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<document id="C42F60E80D4B91E86F0EA7B30A941EF7" ID-DOI="10.5281/zenodo.5352924" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352924" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630209239027" checkinUser="felipe" docAuthor="Edmonds, W. D. &amp; Zidek, J." docDate="2010" docId="3A7F7B2C7F58885AFF34FAB22341E9F2" docLanguage="en" docName="0129.pdf" docOrigin="Insecta Mundi 2010 (129)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Coprophanaeus (Metallophanaeus) pertyi" docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="C64603547F7C887DFFF4FFB02050E849" masterDocTitle="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="111" masterPageNumber="1" pageNumber="33" updateTime="1699186523840" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="DB71452F72495AFBD1AAF1E4564147F6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="25D2E53F74320DF29536AC8BFA8809F5">
<mods:title id="16B491474ABD9B719E7C2CDAD87FEA73">A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="D96F81D54C4597F5231D964FE636BB55" type="personal">
<mods:role id="879A23947A094296AA010D6F2A0068C8">
<mods:roleTerm id="E6284B45A4A38D16898D17192A62F7AA">Author</mods:roleTerm>
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<mods:namePart id="F1922EC00C7B9E5A3587DA42E9317BB9">Edmonds, W. D.</mods:namePart>
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<mods:name id="8E8F23CE5146A67E4FA622282365CED4" type="personal">
<mods:role id="2A04922F1AD6C20EF74AEF5B5AA5DCF4">
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</mods:role>
<mods:namePart id="888CED333DCF55583EBB28EA37E55731">Zidek, J.</mods:namePart>
<mods:affiliation id="EFF21B3A5ACD2417BAD2566860147973">Coprophanaeus bonariensis (Gory)</mods:affiliation>
</mods:name>
<mods:typeOfResource id="225121C014BB484B0124B1BD10B169C2">text</mods:typeOfResource>
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<mods:title id="8968411AA611C73944288E09C96137AE">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="A94891511358BC24D7297EF3D3CC0101">
<mods:date id="120E95E20BE74D0C65BBC3AD0171DA02">2010</mods:date>
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<mods:number id="56E60E366F4A31CCE6E85026967E741C">2010-07-02</mods:number>
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<mods:detail id="5ECBDDADAD654622848AE63FB8F924F0" type="volume">
<mods:number id="7D272965BCA779D9D3A099CADE4DEBE3">2010</mods:number>
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<mods:detail id="DDCB6E1706FA07AF8FF710F2F913AC22" type="issue">
<mods:number id="9A45B753DEFA759DF161BD4E40999211">129</mods:number>
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<mods:start id="2428EB60CB2FE1642CD776177625E425">1</mods:start>
<mods:end id="A9472264BC04585D43AC3CBDCDA49110">111</mods:end>
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<mods:classification id="15F44A64643601EAF1500DD9F7C9F140">journal article</mods:classification>
<mods:identifier id="4F6D784EFE7600B904AE0AD32035F436" type="DOI">10.5281/zenodo.5352924</mods:identifier>
<mods:identifier id="16643C9FCCCC2E818705AC7A826CE2D9" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="A27B8E361BAA4FAF8E88CAACA5C3F4B0" type="Zenodo-Dep">5352924</mods:identifier>
</mods:mods>
<treatment id="3A7F7B2C7F58885AFF34FAB22341E9F2" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2" httpUri="http://treatment.plazi.org/id/3A7F7B2C7F58885AFF34FAB22341E9F2" lastPageId="39" lastPageNumber="35" pageId="36" pageNumber="33">
<subSubSection id="FACC99B17F588859FF34FAB22386ED52" box="[192,982,1282,1307]" pageId="36" pageNumber="33" type="nomenclature">
<paragraph id="B269CA3A7F588859FF34FAB22386ED52" blockId="36.[192,982,1282,1339]" box="[192,982,1282,1307]" pageId="36" pageNumber="33">
<heading id="E9217D567F588859FF34FAB22386ED52" bold="true" box="[192,982,1282,1307]" fontSize="10" level="2" pageId="36" pageNumber="33" reason="3">
<taxonomicName id="75D6B1B97F588859FF34FAB22386ED52" ID-CoL="Y6CG" authority="(Olsoufieff, 1924)" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[192,982,1282,1307]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="pertyi" subGenus="Metallophanaeus">
<emphasis id="80A216287F588859FF34FAB22386ED52" bold="true" box="[192,982,1282,1307]" pageId="36" pageNumber="33">
<emphasis id="80A216287F588859FF34FAB221C6ED52" bold="true" box="[192,406,1282,1307]" italics="true" pageId="36" pageNumber="33">Coprophanaeus</emphasis>
(
<emphasis id="80A216287F588859FE51FAB222C1ED52" bold="true" box="[421,657,1282,1307]" italics="true" pageId="36" pageNumber="33">Metallophanaeus</emphasis>
)
<emphasis id="80A216287F588859FD6BFAB322BEED52" bold="true" box="[671,750,1283,1307]" italics="true" pageId="36" pageNumber="33">pertyi</emphasis>
(
<bibRefCitation id="D647B7CB7F588859FD0FFAB2239DED52" author="Olsoufieff, G." box="[763,973,1282,1307]" pageId="36" pageNumber="33" pagination="4 - 201" refId="ref59457" refString="Olsoufieff, G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F588859FF34FA9321EEED72" box="[192,446,1315,1339]" pageId="36" pageNumber="33" type="description">
<paragraph id="B269CA3A7F588859FF34FA9321EEED72" blockId="36.[192,982,1282,1339]" box="[192,446,1315,1339]" pageId="36" pageNumber="33">
<figureCitation id="2AEDD6BF7F588859FF34FA932112ED72" box="[192,322,1315,1339]" captionStart="Figure 50-61" captionStartId="29.[192,276,1484,1506]" captionTargetBox="[191,1440,190,1454]" captionTargetId="figure-130@29.[191,1440,190,1454]" captionTargetPageId="29" captionText="Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai." figureDoi="http://doi.org/10.5281/zenodo.5352946" httpUri="https://zenodo.org/record/5352946/files/figure.png" pageId="36" pageNumber="33">Fig. 51, 57</figureCitation>
,
<figureCitation id="2AEDD6BF7F588859FEBAFA93213FED72" box="[334,367,1315,1339]" captionStart="Figure 62" captionStartId="30.[191,275,1894,1916]" captionTargetBox="[247,1385,192,1865]" captionTargetId="figure-18@30.[247,1386,191,1865]" captionTargetPageId="30" captionText="Figure 62. Approximate geographic distribution of Coprophanaeus (Metallophanaeus) species." figureDoi="http://doi.org/10.5281/zenodo.5352948" httpUri="https://zenodo.org/record/5352948/files/figure.png" pageId="36" pageNumber="33">62</figureCitation>
,
<figureCitation id="2AEDD6BF7F588859FE8DFA9321EEED72" box="[377,446,1315,1339]" captionStart="Figure 80-84" captionStartId="37.[192,276,1504,1526]" captionTargetBox="[271,1356,207,1484]" captionTargetId="figure-112@37.[263,1374,192,1488]" captionTargetPageId="37" captionText="Figure 80-84. Coprophanaeus pertyi. 80-81) Male habitus. 82) Aedeagus (dorsal view on left; lateral view on right). 83-84) Female habitus." figureDoi="http://doi.org/10.5281/zenodo.5352956" httpUri="https://zenodo.org/record/5352956/files/figure.png" pageId="36" pageNumber="33">80-84</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F588859FF34FAD323B3ED92" pageId="36" pageNumber="33" type="reference_group">
<paragraph id="B269CA3A7F588859FF34FAD32225ED32" blockId="36.[192,1396,1379,1499]" box="[192,629,1379,1403]" pageId="36" pageNumber="33">
<treatmentCitationGroup id="92C6ED147F588859FF34FAD32225ED32" box="[192,629,1379,1403]" pageId="36" pageNumber="33">
<treatmentCitation id="3377EC2B7F588859FF34FAD32225ED32" author="Olsoufieff, G." box="[192,629,1379,1403]" page="71" pageId="36" pageNumber="33" year="1924">
<taxonomicName id="75D6B1B97F588859FF34FAD32225ED32" ID-CoL="4FSJ4" authority="Olsoufieff, 1924: 71" authorityName="Olsoufieff" authorityPageNumber="71" authorityYear="1924" box="[192,629,1379,1403]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="pertyi">
<emphasis id="80A216287F588859FF34FAD321D4ED32" box="[192,388,1379,1403]" italics="true" pageId="36" pageNumber="33">Phanaeus pertyi</emphasis>
<bibRefCitation id="D647B7CB7F588859FE7EFAD32225ED32" author="Olsoufieff, G." box="[394,629,1379,1403]" pageId="36" pageNumber="33" pagination="4 - 201" refId="ref59457" refString="Olsoufieff, G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">Olsoufieff, 1924: 71</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B269CA3A7F588859FF34FA3323B3ED92" blockId="36.[192,1396,1379,1499]" pageId="36" pageNumber="33">
<treatmentCitationGroup id="92C6ED147F588859FF34FA3323B3ED92" pageId="36" pageNumber="33">
<taxonomicName id="75D6B1B97F588859FF34FA332297EDD2" ID-CoL="4FSP2" authority="sensu Felsche, 1901" authorityName="Felsche" authorityYear="1901" box="[192,711,1411,1435]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis id="80A216287F588859FF34FA332249EDD2" box="[192,537,1411,1435]" italics="true" pageId="36" pageNumber="33">Phanaeus thalassinus sensu</emphasis>
<bibRefCitation id="D647B7CB7F588859FDEAFA332297EDD2" author="Felsche, C." box="[542,711,1411,1435]" pageId="36" pageNumber="33" pagination="135 - 160" refId="ref57396" refString="Felsche, C. 1901. Beschreibungen coprophager Scarabaeiden. Deutsche Entomologische Zeitschrift 1901: 135 - 160." type="journal article" year="1901">Felsche, 1901</bibRefCitation>
</taxonomicName>
,
<emphasis id="80A216287F588859FD26FA3422AAEDD2" box="[722,762,1412,1435]" italics="true" pageId="36" pageNumber="33">nec</emphasis>
<bibRefCitation id="D647B7CB7F588859FCF4FA3423DDEDD2" author="Perty, M." box="[768,909,1411,1436]" pageId="36" pageNumber="33" refId="ref59736" refString="Perty, M. 1830. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII- MDCCCXX jussu et auspiciis Maximiliani Josephi I … fasc. 1. Impensis Editoris; Munich. 44 p., 12 plates." type="book" year="1830">Perty, 1830</bibRefCitation>
(syn. by
<treatmentCitation id="3377EC2B7F588859FC03FA33253BEDD2" author="Martinez, A. &amp; F. S. Pereira" box="[1015,1387,1411,1435]" page="66" pageId="36" pageNumber="33" year="1967">
<bibRefCitation id="D647B7CB7F588859FC03FA33253BEDD2" author="Martinez, A. &amp; F. S. Pereira" box="[1015,1387,1411,1435]" pageId="36" pageNumber="33" pagination="53 - 69" refId="ref59045" refString="Martinez, A., and F. S. Pereira. 1967. Notas escarabeidologicas III. (Col. Scarabaeidae - Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez and Pereira 1967: 66</bibRefCitation>
</treatmentCitation>
)
<treatmentCitation id="3377EC2B7F588859FF34FA132331EDF2" author="Pereira, F. S. &amp; M. A. V. d'Andretta" box="[192,865,1443,1467]" page="253" pageId="36" pageNumber="33" year="1955">
<taxonomicName id="75D6B1B97F588859FF34FA132331EDF2" ID-CoL="4FSLG" authority="Pereira and d'Andretta, 1955: 253" authorityName="Pereira and d'Andretta" authorityPageNumber="253" authorityYear="1955" box="[192,865,1443,1467]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="80A216287F588859FF34FA132192EDF2" box="[192,450,1443,1467]" italics="true" pageId="36" pageNumber="33">Phanaeus alvarengai</emphasis>
<bibRefCitation id="D647B7CB7F588859FE33FA132331EDF2" author="Pereira, F. S. &amp; M. A. V. d'Andretta" box="[455,865,1443,1467]" pageId="36" pageNumber="33" pagination="247 - 264" refId="ref59634" refString="Pereira, F. S., and M. A. V. d'Andretta. 1955. Novos escarabeideos e novas sinonimias. Papeis Avulsos do Departamento de Zoologia 12: 247 - 264." type="journal article" year="1955">Pereira and d’Andretta, 1955: 253</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(syn. by
<treatmentCitation id="3377EC2B7F588859FC3FFA13256CEDF2" author="Martinez, A. &amp; F. S. Pereira" box="[971,1340,1443,1467]" page="67" pageId="36" pageNumber="33" year="1967">
<bibRefCitation id="D647B7CB7F588859FC3FFA13256CEDF2" author="Martinez, A. &amp; F. S. Pereira" box="[971,1340,1443,1467]" pageId="36" pageNumber="33" pagination="53 - 69" refId="ref59045" refString="Martinez, A., and F. S. Pereira. 1967. Notas escarabeidologicas III. (Col. Scarabaeidae - Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez and Pereira 1967: 67</bibRefCitation>
</treatmentCitation>
)
<taxonomicName id="75D6B1B97F588859FF34FA732208ED92" ID-CoL="Y6CG" authority="(Olsoufieff)" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[192,600,1475,1499]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="pertyi">
<emphasis id="80A216287F588859FF34FA73219AED92" box="[192,458,1475,1499]" italics="true" pageId="36" pageNumber="33">Coprophanaeus pertyi</emphasis>
(Olsoufieff)
</taxonomicName>
(recomb. by
<treatmentCitation id="3377EC2B7F588859FD19FA73238AED92" author="Edmonds, W. D." box="[749,986,1475,1499]" page="841" pageId="36" pageNumber="33" year="1972">
<bibRefCitation id="D647B7CB7F588859FD19FA73238AED92" author="Edmonds, W. D." box="[749,986,1475,1499]" pageId="36" pageNumber="33" pagination="731 - 874" refId="ref56889" refString="Edmonds, W. D. 1972. Comparative skeletal morphology, systematic and evolution of the phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds 1972: 841</bibRefCitation>
</treatmentCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F588859FF34F9B3249EEE72" pageId="36" pageNumber="33" type="materials_examined">
<paragraph id="B269CA3A7F588859FF34F9B3249EEE72" blockId="36.[192,1440,1539,1595]" pageId="36" pageNumber="33">
<emphasis id="80A216287F588859FF34F9B3215DEE52" bold="true" box="[192,269,1539,1563]" pageId="36" pageNumber="33">Type.</emphasis>
<taxonomicName id="75D6B1B97F588859FEE3F9B421D1EE52" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[279,385,1540,1563]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="pertyi">
<emphasis id="80A216287F588859FEE3F9B421D1EE52" box="[279,385,1540,1563]" italics="true" pageId="36" pageNumber="33">P. pertyi</emphasis>
</taxonomicName>
–
<materialsCitation id="02BEC0677F588859FE51F9B32527EE52" box="[421,1399,1539,1563]" location="Munich" municipality="Zoologische Staatssammlung" pageId="36" pageNumber="33" specimenCount="1" typeStatus="lectotype">
<typeStatus id="6D6D74987F588859FE51F9B32245EE52" box="[421,533,1539,1563]" pageId="36" pageNumber="33" type="lectotype">lectotype</typeStatus>
male,
<collectingMunicipality id="520D50407F588859FD99F9B3238CEE52" box="[621,988,1539,1563]" pageId="36" pageNumber="33">Zoologische Staatssammlung</collectingMunicipality>
,
<location id="B7099CE17F588859FC1FF9B3241AEE52" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F588859FC1FF9B3241AEE52" box="[1003,1098,1539,1563]" municipality="Zoologische Staatssammlung" name="Munich" pageId="36" pageNumber="33">Munich</location>
(des.
<bibRefCitation id="D647B7CB7F588859FB6CF9B3253DEE52" author="Arnaud, P." box="[1176,1389,1539,1563]" pageId="36" pageNumber="33" pagination="11 - 12" refId="ref56478" refString="Arnaud, P. 2002 a. Description de nouvelles especes de Phanaeides (Col. Scarabaeidae). Besoiro 7: 2 - 9, 11 - 12." type="journal article" year="2002">Arnaud 2002a: 6</bibRefCitation>
)
</materialsCitation>
;
<materialsCitation id="02BEC0677F588859FA73F9B4249BEE72" location="Universidade" municipality="Museo de Zoologia" pageId="36" pageNumber="33" specimenCount="1" stateProvince="Sao Paulo, Sao Paulo" typeStatus="holotype">
<taxonomicName id="75D6B1B97F588859FA73F9B42113EE72" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="80A216287F588859FA73F9B42113EE72" italics="true" pageId="36" pageNumber="33">P. alvarengai</emphasis>
</taxonomicName>
–
<typeStatus id="6D6D74987F588859FEA9F9932195EE72" box="[349,453,1571,1595]" pageId="36" pageNumber="33" type="holotype">holotype</typeStatus>
male,
<collectingMunicipality id="520D50407F588859FDE7F99422A4EE72" box="[531,756,1571,1595]" pageId="36" pageNumber="33">Museo de Zoologia</collectingMunicipality>
,
<location id="B7099CE17F588859FD09F99323F2EE72" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F588859FD09F99323F2EE72" box="[765,930,1571,1595]" municipality="Museo de Zoologia" name="Universidade" pageId="36" pageNumber="33" stateProvince="Sao Paulo, Sao Paulo">Universidade</location>
de
<collectingRegion id="701204D87F588859FC3DF993249BEE72" box="[969,1227,1571,1595]" isEnumeration="true" pageId="36" pageNumber="33">São Paulo, São Paulo</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F588858FF34F9D32260EF1E" lastPageId="37" lastPageNumber="34" pageId="36" pageNumber="33" type="diagnosis">
<paragraph id="B269CA3A7F588858FF34F9D321A9EE3E" blockId="36.[192,1440,1635,1883]" lastBlockId="37.[192,1440,1599,1879]" lastPageId="37" lastPageNumber="34" pageId="36" pageNumber="33">
<emphasis id="80A216287F588859FF34F9D32101EE32" bold="true" box="[192,337,1635,1659]" pageId="36" pageNumber="33">Diagnosis.</emphasis>
<emphasis id="80A216287F588859FEA3F9D321E9EE32" box="[343,441,1635,1659]" italics="true" pageId="36" pageNumber="33">General</emphasis>
– Paraocular areas (genae) nearly flat, strongly and densely punctate, with short carina lateral to eyes. Posteromedian portion of pronotum appearing smoother and shinier than remain- der of disk, simply punctate, less strongly and densely so near basal fossae; posterior angle normal, basal fossae small, incorporated into inner ends of distinct sulcus paralleling posterior margin. Salient anteromedian angle of metasternum (seen in profile,
<figureCitation id="2AEDD6BF7F588859FCA0F95323FAEEB2" box="[852,938,1763,1787]" captionStart="Figure 50-61" captionStartId="29.[192,276,1484,1506]" captionTargetBox="[191,1440,190,1454]" captionTargetId="figure-130@29.[191,1440,190,1454]" captionTargetPageId="29" captionText="Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai." figureDoi="http://doi.org/10.5281/zenodo.5352946" httpUri="https://zenodo.org/record/5352946/files/figure.png" pageId="36" pageNumber="33">Fig. 57</figureCitation>
) rounded. Striae (x5) weakly impressed, simple; bases of striae 1-4 impressed, not distinctly fossate. Elytral interstriae weakly convex. Ventral surface of protibia lateral to longitudinal carina mostly smooth, usually with single or double row of shallow, confluent punctures paralleling longitudinal carina (
<figureCitation id="2AEDD6BF7F588859FC4EF8F32441EF12" box="[954,1041,1859,1883]" captionStart="Figure 50-61" captionStartId="29.[192,276,1484,1506]" captionTargetBox="[191,1440,190,1454]" captionTargetId="figure-130@29.[191,1440,190,1454]" captionTargetPageId="29" captionText="Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai." figureDoi="http://doi.org/10.5281/zenodo.5352946" httpUri="https://zenodo.org/record/5352946/files/figure.png" pageId="36" pageNumber="33">Fig. 51</figureCitation>
, arrow). Pygidium lacking basal groove. Length
<quantity id="752E67DF7F598858FE8AF98F21ABEE1E" box="[382,507,1599,1623]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="37" pageNumber="34" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14-20 mm</quantity>
. Elytra dull black; pronotum, head, pygidium and venter with strong bluishgreen metallic reflections.
</paragraph>
<caption id="E6A99AB27F598858FF34FA502247EE5A" ID-DOI="http://doi.org/10.5281/zenodo.5352956" ID-Zenodo-Dep="5352956" httpUri="https://zenodo.org/record/5352956/files/figure.png" pageId="37" pageNumber="34" startId="37.[192,276,1504,1526]" targetBox="[271,1356,207,1484]" targetPageId="37">
<paragraph id="B269CA3A7F598858FF34FA502247EE5A" blockId="37.[192,1440,1504,1555]" pageId="37" pageNumber="34">
<emphasis id="80A216287F598858FF34FA502134EDBC" bold="true" box="[192,356,1504,1526]" pageId="37" pageNumber="34">Figure 80-84.</emphasis>
<taxonomicName id="75D6B1B97F598858FE98FA512233EDBF" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[364,611,1505,1526]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="34" phylum="Arthropoda" rank="species" species="pertyi">
<emphasis id="80A216287F598858FE98FA512233EDBF" box="[364,611,1505,1526]" italics="true" pageId="37" pageNumber="34">Coprophanaeus pertyi</emphasis>
</taxonomicName>
.
<emphasis id="80A216287F598858FD87FA5022E3EDBC" bold="true" box="[627,691,1504,1525]" pageId="37" pageNumber="34">80-81</emphasis>
) Male habitus.
<emphasis id="80A216287F598858FC9CFA5023DDEDBF" bold="true" box="[872,909,1504,1526]" pageId="37" pageNumber="34">82)</emphasis>
Aedeagus (dorsal view on left; lateral view on right).
<emphasis id="80A216287F598858FEFBFA4D2108EE5A" bold="true" box="[271,344,1533,1555]" pageId="37" pageNumber="34">83-84)</emphasis>
Female habitus.
</paragraph>
</caption>
<paragraph id="B269CA3A7F598858FF04F9CF2307EE9E" blockId="37.[192,1440,1599,1879]" pageId="37" pageNumber="34">
<emphasis id="80A216287F598858FF04F9CF217DEEDE" box="[240,301,1663,1687]" italics="true" pageId="37" pageNumber="34">Male</emphasis>
(
<figureCitation id="2AEDD6BF7F598858FECFF9CF21E4EEDE" box="[315,436,1663,1687]" captionStart="Figure 80-84" captionStartId="37.[192,276,1504,1526]" captionTargetBox="[271,1356,207,1484]" captionTargetId="figure-112@37.[263,1374,192,1488]" captionTargetPageId="37" captionText="Figure 80-84. Coprophanaeus pertyi. 80-81) Male habitus. 82) Aedeagus (dorsal view on left; lateral view on right). 83-84) Female habitus." figureDoi="http://doi.org/10.5281/zenodo.5352956" httpUri="https://zenodo.org/record/5352956/files/figure.png" pageId="37" pageNumber="34">Fig. 80-82</figureCitation>
) – Head with evenly tapering long horn, base of horn abruptly swollen. Pronotum of large male with deep anteromedian concavity at summit marked by a lobate tumosity. Tips of parameres only slightly swollen laterally, lacking distinct flanges.
</paragraph>
<paragraph id="B269CA3A7F598858FF04F96F221EEF7E" blockId="37.[192,1440,1599,1879]" pageId="37" pageNumber="34">
<emphasis id="80A216287F598858FF04F96F211AEEBE" box="[240,330,1759,1783]" italics="true" pageId="37" pageNumber="34">Female</emphasis>
(
<figureCitation id="2AEDD6BF7F598858FEACF96F2182EEBE" box="[344,466,1759,1783]" captionStart="Figure 80-84" captionStartId="37.[192,276,1504,1526]" captionTargetBox="[271,1356,207,1484]" captionTargetId="figure-112@37.[263,1374,192,1488]" captionTargetPageId="37" captionText="Figure 80-84. Coprophanaeus pertyi. 80-81) Male habitus. 82) Aedeagus (dorsal view on left; lateral view on right). 83-84) Female habitus." figureDoi="http://doi.org/10.5281/zenodo.5352956" httpUri="https://zenodo.org/record/5352956/files/figure.png" pageId="37" pageNumber="34">Fig. 83-84</figureCitation>
) – Cephalic carina simple, scarcely elevated, at most only weakly trituberculate. Pronotum with deep, oval anteromedian concavity, bearing strongly raised, trituberculate, quadrate process adjacent to anterior margin.
</paragraph>
<paragraph id="B269CA3A7F598858FF04F88F2260EF1E" blockId="37.[192,1440,1599,1879]" box="[240,560,1855,1879]" pageId="37" pageNumber="34">
<emphasis id="80A216287F598858FF04F88F21A0EF1E" box="[240,496,1855,1879]" italics="true" pageId="37" pageNumber="34">Specimens examined</emphasis>
– 48.
</paragraph>
</subSubSection>
<caption id="E6A99AB27F5A885BFF34FA2B2257ED86" ID-DOI="http://doi.org/10.5281/zenodo.5352958" ID-Zenodo-Dep="5352958" httpUri="https://zenodo.org/record/5352958/files/figure.png" pageId="38" pageNumber="35" startId="38.[192,276,1435,1457]" targetBox="[232,1409,206,1401]" targetPageId="38">
<paragraph id="B269CA3A7F5A885BFF34FA2B2257ED86" blockId="38.[192,1440,1435,1487]" pageId="38" pageNumber="35">
<emphasis id="80A216287F5A885BFF34FA2B210BEDF8" bold="true" box="[192,347,1435,1457]" pageId="38" pageNumber="35">Figure 85-89</emphasis>
.
<taxonomicName id="75D6B1B97F5A885BFE9DFA2C223FEDFB" baseAuthorityName="Pereira" baseAuthorityYear="1949" box="[361,623,1436,1458]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="35" phylum="Arthropoda" rank="species" species="pessoai">
<emphasis id="80A216287F5A885BFE9DFA2C223FEDFB" box="[361,623,1436,1458]" italics="true" pageId="38" pageNumber="35">Coprophanaeus pessoai</emphasis>
</taxonomicName>
.
<emphasis id="80A216287F5A885BFD88FA2B2295EDF8" bold="true" box="[636,709,1435,1457]" pageId="38" pageNumber="35">85-86)</emphasis>
Female habitus.
<emphasis id="80A216287F5A885BFC7CFA2B2381EDF8" bold="true" box="[904,977,1435,1457]" pageId="38" pageNumber="35">87-88)</emphasis>
Male habitus.
<emphasis id="80A216287F5A885BFB8DFA2B24CEEDF8" bold="true" box="[1145,1182,1435,1457]" pageId="38" pageNumber="35">89)</emphasis>
Aedeagus (dorsal view on left; lateral view on right).
</paragraph>
</caption>
<subSubSection id="FACC99B17F5A885BFF32FA41254CEF40" pageId="38" pageNumber="35" type="distribution">
<paragraph id="B269CA3A7F5A885BFF32FA412286EE40" blockId="38.[198,726,1521,1545]" box="[198,726,1521,1545]" pageId="38" pageNumber="35">
<emphasis id="80A216287F5A885BFF32FA41212BEE40" bold="true" box="[198,379,1521,1545]" pageId="38" pageNumber="35">Distribution.</emphasis>
Caatinga province (
<figureCitation id="2AEDD6BF7F5A885BFD85FA412299EE40" box="[625,713,1521,1545]" captionStart="Figure 62" captionStartId="30.[191,275,1894,1916]" captionTargetBox="[247,1385,192,1865]" captionTargetId="figure-18@30.[247,1386,191,1865]" captionTargetPageId="30" captionText="Figure 62. Approximate geographic distribution of Coprophanaeus (Metallophanaeus) species." figureDoi="http://doi.org/10.5281/zenodo.5352948" httpUri="https://zenodo.org/record/5352948/files/figure.png" pageId="38" pageNumber="35">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="B269CA3A7F5A885BFF32F981254CEF40" blockId="38.[198,1447,1585,1801]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFF32F9812181EE00" box="[198,465,1585,1609]" pageId="38" pageNumber="35">
<emphasis id="80A216287F5A885BFF32F9812181EE00" bold="true" box="[198,465,1585,1609]" pageId="38" pageNumber="35">Collection Records.</emphasis>
</collectingRegion>
<materialsCitation id="02BEC0677F5A885BFE21F9812393EE00" box="[469,963,1585,1609]" collectorName="Cruz das Almas" country="Brazil" location="Bahia" pageId="38" pageNumber="35" specimenCount="1" stateProvince="Bahia">
<emphasis id="80A216287F5A885BFE21F98122F2EE00" bold="true" box="[469,674,1585,1609]" pageId="38" pageNumber="35">
<collectingCountry id="CAC18AAA7F5A885BFE21F9812216EE00" box="[469,582,1585,1609]" name="Brazil" pageId="38" pageNumber="35">BRAZIL</collectingCountry>
:
<collectingRegion id="701204D87F5A885BFDA5F98122F2EE00" box="[593,674,1585,1609]" pageId="38" pageNumber="35">Bahia</collectingRegion>
</emphasis>
–
<collectorName id="1F23AFEC7F5A885BFD48F981232BEE00" box="[700,891,1585,1609]" pageId="38" pageNumber="35">Cruz das Almas</collectorName>
(Apr)
</materialsCitation>
; Maracas (Feb);
<materialsCitation id="02BEC0677F5A885BFB7CF98122A1EE20" collectorName="Jun &amp; Maranguape &amp; Tabatinga" country="Brazil" location="Feira de Santana" longitude="-38.73" pageId="38" pageNumber="35" specimenCount="1" stateProvince="Ceara">
<location id="B7099CE17F5A885BFB7CF9812509EE00" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F5A885BFB7CF9812509EE00" box="[1160,1369,1585,1609]" country="Brazil" longitude="-38.73" name="Feira de Santana" pageId="38" pageNumber="35" stateProvince="Ceara">Feira de Santana</location>
(
<collectorName id="1F23AFEC7F5A885BFA93F98125C8EE00" box="[1383,1432,1585,1609]" pageId="38" pageNumber="35">Jun</collectorName>
).
<emphasis id="80A216287F5A885BFF32F9E02148EE20" bold="true" box="[198,280,1616,1641]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFF32F9E02148EE20" box="[198,280,1616,1641]" country="Brazil" name="Ceara" pageId="38" pageNumber="35">Ceará</collectingRegion>
</emphasis>
–
<collectorName id="1F23AFEC7F5A885BFEC6F9E22181EE23" box="[306,465,1618,1642]" pageId="38" pageNumber="35">Maranguape</collectorName>
,
<collectorName id="1F23AFEC7F5A885BFE2EF9E1220AEE20" box="[474,602,1617,1641]" pageId="38" pageNumber="35">Tabatinga</collectorName>
,
<geoCoordinate id="D7E2ACFD7F5A885BFD91F9E122A1EE20" box="[613,753,1615,1641]" degrees="38" direction="west" minutes="43" orientation="longitude" pageId="38" pageNumber="35" precision="15" seconds="48" value="-38.73">
38
<superScript id="45A367727F5A885BFD76F9FF22DAEE14" attach="none" box="[642,650,1615,1629]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
43’48&quot;W
</geoCoordinate>
</materialsCitation>
<materialsCitation id="02BEC0677F5A885BFD05F9E12452EE20" box="[753,1026,1615,1641]" country="Brazil" latitude="-4.011667" location="Mar" pageId="38" pageNumber="35" specimenCount="1" stateProvince="Bahia">
;
<geoCoordinate id="D7E2ACFD7F5A885BFD0FF9E1232EEE20" box="[763,894,1615,1641]" degrees="04" direction="south" minutes="00" orientation="latitude" pageId="38" pageNumber="35" precision="15" seconds="42" value="-4.011667">
04
<superScript id="45A367727F5A885BFCECF9FF2370EE14" attach="none" box="[792,800,1615,1629]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
00’42&quot;S
</geoCoordinate>
, 126 (
<location id="B7099CE17F5A885BFC36F9E123A9EE20" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F5A885BFC36F9E123A9EE20" box="[962,1017,1617,1641]" country="Brazil" latitude="-4.011667" name="Mar" pageId="38" pageNumber="35" stateProvince="Bahia">Mar</location>
)
</materialsCitation>
;
<materialsCitation id="02BEC0677F5A885BFBFFF9E221A2EEE0" collectorName="Maranguape &amp; Cagado &amp; Sao Benedito" country="Brazil" elevation="114" latitude="-3.8747222" location="Mar" longLatPrecision="21" longitude="-38.68" municipality="Apr" pageId="38" pageNumber="35" specimenCount="1" stateProvince="Bahia">
<collectorName id="1F23AFEC7F5A885BFBFFF9E224FAEE23" box="[1035,1194,1618,1642]" pageId="38" pageNumber="35">Maranguape</collectorName>
,
<collectorName id="1F23AFEC7F5A885BFB40F9E12542EE20" box="[1204,1298,1617,1641]" pageId="38" pageNumber="35">Cagado</collectorName>
,
<geoCoordinate id="D7E2ACFD7F5A885BFAE8F9E125F6EE20" box="[1308,1446,1615,1641]" degrees="38" direction="west" minutes="45" orientation="longitude" pageId="38" pageNumber="35" precision="15" seconds="05" value="-38.75139">
38
<superScript id="45A367727F5A885BFACEF9FF2512EE14" attach="none" box="[1338,1346,1615,1629]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
45’05&quot;W
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F5A885BFF32F9C12118EEC0" box="[198,328,1647,1673]" degrees="04" direction="south" minutes="01" orientation="latitude" pageId="38" pageNumber="35" precision="15" seconds="57" value="-4.0325003">
04
<superScript id="45A367727F5A885BFF17F9DF20BBEE34" attach="none" box="[227,235,1647,1661]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
01’57&quot;S
</geoCoordinate>
,
<quantity id="752E67DF7F5A885BFEA6F9C121CBEEC0" box="[338,411,1649,1673]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="38" pageNumber="35" unit="m" value="114.0">
<elevation id="39FB2D097F5A885BFEA6F9C121CBEEC0" box="[338,411,1649,1673]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="38" pageNumber="35" unit="m" value="114.0">114 m</elevation>
</quantity>
(
<collectingMunicipality id="520D50407F5A885BFE5CF9C1218AEEC0" box="[424,474,1649,1673]" pageId="38" pageNumber="35">Apr</collectingMunicipality>
).
<collectorName id="1F23AFEC7F5A885BFE1FF9C222D9EEC3" box="[491,649,1650,1674]" pageId="38" pageNumber="35">Maranguape</collectorName>
,
<collectorName id="1F23AFEC7F5A885BFD66F9C12363EEC0" box="[658,819,1649,1673]" pageId="38" pageNumber="35">Sao Benedito</collectorName>
, Sanctuario da Nossa Senhora de Penha,
<geoCoordinate id="D7E2ACFD7F5A885BFAE9F9C125F7EEC0" box="[1309,1447,1647,1673]" degrees="38" direction="west" minutes="40" orientation="longitude" pageId="38" pageNumber="35" precision="15" seconds="48" value="-38.68">
38
<superScript id="45A367727F5A885BFACFF9DF2513EE34" attach="right" box="[1339,1347,1647,1661]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
40’48&quot;W
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F5A885BFF32F921211CEEE0" box="[198,332,1679,1705]" degrees="03" direction="south" minutes="52" orientation="latitude" pageId="38" pageNumber="35" precision="15" seconds="29" value="-3.8747222">
03
<superScript id="45A367727F5A885BFF10F93F20BCEED4" attach="none" box="[228,236,1679,1693]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
52’29&quot;S
</geoCoordinate>
,
<quantity id="752E67DF7F5A885BFEA8F92121CDEEE0" box="[348,413,1681,1705]" metricMagnitude="1" metricUnit="m" metricValue="9.6" pageId="38" pageNumber="35" unit="m" value="96.0">
<elevation id="39FB2D097F5A885BFEA8F92121CDEEE0" box="[348,413,1681,1705]" metricMagnitude="1" metricUnit="m" metricValue="9.6" pageId="38" pageNumber="35" unit="m" value="96.0">96 m</elevation>
</quantity>
(
<location id="B7099CE17F5A885BFE45F92121B9EEE0" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F5A885BFE45F92121B9EEE0" box="[433,489,1681,1705]" country="Brazil" latitude="-3.8747222" longLatPrecision="21" longitude="-38.68" municipality="Apr" name="Mar" pageId="38" pageNumber="35" stateProvince="Bahia">Mar</location>
)
</materialsCitation>
; Juazeiro do Norte.
<emphasis id="80A216287F5A885BFD0FF92123E8EEE0" bold="true" box="[763,952,1681,1705]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFD0FF92123E8EEE0" box="[763,952,1681,1705]" country="Brazil" name="Minas Gerais" pageId="38" pageNumber="35">Minas Gerais</collectingRegion>
</emphasis>
– Aguas Vermelhas (Dec).
<emphasis id="80A216287F5A885BFAE8F92025DBEEE0" bold="true" box="[1308,1419,1680,1705]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFAE8F92025DBEEE0" box="[1308,1419,1680,1705]" country="Brazil" name="Paraiba" pageId="38" pageNumber="35">Paraíba</collectingRegion>
</emphasis>
– Mamanguape (Jul); Soledade (Mar);
<materialsCitation id="02BEC0677F5A885BFD80F9012236EEA0" collectorName="Sao Jose dos Cordeiros &amp; Fazenda Almas" country="Brazil" county="Feb" elevation="650" latitude="-7.4666667" location="Mar" longLatPrecision="1303" longitude="-36.883335" municipality="Brejo Novo" pageId="38" pageNumber="35" specimenCount="1" stateProvince="Pernambuco">
<collectorName id="1F23AFEC7F5A885BFD80F90123D0EE80" box="[628,896,1713,1737]" pageId="38" pageNumber="35">São José dos Cordeiros</collectorName>
[
<collectorName id="1F23AFEC7F5A885BFC7FF901241AEE80" box="[907,1098,1713,1737]" pageId="38" pageNumber="35">Fazenda Almas</collectorName>
],
<geoCoordinate id="D7E2ACFD7F5A885BFBAEF90224F3EE80" box="[1114,1187,1711,1737]" degrees="7" direction="south" minutes="28" orientation="latitude" pageId="38" pageNumber="35" precision="925" value="-7.4666667">
7
<superScript id="45A367727F5A885BFB9DF91F2420EEF4" attach="none" box="[1129,1136,1711,1725]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
28’S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F5A885BFB5CF901255BEE80" box="[1192,1291,1711,1737]" degrees="36" direction="west" minutes="53" orientation="longitude" pageId="38" pageNumber="35" precision="925" value="-36.883335">
36
<superScript id="45A367727F5A885BFB30F91F249BEEF4" attach="none" box="[1220,1227,1711,1725]" fontSize="6" pageId="38" pageNumber="35">o</superScript>
53’W
</geoCoordinate>
,
<quantity id="752E67DF7F5A885BFAE0F901250CEE80" box="[1300,1372,1713,1737]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="38" pageNumber="35" unit="m" value="650.0">
<elevation id="39FB2D097F5A885BFAE0F901250CEE80" box="[1300,1372,1713,1737]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="38" pageNumber="35" unit="m" value="650.0">650 m</elevation>
</quantity>
(
<collectingCounty id="5B08B2B67F5A885BFA9DF90125CAEE80" box="[1385,1434,1713,1737]" pageId="38" pageNumber="35">Feb</collectingCounty>
).
<emphasis id="80A216287F5A885BFF32F9612127EEA0" bold="true" box="[198,375,1745,1769]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFF32F9612127EEA0" box="[198,375,1745,1769]" country="Brazil" name="Pernambuco" pageId="38" pageNumber="35">Pernambuco</collectingRegion>
</emphasis>
–
<collectingMunicipality id="520D50407F5A885BFE65F9612247EEA0" box="[401,535,1745,1769]" pageId="38" pageNumber="35">Brejo Novo</collectingMunicipality>
(
<location id="B7099CE17F5A885BFDD1F961220CEEA0" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F5A885BFDD1F961220CEEA0" box="[549,604,1745,1769]" country="Brazil" county="Feb" latitude="-7.4666667" longLatPrecision="1303" longitude="-36.883335" municipality="Brejo Novo" name="Mar" pageId="38" pageNumber="35" stateProvince="Pernambuco">Mar</location>
)
</materialsCitation>
;
<materialsCitation id="02BEC0677F5A885BFD84F961246DEF40" country="Brazil" location="Parque Nacional Serra da Capivara" pageId="38" pageNumber="35" specimenCount="1" stateProvince="Piaui">
Garanhuns (Mar).
<emphasis id="80A216287F5A885BFCA0F96023CFEEA0" bold="true" box="[852,927,1744,1769]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFCA0F96023CFEEA0" box="[852,927,1744,1769]" country="Brazil" name="Piaui" pageId="38" pageNumber="35">Piauí</collectingRegion>
</emphasis>
– São Raimundo Nonato [
<location id="B7099CE17F5A885BFB2CF96221F9EF40" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F58885AFF34FAB22341E9F2:B7099CE17F5A885BFB2CF96221F9EF40" country="Brazil" name="Parque Nacional Serra da Capivara" pageId="38" pageNumber="35" stateProvince="Piaui">Parque Nacional Serra da Capivara</location>
] (Jan).
<emphasis id="80A216287F5A885BFDF7F941234DEF40" bold="true" box="[515,797,1777,1801]" pageId="38" pageNumber="35">
<collectingRegion id="701204D87F5A885BFDF7F941234DEF40" box="[515,797,1777,1801]" country="Brazil" name="Rio Grande do Norte" pageId="38" pageNumber="35">Rio Grande do Norte</collectingRegion>
</emphasis>
– Canguaretama (Aug)
</materialsCitation>
; Natal (Jun, Nov).
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F5A885AFF32F8812341E9F2" lastPageId="39" lastPageNumber="36" pageId="38" pageNumber="35" type="discussion">
<paragraph id="B269CA3A7F5A885AFF32F8812341E9F2" blockId="38.[198,1447,1841,1898]" lastBlockId="39.[192,1441,195,443]" lastPageId="39" lastPageNumber="36" pageId="38" pageNumber="35">
<emphasis id="80A216287F5A885BFF32F8812132EF00" bold="true" box="[198,354,1841,1865]" pageId="38" pageNumber="35">Comments.</emphasis>
<taxonomicName id="75D6B1B97F5A885BFE9EF8812263EF00" authorityName="Olsoufieff" authorityYear="1924" box="[362,563,1841,1865]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="35" phylum="Arthropoda" rank="species" species="pertyi">
<emphasis id="80A216287F5A885BFE9EF8812263EF00" box="[362,563,1841,1865]" italics="true" pageId="38" pageNumber="35">Phanaeus pertyi</emphasis>
</taxonomicName>
was proposed as a new name for the taxon
<bibRefCitation id="D647B7CB7F5A885BFBA3F881255CEF00" author="Felsche, C." box="[1111,1292,1841,1865]" pageId="38" pageNumber="35" pagination="135 - 160" refId="ref57396" refString="Felsche, C. 1901. Beschreibungen coprophager Scarabaeiden. Deutsche Entomologische Zeitschrift 1901: 135 - 160." type="journal article" year="1901">Felsche (1901)</bibRefCitation>
erroneously regarded as
<taxonomicName id="75D6B1B97F5A885BFEAFF8E123D4EF20" authority="Perty (Olsoufieff 1924)" authorityName="Perty (Olsoufieff" authorityYear="1924" box="[347,900,1873,1898]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="35" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis id="80A216287F5A885BFEAFF8E12239EF20" box="[347,617,1873,1897]" italics="true" pageId="38" pageNumber="35">Phanaeus thalassinus</emphasis>
Perty (
<bibRefCitation id="D647B7CB7F5A885BFD34F8E1232CEF20" author="Olsoufieff, G." box="[704,892,1873,1897]" pageId="38" pageNumber="35" pagination="4 - 201" refId="ref59457" refString="Olsoufieff, G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">Olsoufieff 1924</bibRefCitation>
)
</taxonomicName>
(see Comments under
<taxonomicName id="75D6B1B97F5A885BFB6DF8E12519EF20" authorityName="Perty (Olsoufieff" authorityYear="1924" box="[1177,1353,1873,1897]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="35" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis id="80A216287F5A885BFB6DF8E12519EF20" box="[1177,1353,1873,1897]" italics="true" pageId="38" pageNumber="35">C. thalassinus</emphasis>
</taxonomicName>
below). Subsequently,
<bibRefCitation id="D647B7CB7F5B885AFE8CFF732287E892" author="Martinez, A. &amp; F. S. Pereira" box="[376,727,195,219]" pageId="39" pageNumber="36" pagination="53 - 69" refId="ref59045" refString="Martinez, A., and F. S. Pereira. 1967. Notas escarabeidologicas III. (Col. Scarabaeidae - Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez and Pereira (1967)</bibRefCitation>
synonymized
<taxonomicName id="75D6B1B97F5B885AFC7CFF73215CE8B2" authority="Martinez and Pereira (1955)" authorityName="Martinez and Pereira" authorityYear="1955" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="80A216287F5B885AFC7CFF7324DEE892" box="[904,1166,195,219]" italics="true" pageId="39" pageNumber="36">Phanaeus alvarengai</emphasis>
Martínez and
<bibRefCitation id="D647B7CB7F5B885AFAB0FF73215CE8B2" author="Pereira, F. S. &amp; M. A. V. d'Andretta" pageId="39" pageNumber="36" pagination="247 - 264" refId="ref59634" refString="Pereira, F. S., and M. A. V. d'Andretta. 1955. Novos escarabeideos e novas sinonimias. Papeis Avulsos do Departamento de Zoologia 12: 247 - 264." type="journal article" year="1955">Pereira (1955)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="75D6B1B97F5B885AFEB0FF5421F7E8B2" authorityName="Olsoufieff" authorityYear="1924" box="[324,423,228,251]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="species" species="pertyi">
<emphasis id="80A216287F5B885AFEB0FF5421F7E8B2" box="[324,423,228,251]" italics="true" pageId="39" pageNumber="36">P. pertyi</emphasis>
</taxonomicName>
. In accordance with Article 53.3 of the International Code of Zoological Nomenclature (
<bibRefCitation id="D647B7CB7F5B885AFF3CFEB32103E952" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[200,339,259,283]" pageId="39" pageNumber="36" refId="ref58443" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
),
<taxonomicName id="75D6B1B97F5B885AFE92FEB3232BE952" authority="Martinez and Pereira" authorityName="Martinez and Pereira" authorityYear="1955" box="[358,891,259,283]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="80A216287F5B885AFE92FEB3223CE952" box="[358,620,259,283]" italics="true" pageId="39" pageNumber="36">Phanaeus alvarengai</emphasis>
Martínez and Pereira
</taxonomicName>
is the senior primary homonym of
<taxonomicName id="75D6B1B97F5B885AFAD2FEB321ACE972" authority="Arnaud (1984)" authorityName="sensu Arnaud" authorityYear="1984" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="80A216287F5B885AFAD2FEB32113E972" italics="true" pageId="39" pageNumber="36">Phanaeus alvarengai</emphasis>
<bibRefCitation id="D647B7CB7F5B885AFEBDFE9321ACE972" author="Arnaud, P." box="[329,508,291,315]" pageId="39" pageNumber="36" pagination="59 - 64" refId="ref56412" refString="Arnaud, P. 1984. Nouveaux Phanaeini (Col. Scarabaeidae). Miscellanea Entomologica 50: 59 - 64." type="journal article" year="1984">Arnaud (1984)</bibRefCitation>
</taxonomicName>
, currently still placed in the genus
<taxonomicName id="75D6B1B97F5B885AFC6BFE932447E972" authorityName="MacLeay" authorityYear="1819" box="[927,1047,291,315]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="80A216287F5B885AFC6BFE932447E972" box="[927,1047,291,315]" italics="true" pageId="39" pageNumber="36">Phanaeus</emphasis>
</taxonomicName>
(
<bibRefCitation id="D647B7CB7F5B885AFBD1FE93248CE972" author="Edmonds, W. D." box="[1061,1244,291,315]" pageId="39" pageNumber="36" pagination="1 - 105" refId="ref56927" refString="Edmonds, W. D. 1994. Revision of Phanaeus MacLeay, a New World genus of scarabaeine dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Natural History Museum of Los Angeles County Contributions in Science 443: 1 - 105." type="journal article" year="1994">Edmonds 1994</bibRefCitation>
;
<bibRefCitation id="D647B7CB7F5B885AFB12FE9325C4E972" author="Arnaud, P." box="[1254,1428,291,315]" pageId="39" pageNumber="36" refId="ref56534" refString="Arnaud, P. 2002 c. Phanaeini. Les coleopteres du monde, vol. 28. Hillside Books; Canterbury. 151 p." type="book" year="2002">Arnaud 2002c</bibRefCitation>
). The fact that Martínez and Pereira placed their taxon in the subgenus
<taxonomicName id="75D6B1B97F5B885AFBF9FEF3249BE912" authorityName="Olsoufieff" authorityYear="1924" box="[1037,1227,323,347]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="80A216287F5B885AFBF9FEF3249BE912" box="[1037,1227,323,347]" italics="true" pageId="39" pageNumber="36">Coprophanaeus</emphasis>
</taxonomicName>
of
<taxonomicName id="75D6B1B97F5B885AFB18FEF32534E912" authorityName="MacLeay" authorityYear="1819" box="[1260,1380,323,347]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="80A216287F5B885AFB18FEF32534E912" box="[1260,1380,323,347]" italics="true" pageId="39" pageNumber="36">Phanaeus</emphasis>
</taxonomicName>
does not, in accordance with Article 57.4 of the Code (
<bibRefCitation id="D647B7CB7F5B885AFCF7FED323DCE932" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[771,908,355,379]" pageId="39" pageNumber="36" refId="ref58443" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
), shield Arnaud’s name from unavailability, and, therefore, the name
<taxonomicName id="75D6B1B97F5B885AFE00FE332379E9D2" authority="sensu Arnaud" authorityName="sensu Arnaud" authorityYear="1984" box="[500,809,387,411]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="36" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="80A216287F5B885AFE00FE332295E9D2" box="[500,709,387,411]" italics="true" pageId="39" pageNumber="36">alvarengai sensu</emphasis>
Arnaud
</taxonomicName>
must be replaced. We have referred the issue to the species’ author, Patrick Arnaud, for rectification.
</paragraph>
</subSubSection>
</treatment>
</document>
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