670 lines
163 KiB
XML
670 lines
163 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7867FFA5FF013FD2AB1DECD9" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Hylekobolus Wesener & Enghoff & Sierwald 2009, gen. n." docType="treatment" docUuid="A90561DE-4844-47EF-8654-19C38A402BA5" docUuidSource="ZooBank" docVersion="6" lastPageNumber="100" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="97" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791471" ID-GBIF-Taxon="164184217" ID-Zenodo-Dep="3791471" LSID="urn:lsid:zoobank.org:act:A90561DE-4844-47EF-8654-19C38A402BA5" httpUri="http://treatment.plazi.org/id/1772122E7867FFA5FF013FD2AB1DECD9" lastPageId="99" lastPageNumber="100" pageId="96" pageNumber="97">
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<subSubSection box="[140,486,166,193]" pageId="96" pageNumber="97" type="nomenclature">
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<paragraph blockId="96.[140,962,166,229]" box="[140,486,166,193]" pageId="96" pageNumber="97">
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<heading box="[140,486,166,193]" fontSize="11" level="2" pageId="96" pageNumber="97" reason="3">
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<emphasis bold="true" box="[140,486,166,193]" pageId="96" pageNumber="97">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,393,166,193]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[140,285,166,193]" italics="true" pageId="96" pageNumber="97">Hylekobolus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[405,486,167,193]" pageId="96" pageNumber="97" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,962,202,229]" pageId="96" pageNumber="97" type="description">
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<paragraph blockId="96.[140,962,166,229]" box="[140,962,202,229]" pageId="96" pageNumber="97">
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<uri box="[140,962,202,229]" pageId="96" pageNumber="97">
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urn:lsid:zoobank.org:act:
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<uuid box="[417,962,202,229]" pageId="96" pageNumber="97">A90561DE-4844-47EF-8654-19C38A402BA5</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSection lastPageId="99" lastPageNumber="100" pageId="96" pageNumber="97" type="multiple">
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<subSubSection box="[140,684,272,299]" pageId="96" pageNumber="97" type="type_taxon">
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<paragraph blockId="96.[140,684,272,299]" box="[140,684,272,299]" pageId="96" pageNumber="97">
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<emphasis bold="true" box="[140,287,272,298]" pageId="96" pageNumber="97">
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<typeStatus box="[140,198,272,298]" pageId="96" pageNumber="97">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[302,616,272,298]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[302,616,272,298]" italics="true" pageId="96" pageNumber="97">Hylekobolus brachiosauroides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[623,684,273,299]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="96" pageNumber="97" type="reference_group">
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,419,342,369]" pageId="96" pageNumber="97">
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||
<emphasis bold="true" box="[140,413,342,368]" pageId="96" pageNumber="97">
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<emphasis box="[140,212,342,368]" inLineHeading="true" pageId="96" pageNumber="97" reason="1">Other</emphasis>
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species included
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</emphasis>
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||
:
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</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,300,378,404]" pageId="96" pageNumber="97">
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,232,378,404]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,232,378,404]" italics="true" pageId="96" pageNumber="97">H. rufus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[239,300,378,404]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,315,414,440]" pageId="96" pageNumber="97">
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,246,414,439]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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||
<emphasis box="[140,246,414,439]" italics="true" pageId="96" pageNumber="97">H. griseus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[254,315,414,440]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="96.[140,419,342,721]" box="[140,362,448,475]" pageId="96" pageNumber="97">
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,294,448,474]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,294,448,474]" italics="true" pageId="96" pageNumber="97">H. albicollaris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[301,362,449,475]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,351,483,510]" pageId="96" pageNumber="97">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,283,483,509]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,283,483,509]" italics="true" pageId="96" pageNumber="97">H. goodmani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[290,351,484,510]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,351,519,545]" pageId="96" pageNumber="97">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,283,519,544]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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||
<emphasis box="[140,283,519,544]" italics="true" pageId="96" pageNumber="97">H. montanus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel box="[290,351,519,545]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,375,554,580]" pageId="96" pageNumber="97">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,307,554,580]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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||
<emphasis box="[140,307,554,580]" italics="true" pageId="96" pageNumber="97">H. analavelona</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel box="[314,375,554,580]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,334,589,616]" pageId="96" pageNumber="97">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,266,589,615]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,266,589,615]" italics="true" pageId="96" pageNumber="97">H. latifrons</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel box="[273,334,590,616]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
<paragraph blockId="96.[140,419,342,721]" box="[140,378,624,651]" pageId="96" pageNumber="97">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,310,624,650]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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||
