392 lines
49 KiB
XML
392 lines
49 KiB
XML
<document id="5CF20CDC6A1B933967A41B089CC4008F" ID-CLB-Dataset="32655" ID-DOI="10.11646/zootaxa.4294.2.10" ID-GBIF-Dataset="ea7cbde6-2399-4d12-9793-d6321bbea6a9" ID-ISSN="1175-5326" ID-Zenodo-Dep="831730" ID-ZooBank="332D0F30-1998-426D-AD64-4C23CB292B22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500398718492" checkinUser="plazi" docAuthor="Dantas, Galileu P. S. & Giłka, Wojciech" docDate="2017" docId="039B9E54FFB0FF90FF4AFCD0FE6F36DA" docLanguage="en" docName="zootaxa.4294.2.10.pdf" docOrigin="Zootaxa 4294 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus insolens Dantas & Giłka 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="290" masterDocId="FFA2E62CFFB6FF99FFDDFFDEFFDE3472" masterDocTitle="New Tanytarsus van der Wulp from the Brazilian Amazonia indicate clues to intrageneric relations (Diptera: Chironomidae)" masterLastPageNumber="291" masterPageNumber="281" pageNumber="287" updateTime="1698465757151" updateUser="ExternalLinkService">
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<mods:mods id="A576524C5F14C0D2D563CC255E7A4F82" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="75F07D6E9B1B873964D2563228B7C307">
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<mods:title id="49DA0C2E645C7597F7C3DD02F9DA5251">New Tanytarsus van der Wulp from the Brazilian Amazonia indicate clues to intrageneric relations (Diptera: Chironomidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C62D97E2AD30B7A937534FB1396677CA" type="personal">
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<mods:role id="65DD64ED55EE4786244905F05812B802">
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<mods:roleTerm id="1390AB571507771B726380A0EDBCC281">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F60695DCF52232FC992059F23F13CDAA">Dantas, Galileu P. S.</mods:namePart>
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</mods:name>
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<mods:name id="30DC5C46CF43396084DC0C5BAF21FC33" type="personal">
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<mods:role id="1FA7ED152534415D9B54FB23431F2FE7">
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<mods:roleTerm id="9D8B51C946BCB7CA56D312640920AAA4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3FD05843BA68485913B567B1E48943E8">Giłka, Wojciech</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="061F132578D5E2BFCBF97FC0FD72C641">text</mods:typeOfResource>
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<mods:titleInfo id="CE078A7C29019D9EE3C624BAB5729BA7">
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<mods:title id="0CA627977CFC27FC543BF0A652921F73">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0C00E1BA346181FA70601427B1F455B8">
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<mods:date id="FD1C49C6639146B91C71D4101220627A">2017</mods:date>
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<mods:detail id="A1F4AEAF5693272B9D49911ADA777D49" type="volume">
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<mods:number id="D5BCA1790F82A89F0699CD86DB9930D7">4294</mods:number>
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</mods:detail>
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<mods:detail id="596AA367F09AAB2FBED510090CCBFB63" type="issue">
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<mods:number id="8D605984512D732B1ECD65D1F7B1629E">2</mods:number>
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</mods:detail>
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<mods:extent id="2D7437ACA2EF79E0235F10690E096B12" unit="page">
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<mods:start id="D8C7D4C23B6FBB45DFC1DFC2373FA83A">281</mods:start>
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<mods:end id="2B1B00C2CBB48F81DF41D3BCFE00C7D1">291</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="719F44C80E43604EED82E3DCE8359002">journal article</mods:classification>
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<mods:identifier id="9159AC543A0DB39CCC97371AADFAC586" type="CLB-Dataset">32655</mods:identifier>
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<mods:identifier id="89A8964E5710160F3877A8F1367C53D9" type="DOI">10.11646/zootaxa.4294.2.10</mods:identifier>
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<mods:identifier id="1991D39AA79E693337BD341E7BE75F0D" type="GBIF-Dataset">ea7cbde6-2399-4d12-9793-d6321bbea6a9</mods:identifier>
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<mods:identifier id="72C48849F79AFA7421A2BEB2B9B09BEC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="918D1BAAC927914AF19CCE5E92F56864" type="Zenodo-Dep">831730</mods:identifier>
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<mods:identifier id="C52102C1048C541C792BDF3630C6A37F" type="ZooBank">332D0F30-1998-426D-AD64-4C23CB292B22</mods:identifier>
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</mods:mods>
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<treatment id="039B9E54FFB0FF90FF4AFCD0FE6F36DA" ID-DOI="http://doi.org/10.5281/zenodo.6004198" ID-GBIF-Taxon="132550169" ID-Zenodo-Dep="6004198" LSID="urn:lsid:plazi:treatment:039B9E54FFB0FF90FF4AFCD0FE6F36DA" httpUri="http://treatment.plazi.org/id/039B9E54FFB0FF90FF4AFCD0FE6F36DA" lastPageId="9" lastPageNumber="290" pageId="6" pageNumber="287">
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<subSubSection id="C3287CC9FFB0FF9FFF4AFCD0FE8E3738" pageId="6" pageNumber="287" type="nomenclature">
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||
<paragraph id="8B8D2F42FFB0FF9FFF4AFCD0FE34375A" blockId="6.[151,490,782,842]" box="[151,490,782,808]" pageId="6" pageNumber="287">
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<heading id="D0C5982EFFB0FF9FFF4AFCD0FE34375A" bold="true" box="[151,490,782,808]" fontSize="11" level="1" pageId="6" pageNumber="287" reason="1">
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||
<emphasis id="B946F350FFB0FF9FFF4AFCD0FE34375A" bold="true" box="[151,490,782,808]" pageId="6" pageNumber="287">
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<taxonomicName id="4C3254C1FFB0FF9FFF4AFCD0FE5B375A" ID-CoL="9HS4K" authority="Dantas & Giłka, 2017" authorityName="Dantas & Giłka" authorityYear="2017" box="[151,389,782,808]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="insolens" status="sp. nov.">
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<emphasis id="B946F350FFB0FF9FFF4AFCD0FE5B375A" bold="true" box="[151,389,782,808]" italics="true" pageId="6" pageNumber="287">Tanytarsus insolens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2754E2BFFB0FF9FFE50FCD0FE34375A" box="[397,490,782,808]" pageId="6" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8D2F42FFB0FF9FFF4AFCEFFE8E3738" blockId="6.[151,490,782,842]" box="[151,336,817,842]" pageId="6" pageNumber="287">
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(
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<figureCitation id="130933C7FFB0FF9FFF42FCEFFF3D3738" box="[159,227,817,842]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="6" pageNumber="287">Figs 3</figureCitation>
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A–F; 4C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3287CC9FFB0FF9FFF4AFCA7FD4637A8" pageId="6" pageNumber="287" type="materials_examined">
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<paragraph id="8B8D2F42FFB0FF9FFF4AFCA7FD4637A8" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
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<materialsCitation id="3B5A251FFFB0FF9FFF4AFCA7FD4637A8" ID-GBIF-Occurrence="1576154701" collectingDate="2015-06-19" collectingDateMax="2015-07-07" collectingDateMin="2015-06-19" collectionCode="INPA, DIZP" collectorName="G. P. S. Dantas" country="Brazil" latitude="-2.7172222" location="Manaus" longLatPrecision="21" longitude="-59.90111" municipality="Puraquequara" pageId="6" pageNumber="287" specimenCount="4" specimenCount-male="4" stateProvince="Amazonas" typeStatus="holotype">
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<emphasis id="B946F350FFB0FF9FFF4AFCA7FE9237E0" bold="true" box="[151,332,889,914]" pageId="6" pageNumber="287">
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<typeStatus id="548991E0FFB0FF9FFF4AFCA7FF0C37E0" box="[151,210,889,914]" pageId="6" pageNumber="287">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="548991E0FFB0FF9FFE87FCA7FE1B37E0" box="[346,453,889,914]" pageId="6" pageNumber="287" type="holotype">Holotype</typeStatus>
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, adult male:
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<collectingCountry id="F3256FD2FFB0FF9FFDB4FCA4FD0F37E0" box="[617,721,890,914]" name="Brazil" pageId="6" pageNumber="287">BRAZIL</collectingCountry>
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,
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<collectingRegion id="49F6E1A0FFB0FF9FFD39FCA4FC8437E0" box="[740,858,890,914]" country="Brazil" name="Amazonas" pageId="6" pageNumber="287">Amazonas</collectingRegion>
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state,
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<collectingMunicipality id="6BE9B538FFB0FF9FFC6DFCA4FB9437E3" box="[944,1098,890,913]" pageId="6" pageNumber="287">Puraquequara</collectingMunicipality>
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near
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<location id="8EED7999FFB0FF9FFB4AFCA4FB2E37E3" LSID="urn:lsid:plazi:treatment:039B9E54FFB0FF90FF4AFCD0FE6F36DA:8EED7999FFB0FF9FFB4AFCA4FB2E37E3" box="[1175,1264,890,913]" country="Brazil" latitude="-2.7172222" longLatPrecision="21" longitude="-59.90111" municipality="Puraquequara" name="Manaus" pageId="6" pageNumber="287" stateProvince="Amazonas">Manaus</location>
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(
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<geoCoordinate id="EE064985FFB0FF9FFADBFCA4FA5937E0" box="[1286,1415,890,914]" direction="south" orientation="latitude" pageId="6" pageNumber="287" precision="15" value="-2.7172222">02°43'02"S</geoCoordinate>
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/
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||
<geoCoordinate id="EE064985FFB0FF9FFF4AFC43FEFB37C4" box="[151,293,925,950]" direction="west" orientation="longitude" pageId="6" pageNumber="287" precision="15" value="-59.90111">59°54'04"W</geoCoordinate>
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), 0
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<collectingDate id="EFC8F06AFFB0FF9FFE95FC40FE1137C4" box="[328,463,925,950]" pageId="6" pageNumber="287" value="2015-07-07">
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<date id="FF8C0982FFB0FF9FFE95FC40FE1137C4" box="[328,463,925,950]" pageId="6" pageNumber="287" value="2015-07-07">7 July 2015</date>
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</collectingDate>
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,
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<collectingMethod id="52735755FFB0FF9FFE06FC43FDB437C7" box="[475,618,925,950]" pageId="6" pageNumber="287">Malaise trap</collectingMethod>
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,
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<collectorName id="26C74A94FFB0FF9FFDABFC40FCD037C7" box="[630,782,926,950]" pageId="6" pageNumber="287">G.P.S. Dantas</collectorName>
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(
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<collectionCode id="ED23B787FFB0FF9FFCC3FC40FCBA37C4" box="[798,868,926,950]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="6" pageNumber="287">INPA</collectionCode>
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).
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<typeStatus id="548991E0FFB0FF9FFCABFC40FC3637C4" box="[886,1000,926,950]" pageId="6" pageNumber="287" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D34E4CBFFB0FF9FFC2EFC40FB9437C4" box="[1011,1098,925,950]" pageId="6" pageNumber="287" type="male">4 males</specimenCount>
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(1
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<collectionCode id="ED23B787FFB0FF9FFBACFC40FB6D37C4" box="[1137,1203,926,950]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="6" pageNumber="287">INPA</collectionCode>
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, 3
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<collectionCode id="ED23B787FFB0FF9FFB08FC40FAC637C4" box="[1237,1304,926,950]" pageId="6" pageNumber="287">DIZP</collectionCode>
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), same data as for
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<typeStatus id="548991E0FFB0FF9FFF00FC1FFEE137A8" box="[221,319,961,986]" pageId="6" pageNumber="287" type="holotype">holotype</typeStatus>
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except for date:
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<collectingDate id="EFC8F06AFFB0FF9FFE26FC1CFD4A37A8" box="[507,660,961,986]" pageId="6" pageNumber="287" value="2015-06-19">
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<date id="FF8C0982FFB0FF9FFE26FC1CFD4A37A8" box="[507,660,961,986]" pageId="6" pageNumber="287" value="2015-06-19">19 June 2015</date>
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</collectingDate>
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.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3287CC9FFB0FF9FFF1AFC3BFC77378C" box="[199,937,997,1022]" pageId="6" pageNumber="287" type="etymology">
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<paragraph id="8B8D2F42FFB0FF9FFF1AFC3BFC77378C" blockId="6.[151,1437,889,1925]" box="[199,937,997,1022]" pageId="6" pageNumber="287">
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<emphasis id="B946F350FFB0FF9FFF1AFC3BFE7A378C" bold="true" box="[199,420,997,1022]" pageId="6" pageNumber="287">Derivatio nominis.</emphasis>
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From Latin
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<taxonomicName id="4C3254C1FFB0FF9FFDEDFC3BFD52378C" authorityName="Dantas & Giłka" authorityYear="2017" box="[560,652,997,1022]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="insolens">
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<emphasis id="B946F350FFB0FF9FFDEDFC3BFD52378C" box="[560,652,997,1022]" italics="true" pageId="6" pageNumber="287">insolens</emphasis>
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</taxonomicName>
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(unusual, extraordinary).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3287CC9FFB0FF9FFF1AFBD7FB5630FC" pageId="6" pageNumber="287" type="diagnosis">
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<paragraph id="8B8D2F42FFB0FF9FFF1AFBD7FB5630FC" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
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<emphasis id="B946F350FFB0FF9FFF1AFBD7FE9F3050" bold="true" box="[199,321,1033,1058]" pageId="6" pageNumber="287">Diagnosis.</emphasis>
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Tergite IX with sparse long hair-like macrotrichia. Anal point triangular, bearing several spinulae, with the most distal one enlarged, bar-like. Anterior (ventral) projection of superior volsella finger-shaped, posterior (dorsal) projection narrowly triangular, pointed. Digitus large, lanceolate or knife-shaped, broadly fused with posterior projection of superior volsella. Median volsella with 4–5 subulate lamellae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3287CC9FFB0FF90FF1AFB47FD1C348A" lastPageId="9" lastPageNumber="290" pageId="6" pageNumber="287" type="description">
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<paragraph id="8B8D2F42FFB0FF9FFF1AFB47FDE030C0" blockId="6.[151,1437,889,1925]" box="[199,574,1177,1202]" pageId="6" pageNumber="287">
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<emphasis id="B946F350FFB0FF9FFF1AFB47FE3B30C0" bold="true" box="[199,485,1177,1202]" pageId="6" pageNumber="287">Description. Adult male</emphasis>
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(n = 5).
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</paragraph>
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<paragraph id="8B8D2F42FFB0FF9FFF1AFB63FD273088" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
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<emphasis id="B946F350FFB0FF9FFF1AFB63FE2930A7" box="[199,503,1213,1238]" italics="true" pageId="6" pageNumber="287">Body size and proportions.</emphasis>
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Total length
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<quantity id="4CCA82A7FFB0FF9FFD53FB60FCEB30A7" box="[654,821,1214,1238]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.49" metricValueMin="2.31" pageId="6" pageNumber="287" unit="mm" value="2.4" valueMax="2.49" valueMin="2.31">2.31–2.49 mm</quantity>
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. Wing length
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<quantity id="4CCA82A7FFB0FF9FFC09FB63FBA530A7" box="[980,1147,1213,1238]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" metricValueMax="1.31" metricValueMin="1.25" pageId="6" pageNumber="287" unit="mm" value="1.28" valueMax="1.31" valueMin="1.25">1.25–1.31 mm</quantity>
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. Total length/wing length 1.80–1.98. Wing length/length of profemur 1.58–1.68.
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</paragraph>
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<paragraph id="8B8D2F42FFB0FF9FFF1AFADBFEFD31A0" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
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||
<emphasis id="B946F350FFB0FF9FFF1AFADBFE86316C" box="[199,344,1285,1310]" italics="true" pageId="6" pageNumber="287">Colouration.</emphasis>
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Eyes black. Antenna, palp, scutal vittae and postnotum brown. Head capsule, ground colour of thorax, scutellum, sternum and haltere yellow to light brown. Fore leg: coxa and trochanter yellow, femur and tibia brown. Mid and hind legs: coxae, trochanters and proximal half of femora yellow, distal half of femora, tibiae and tarsi brown. Wing veins brown, membrane with brownish undertone. Abdomen yellow, gradually becoming brown posteriorly; segments with darker margins dorsally (transversal bands), distinctly standing out from lighter background.
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</paragraph>
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<paragraph id="8B8D2F42FFB0FF9FFF1AFA03FEC132F4" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
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||
<emphasis id="B946F350FFB0FF9FFF1AFA03FED53184" box="[199,267,1501,1526]" italics="true" pageId="6" pageNumber="287">Head.</emphasis>
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Eyes bare, with well developed dorsomedian extensions. Antenna with 13 flagellomeres; ultimate flagellomere 405–440 µm long; AR 0.85–0.88. Frontal tubercles absent or in shape of minute swellings (
|
||
<emphasis id="B946F350FFB0FF9FFAF7F9DCFA95326B" box="[1322,1355,1538,1561]" italics="true" pageId="6" pageNumber="287">ca.</emphasis>
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2 µm). Tentorium 118–125 µm long. Temporal setae 7–10 on each side. Clypeus with 11–14 setae. Lengths of palpomeres 1–5 (in µm): 30–40, 36–48, 115–127, 107–119, 195–226; third palpomere with 2 sensilla clavata subapically, 10– 12 µm long.
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</paragraph>
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<paragraph id="8B8D2F42FFB0FF9FFF1AF94FFBC632BF" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
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||
<emphasis id="B946F350FFB0FF9FFF1AF94FFEFE32D8" box="[199,288,1681,1706]" italics="true" pageId="6" pageNumber="287">Thorax.</emphasis>
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Ac 11–20, restricted to anterior region of scutum; Dc 5–7 on each side, uniserial; Pa 1 on each side; Scts 5–6. Scutum projected and rounded anteriorly, overreaching antepronotum.
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</paragraph>
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<paragraph id="8B8D2F42FFB0FF9FFF1AF904FBB73364" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
|
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<emphasis id="B946F350FFB0FF9FFF1AF904FED93283" box="[199,263,1754,1777]" italics="true" pageId="6" pageNumber="287">Wing.</emphasis>
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Obovate, with anal lobe strongly reduced, as shown in
|
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<figureCitation id="130933C7FFB0FF9FFC58F907FC163280" box="[901,968,1753,1778]" captionStart="FIGURE 4" captionStartId="8.[151,250,1658,1680]" captionTargetBox="[151,1436,435,1636]" captionTargetId="figure@8.[151,1436,435,1636]" captionTargetPageId="8" captionText="FIGURE 4. Wing of male: Tanytarsus illustris sp. nov. (A), Tanytarsus insignis sp. nov. (B), Tanytarsus insolens sp. nov. (C)." httpUri="https://zenodo.org/record/831738/files/figure.png" pageId="6" pageNumber="287">Fig. 4</figureCitation>
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C. Almost all veins and entire membrane below radial veins covered with macrotrichia. Brachiolum with 1 seta. VR Cu 1.27–1.35.
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||
</paragraph>
|
||
<paragraph id="8B8D2F42FFB0FF9FFF1AF8FBFC3133F6" blockId="6.[151,1437,889,1925]" pageId="6" pageNumber="287">
|
||
<emphasis id="B946F350FFB0FF9FFF1AF8FBFEDD334E" box="[199,259,1829,1852]" italics="true" pageId="6" pageNumber="287">Legs.</emphasis>
|
||
Fore leg tibia with short lanceolate spur 8–15 µm long. Tibial combs of mid and hind legs separated, only one comb bearing slightly bent apically spur, 32–42 µm (mid leg) to 37–52 µm long (hind leg). Basitarsus of mid leg with 1–2 sensilla chaetica. Lengths and proportions of legs as in
|
||
<tableCitation id="C6B01AF9FFB0FF9FFC48F8B2FC3433F6" box="[917,1002,1900,1925]" captionStart="TABLE 3" captionStartId="8.[151,239,152,175]" captionText="TABLE 3. Lengths of leg segments (µm) and leg ratios of male Tanytarsus insolens sp. nov." pageId="6" pageNumber="287">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4D7FCAFFB1FF9EFF4AF890FE723392" httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="7" pageNumber="288" targetBox="[180,1412,184,1823]" targetPageId="7">
|
||
<paragraph id="8B8D2F42FFB1FF9EFF4AF890FE723392" blockId="7.[151,1436,1870,2016]" pageId="7" pageNumber="288">
|
||
<emphasis id="B946F350FFB1FF9EFF4AF890FECD3311" bold="true" box="[151,275,1870,1892]" pageId="7" pageNumber="288">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3254C1FFB1FF9EFEC7F891FE3C3316" authority="Dantas & Giłka, 2017" authorityName="Dantas & Giłka" authorityYear="2017" box="[282,482,1870,1892]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="insolens" status="sp. nov.">
|
||
<emphasis id="B946F350FFB1FF9EFEC7F891FE3C3316" box="[282,482,1870,1892]" italics="true" pageId="7" pageNumber="288">Tanytarsus insolens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946F350FFB1FF9EFE34F891FDE43316" bold="true" box="[489,570,1871,1892]" pageId="7" pageNumber="288">
|
||
<taxonomicNameLabel id="A2754E2BFFB1FF9EFE34F891FDE43316" box="[489,570,1871,1892]" pageId="7" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B946F350FFB1FF9EFD97F890FDA63316" bold="true" box="[586,632,1870,1892]" pageId="7" pageNumber="288">A–F</emphasis>
|
||
) and
|
||
<taxonomicName id="4C3254C1FFB1FF9EFD6EF891FB853316" authority="Vinogradova, Riss et Spies, 2009" authorityName="Vinogradova, Riss et Spies" authorityYear="2009" box="[691,1115,1870,1892]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="kiche">
|
||
<emphasis id="B946F350FFB1FF9EFD6EF891FCDF3316" box="[691,769,1870,1892]" italics="true" pageId="7" pageNumber="288">T. kiche</emphasis>
|
||
Vinogradova, Riss
|
||
<emphasis id="B946F350FFB1FF9EFC16F891FC033316" box="[971,989,1871,1892]" italics="true" pageId="7" pageNumber="288">et</emphasis>
|
||
Spies, 2009
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B946F350FFB1FF9EFBB7F890FBA03316" bold="true" box="[1130,1150,1870,1892]" pageId="7" pageNumber="288">G</emphasis>
|
||
), male.
|
||
<emphasis id="B946F350FFB1FF9EFB0FF890FB3A3311" bold="true" box="[1234,1252,1870,1891]" pageId="7" pageNumber="288">A</emphasis>
|
||
,
|
||
<emphasis id="B946F350FFB1FF9EFB2FF891FADD3316" bold="true" box="[1266,1283,1871,1892]" pageId="7" pageNumber="288">B</emphasis>
|
||
—hypopygium in dorsal (
|
||
<emphasis id="B946F350FFB1FF9EFEDCF8B3FECD33F0" bold="true" box="[257,275,1901,1922]" pageId="7" pageNumber="288">A</emphasis>
|
||
) and ventral aspect (
|
||
<emphasis id="B946F350FFB1FF9EFE33F8B3FE2133F0" bold="true" box="[494,511,1901,1922]" pageId="7" pageNumber="288">B</emphasis>
|
||
);
|
||
<emphasis id="B946F350FFB1FF9EFDCAF8B3FDF733F1" bold="true" box="[535,553,1901,1923]" pageId="7" pageNumber="288">C</emphasis>
|
||
—anterior region of anal tergite covered with microtrichia and hair-like macrotrichia;
|
||
<emphasis id="B946F350FFB1FF9EFF4AF852FF7733D3" bold="true" box="[151,169,1932,1953]" pageId="7" pageNumber="288">D</emphasis>
|
||
—anal point (variation);
|
||
<emphasis id="B946F350FFB1FF9EFE76F852FE6233D3" bold="true" box="[427,444,1932,1953]" pageId="7" pageNumber="288">E</emphasis>
|
||
,
|
||
<emphasis id="B946F350FFB1FF9EFE17F852FE0033D0" bold="true" box="[458,478,1932,1954]" pageId="7" pageNumber="288">G</emphasis>
|
||
—superior volsella: anterior/ventral projection (av, yellow arrow), posterior/dorsal projection (pd, red) and digitus (di, green);
|
||
<emphasis id="B946F350FFB1FF9EFE37F874FE2733CD" bold="true" box="[490,505,1962,1983]" pageId="7" pageNumber="288">F</emphasis>
|
||
—median volsella [C–F magnified
|
||
<emphasis id="B946F350FFB1FF9EFCBBF875FC5B33B2" box="[870,901,1963,1984]" italics="true" pageId="7" pageNumber="288">ca.</emphasis>
|
||
2–3 times relative to A and B; G out of scale, after Vinogradova
|
||
<emphasis id="B946F350FFB1FF9EFEFDF815FE8D3392" box="[288,339,1994,2016]" italics="true" pageId="7" pageNumber="288">et al.</emphasis>
|
||
(2009)].
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D7FCAFFBEFF91FF4AFF46FBAC34DD" box="[151,1138,152,175]" pageId="8" pageNumber="289">
|
||
<paragraph id="8B8D2F42FFBEFF91FF4AFF46FBAC34DD" blockId="8.[151,1138,152,175]" box="[151,1138,152,175]" pageId="8" pageNumber="289">
|
||
<emphasis id="B946F350FFBEFF91FF4AFF46FEDD34DD" bold="true" box="[151,259,152,175]" pageId="8" pageNumber="289">TABLE 3</emphasis>
|
||
. Lengths of leg segments (µm) and leg ratios of male
|
||
<taxonomicName id="4C3254C1FFBEFF91FC99FF46FBCB34DD" authority="Dantas & Giłka, 2017" authorityName="Dantas & Giłka" authorityYear="2017" box="[836,1045,152,175]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="insolens" status="sp. nov.">
|
||
<emphasis id="B946F350FFBEFF91FC99FF46FBCB34DD" box="[836,1045,152,175]" italics="true" pageId="8" pageNumber="289">Tanytarsus insolens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2754E2BFFBEFF91FBC0FF46FBAC34DD" box="[1053,1138,152,175]" pageId="8" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D2F42FFBEFF91FF00FF14FA8B3516" pageId="8" pageNumber="289">
|
||
<table id="F932DDE2FFBE0066FF7DFF14FA513515" box="[160,1423,202,359]" gridcols="11" gridrows="4" pageId="8" pageNumber="289">
|
||
<tr id="35022D00FFBE0066FF7DFF14FA513496" box="[160,1423,202,228]" gridrow="0" pageId="8" pageNumber="289" rowspan-0="1">
|
||
<th id="76D3447CFFBE0066FF00FF14FEE83496" box="[221,310,202,228]" gridcol="1" gridrow="0" pageId="8" pageNumber="289">fe</th>
|
||
<th id="76D3447CFFBE0066FE8AFF14FE6E3496" box="[343,432,202,228]" gridcol="2" gridrow="0" pageId="8" pageNumber="289">ti</th>
|
||
<th id="76D3447CFFBE0066FE0CFF14FDF43496" box="[465,554,202,228]" gridcol="3" gridrow="0" pageId="8" pageNumber="289">ta1</th>
|
||
<th id="76D3447CFFBE0066FD91FF14FD7A3496" box="[588,676,202,228]" gridcol="4" gridrow="0" pageId="8" pageNumber="289">ta2</th>
|
||
<th id="76D3447CFFBE0066FD1BFF14FCC13496" box="[710,799,202,228]" gridcol="5" gridrow="0" pageId="8" pageNumber="289">ta3</th>
|
||
<th id="76D3447CFFBE0066FC9DFF14FC5E3496" box="[832,896,202,228]" gridcol="6" gridrow="0" pageId="8" pageNumber="289">ta4</th>
|
||
<th id="76D3447CFFBE0066FC67FF14FC243496" box="[954,1018,202,228]" gridcol="7" gridrow="0" pageId="8" pageNumber="289">ta5</th>
|
||
<th id="76D3447CFFBE0066FBE9FF14FB443496" box="[1076,1178,202,228]" gridcol="8" gridrow="0" pageId="8" pageNumber="289">LR</th>
|
||
<th id="76D3447CFFBE0066FB72FF14FACA3496" box="[1199,1300,202,228]" gridcol="9" gridrow="0" pageId="8" pageNumber="289">BV</th>
|
||
<th id="76D3447CFFBE0066FAF4FF14FA513496" box="[1321,1423,202,228]" gridcol="10" gridrow="0" pageId="8" pageNumber="289">SV</th>
|
||
</tr>
|
||
<tr id="35022D00FFBE0066FF7DFF28FA513562" box="[160,1423,246,272]" gridrow="1" pageId="8" pageNumber="289">
|
||
<th id="76D3447CFFBE0066FF7DFF28FF6D3562" box="[160,179,246,272]" gridcol="0" gridrow="1" pageId="8" pageNumber="289">p1</th>
|
||
<td id="76D3447CFFBE0066FF00FF28FEE83562" box="[221,310,246,272]" gridcol="1" gridrow="1" pageId="8" pageNumber="289">7 5 5–8 0 8</td>
|
||
<td id="76D3447CFFBE0066FE8AFF28FE6E3562" box="[343,432,246,272]" gridcol="2" gridrow="1" pageId="8" pageNumber="289">3 6 0–3 9 7</td>
|
||
<td id="76D3447CFFBE0066FE0CFF28FDF43562" box="[465,554,246,272]" gridcol="3" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FD91FF28FD7A3562" box="[588,676,246,272]" gridcol="4" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FD1BFF28FCC13562" box="[710,799,246,272]" gridcol="5" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FC9DFF28FC5E3562" box="[832,896,246,272]" gridcol="6" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FC67FF28FC243562" box="[954,1018,246,272]" gridcol="7" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FBE9FF28FB443562" box="[1076,1178,246,272]" gridcol="8" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FB72FF28FACA3562" box="[1199,1300,246,272]" gridcol="9" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
<td id="76D3447CFFBE0066FAF4FF28FA513562" box="[1321,1423,246,272]" gridcol="10" gridrow="1" pageId="8" pageNumber="289">–</td>
|
||
</tr>
|
||
<tr id="35022D00FFBE0066FF7DFEFCFA51354E" box="[160,1423,290,316]" gridrow="2" pageId="8" pageNumber="289">
|
||
<th id="76D3447CFFBE0066FF7DFEFCFF6D354E" box="[160,179,290,316]" gridcol="0" gridrow="2" pageId="8" pageNumber="289">p2</th>
|
||
<td id="76D3447CFFBE0066FF00FEFCFEE8354E" box="[221,310,290,316]" gridcol="1" gridrow="2" pageId="8" pageNumber="289">742–772</td>
|
||
<td id="76D3447CFFBE0066FE8AFEFCFE6E354E" box="[343,432,290,316]" gridcol="2" gridrow="2" pageId="8" pageNumber="289">570–610</td>
|
||
<td id="76D3447CFFBE0066FE0CFEFCFDF4354E" box="[465,554,290,316]" gridcol="3" gridrow="2" pageId="8" pageNumber="289">382–397</td>
|
||
<td id="76D3447CFFBE0066FD91FEFCFD7A354E" box="[588,676,290,316]" gridcol="4" gridrow="2" pageId="8" pageNumber="289">198–206</td>
|
||
<td id="76D3447CFFBE0066FD1BFEFCFCC1354E" box="[710,799,290,316]" gridcol="5" gridrow="2" pageId="8" pageNumber="289">147–154</td>
|
||
<td id="76D3447CFFBE0066FC9DFEFCFC5E354E" box="[832,896,290,316]" gridcol="6" gridrow="2" pageId="8" pageNumber="289">88–92</td>
|
||
<td id="76D3447CFFBE0066FC67FEFCFC24354E" box="[954,1018,290,316]" gridcol="7" gridrow="2" pageId="8" pageNumber="289">59–62</td>
|
||
<td id="76D3447CFFBE0066FBE9FEFCFB44354E" box="[1076,1178,290,316]" gridcol="8" gridrow="2" pageId="8" pageNumber="289">0.64–0.65</td>
|
||
<td id="76D3447CFFBE0066FB72FEFCFACA354E" box="[1199,1300,290,316]" gridcol="9" gridrow="2" pageId="8" pageNumber="289">3.45–3.56</td>
|
||
<td id="76D3447CFFBE0066FAF4FEFCFA51354E" box="[1321,1423,290,316]" gridcol="10" gridrow="2" pageId="8" pageNumber="289">3.48–3.58</td>
|
||
</tr>
|
||
<tr id="35022D00FFBE0066FF7DFE90FA513515" box="[160,1423,334,359]" gridrow="3" pageId="8" pageNumber="289">
|
||
<th id="76D3447CFFBE0066FF7DFE90FF6D3515" box="[160,179,334,359]" gridcol="0" gridrow="3" pageId="8" pageNumber="289">p3</th>
|
||
<td id="76D3447CFFBE0066FF00FE90FEE83515" box="[221,310,334,359]" gridcol="1" gridrow="3" pageId="8" pageNumber="289">778–823</td>
|
||
<td id="76D3447CFFBE0066FE8AFE90FE6E3515" box="[343,432,334,359]" gridcol="2" gridrow="3" pageId="8" pageNumber="289">706–750</td>
|
||
<td id="76D3447CFFBE0066FE0CFE90FDF43515" box="[465,554,334,359]" gridcol="3" gridrow="3" pageId="8" pageNumber="289">457</td>
|
||
<td id="76D3447CFFBE0066FD91FE90FD7A3515" box="[588,676,334,359]" gridcol="4" gridrow="3" pageId="8" pageNumber="289">305</td>
|
||
<td id="76D3447CFFBE0066FD1BFE90FCC13515" box="[710,799,334,359]" gridcol="5" gridrow="3" pageId="8" pageNumber="289">281</td>
|
||
<td id="76D3447CFFBE0066FC9DFE90FC5E3515" box="[832,896,334,359]" gridcol="6" gridrow="3" pageId="8" pageNumber="289">165</td>
|
||
<td id="76D3447CFFBE0066FC67FE90FC243515" box="[954,1018,334,359]" gridcol="7" gridrow="3" pageId="8" pageNumber="289">81</td>
|
||
<td id="76D3447CFFBE0066FBE9FE90FB443515" box="[1076,1178,334,359]" gridcol="8" gridrow="3" pageId="8" pageNumber="289">0.64</td>
|
||
<td id="76D3447CFFBE0066FB72FE90FACA3515" box="[1199,1300,334,359]" gridcol="9" gridrow="3" pageId="8" pageNumber="289">2.36</td>
|
||
<td id="76D3447CFFBE0066FAF4FE90FA513515" box="[1321,1423,334,359]" gridcol="10" gridrow="3" pageId="8" pageNumber="289">3.30</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF4D7FCAFFBEFF91FF4AF9A4FA4532E2" box="[151,1435,1658,1680]" httpUri="https://zenodo.org/record/831738/files/figure.png" pageId="8" pageNumber="289" targetBox="[151,1436,435,1636]" targetPageId="8">
|
||
<paragraph id="8B8D2F42FFBEFF91FF4AF9A4FA4532E2" blockId="8.[151,1435,1658,1680]" box="[151,1435,1658,1680]" pageId="8" pageNumber="289">
|
||
<emphasis id="B946F350FFBEFF91FF4AF9A4FECC32FD" bold="true" box="[151,274,1658,1680]" pageId="8" pageNumber="289">FIGURE 4.</emphasis>
|
||
Wing of male:
|
||
<taxonomicName id="4C3254C1FFBEFF91FE6DF9A5FDAF32E2" authority="Dantas & Giłka, 2017" authorityName="Dantas & Giłka" authorityYear="2017" box="[432,625,1658,1680]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="illustris" status="sp. nov.">
|
||
<emphasis id="B946F350FFBEFF91FE6DF9A5FDAF32E2" box="[432,625,1658,1680]" italics="true" pageId="8" pageNumber="289">Tanytarsus illustris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2754E2BFFBEFF91FDA5F9A4FD1632FD" box="[632,712,1658,1679]" pageId="8" pageNumber="289" rank="species">
|
||
<emphasis id="B946F350FFBEFF91FDA5F9A4FD1632FD" bold="true" box="[632,712,1658,1679]" pageId="8" pageNumber="289">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B946F350FFBEFF91FD05F9A4FD3432FD" bold="true" box="[728,746,1658,1679]" pageId="8" pageNumber="289">A</emphasis>
|
||
),
|
||
<taxonomicName id="4C3254C1FFBEFF91FD22F9A5FC1C32E2" authority="Dantas & Giłka, 2017" authorityName="Dantas & Giłka" authorityYear="2017" box="[767,962,1659,1680]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="insignis" status="sp. nov.">
|
||
<emphasis id="B946F350FFBEFF91FD22F9A5FC1C32E2" box="[767,962,1659,1680]" italics="true" pageId="8" pageNumber="289">Tanytarsus insignis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946F350FFBEFF91FC15F9A4FBC732FD" bold="true" box="[968,1049,1658,1679]" pageId="8" pageNumber="289">
|
||
<taxonomicNameLabel id="A2754E2BFFBEFF91FC15F9A4FBC732FD" box="[968,1049,1658,1679]" pageId="8" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B946F350FFBEFF91FBF5F9A4FBE732FD" bold="true" box="[1064,1081,1658,1679]" pageId="8" pageNumber="289">B</emphasis>
|
||
),
|
||
<taxonomicName id="4C3254C1FFBEFF91FB93F9A5FACB32E2" authority="Dantas & Giłka, 2017" authorityName="Dantas & Giłka" authorityYear="2017" box="[1102,1301,1658,1680]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="insolens" status="sp. nov.">
|
||
<emphasis id="B946F350FFBEFF91FB93F9A5FACB32E2" box="[1102,1301,1658,1680]" italics="true" pageId="8" pageNumber="289">Tanytarsus insolens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946F350FFBEFF91FAC6F9A4FAB232FD" bold="true" box="[1307,1388,1658,1679]" pageId="8" pageNumber="289">
|
||
<taxonomicNameLabel id="A2754E2BFFBEFF91FAC6F9A4FAB232FD" box="[1307,1388,1658,1679]" pageId="8" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B946F350FFBEFF91FAA6F9A4FA5332E2" bold="true" box="[1403,1421,1658,1680]" pageId="8" pageNumber="289">C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D2F42FFBEFF90FF1AF963FD1C348A" blockId="8.[151,1437,1724,2037]" lastBlockId="9.[151,1437,151,680]" lastPageId="9" lastPageNumber="290" pageId="8" pageNumber="289">
|
||
<emphasis id="B946F350FFBEFF91FF1AF963FE8832A6" box="[199,342,1725,1748]" italics="true" pageId="8" pageNumber="289">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="130933C7FFBEFF91FEBBF962FE7732A7" box="[358,425,1724,1749]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="8" pageNumber="289">Fig. 3</figureCitation>
|
||
A–F). Tergite IX covered with dense short microtrichia on entire surface, sparse hair-like 10–15 µm long macrotrichia in anterior part (
|
||
<figureCitation id="130933C7FFBEFF91FD44F93EFD05328B" box="[665,731,1760,1785]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="8" pageNumber="289">Fig. 3</figureCitation>
|
||
A, C) and 20–25 stout setae around base of anal point. Lateral teeth absent. Anal tergite bands Y-shaped, fading at base of anal point. Anal point triangular, with short round crests flanking 4–7 spinulae, with the most distal one enlarged, bar-like (
|
||
<figureCitation id="130933C7FFBEFF91FC02F8F6FBFD3333" box="[991,1059,1832,1857]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="8" pageNumber="289">Fig. 3</figureCitation>
|
||
A, D). Superior volsella bilobed: anterior (ventral) projection finger-shaped, with 3–4 apical/subapical setae, posterior (dorsal) projection narrowly triangular, pointed, bearing 4 short setae dorsally (2 proximal + 2 distal); digitus large, extending far beyond superior volsella, somewhat lanceolate or knife-shaped, broadly fused with posterior projection of superior volsella as shown in
|
||
<figureCitation id="130933C7FFBEFF91FEF8F866FEB933A3" box="[293,359,1976,2001]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="8" pageNumber="289">Fig. 3</figureCitation>
|
||
A, B, E. Stem of median volsella simple, 12–15 µm long, with 4–5 subulate lamellae (
|
||
<figureCitation id="130933C7FFBEFF91FAE2F866FA5D33A3" box="[1343,1411,1976,2001]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="8" pageNumber="289">Fig. 3</figureCitation>
|
||
B, F). Inferior volsella 70–85 µm long, evenly curved and posteromedially directed, with distinct longitudinal ridge dorsally. Phallapodeme 65–75 µm long; transverse sternapodeme 53–64 µm long, with well developed oral projections. Gonocoxite 102–109 µm long. Gonostylus 98–107 µm long, narrow, evenly curved and tapering toward blunt apex. HR 1.02–1.08, HV 2.17–2.52.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3287CC9FFBFFF90FF1AFEDAFE6F36DA" pageId="9" pageNumber="290" type="discussion">
|
||
<paragraph id="8B8D2F42FFBFFF90FF1AFEDAFE6F36DA" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="290">
|
||
<emphasis id="B946F350FFBFFF90FF1AFEDAFE92356F" bold="true" box="[199,332,260,285]" pageId="9" pageNumber="290">Discussion.</emphasis>
|
||
The adult male of
|
||
<taxonomicName id="4C3254C1FFBFFF90FDFDFEDBFD23356F" authorityName="Dantas & Giłka" authorityYear="2017" box="[544,765,260,285]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B946F350FFBFFF90FDFDFEDBFD23356F" box="[544,765,260,285]" italics="true" pageId="9" pageNumber="290">Tanytarsus insolens</emphasis>
|
||
</taxonomicName>
|
||
displays several distinct characters treated as diagnostic for a new group here proposed for this new species and
|
||
<taxonomicName id="4C3254C1FFBFFF90FD39FEF7FCE53532" authorityName="Vinogradova, Riss et Spies" authorityYear="2009" box="[740,827,295,320]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="kiche">
|
||
<emphasis id="B946F350FFBFFF90FD39FEF7FCE53532" box="[740,827,295,320]" italics="true" pageId="9" pageNumber="290">T. kiche</emphasis>
|
||
</taxonomicName>
|
||
. The two species are unique in having the hypopygial superior volsella divided into two lobes or projections: anterior (ventral) and posterior (dorsal), and the stout digitus, extending far beyond the superior volsella. For homology of these structures in
|
||
<taxonomicName id="4C3254C1FFBFFF90FBAEFEAFFA8E35FA" authorityName="Dantas & Giłka" authorityYear="2017" box="[1139,1360,367,392]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B946F350FFBFFF90FBAEFEAFFA8E35FA" box="[1139,1360,367,392]" italics="true" pageId="9" pageNumber="290">Tanytarsus insolens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3254C1FFBFFF90FA5AFEAFFF0C35DF" authorityName="Vinogradova, Riss et Spies" authorityYear="2009" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="kiche">
|
||
<emphasis id="B946F350FFBFFF90FA5AFEAFFF0C35DF" italics="true" pageId="9" pageNumber="290">T. kiche</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<figureCitation id="130933C7FFBFFF90FED6FE4AFE9435DE" box="[267,330,404,429]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="9" pageNumber="290">Fig. 3</figureCitation>
|
||
E and G. The round anal point crests flanking the spinulae, with the most distal one separated from the others or enlarged and bar-shaped are also characters common for the two species, so treated as supporting the group concept (
|
||
<emphasis id="B946F350FFBFFF90FE9BFE02FEBC3586" box="[326,354,476,500]" italics="true" pageId="9" pageNumber="290">cf.</emphasis>
|
||
<figureCitation id="130933C7FFBFFF90FEB4FE02FE773586" box="[361,425,476,501]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="9" pageNumber="290">Fig. 3</figureCitation>
|
||
A, D and
|
||
<bibRefCitation id="EFA352B3FFBFFF90FDCAFE02FCF43586" author="Vinogradova" box="[535,810,476,501]" pageId="9" pageNumber="290" refString="Vinogradova, E. M., Riss, H. W. & Spies, M. (2009) New species of Tanytarsus van der Wulp, 1874 (Diptera: Chironomidae) from Central America. Aquatic Insects, 31, 11 - 17." type="journal article" year="2009">
|
||
Vinogradova
|
||
<emphasis id="B946F350FFBFFF90FD72FE03FD393587" box="[687,743,476,501]" italics="true" pageId="9" pageNumber="290">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
: fig. 2). Another extraordinary character found in males of
|
||
<taxonomicName id="4C3254C1FFBFFF90FF6BFDDFFE4A366A" authorityName="Dantas & Giłka" authorityYear="2017" box="[182,404,511,536]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B946F350FFBFFF90FF6BFDDFFE4A366A" box="[182,404,511,536]" italics="true" pageId="9" pageNumber="290">Tanytarsus insolens</emphasis>
|
||
</taxonomicName>
|
||
are the long hair-like macrotrichia on the anal tergite (
|
||
<figureCitation id="130933C7FFBFFF90FBDFFE21FB9B366A" box="[1026,1093,511,536]" captionStart="FIGURE 3" captionStartId="7.[151,250,1870,1892]" captionTargetBox="[180,1412,184,1823]" captionTargetId="figure@7.[180,1412,184,1851]" captionTargetPageId="7" captionText="FIGURE 3. Tanytarsus insolens sp. nov. (A – F) and T. kiche Vinogradova, Riss et Spies, 2009 (G), male. A, B — hypopygium in dorsal (A) and ventral aspect (B); C — anterior region of anal tergite covered with microtrichia and hair-like macrotrichia; D — anal point (variation); E, G — superior volsella: anterior / ventral projection (av, yellow arrow), posterior / dorsal projection (pd, red) and digitus (di, green); F — median volsella [C – F magnified ca. 2 – 3 times relative to A and B; G out of scale, after Vinogradova et al. (2009)]." httpUri="https://zenodo.org/record/831736/files/figure.png" pageId="9" pageNumber="290">Fig. 3</figureCitation>
|
||
A, C). This peculiar character, however, does not appear in males of
|
||
<taxonomicName id="4C3254C1FFBFFF90FDB0FDFBFCF1364F" authorityName="Vinogradova, Riss et Spies" authorityYear="2009" box="[621,815,548,573]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="kiche">
|
||
<emphasis id="B946F350FFBFFF90FDB0FDFBFCF1364F" box="[621,815,548,573]" italics="true" pageId="9" pageNumber="290">Tanytarsus kiche</emphasis>
|
||
</taxonomicName>
|
||
(Spies, pers. comm.), thus should be treated as an autapomorphy as long as it is known from
|
||
<taxonomicName id="4C3254C1FFBFFF90FDA2FD97FD263612" authorityName="Dantas & Giłka" authorityYear="2017" box="[639,760,583,608]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B946F350FFBFFF90FDA2FD97FD263612" box="[639,760,583,608]" italics="true" pageId="9" pageNumber="290">T. insolens</emphasis>
|
||
</taxonomicName>
|
||
exclusively. The two representatives of the
|
||
<taxonomicName id="4C3254C1FFBFFF90FB30FD99FAF63612" authorityName="Vinogradova, Riss et Spies" authorityYear="2009" box="[1261,1320,583,608]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="kiche">
|
||
<emphasis id="B946F350FFBFFF90FB30FD99FAF63612" box="[1261,1320,583,608]" italics="true" pageId="9" pageNumber="290">kiche</emphasis>
|
||
</taxonomicName>
|
||
group are distributed across Central America, from
|
||
<collectingRegion id="49F6E1A0FFBFFF90FDBBFDB2FD6536F7" box="[614,699,620,645]" country="Mexico" name="Mexico" pageId="9" pageNumber="290">
|
||
<collectingCountry id="F3256FD2FFBFFF90FDBBFDB2FD6536F7" box="[614,699,620,645]" name="Mexico" pageId="9" pageNumber="290">Mexico</collectingCountry>
|
||
</collectingRegion>
|
||
in the north through
|
||
<collectingRegion id="49F6E1A0FFBFFF90FC7AFDB3FBDE36F6" box="[935,1024,621,644]" country="Panama" name="Panama" pageId="9" pageNumber="290">
|
||
<collectingCountry id="F3256FD2FFBFFF90FC7AFDB3FBDE36F6" box="[935,1024,621,644]" name="Panama" pageId="9" pageNumber="290">Panama</collectingCountry>
|
||
</collectingRegion>
|
||
(
|
||
<taxonomicName id="4C3254C1FFBFFF90FBCDFDB3FBBA36F7" authorityName="Vinogradova, Riss et Spies" authorityYear="2009" box="[1040,1124,620,645]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="kiche">
|
||
<emphasis id="B946F350FFBFFF90FBCDFDB3FBBA36F7" box="[1040,1124,620,645]" italics="true" pageId="9" pageNumber="290">T. kiche</emphasis>
|
||
</taxonomicName>
|
||
) to the Brazilian Amazonia in the south (
|
||
<taxonomicName id="4C3254C1FFBFFF90FEF7FD4FFE7F36DA" authorityName="Dantas & Giłka" authorityYear="2017" box="[298,417,655,680]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B946F350FFBFFF90FEF7FD4FFE7F36DA" box="[298,417,655,680]" italics="true" pageId="9" pageNumber="290">T. insolens</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |