1" gridrow="11" pageId="3" pageNumber="20">0.32 0.39 0.53</td>
<td id="E30C5DA0FF8FB7894506FA42FBE6F9E1" box="[847,1120,1535,1638]" gridcol="2" gridrow="11" pageId="3" pageNumber="20">0.29 0.52 0.40</td>
<td id="E30C5DA0FF8FB789423EFA42FA02F9E1" box="[1143,1412,1535,1638]" gridcol="3" gridrow="11" pageId="3" pageNumber="20">0.35 0.69 0.67</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3F9CAFA02F931" box="[154,1412,1655,1718]" gridrow="12" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3F9CAFD96F931" box="[154,528,1655,1718]" gridcol="0" gridrow="12" pageId="3" pageNumber="20">Ratios of lengths of stylus II (without spine) and urocoxite VII</th>
<td id="E30C5DA0FF8FB7894461F9CAFCA9F931" box="[552,815,1655,1718]" gridcol="1" gridrow="12" pageId="3" pageNumber="20">0.91 0.77</td>
<td id="E30C5DA0FF8FB7894506F9CAFBE6F931" box="[847,1120,1655,1718]" gridcol="2" gridrow="12" pageId="3" pageNumber="20">0.77 0.61</td>
<td id="E30C5DA0FF8FB789423EF9CAFA02F931" box="[1143,1412,1655,1718]" gridcol="3" gridrow="12" pageId="3" pageNumber="20">0.85 0.74</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3F975FA02F958" box="[154,1412,1736,1759]" gridrow="13" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3F975FD96F958" box="[154,528,1736,1759]" gridcol="0" gridrow="13" pageId="3" pageNumber="20">Posterior angle of urosternite II</th>
<td id="E30C5DA0FF8FB7894461F975FCA9F958" box="[552,815,1736,1759]" gridcol="1" gridrow="13" pageId="3" pageNumber="20">128°</td>
<td id="E30C5DA0FF8FB7894506F975FBE6F958" box="[847,1120,1736,1759]" gridcol="2" gridrow="13" pageId="3" pageNumber="20">108°</td>
<td id="E30C5DA0FF8FB789423EF975FA02F958" box="[1143,1412,1736,1759]" gridcol="3" gridrow="13" pageId="3" pageNumber="20">118°</td>
</tr>
<tr id="A0DD34DCFF8FB78946D3F94DFA02F880" box="[154,1412,1776,1799]" gridrow="14" pageId="3" pageNumber="20">
<th id="E30C5DA0FF8FB78946D3F94DFD96F880" box="[154,528,1776,1799]" gridcol="0" gridrow="14" pageId="3" pageNumber="20">Number of ovipositor divisions</th>
<td id="E30C5DA0FF8FB7894461F94DFCA9F880" box="[552,815,1776,1799]" gridcol="1" gridrow="14" pageId="3" pageNumber="20">55</td>
<td id="E30C5DA0FF8FB7894506F94DFBE6F880" box="[847,1120,1776,1799]" gridcol="2" gridrow="14" pageId="3" pageNumber="20">?</td>
<td id="E30C5DA0FF8FB789423EF94DFA02F880" box="[1143,1412,1776,1799]" gridcol="3" gridrow="14" pageId="3" pageNumber="20">52–53</td>
</tr>
</table>
</paragraph>
<paragraph id="1E52369EFF8F487546D9F899FEB3F8F4" blockId="3.[144,1429,1826,1907]" pageId="3" pageNumber="20">
<tableNote id="E30B3710FF8F487546D9F899FEB3F8F4" pageId="3" pageNumber="20" targetBox="[154,1412,720,1799]" targetPageId="3">
<emphasis id="2C99EA8CFF8F487546D9F899FF39F8BE" box="[144,191,1828,1849]" italics="true" pageId="3" pageNumber="20">Note</emphasis>
.
<taxonomicName id="D9ED4D1DFF8F48754682F89FFDCFF8BF" authority="Silvestri, 1943" authorityName="Silvestri" authorityYear="1943" box="[203,585,1826,1849]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="takunagae">
<emphasis id="2C99EA8CFF8F48754682F89FFE28F8BE" box="[203,430,1826,1849]" italics="true" pageId="3" pageNumber="20">Petrobiellus takunagae</emphasis>
<bibRefCitation id="7A7C4B6FFF8F487547FCF89FFDCFF8BF" author="Silvestri F." box="[437,585,1826,1849]" pageId="3" pageNumber="20" pagination="283 - 306" refId="ref2458" refString="Silvestri F. 1943. Contributto alla conoscenza dei Machilidae (Insecta, Thysanura) del Giappone. Bollettino del Laboratorio di Zoologia, generale e agraria della R. Scuola superiore d'Agricoltura in Portici, 32: 283 - 306." type="journal article" year="1943">Silvestri, 1943</bibRefCitation>
</taxonomicName>
is not included in the table due to insufficient data in its original description; the characters distinguishing this species from
<taxonomicName id="D9ED4D1DFF8F4875440EF8FDFD67F8D1" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[583,737,1855,1878]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8F4875440EF8FDFD67F8D1" box="[583,737,1855,1878]" italics="true" pageId="3" pageNumber="20">P. sachalinensis</emphasis>
</taxonomicName>
<emphasis id="2C99EA8CFF8F487544AFF8FDFCB1F8D2" bold="true" box="[742,823,1856,1877]" pageId="3" pageNumber="20">
<taxonomicNameLabel id="37AA57F7FF8F487544AFF8FDFCB1F8D2" box="[742,823,1856,1877]" pageId="3" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are given in the
<emphasis id="2C99EA8CFF8F487545A9F8FDFBDEF8D2" box="[992,1112,1856,1877]" italics="true" pageId="3" pageNumber="20">Comparison</emphasis>
section after the description of the new species.
</tableNote>
</paragraph>
<caption id="4A926616FF88487246D6F9A0FCF0F912" ID-DOI="http://doi.org/10.5281/zenodo.10135595" ID-Zenodo-Dep="10135595" httpUri="https://zenodo.org/record/10135595/files/figure.png" pageId="4" pageNumber="21" startId="4.[159,215,1565,1589]" targetBox="[159,1000,181,1539]" targetPageId="4">
<paragraph id="1E52369EFF88487246D6F9A0FCF0F912" blockId="4.[159,990,1564,1685]" pageId="4" pageNumber="21">
<emphasis id="2C99EA8CFF88487246D6F9A0FEB3F9B2" bold="true" box="[159,309,1565,1589]" pageId="4" pageNumber="21">Figs 10–13.</emphasis>
<emphasis id="2C99EA8CFF8848724774F9A1FD06F9B2" box="[317,640,1564,1589]" italics="true" pageId="4" pageNumber="21">Petrobiellus sachalinensis</emphasis>
<emphasis id="2C99EA8CFF88487244CEF9A0FD6CF9B2" bold="true" box="[647,746,1565,1589]" pageId="4" pageNumber="21">sp. nov.</emphasis>
, holotype (female).
<emphasis id="2C99EA8CFF88487246D6F980FF3DF9D2" bold="true" box="[159,187,1597,1621]" pageId="4" pageNumber="21">10</emphasis>
, urosternite and urocoxites (part) V;
<emphasis id="2C99EA8CFF8848724435F980FD11F9D2" bold="true" box="[636,663,1597,1621]" pageId="4" pageNumber="21">11</emphasis>
, urocoxites VIII (part) and IX, with posterior gonapophyses;
<emphasis id="2C99EA8CFF8848724461F9E0FDC2F9F2" bold="true" box="[552,580,1629,1653]" pageId="4" pageNumber="21">12</emphasis>
, distal part of anterior gonapophy- ses;
<emphasis id="2C99EA8CFF8848724684F9C0FF6FF912" bold="true" box="[205,233,1661,1685]" pageId="4" pageNumber="21">13</emphasis>
, distal part of posterior gonapophyses. Scale bar: 0.1 mm
</paragraph>
</caption>
<paragraph id="1E52369EFF884872453CF9C3FCFFF912" blockId="4.[159,990,1564,1685]" box="[885,889,1662,1685]" pageId="4" pageNumber="21">.</paragraph>
<paragraph id="1E52369EFF884872468CF974FAC2FCCA" blockId="4.[159,779,1736,1906]" lastBlockId="4.[1013,1445,185,1729]" pageId="4" pageNumber="21">
Apical article of maxillary palp 0.60–0.62 times as long as preceding one. Fifth article 1.04– 1.06 times as long as 4th article (
<figureCitation id="86D62A1BFF8848724402F8ADFD1AF8AC" box="[587,668,1807,1835]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="4" pageNumber="21">Fig. 6</figureCitation>
). Dorsal surface of 7th, 6th and 5th articles of maxillary palp with 13–16, 10–13 and 3 hyaline spines, respectively. Apical article of labial palp fingerlike, 3.3–3.4 times as long as wide (
<figureCitation id="86D62A1BFF88487242F8FF42FA82FE9E" box="[1201,1284,255,281]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="4" pageNumber="21">Fig. 7</figureCitation>
). Mandibles with 2–3 distal teeth (
<figureCitation id="86D62A1BFF8848724346FE9FFAD3FEBB" box="[1295,1365,290,316]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="4" pageNumber="21">Fig 8</figureCitation>
). Femora slightly widened. Ratios of length to width of femur, tibia and tarsus as shown in
<tableCitation id="536F0325FF8848724333FE36FBB3FE4F" captionStart="Table 1" captionStartId="2.[824,884,1649,1670]" captionTargetBox="[835,1412,1740,1913]" captionTargetId="graphics-154@2.[824,1443,1728,1919]" captionTargetPageId="2" captionText="Table 1. Ratios of the length to width of the leg parts in females of Petrobiellus sachalinensis sp. nov." httpUri="http://table.plazi.org/id/4A926616FF8E48744571F9CCFB74F924" pageId="4" pageNumber="21" tableUuid="4A926616FF8E48744571F9CCFB74F924">Table 1</tableCitation>
. Fore and middle legs shorter than hind legs. Fore and middle tarsus and tibia 0.77 and 0.83 times as long as those of hind legs, respectively. Ratio of length of apical hind tarsomere to total length of hind tarsus about 0.32. Ventral surface of legs without long thin bristles. Fore, middle and hind tarsi and hind tibia with slightly pigmented spines (
<figureCitation id="86D62A1BFF88487242F0FCB2FA81FCAD" box="[1209,1287,783,810]" captionStart="Figs 2–9" captionStartId="2.[159,213,1474,1498]" captionTargetBox="[159,1444,181,1447]" captionTargetId="figure-79@2.[159,1444,181,1447]" captionTargetPageId="2" captionText="Figs 2–9. Petrobiellus sachalinensis sp. nov., holotype (female). 2, distal chain of flagellum; 3, apex of cercus; 4, divisions in middle part of cercus; 5, anterior part of head (vertex, compound eyes, paired ocelli, frons, antennae bases, median ocellus, clypeus, and labrum); 6, maxillary palpus; 7, labial palpus and labium (part); 8, apex of mandible; 9, fore leg (part). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135593" httpUri="https://zenodo.org/record/10135593/files/figure.png" pageId="4" pageNumber="21">Fig. 9</figureCitation>
). Number of spines as shown in
<tableCitation id="536F0325FF88487242ABFC8EFAB9FCCA" box="[1250,1343,819,845]" captionStart="Table 2" captionStartId="3.[144,203,184,205]" captionTargetBox="[154,712,270,558]" captionTargetId="graphics-35@3.[144,762,255,566]" captionTargetPageId="3" captionText="Table 2. Number of spines at the legs in females of Petrobiellus sachalinensis sp. nov." httpUri="http://table.plazi.org/id/4A926616FF8F487546D9FF05FE0CFF6D" pageId="4" pageNumber="21" tableUuid="4A926616FF8F487546D9FF05FE0CFF6D">Table 2</tableCitation>
.
</paragraph>
<paragraph id="1E52369EFF8848724255FCEBFA0BFC5D" blockId="4.[1013,1445,185,1729]" pageId="4" pageNumber="21">Middle and hind legs with coxal styli. Ratio of length of styli to width of middle and hind coxae 1.03–1.06 and 1.09, respectively.</paragraph>
<paragraph id="1E52369EFF8848724255FC5EFB56F988" blockId="4.[1013,1445,185,1729]" pageId="4" pageNumber="21">
Urites I, VI and VII with 1 + 1, urites II–V with 2 + 2 eversible vesicles (
<figureCitation id="86D62A1BFF8848724221FB97FB4EFBC3" box="[1128,1224,1066,1092]" captionStart="Figs 10–13" captionStartId="4.[159,215,1565,1589]" captionTargetBox="[159,1000,181,1539]" captionTargetId="figure-367@4.[159,1000,181,1539]" captionTargetPageId="4" captionText="Figs 10–13. Petrobiellus sachalinensis sp. nov., holotype (female). 10, urosternite and urocoxites (part) V; 11, urocoxites VIII (part) and IX, with posterior gonapophyses; 12, distal part of anterior gonapophy- ses; 13, distal part of posterior gonapophyses. Scale bar: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.10135595" httpUri="https://zenodo.org/record/10135595/files/figure.png" pageId="4" pageNumber="21">Fig. 10</figureCitation>
). Posterior angle of urosternites II, III–V, VI, and VII approximately 118°, 110–112°, 107°, and 120°, respectively. Inner posterior lobes of urocoxites VII protruding. Ratio width to length of protruding lobe about 1.0. Ratios of lengths of stylus (without apical spine), urosternites and urocoxites II–IX as shown in
<tableCitation id="536F0325FF8848724333FAD5FBB3FA22" captionStart="Table 3" captionStartId="3.[809,868,184,205]" captionTargetBox="[820,1418,268,560]" captionTargetId="graphics-127@3.[809,1429,255,566]" captionTargetPageId="3" captionText="Table 3. Ratios of the lengths of some abdominal structures in females of Petrobiellus sachalinensis sp. nov." httpUri="http://table.plazi.org/id/4A926616FF8F48754560FF05FA86FF6D" pageId="4" pageNumber="21" tableUuid="4A926616FF8F48754560FF05FA86FF6D">Table 3</tableCitation>
. Ratio of lengths of urosternite and urocoxites I about 0.35. Urocoxites IX with 1 + 1 outer sublateral spines.
</paragraph>
<paragraph id="1E52369EFF8848734252F9A5FD9FFF71" blockId="4.[1013,1445,185,1729]" lastBlockId="5.[144,764,185,1607]" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21">
Ovipositor slender, elongate, significantly protruding beyond apices of styli IX. Anterior and posterior gonapophyses with 52 or 53 divisions (
<figureCitation id="86D62A1BFF884872428BF91BFA99F947" box="[1218,1311,1702,1728]" captionStart="Figs 10–13" captionStartId="4.[159,215,1565,1589]" captionTargetBox="[159,1000,181,1539]" captionTargetId="figure-367@4.[159,1000,181,1539]" captionTargetPageId="4" captionText="Figs 10–13. Petrobiellus sachalinensis sp. nov., holotype (female). 10, urosternite and urocoxites (part) V; 11, urocoxites VIII (part) and IX, with posterior gonapophyses; 12, distal part of anterior gonapophy- ses; 13, distal part of posterior gonapophyses. Scale bar: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.10135595" httpUri="https://zenodo.org/record/10135595/files/figure.png" pageId="4" pageNumber="21">Fig. 11</figureCitation>
). Four basal divisions of anterior gonapophyses and about 11 basal divisions of posterior gonapophyses glabrous. Apical spines of gonapophyses as long as three apical divisions combined. Distal divisions of anterior and posterior gonapophyses with seven or eight setae (not counting sensory setae and apical spines) (
<figureCitation id="86D62A1BFF8948734732FF61FD8FFF71" box="[379,521,220,246]" captionStart="Figs 10–13" captionStartId="4.[159,215,1565,1589]" captionTargetBox="[159,1000,181,1539]" captionTargetId="figure-367@4.[159,1000,181,1539]" captionTargetPageId="4" captionText="Figs 10–13. Petrobiellus sachalinensis sp. nov., holotype (female). 10, urosternite and urocoxites (part) V; 11, urocoxites VIII (part) and IX, with posterior gonapophyses; 12, distal part of anterior gonapophy- ses; 13, distal part of posterior gonapophyses. Scale bar: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.10135595" httpUri="https://zenodo.org/record/10135595/files/figure.png" pageId="5" pageNumber="22">Figs 12, 13</figureCitation>
).
</paragraph>
<paragraph id="1E52369EFF89487346FCFEBDFEE7FEBA" blockId="5.[144,764,185,1607]" pageId="5" pageNumber="22">Males are unknown; probably the species is parthenogenetic.</paragraph>
</subSubSection>
<subSubSection id="56F76515FF89487346FCFEF5FEEFFA14" pageId="5" pageNumber="22" type="diagnosis">
<paragraph id="1E52369EFF89487346FCFEF5FEEFFA14" blockId="5.[144,764,185,1607]" pageId="5" pageNumber="22">
<emphasis id="2C99EA8CFF89487346FCFEF5FD15FEE6" box="[181,659,327,354]" italics="true" pageId="5" pageNumber="22">
Comparison.
<taxonomicName id="D9ED4D1DFF8948734711FEFAFD15FEE6" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[344,659,327,353]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">Petrobiellus sachalinensis</taxonomicName>
</emphasis>
<emphasis id="2C99EA8CFF89487344D0FEFAFD7AFEE6" bold="true" box="[665,764,327,353]" pageId="5" pageNumber="22">
<taxonomicNameLabel id="37AA57F7FF89487344D0FEFAFD7AFEE6" box="[665,764,327,353]" pageId="5" pageNumber="22" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from its congeners by the following characters: the general body colour whitish or slightly yellowish (
<emphasis id="2C99EA8CFF894873479DFE0EFE6BFE4B" box="[468,493,435,460]" italics="true" pageId="5" pageNumber="22">vs</emphasis>
yellowish or rufous); colour of scales whitish or light gray (
<emphasis id="2C99EA8CFF8948734437FE6AFD11FE77" box="[638,663,471,496]" italics="true" pageId="5" pageNumber="22">vs</emphasis>
brownish or dark brownish); body (excluding head and tarsi) almost without pigment (
<emphasis id="2C99EA8CFF8948734471FDA3FDD7FDB0" box="[568,593,542,567]" italics="true" pageId="5" pageNumber="22">vs</emphasis>
significantly pigmented); and paired ocelli almost sole-shaped (
<emphasis id="2C99EA8CFF89487346D3FDD8FF35FDF9" box="[154,179,613,638]" italics="true" pageId="5" pageNumber="22">vs</emphasis>
pistiliform, strongly narrowed in middle parts). The new species also differs from all other species of the genus in the most elongated compound eyes; relatively short coxal styli; long 4th article of maxillary palp; small number of short supporting spines in ventral part of tarsi; ratios of lengths of urosternite and urocoxite, stylus and urocoxite; posterior angle of urosternite; and number of divisions of ovipositor. The main morphological differences of females of
<taxonomicName id="D9ED4D1DFF894873478EFC1BFD0AFC47" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[455,652,934,960]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF894873478EFC1BFD0AFC47" box="[455,652,934,960]" italics="true" pageId="5" pageNumber="22">P. sachalinensis</emphasis>
</taxonomicName>
<emphasis id="2C99EA8CFF89487344DCFC1BFD7AFC47" bold="true" box="[661,764,934,960]" pageId="5" pageNumber="22">
<taxonomicNameLabel id="37AA57F7FF89487344DCFC1BFD7AFC47" box="[661,764,934,960]" pageId="5" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from those of the other species of
<taxonomicName id="D9ED4D1DFF8948734472FC74FD4AFC64" authorityName="Silvestri" authorityYear="1943" box="[571,716,969,995]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="2C99EA8CFF8948734472FC74FD4AFC64" box="[571,716,969,995]" italics="true" pageId="5" pageNumber="22">Petrobiellus</emphasis>
</taxonomicName>
are given in
<tableCitation id="536F0325FF8948734748FC50FEE4FB8F" box="[257,354,1005,1032]" captionStart="Table 4" captionStartId="3.[144,203,665,686]" captionTargetBox="[154,1412,720,1799]" captionTargetId="graphics-220@3.[144,1428,707,1804]" captionTargetPageId="3" captionText="Table 4. Main morphological differences between females of the species of Petrobiellus." httpUri="http://table.plazi.org/id/4A926616FF8F487546D9FD24FB84FD29" pageId="5" pageNumber="22" tableUuid="4A926616FF8F487546D9FD24FB84FD29">Table 4</tableCitation>
. According to the brief description of
<taxonomicName id="D9ED4D1DFF89487346A4FBAFFE17FBAC" authorityName="Silvestri" authorityYear="1943" box="[237,401,1041,1067]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="takunagae">
<emphasis id="2C99EA8CFF89487346A4FBAFFE17FBAC" box="[237,401,1041,1067]" italics="true" pageId="5" pageNumber="22">P. takunagae</emphasis>
</taxonomicName>
by
<bibRefCitation id="7A7C4B6FFF894873478DFBACFD0DFBAC" author="Silvestri F." box="[452,651,1041,1067]" pageId="5" pageNumber="22" pagination="283 - 306" refId="ref2458" refString="Silvestri F. 1943. Contributto alla conoscenza dei Machilidae (Insecta, Thysanura) del Giappone. Bollettino del Laboratorio di Zoologia, generale e agraria della R. Scuola superiore d'Agricoltura in Portici, 32: 283 - 306." type="journal article" year="1943">Silvestri (1943)</bibRefCitation>
, the distance between the paired ocelli in this species is about half the length of ocellus, and in
<taxonomicName id="D9ED4D1DFF8948734432FBE7FF5BFB10" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sachalinensis" status="sp. nov.">
<emphasis id="2C99EA8CFF8948734432FBE7FF5BFB10" italics="true" pageId="5" pageNumber="22">P. sachalinensis</emphasis>
</taxonomicName>
<emphasis id="2C99EA8CFF89487346A0FBC0FED4FB10" bold="true" box="[233,338,1149,1175]" pageId="5" pageNumber="22">
<taxonomicNameLabel id="37AA57F7FF89487346A0FBC0FED4FB10" box="[233,338,1149,1175]" pageId="5" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, this distance is almost equal to the length of ocellus. The new species differs from
<taxonomicName id="D9ED4D1DFF89487346D9FB7BFEB5FB58" authorityName="Silvestri" authorityYear="1943" box="[144,307,1221,1247]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="takunagae">
<emphasis id="2C99EA8CFF89487346D9FB7BFEB5FB58" box="[144,307,1221,1247]" italics="true" pageId="5" pageNumber="22">P. takunagae</emphasis>
</taxonomicName>
also in femora without pigment and tibiae poorly pigmented (
<emphasis id="2C99EA8CFF894873478EFB57FE66FA84" box="[455,480,1258,1283]" italics="true" pageId="5" pageNumber="22">vs</emphasis>
femora and tibiae with intensive blackish-violet pigment); and four basal divisions of anterior gonapophyses glabrous (
<emphasis id="2C99EA8CFF89487346D5FAE8FF33FAE9" box="[156,181,1365,1390]" italics="true" pageId="5" pageNumber="22">vs</emphasis>
about 12 basal divisions of anterior gonapophyses glabrous).
</paragraph>
</subSubSection>
<subSubSection id="56F76515FF89487346FCFA21FE4AFA5D" pageId="5" pageNumber="22" type="etymology">
<paragraph id="1E52369EFF89487346FCFA21FE4AFA5D" blockId="5.[144,764,185,1607]" pageId="5" pageNumber="22">
<emphasis id="2C99EA8CFF89487346FCFA21FEB1FA31" box="[181,311,1436,1462]" italics="true" pageId="5" pageNumber="22">Etymology</emphasis>
. The species name is derived from the toponym of
<collectingRegion id="DC29F87CFF894873471DFA7DFE4EFA5D" box="[340,456,1472,1498]" country="Russia" name="Sakhalin" pageId="5" pageNumber="22">Sakhalin</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="56F76515FF89487346FCFA59FDF8F9C0" pageId="5" pageNumber="22" type="biology_ecology">
<paragraph id="1E52369EFF89487346FCFA59FDF8F9C0" blockId="5.[144,764,185,1607]" pageId="5" pageNumber="22">
<emphasis id="2C99EA8CFF89487346FCFA59FEAFFA79" box="[181,297,1508,1534]" italics="true" pageId="5" pageNumber="22">Habitats.</emphasis>
Western shores of the northern part of
<collectingRegion id="DC29F87CFF89487346F8F9B5FEA4F9A5" box="[177,290,1544,1570]" country="Russia" name="Sakhalin" pageId="5" pageNumber="22">Sakhalin</collectingRegion>
Island. Under stones near the line of seashore (Fig. 14; see Addenda section).
</paragraph>
</subSubSection>
</treatment>
</document>
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