<emphasis box="[140,310,624,650]" italics="true" pageId="96" pageNumber="97">H. andasibensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[317,378,625,651]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="96.[140,419,342,721]" box="[140,337,660,686]" pageId="96" pageNumber="97">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,269,660,685]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,269,660,685]" italics="true" pageId="96" pageNumber="97">H. marojejy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[276,337,660,686]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="96.[140,419,342,721]" box="[140,390,695,721]" pageId="96" pageNumber="97">
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,322,695,721]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,322,695,721]" italics="true" pageId="96" pageNumber="97">H. anjanaharibe</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[329,390,695,721]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97" type="diagnosis">
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||
<paragraph blockId="96.[140,1108,765,1672]" pageId="96" pageNumber="97">
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<emphasis bold="true" box="[140,258,765,791]" inLineHeading="true" pageId="96" pageNumber="97" reason="1">Diagnosis</emphasis>
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: legs without coxal processes, but coxa
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<quantity box="[714,762,765,792]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="96" pageNumber="97" unit="in" value="7.0">7 in</quantity>
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males always twice as wide as coxa 6 (
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<figureCitation box="[227,328,800,827]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Fig. 55B</figureCitation>
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), a unique feature for
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<taxonomicName box="[578,748,800,826]" class="Diplopoda" family="Spirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="family">Spirobolellidae</taxonomicName>
|
||
. The anterior (
|
||
<figureCitation box="[912,1049,800,827]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55L, N</figureCitation>
|
||
) and posterior gonopods (
|
||
<figureCitation box="[369,513,835,862]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55M, O</figureCitation>
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||
) of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[558,686,835,861]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[558,686,835,861]" italics="true" pageId="96" pageNumber="97">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
are unique within the order
|
||
<taxonomicName authorityName="Cook" authorityYear="1895" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
|
||
. Coxite of posterior gonopods basally at mesal margin with a triangular sclerite (
|
||
<figureCitation captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55O</figureCitation>
|
||
,
|
||
<figureCitation box="[208,261,906,933]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="96" pageNumber="97">56G</figureCitation>
|
||
,
|
||
<figureCitation box="[275,329,906,933]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="96" pageNumber="97">57H</figureCitation>
|
||
,
|
||
<figureCitation box="[343,397,906,933]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="96" pageNumber="97">58G</figureCitation>
|
||
), a potential apomorphy of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[718,846,906,932]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[718,846,906,932]" italics="true" pageId="96" pageNumber="97">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
. Species medium-sized (
|
||
<quantity box="[148,236,941,968]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="96" pageNumber="97" unit="mm" value="30.0">30 mm</quantity>
|
||
) to quite large (
|
||
<quantity box="[427,515,941,968]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="96" pageNumber="97" unit="mm" value="75.0">75 mm</quantity>
|
||
) for species of the
|
||
<taxonomicName box="[741,911,941,967]" class="Diplopoda" family="Spirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="family">Spirobolellidae</taxonomicName>
|
||
. Anterior part of body rings usually black, dark grey or reddish, posterior third light brown to white (
|
||
<figureCitation captionStart="Figure 53" captionStartId="97.[140,212,848,870]" captionTargetBox="[140,1108,165,829]" captionTargetId="figure@97.[140,1108,165,829]" captionTargetPageId="97" captionText="Figure 53. Living specimens of A Hylekobolus brachiosauroides sp. n., male B H. griseus sp. n., mating pair C H. albicollaris sp. n." figureDoi="http://doi.org/10.5281/zenodo.3820856" httpUri="https://zenodo.org/record/3820856/files/figure.png" pageId="96" pageNumber="97">Figs 53</figureCitation>
|
||
A–C). Telson with a well-rounded preanal process (
|
||
<figureCitation box="[752,859,1011,1038]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55C</figureCitation>
|
||
,
|
||
<figureCitation box="[873,925,1011,1038]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="96" pageNumber="97">56C</figureCitation>
|
||
), which is however much shorter than in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[441,580,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[441,580,1047,1073]" italics="true" pageId="96" pageNumber="97">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[620,811,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[620,811,1047,1073]" italics="true" pageId="96" pageNumber="97">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
species. Anal valves small, lips absent (
|
||
<figureCitation box="[272,377,1082,1109]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55C</figureCitation>
|
||
,
|
||
<figureCitation box="[389,440,1082,1109]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="96" pageNumber="97">56C</figureCitation>
|
||
,
|
||
<figureCitation box="[453,502,1082,1109]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="96" pageNumber="97">57E</figureCitation>
|
||
,
|
||
<figureCitation box="[513,566,1082,1109]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="96" pageNumber="97">58C</figureCitation>
|
||
,
|
||
<figureCitation box="[577,629,1082,1109]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="96" pageNumber="97">59C</figureCitation>
|
||
,
|
||
<figureCitation box="[641,694,1082,1109]" captionStart="Figure 60" captionStartId="111.[140,212,844,866]" captionTargetBox="[140,1108,165,819]" captionTargetId="figure@111.[140,1108,438,824]" captionTargetPageId="111" captionText="Figure 60. Hylekobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767358" httpUri="https://zenodo.org/record/3767358/files/figure.png" pageId="96" pageNumber="97">60C</figureCitation>
|
||
). Collum ventrally not protruding as far as body ring 2, but anteriorly covering first mandibular basal joint as well as part of antenna (
|
||
<figureCitation box="[280,426,1152,1179]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55A, D</figureCitation>
|
||
). Part of head projecting slightly above second mandibular basal joint (
|
||
<figureCitation box="[269,375,1187,1214]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Fig. 55D</figureCitation>
|
||
). Such a modified head capsule (
|
||
<figureCitation box="[738,845,1187,1214]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55D</figureCitation>
|
||
,
|
||
<figureCitation box="[857,907,1187,1214]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="96" pageNumber="97">57C</figureCitation>
|
||
) and collum (
|
||
<figureCitation captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Fig. 55A</figureCitation>
|
||
) are similar to the conditions observed in the South American genus
|
||
<taxonomicName authority="(Hoffman 1979)" baseAuthorityName="Hoffman" baseAuthorityYear="1979" class="Diplopoda" family="Pachybolidae" genus="Atlanticobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[955,1107,1223,1249]" italics="true" pageId="96" pageNumber="97">Atlanticobolus</emphasis>
|
||
(
|
||
<bibRefCitation author="Hoffman" box="[147,322,1258,1284]" firstAuthor="Hoffman" pageId="96" pageNumber="97" pagination="930 - 934" refId="ref57195" refString="Hoffman RL (1979) Studies on spiroboloid millipeds. XII. The status of Spirobolus noronhensis Pocock, 1890, and some related species (Pachybolidae). Proceedings of the Biological Society of Washington 91 (4): 930 - 934." type="journal article" year="1979">Hoffman 1979</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, as well as the West African genus
|
||
<taxonomicName authority="(Keeton 1964)" baseAuthorityName="Keeton" baseAuthorityYear="1964" box="[725,1025,1258,1285]" class="Diplopoda" genus="Amblybolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[725,849,1258,1284]" italics="true" pageId="96" pageNumber="97">Amblybolus</emphasis>
|
||
(
|
||
<bibRefCitation author="Keeton" box="[864,1016,1258,1285]" firstAuthor="Keeton" pageId="96" pageNumber="97" pagination="5 - 6" refId="ref57322" refString="Keeton WT (1964) Amblybolus genus nov. (Diplopoda: Spirobolida: Trigoniuloidea). Pilot Register of Zoology, card Nos 5 - 6." type="book chapter" year="1964">Keeton 1964</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, whose family positions are currently unclear (
|
||
<bibRefCitation author="Hoffman" box="[577,753,1293,1319]" firstAuthor="Hoffman" pageId="96" pageNumber="97" refId="ref57240" refString="Hoffman RL (1980) Classification of the Diplopoda. Musee d'Histoire Naturelle de Geneve, Geneve, Switzerland, 237 pp." type="book" year="1980">Hoffman 1980</bibRefCitation>
|
||
). However, in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[921,1049,1293,1319]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[921,1049,1293,1319]" italics="true" pageId="96" pageNumber="97">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
only the second mandibular joint and the collum form a shallow cavity for the short antenna (
|
||
<figureCitation box="[148,254,1363,1390]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="96" pageNumber="97">Fig. 56A</figureCitation>
|
||
), while in
|
||
<taxonomicName baseAuthorityName="Keeton" baseAuthorityYear="1964" box="[381,505,1363,1389]" class="Diplopoda" genus="Amblybolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[381,505,1363,1389]" italics="true" pageId="96" pageNumber="97">Amblybolus</emphasis>
|
||
</taxonomicName>
|
||
the area around the ocelli is strongly modified, too (
|
||
<bibRefCitation author="Keeton" box="[147,297,1399,1425]" firstAuthor="Keeton" pageId="96" pageNumber="97" pagination="5 - 6" refId="ref57322" refString="Keeton WT (1964) Amblybolus genus nov. (Diplopoda: Spirobolida: Trigoniuloidea). Pilot Register of Zoology, card Nos 5 - 6." type="book chapter" year="1964">Keeton 1964</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Demange and Mauries" box="[308,631,1398,1425]" firstAuthor="Demange" pageId="96" pageNumber="97" pagination="1 - 192" refId="ref56869" refString="Demange J-M, Mauries J-P (1975) Myriapodes - Diplopodes des Monts Nimba et Tonkoui (Cote d'Ivoire, Guinee) recoltes par M. Lamotte et ses collaborateurs de 1942 a 1961. Annalen, Koninklijk Museum voor Midden-Afrika - Zoologische wetenschappen 212: 1 - 192." type="journal article" year="1975">Demange and Mauriès 1975</bibRefCitation>
|
||
). Similar modifications of the head capsule also appear in numerous species of the family
|
||
<taxonomicName box="[652,821,1434,1460]" class="Diplopoda" family="Spirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="family">Spirobolellidae</taxonomicName>
|
||
, for example in the genus
|
||
<taxonomicName authorityName="Mauries" authorityYear="1980" box="[140,250,1469,1495]" class="Diplopoda" family="Pseudospirobolellidae" genus="Benoitolus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,250,1469,1495]" italics="true" pageId="96" pageNumber="97">Benoitolus</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingCountry box="[365,476,1469,1495]" name="Seychelles" pageId="96" pageNumber="97">Seychelles</collectingCountry>
|
||
Islands (
|
||
<bibRefCitation author="Mauries" box="[576,739,1469,1495]" firstAuthor="Mauries" pageId="96" pageNumber="97" pagination="138 - 168" refId="ref57430" refString="Mauries J-P (1980) Contributions a l'etude de la faune terrestre des ile granitiques de ' archipel des Sechelles (Mission P. L. G. Benoit - J. J. Van Mol 1972). Myriapoda - Diplopoda. Revue Zoologique Africaine 94 (1): 138 - 168." type="journal article" year="1980">Mauriès 1980</bibRefCitation>
|
||
). Male legs with tarsal pads only on legs 3–7 (
|
||
<figureCitation box="[283,384,1504,1531]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Fig. 55B</figureCitation>
|
||
). Short and wide tarsus with a single pair of long median setae in legs 8 and beyond (
|
||
<figureCitation box="[356,488,1539,1566]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Figs 55H, I</figureCitation>
|
||
), unique for Malagasy millipedes. Body ring
|
||
<quantity box="[993,1038,1539,1566]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="96" pageNumber="97" unit="in" value="7.0">7 in</quantity>
|
||
males greatly enlarged, ventrally as wide as two regular body rings (
|
||
<figureCitation box="[855,959,1574,1601]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="96" pageNumber="97">Fig. 57B</figureCitation>
|
||
), a character shared only with the Malagasy genus
|
||
<taxonomicName authorityName="de Saussure & Zehntner" authorityYear="1901" box="[548,676,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[548,676,1610,1636]" italics="true" pageId="96" pageNumber="97">Spiromimus</emphasis>
|
||
</taxonomicName>
|
||
. Anterior gonopods prominently elongated (
|
||
<figureCitation box="[214,315,1645,1672]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="96" pageNumber="97">Figs 57E</figureCitation>
|
||
,
|
||
<figureCitation box="[327,381,1645,1672]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="96" pageNumber="97">58D</figureCitation>
|
||
), especially telopodites (
|
||
<figureCitation box="[650,755,1645,1672]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="96" pageNumber="97">Fig. 55N</figureCitation>
|
||
), tips of anterior gonopods well
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820856" ID-Zenodo-Dep="3820856" httpUri="https://zenodo.org/record/3820856/files/figure.png" pageId="97" pageNumber="98" targetBox="[140,1108,165,829]" targetPageId="97">
|
||
<paragraph blockId="97.[140,1107,848,903]" pageId="97" pageNumber="98">
|
||
<emphasis bold="true" box="[140,252,848,870]" pageId="97" pageNumber="98">Figure 53.</emphasis>
|
||
Living specimens of
|
||
<emphasis bold="true" box="[452,471,848,870]" pageId="97" pageNumber="98">A</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[477,735,849,870]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[477,735,849,870]" italics="true" pageId="97" pageNumber="98">Hylekobolus brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[742,795,849,870]" pageId="97" pageNumber="98" rank="species">
|
||
<emphasis bold="true" box="[742,789,849,870]" pageId="97" pageNumber="98">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male
|
||
<emphasis bold="true" box="[859,876,848,870]" pageId="97" pageNumber="98">B</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[883,971,849,870]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[883,971,849,870]" italics="true" pageId="97" pageNumber="98">H. griseus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[978,1030,849,870]" pageId="97" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
|
||
, mating pair
|
||
<emphasis bold="true" box="[182,201,880,902]" pageId="97" pageNumber="98">C</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[206,333,881,902]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[206,333,881,902]" italics="true" pageId="97" pageNumber="98">H. albicollaris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[338,389,881,902]" pageId="97" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767346" ID-Zenodo-Dep="3767346" box="[138,719,1653,1675]" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="97" pageNumber="98" startId="97.[138,210,1653,1675]" targetBox="[130,1107,1100,1636]" targetPageId="97">
|
||
<paragraph blockId="97.[138,719,1653,1675]" box="[138,719,1653,1675]" pageId="97" pageNumber="98">
|
||
<emphasis bold="true" box="[138,250,1653,1675]" pageId="97" pageNumber="98">Figure 54.</emphasis>
|
||
Distribution map of the genus
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[543,649,1653,1674]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[543,649,1653,1674]" italics="true" pageId="97" pageNumber="98">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[655,719,1653,1674]" pageId="97" pageNumber="98" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
visible, even in non-dissected specimens (
|
||
<figureCitation box="[597,699,166,193]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55B</figureCitation>
|
||
,
|
||
<figureCitation box="[711,762,166,193]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="98" pageNumber="99">57B</figureCitation>
|
||
). Posterior gonopod coxite and telopodite fused, slender and conspicuously elongated (
|
||
<figureCitation box="[746,853,202,229]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55O</figureCitation>
|
||
,
|
||
<figureCitation box="[864,918,202,229]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="98" pageNumber="99">56G</figureCitation>
|
||
), which is typical for the species of the family. Telopodites arranged face-to-face with one another. Tip of telopodite bent mesally by 60° (
|
||
<figureCitation box="[494,597,272,298]" captionStart="Figure 34" captionStartId="61.[140,212,641,663]" captionTargetBox="[140,1108,165,621]" captionTargetId="figure@61.[140,1108,165,621]" captionTargetPageId="61" captionText="Figure 34. Pseudocentrobolus aureus sp. n., male holotype, anterior gonopod A anterior view B posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="98" pageNumber="99">Fig. 34D</figureCitation>
|
||
) or (more often) 90° (
|
||
<figureCitation box="[845,951,272,299]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Fig. 55O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="98" pageNumber="99" type="distribution">
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
<emphasis bold="true" box="[185,489,307,333]" pageId="98" pageNumber="99">
|
||
<emphasis box="[185,336,307,333]" inLineHeading="true" pageId="98" pageNumber="99" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: all species of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[655,785,307,333]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[655,785,307,333]" italics="true" pageId="98" pageNumber="99">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
were collected in rainforest. Most of the species (6 of 11) were found in Southeastern
|
||
<collectingCountry box="[802,935,343,369]" name="Madagascar" pageId="98" pageNumber="99">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation box="[952,1038,342,369]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="98" pageNumber="99">Fig. 54</figureCitation>
|
||
). This could represent a collection artefact since a larger amount of inventory work was conducted around Tolagnaro (Fort Dauphin). That only few species of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[907,1037,413,439]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[907,1037,413,439]" italics="true" pageId="98" pageNumber="99">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
occur north of Andasibe is puzzling, as it can hardly be accounted for by the lack of more exhaustive inventories. Of special interest is the occurrence of
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[840,1107,483,509]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[840,1107,483,509]" italics="true" pageId="98" pageNumber="99">Hylekobolus analavelona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[140,202,519,545]" pageId="98" pageNumber="99" rank="species">sp. n.</taxonomicNameLabel>
|
||
in the isolated western rainforest of Analavelona. The Analavelona rainforest is isolated by large areas of dry and spiny forest (in which no
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[808,938,554,580]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[808,938,554,580]" italics="true" pageId="98" pageNumber="99">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
species was recorded yet) from other humid forests (
|
||
<bibRefCitation author="Moat and Smith" box="[573,831,589,615]" firstAuthor="Moat" pageId="98" pageNumber="99" refId="ref57536" refString="Moat J, Smith P (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat and Smith 2007</bibRefCitation>
|
||
). Numerous
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[978,1108,589,615]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[978,1108,589,615]" italics="true" pageId="98" pageNumber="99">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
species are obviously limited in their distribution areas, which have very sharp boundaries (
|
||
<figureCitation box="[194,279,659,686]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="98" pageNumber="99">Fig. 54</figureCitation>
|
||
). The mechanisms explaining those boundaries are as little understood as in other Malagasy millipedes.
|
||
</paragraph>
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
Large numbers of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[404,534,730,756]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[404,534,730,756]" italics="true" pageId="98" pageNumber="99">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
specimens were found inside dead wood. Single specimens walking on the forest floor were rarely discovered. Two
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[893,1023,765,791]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[893,1023,765,791]" italics="true" pageId="98" pageNumber="99">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
species were often sympatric (
|
||
<figureCitation box="[393,478,800,827]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="98" pageNumber="99">Fig. 54</figureCitation>
|
||
). In such instances, one species was always significantly larger than the other (like
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[437,529,836,862]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[437,529,836,862]" italics="true" pageId="98" pageNumber="99">H. rufus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[536,596,836,862]" pageId="98" pageNumber="99" rank="species">sp. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[654,808,835,861]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[654,808,835,861]" italics="true" pageId="98" pageNumber="99">H. albicollaris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[815,876,836,862]" pageId="98" pageNumber="99" rank="species">sp. n.</taxonomicNameLabel>
|
||
in Ivorona).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99" type="description">
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
<emphasis bold="true" box="[185,335,870,896]" pageId="98" pageNumber="99">
|
||
<emphasis box="[185,330,870,896]" inLineHeading="true" pageId="98" pageNumber="99" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[343,619,871,897]" italics="true" pageId="98" pageNumber="99">Size of males and females</emphasis>
|
||
: species-specific, species with 42–51 rings, no apodous ring.
|
||
</paragraph>
|
||
<paragraph blockId="98.[140,1108,166,1672]" box="[185,1108,941,967]" pageId="98" pageNumber="99">
|
||
<emphasis box="[185,260,941,967]" italics="true" pageId="98" pageNumber="99">Colour</emphasis>
|
||
highly species-specific, but posterior margin of body rings always of lighter.
|
||
</paragraph>
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
<emphasis box="[185,242,976,1002]" italics="true" pageId="98" pageNumber="99">Head</emphasis>
|
||
: each eye with circa 32–48 ocelli arranged in seven vertical rows (
|
||
<figureCitation box="[998,1103,976,1003]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55A</figureCitation>
|
||
,
|
||
<figureCitation box="[140,189,1011,1038]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="98" pageNumber="99">56A</figureCitation>
|
||
,
|
||
<figureCitation box="[200,252,1011,1038]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="98" pageNumber="99">57D</figureCitation>
|
||
,
|
||
<figureCitation box="[264,315,1011,1038]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="98" pageNumber="99">58A</figureCitation>
|
||
). Labrum with standard three irregular teeth and one row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side (
|
||
<figureCitation box="[883,1025,1046,1073]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="98" pageNumber="99">Figs 57B, D</figureCitation>
|
||
). Head around incisura lateralis projecting slightly above basal joints of mandible (
|
||
<figureCitation box="[993,1103,1082,1109]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55D</figureCitation>
|
||
,
|
||
<figureCitation box="[140,192,1117,1144]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="98" pageNumber="99">57C</figureCitation>
|
||
). Head projection and basal mandible joint forming antennal cavity. Antennae short, protruding back to body ring 2 (
|
||
<figureCitation box="[496,597,1152,1179]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55A</figureCitation>
|
||
,
|
||
<figureCitation box="[609,658,1152,1179]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="98" pageNumber="99">56A</figureCitation>
|
||
,
|
||
<figureCitation box="[670,720,1152,1179]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="98" pageNumber="99">57A</figureCitation>
|
||
). Relative lengths of antennomeres: 1<<2>3=4=5<6 (
|
||
<figureCitation box="[328,432,1187,1214]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55A</figureCitation>
|
||
,
|
||
<figureCitation box="[446,496,1187,1214]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">61A</figureCitation>
|
||
). Terminal antennomere with four large sensory cones located together inside a membranous area (
|
||
<figureCitation box="[643,746,1223,1250]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61A</figureCitation>
|
||
). Antennomere 5 latero-apically with four rows (
|
||
<figureCitation box="[312,408,1258,1285]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61B</figureCitation>
|
||
), antennomere 6 with two rows (
|
||
<figureCitation box="[766,863,1258,1285]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61C</figureCitation>
|
||
) of sensilla basiconica.
|
||
</paragraph>
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
<emphasis box="[185,373,1293,1319]" italics="true" pageId="98" pageNumber="99">Gnathochilarium</emphasis>
|
||
lamellae linguales each with two standard setae located behind one another (
|
||
<figureCitation box="[291,393,1328,1355]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Fig. 55E</figureCitation>
|
||
). Stipites each with three apical setae. Mentum not subdivided, basally with several sclerotized ridges (
|
||
<figureCitation box="[561,693,1363,1390]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Figs 55E, F</figureCitation>
|
||
). Stipites towards mentum with circa three weakly-developed, sclerotized ledges (
|
||
<figureCitation box="[640,750,1398,1425]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="98" pageNumber="99">Fig. 55G</figureCitation>
|
||
). Hypopharyngeal crest with a field of spine-like structures (
|
||
<figureCitation box="[468,574,1434,1461]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61D</figureCitation>
|
||
). Central pads of endochilarium separated into two areas, apically with circa eight, basally with>20 sensory cones (
|
||
<figureCitation box="[901,1008,1469,1496]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="98.[140,1108,166,1672]" pageId="98" pageNumber="99">
|
||
<emphasis box="[185,291,1504,1530]" italics="true" pageId="98" pageNumber="99">Mandible</emphasis>
|
||
: external tooth simple, rounded; mesal tooth with three well-developed cusps and in some species with an additional fourth, shallow cusp (
|
||
<figureCitation box="[923,1027,1539,1566]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61E</figureCitation>
|
||
). Basal most cusp extraordinarily large and sharp-edged (
|
||
<figureCitation box="[722,828,1575,1602]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61E</figureCitation>
|
||
). Four or five pectinate lamellae. Molar plate towards pectinate lamellae with a large furrow and circa eight smaller posterior furrows (
|
||
<figureCitation box="[436,538,1645,1672]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="98" pageNumber="99">Fig. 61E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,274,167,193]" italics="true" pageId="99" pageNumber="100">Collum:</emphasis>
|
||
smooth, ventrally not protruding as far as ring 2 (
|
||
<figureCitation box="[826,927,166,193]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">Fig. 57B</figureCitation>
|
||
). Collum covering basal mandibular joint anteriorly (
|
||
<figureCitation box="[570,672,202,229]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55A</figureCitation>
|
||
), as well as part of antenna (
|
||
<figureCitation box="[992,1095,202,229]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">Fig. 56A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,306,237,263]" italics="true" pageId="99" pageNumber="100">Body rings:</emphasis>
|
||
dorsally and laterally with micropunctation, meso- and metazona ventrally with numerous transverse impressions. Ozopores starting at ring 6, touching suture between mesozona and metazona (
|
||
<figureCitation box="[608,712,307,334]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,251,343,369]" italics="true" pageId="99" pageNumber="100">Telson</emphasis>
|
||
: preanal process protruding, well-rounded, slightly curved ventrally (
|
||
<figureCitation captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55C</figureCitation>
|
||
,
|
||
<figureCitation box="[205,258,378,405]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56C</figureCitation>
|
||
,
|
||
<figureCitation box="[271,320,378,405]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">57E</figureCitation>
|
||
,
|
||
<figureCitation box="[333,386,378,405]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="99" pageNumber="100">58C</figureCitation>
|
||
). Anal valves short, smooth, with neither grooves, nor lips, nor micropunctation (
|
||
<figureCitation box="[346,450,413,440]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55C</figureCitation>
|
||
). Subanal scale inconspicuous (
|
||
<figureCitation box="[804,909,413,440]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,228,449,474]" italics="true" pageId="99" pageNumber="100">Legs</emphasis>
|
||
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs in males 0.6–0.7 (
|
||
<figureCitation box="[241,345,518,545]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55H</figureCitation>
|
||
), in females 0.4–0.5 times body diameter (
|
||
<figureCitation box="[822,913,518,545]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55I</figureCitation>
|
||
). Each podomere with a long, apical, ventral seta. Coxae 3 and beyond of rectangular shape (
|
||
<figureCitation box="[941,1044,554,581]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55H</figureCitation>
|
||
). Tarsus with a tiny dorso-apical seta and a pair of extraordinary long ventral setae (
|
||
<figureCitation box="[967,1093,589,616]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55H, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,425,624,650]" italics="true" pageId="99" pageNumber="100">Male sexual characters</emphasis>
|
||
: tarsal pads present only on legs 3–7 (
|
||
<figureCitation box="[851,954,624,651]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55B</figureCitation>
|
||
,
|
||
<figureCitation box="[966,1018,624,651]" captionStart="Figure 60" captionStartId="111.[140,212,844,866]" captionTargetBox="[140,1108,165,819]" captionTargetId="figure@111.[140,1108,438,824]" captionTargetPageId="111" captionText="Figure 60. Hylekobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767358" httpUri="https://zenodo.org/record/3767358/files/figure.png" pageId="99" pageNumber="100">60A</figureCitation>
|
||
). Coxae 3–7 without processes, but coxa 7 twice as wide as coxa 6 (
|
||
<figureCitation box="[803,907,659,686]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55B</figureCitation>
|
||
,
|
||
<figureCitation box="[919,969,659,686]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56B</figureCitation>
|
||
,
|
||
<figureCitation box="[981,1033,659,686]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">57B</figureCitation>
|
||
). Ring 7 large, in ventral view twice as wide as normal rings (
|
||
<figureCitation box="[747,851,694,721]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55B</figureCitation>
|
||
,
|
||
<figureCitation box="[863,915,694,721]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">57B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,369,730,756]" italics="true" pageId="99" pageNumber="100">Anterior gonopods</emphasis>
|
||
: prominently elongated, tips projecting from gonopod pouch (
|
||
<figureCitation captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55B</figureCitation>
|
||
,
|
||
<figureCitation box="[197,245,765,792]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">57B</figureCitation>
|
||
). Anterior gonopods basally with a small sternal apodeme (
|
||
<figureCitation box="[856,948,765,792]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55L</figureCitation>
|
||
), coxite and telopodite apodeme absent. Sternite always well-developed, but hyaline (sheer), of species-specific shape, either triangular (
|
||
<figureCitation box="[446,539,835,862]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55L</figureCitation>
|
||
), rectangular (
|
||
<figureCitation box="[690,790,835,862]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">Fig. 56D</figureCitation>
|
||
), or well-rounded (
|
||
<figureCitation box="[994,1095,835,862]" captionStart="Figure 60" captionStartId="111.[140,212,844,866]" captionTargetBox="[140,1108,165,819]" captionTargetId="figure@111.[140,1108,438,824]" captionTargetPageId="111" captionText="Figure 60. Hylekobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767358" httpUri="https://zenodo.org/record/3767358/files/figure.png" pageId="99" pageNumber="100">Fig. 60D</figureCitation>
|
||
). Coxites wide, in some species mesally with a wide, well-rounded process (
|
||
<figureCitation box="[938,1037,870,897]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">Figs 57E</figureCitation>
|
||
,
|
||
<figureCitation box="[1050,1103,870,897]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="99" pageNumber="100">58D</figureCitation>
|
||
,
|
||
<figureCitation box="[140,193,906,933]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="99" pageNumber="100">59D</figureCitation>
|
||
). Telopodites large, elongated (
|
||
<figureCitation box="[514,614,906,933]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55N</figureCitation>
|
||
), mesally protruding into a wide, well-rounded process (
|
||
<figureCitation box="[230,334,941,968]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55N</figureCitation>
|
||
,
|
||
<figureCitation box="[346,393,941,968]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56F</figureCitation>
|
||
). Mesal margins of process often large and widely protruding (
|
||
<figureCitation captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">Figs 56F</figureCitation>
|
||
,
|
||
<figureCitation box="[192,238,976,1003]" captionStart="Figure 58" captionStartId="108.[140,212,1526,1548]" captionTargetBox="[140,1108,793,1496]" captionTargetId="figure@108.[140,1108,1082,1507]" captionTargetPageId="108" captionText="Figure 58. Hylekobolus albicollaris sp. n., male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="99" pageNumber="100">58F</figureCitation>
|
||
). Margins basally of process in some species additionally protruding (
|
||
<figureCitation box="[967,1066,976,1003]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">Fig. 57G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,382,1011,1037]" italics="true" pageId="99" pageNumber="100">Posterior gonopods</emphasis>
|
||
resembling long neck and small head of dinosaurs belonging to the Sauropoda group (
|
||
<figureCitation box="[399,554,1046,1073]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55M, O</figureCitation>
|
||
). Coxite and telopodite fused, a suture partially visible (
|
||
<figureCitation box="[227,337,1082,1109]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55O</figureCitation>
|
||
). Sternite absent (
|
||
<figureCitation box="[543,654,1082,1109]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55O</figureCitation>
|
||
,
|
||
<figureCitation box="[667,722,1082,1109]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56G</figureCitation>
|
||
), posterior gonopods nevertheless connected via membranes. Coxite conspicuously elongated (
|
||
<figureCitation box="[822,932,1117,1144]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55O</figureCitation>
|
||
,
|
||
<figureCitation box="[945,999,1117,1144]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56G</figureCitation>
|
||
,
|
||
<figureCitation box="[1011,1093,1117,1144]" captionStart="Figure 57" captionStartId="106.[140,209,1526,1548]" captionTargetBox="[140,1108,399,1489]" captionTargetId="figure@106.[140,1108,399,1063]" captionTargetPageId="106" captionText="Figure 57. Hylekobolus griseus sp. n., A–E male paratype, F–I male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C antennal groove D legs 1–7, ventral view E telson F anterior gonopod, anterior view G left posterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; IL = incisura lateralis; Md = mandible; Pre = preanal ring; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767352" httpUri="https://zenodo.org/record/3767352/files/figure.png" pageId="99" pageNumber="100">57F, H</figureCitation>
|
||
), basally at mesal margin with a triangular sclerite, median part of sclerite membranous (
|
||
<figureCitation box="[147,256,1187,1214]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55O</figureCitation>
|
||
,
|
||
<figureCitation box="[269,324,1187,1214]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56G</figureCitation>
|
||
). Coxite mesally with a single spermatic groove. Telopodites arranged face-to-face with one another, tips bent mesally by 60° (
|
||
<figureCitation box="[767,872,1223,1250]" captionStart="Figure 64" captionStartId="115.[141,213,1590,1612]" captionTargetBox="[151,1099,1082,1569]" captionTargetId="figure@115.[140,1108,1075,1573]" captionTargetPageId="115" captionText="Figure 64. Hylekobolus andasibensis sp. n., male holotype. A anterior gonopod, anterior view B right posterior gonopod, anterior view C anterior gonopod, posterior view D right posterior gonopod, posterior view. Apo = apodeme; Cx = coxite; St = sternite; T = telopodite. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767364" httpUri="https://zenodo.org/record/3767364/files/figure.png" pageId="99" pageNumber="100">Fig. 64D</figureCitation>
|
||
) or (more often) 90° (
|
||
<figureCitation box="[148,255,1258,1285]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55O</figureCitation>
|
||
). Apically at lateral margin often with a well-rounded short projection (
|
||
<emphasis box="[1058,1077,1259,1284]" italics="true" pageId="99" pageNumber="100">w</emphasis>
|
||
in
|
||
<figureCitation box="[140,252,1293,1320]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55M</figureCitation>
|
||
,
|
||
<figureCitation box="[266,317,1293,1320]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56E</figureCitation>
|
||
). Sperm canal apically protruding above telopodite margin (
|
||
<emphasis box="[1010,1022,1294,1319]" italics="true" pageId="99" pageNumber="100">z</emphasis>
|
||
in
|
||
<figureCitation captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55M</figureCitation>
|
||
,
|
||
<figureCitation box="[212,263,1328,1355]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56E</figureCitation>
|
||
). Telopodite often with an apical process protruding as continuance of tip (
|
||
<emphasis box="[149,161,1364,1389]" italics="true" pageId="99" pageNumber="100">x</emphasis>
|
||
in
|
||
<figureCitation box="[199,309,1363,1390]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55M</figureCitation>
|
||
,
|
||
<figureCitation box="[321,372,1363,1390]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56E</figureCitation>
|
||
). A single membranous lobe (
|
||
<emphasis box="[708,719,1364,1389]" italics="true" pageId="99" pageNumber="100">y</emphasis>
|
||
) often present between (
|
||
<emphasis box="[999,1011,1364,1389]" italics="true" pageId="99" pageNumber="100">x</emphasis>
|
||
) and (
|
||
<emphasis box="[1086,1098,1364,1389]" italics="true" pageId="99" pageNumber="100">z</emphasis>
|
||
) (
|
||
<figureCitation box="[147,260,1398,1425]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55M</figureCitation>
|
||
,
|
||
<figureCitation box="[273,323,1398,1425]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56E</figureCitation>
|
||
), lobe in one species greatly enlarged (
|
||
<figureCitation box="[756,860,1399,1426]" captionStart="Figure 66" captionStartId="117.[140,212,1590,1612]" captionTargetBox="[147,1108,1182,1571]" captionTargetId="figure@117.[140,1108,1175,1571]" captionTargetPageId="117" captionText="Figure 66. Hylekobolus anjanaharibe sp. n., male holotype. A anterior gonopod, anterior view B right posterior gonopod, anterior view C anterior gonopod, posterior view D right posterior gonopod, posterior view. Apo = apodeme; Cx = coxite; St = sternite; T = telopodite. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767368" httpUri="https://zenodo.org/record/3767368/files/figure.png" pageId="99" pageNumber="100">Fig. 66B</figureCitation>
|
||
). Apodeme very long, oriented in 90° angle towards gonopod (
|
||
<figureCitation box="[595,704,1434,1461]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Figs 55O</figureCitation>
|
||
,
|
||
<figureCitation box="[716,770,1434,1461]" captionStart="Figure 56" captionStartId="104.[140,212,1526,1548]" captionTargetBox="[140,1108,827,1496]" captionTargetId="figure@104.[140,1108,1090,1507]" captionTargetPageId="104" captionText="Figure 56. Hylekobolus rufus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view.Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledges; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767350" httpUri="https://zenodo.org/record/3767350/files/figure.png" pageId="99" pageNumber="100">56G</figureCitation>
|
||
,
|
||
<figureCitation box="[782,837,1434,1461]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="99" pageNumber="100">59G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis box="[185,456,1469,1495]" italics="true" pageId="99" pageNumber="100">Female sexual characters:</emphasis>
|
||
vulva simple, bivalve-like (
|
||
<figureCitation box="[764,859,1469,1496]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55J</figureCitation>
|
||
), with a small, poorly sclerotized operculum at base (
|
||
<figureCitation box="[491,597,1504,1531]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="99" pageNumber="100">Fig. 55K</figureCitation>
|
||
). Posterior valve apically slightly overlapping anterior one (
|
||
<figureCitation box="[294,395,1539,1566]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="99" pageNumber="100">Fig. 61F</figureCitation>
|
||
). Both valves smooth, sensory cones absent. Two or three rows of setae basally on each valve (
|
||
<figureCitation box="[479,580,1575,1602]" captionStart="Figure 61" captionStartId="112.[140,212,1190,1212]" captionTargetBox="[140,1108,165,1171]" captionTargetId="figure@112.[140,1108,165,1171]" captionTargetPageId="112" captionText="Figure 61. Hylekobolus montanus female paratype, SEM. A antenna, overview B antennomere 5 with sensilla basiconica C antennomere 6 with sensilla basiconica D gnathochilarium, part of endochilarium E left mandible, mesal view F left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="99" pageNumber="100">Fig. 61F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="99" pageNumber="100" type="etymology">
|
||
<paragraph blockId="99.[140,1108,166,1672]" pageId="99" pageNumber="100">
|
||
<emphasis bold="true" box="[185,311,1610,1636]" inLineHeading="true" pageId="99" pageNumber="100" reason="1">Etymology</emphasis>
|
||
:
|
||
<taxonomicName authority=", Latin" authorityName="Latin" box="[325,524,1610,1636]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[325,450,1610,1636]" italics="true" pageId="99" pageNumber="100">Hylekobolus</emphasis>
|
||
, Latin
|
||
</taxonomicName>
|
||
, masculine, consisting of “hyleko”, short form of
|
||
<emphasis italics="true" pageId="99" pageNumber="100">hylekoites</emphasis>
|
||
Greek, m., dweller in wood, which refers to the ecology of this species, and -
|
||
<emphasis box="[1032,1085,1645,1671]" italics="true" pageId="99" pageNumber="100">bolus</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |