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<document ID-DOI="http://doi.org/10.5281/zenodo.3788695" ID-GBIF-Dataset="7faaa125-e85b-4e6a-839f-369f3baa5964" ID-GBIF-Taxon="163755214" ID-ISSN="13132970" ID-Zenodo-Dep="3788695" approvalRequired="285" approvalRequired_for_textStreams="285" checkinTime="1588001517772" checkinUser="plazi" docAuthor="Gall, Lawrence &amp; Hawks, David" docDate="2010" docId="038D2962FF8E1B075EE0FAA5FC61FE2A" docLanguage="en" docName="ZK_article_2167.pdf" docOrigin="ZooKeys 39 (39)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@3faabf8c" docStyleName="zookeys.2008.journal_article" docTitle="Catocala luctuosa Hulst 2010, stat. rev." docType="treatment" docVersion="4" lastPageId="22" lastPageNumber="59" masterDocId="FFB4511AFF9B1B115E6CFF95FFCAFFF0" masterDocTitle="Systematics of moths in the genus Catocala (Lepidoptera, Erebidae) IV. Nomenclatorial stabilization of the Nearctic fauna, with a revised synonymic check list" masterLastPageNumber="83" masterPageNumber="37" pageId="21" pageNumber="58" updateTime="1643559085965" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Systematics of moths in the genus Catocala (Lepidoptera, Erebidae) IV. Nomenclatorial stabilization of the Nearctic fauna, with a revised synonymic check list</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gall, Lawrence</mods:namePart>
<mods:affiliation>Yale University, New Haven ,,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hawks, David</mods:namePart>
<mods:affiliation>University of California, Riverside ,, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-03-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>39</mods:number>
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<mods:detail type="issue">
<mods:number>39</mods:number>
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<mods:start>37</mods:start>
<mods:end>83</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.39.425</mods:identifier>
<mods:identifier type="GBIF-Dataset">7faaa125-e85b-4e6a-839f-369f3baa5964</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576632</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788695" ID-GBIF-Taxon="163755214" ID-Zenodo-Dep="3788695" LSID="urn:lsid:plazi:treatment:038D2962FF8E1B075EE0FAA5FC61FE2A" httpUri="http://treatment.plazi.org/id/038D2962FF8E1B075EE0FAA5FC61FE2A" lastPageId="22" lastPageNumber="59" pageId="21" pageNumber="58">
<subSubSection pageId="21" pageNumber="58" type="nomenclature">
<paragraph blockId="21.[140,539,1328,1390]" box="[140,539,1328,1355]" pageId="21" pageNumber="58">
<heading box="[140,539,1328,1355]" fontSize="11" level="2" pageId="21" pageNumber="58" reason="3">
<emphasis bold="true" box="[140,539,1328,1355]" pageId="21" pageNumber="58">
<taxonomicName authority="Hulst" authorityName="Hulst" authorityYear="2010" box="[140,427,1328,1355]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="species" species="luctuosa" status="stat. rev.">
<emphasis bold="true" box="[140,352,1328,1355]" italics="true" pageId="21" pageNumber="58">Catocala luctuosa</emphasis>
Hulst
</taxonomicName>
,
<taxonomicNameLabel box="[440,539,1328,1354]" pageId="21" pageNumber="58">stat. rev.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[140,539,1328,1390]" box="[140,348,1363,1390]" pageId="21" pageNumber="58">
Fig. 24 (
<typeStatus box="[234,340,1363,1390]" pageId="21" pageNumber="58">lectotype</typeStatus>
)
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="58" type="reference_group">
<paragraph blockId="21.[140,879,1434,1566]" box="[140,517,1434,1461]" pageId="21" pageNumber="58">
<treatmentCitationGroup box="[140,517,1434,1461]" pageId="21" pageNumber="58">
<treatmentCitation author="Hulst GD" box="[140,517,1434,1461]" page="53" pageId="21" pageNumber="58" year="1884">
<taxonomicName authority="Hulst, 1884: 53" authorityName="Hulst" authorityPageNumber="53" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[140,517,1434,1461]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[140,329,1434,1460]" italics="true" pageId="21" pageNumber="58">Catocala luctuosa</emphasis>
<bibRefCitation author="Hulst GD" box="[337,517,1434,1461]" pageId="21" pageNumber="58" pagination="14 - 56" refId="ref17953" refString="Hulst GD (1884) The genus Catocala. Bulletin of the Brooklyn Entomological Society 7: 14 - 56." type="journal article" year="1884">Hulst, 1884: 53</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="21.[140,879,1434,1566]" box="[140,879,1469,1496]" pageId="21" pageNumber="58">
<treatmentCitationGroup box="[140,879,1469,1496]" pageId="21" pageNumber="58">
<taxonomicName authorityName="Grote" authorityYear="1872" box="[140,311,1469,1495]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="species" species="retecta">
<emphasis box="[140,311,1469,1495]" italics="true" pageId="21" pageNumber="58">Catocala retecta</emphasis>
</taxonomicName>
syn.
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[371,458,1469,1495]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[371,458,1469,1495]" italics="true" pageId="21" pageNumber="58">luctuosa</emphasis>
</taxonomicName>
;
<treatmentCitation author="Barnes W &amp; McDunnough J" box="[473,879,1469,1496]" page="13" pageId="21" pageNumber="58" year="1918">
<bibRefCitation author="Barnes W &amp; McDunnough J" box="[473,879,1469,1496]" pageId="21" pageNumber="58" pagination="1 - 47" refId="ref16743" refString="Barnes W, McDunnough J (1918) Illustrations of the North American species of the genus Catocala. Memoirs of the American Museum of Natural History 3: 1 - 47." type="journal article" year="1918">Barnes and McDunnough 1918: 13</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="21.[140,879,1434,1566]" box="[140,757,1504,1531]" pageId="21" pageNumber="58">
<treatmentCitationGroup box="[140,757,1504,1531]" pageId="21" pageNumber="58">
<taxonomicName authorityName="Hulst" authorityYear="1884" box="[140,454,1504,1531]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="retecta" subSpecies="luctuosa">
<emphasis box="[140,311,1504,1530]" italics="true" pageId="21" pageNumber="58">Catocala retecta</emphasis>
ssp.
<emphasis box="[367,454,1504,1530]" italics="true" pageId="21" pageNumber="58">luctuosa</emphasis>
</taxonomicName>
;
<treatmentCitation author="McDunnough JH" box="[469,757,1504,1531]" page="115" pageId="21" pageNumber="58" year="1938">
<bibRefCitation author="McDunnough JH" box="[469,757,1504,1531]" pageId="21" pageNumber="58" pagination="1 - 275" refId="ref18138" refString="McDunnough JH (1938) Check List of the Lepidoptera of Canada and the United States of America. Part I. Macrolepidoptera. Memoirs of the Southern California Academy of Sciences 1: 1 - 275." type="journal article" year="1938">McDunnough 1938: 115</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="21.[140,879,1434,1566]" box="[140,853,1539,1566]" pageId="21" pageNumber="58">
<treatmentCitationGroup box="[140,853,1539,1566]" pageId="21" pageNumber="58">
<taxonomicName authorityName="Hulst" authorityYear="1884" box="[140,454,1539,1566]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="retecta" subSpecies="luctuosa">
<emphasis box="[140,311,1539,1565]" italics="true" pageId="21" pageNumber="58">Catocala retecta</emphasis>
ssp.
<emphasis box="[367,454,1539,1565]" italics="true" pageId="21" pageNumber="58">luctuosa</emphasis>
</taxonomicName>
;
<treatmentCitation author="Franclemont JG &amp; Todd EL" box="[469,853,1539,1566]" page="128" pageId="21" pageNumber="58" year="1983">
<bibRefCitation author="Franclemont JG &amp; Todd EL" box="[469,853,1539,1566]" pageId="21" pageNumber="58" pagination="120 - 159" refId="ref17467" refString="Franclemont JG, Todd EL (1983) Noctuidae. In: Hodges RW (Ed) Check list of the Lepidoptera of America north of Mexico. University Press, Cambidge, 120 - 159." type="book chapter" year="1983">Franclemont and Todd 1983: 128</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="59" pageId="21" pageNumber="58" type="materials_examined">
<paragraph blockId="21.[140,1108,1609,1672]" pageId="21" pageNumber="58">
<emphasis bold="true" box="[140,311,1609,1636]" pageId="21" pageNumber="58">Type material.</emphasis>
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[317,506,1610,1636]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[317,506,1610,1636]" italics="true" pageId="21" pageNumber="58">Catocala luctuosa</emphasis>
</taxonomicName>
: the original description does not state the number of types. Four types are at the
<collectionCode box="[443,538,1645,1671]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="58">AMNH</collectionCode>
, and a male is designated as
<emphasis bold="true" box="[851,960,1645,1671]" pageId="21" pageNumber="58">
<typeStatus box="[851,960,1645,1671]" pageId="21" pageNumber="58">lectotype</typeStatus>
</emphasis>
to clarify ap-
</paragraph>
<paragraph blockId="22.[140,1108,167,475]" pageId="22" pageNumber="59">
plication of the name
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[385,472,167,193]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[385,472,167,193]" italics="true" pageId="22" pageNumber="59">luctuosa</emphasis>
</taxonomicName>
. Th e
<typeStatus box="[535,637,167,193]" pageId="22" pageNumber="59">lectotype</typeStatus>
bears the labels “Ind.”, “Collection / GD- Hulst”, “
<taxonomicName box="[239,340,202,228]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="genus">Catocala</taxonomicName>
/ luctuosa / Type Hulst.” Although Hulst described
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[937,1058,202,228]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[937,1058,202,228]" italics="true" pageId="22" pageNumber="59">C. luctuosa</emphasis>
</taxonomicName>
as a distinct species, the name has been treated in the literature as a synonym or subspecies of
<taxonomicName authority="Grote, 1872" authorityName="Grote" authorityYear="1872" box="[170,423,272,299]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="retecta">
<emphasis box="[170,273,273,299]" italics="true" pageId="22" pageNumber="59">C. retecta</emphasis>
Grote, 1872
</taxonomicName>
.
<bibRefCitation author="Gall LF" box="[434,568,272,298]" pageId="22" pageNumber="59" pagination="173 - 194" refId="ref17585" refString="Gall LF (1991) Evolutionary ecology of sympatric Catocala moths (Lepidoptera: Noctuidae). I. Experiments on larval foodplant specificity. Journal of Research on the Lepidoptera 29: 173 - 194." type="journal article" year="1991">Gall (1991)</bibRefCitation>
established that
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[757,878,272,299]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[757,878,272,299]" italics="true" pageId="22" pageNumber="59">C. luctuosa</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Grote" authorityYear="1872" box="[935,1037,273,299]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="retecta">
<emphasis box="[935,1037,273,299]" italics="true" pageId="22" pageNumber="59">C. retecta</emphasis>
</taxonomicName>
breed true, and that the adult morphologies are consistently distinct. Although
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[963,1083,307,334]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[963,1083,307,334]" italics="true" pageId="22" pageNumber="59">C. luctuosa</emphasis>
</taxonomicName>
is encountered more commonly than
<taxonomicName authorityName="Grote" authorityYear="1872" box="[535,636,343,369]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="retecta">
<emphasis box="[535,636,343,369]" italics="true" pageId="22" pageNumber="59">C. retecta</emphasis>
</taxonomicName>
in the southcentral
<collectingCountry box="[858,1011,343,369]" name="United States of America" pageId="22" pageNumber="59">United States</collectingCountry>
, the area of sympatry between the two is extensive and includes most of the geographic range of
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[140,261,413,440]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[140,261,413,440]" italics="true" pageId="22" pageNumber="59">C. luctuosa</emphasis>
</taxonomicName>
. Accordingly, we reinstate
<taxonomicName authorityName="Hulst" authorityYear="1884" baseAuthorityName="Hulst" baseAuthorityYear="1884" box="[565,686,413,440]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="59" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis box="[565,686,413,440]" italics="true" pageId="22" pageNumber="59">C. luctuosa</emphasis>
</taxonomicName>
to the rank of species. Type Locality: hereby restricted to Ind[iana,
<collectingCountry box="[475,533,448,475]" name="United States of America" pageId="22" pageNumber="59">USA</collectingCountry>
] on the basis of the
<typeStatus box="[760,862,448,474]" pageId="22" pageNumber="59">lectotype</typeStatus>
labels.
</paragraph>
</subSubSection>
</treatment>
</document>