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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7855FF92FF013D72ACC0EB7B" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Alluviobolus tsimelahy Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="EA739EC9-0459-4440-891A-6D703BDE2FCC" docUuidSource="ZooBank" docVersion="11" lastPageNumber="85" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="83" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791447" ID-GBIF-Taxon="163747620" ID-Zenodo-Dep="3791447" LSID="urn:lsid:zoobank.org:act:EA739EC9-0459-4440-891A-6D703BDE2FCC" httpUri="http://treatment.plazi.org/id/1772122E7855FF92FF013D72ACC0EB7B" lastPageId="84" lastPageNumber="85" pageId="82" pageNumber="83">
<subSubSection box="[140,593,518,545]" pageId="82" pageNumber="83" type="nomenclature">
<paragraph blockId="82.[140,981,518,581]" box="[140,593,518,545]" pageId="82" pageNumber="83">
<heading box="[140,593,518,545]" fontSize="11" level="2" pageId="82" pageNumber="83" reason="3">
<emphasis bold="true" box="[140,593,518,545]" pageId="82" pageNumber="83">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,516,518,545]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,408,518,545]" italics="true" pageId="82" pageNumber="83">Alluviobolus tsimelahy</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[528,593,519,545]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,981,554,581]" pageId="82" pageNumber="83" type="description">
<paragraph blockId="82.[140,981,518,581]" box="[140,981,554,581]" pageId="82" pageNumber="83">
<uri box="[140,981,554,581]" pageId="82" pageNumber="83">
urn:lsid:zoobank.org:act:
<uuid box="[418,981,554,581]" pageId="82" pageNumber="83">EA739EC9-0459-4440-891A-6D703BDE2FCC</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="82" pageNumber="83" type="materials_examined">
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis bold="true" box="[140,375,624,650]" pageId="82" pageNumber="83">
<emphasis box="[140,241,624,650]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Material</emphasis>
examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608818278" collectingDate="2007-05-24" collectionCode="FMMC" collectorName="T. Wesener" country="Madagascar" county="Province" latitude="-24.954933" location="Tsimelahy" longLatPrecision="1" longitude="46.62023" municipality="Andohahela" pageId="82" pageNumber="83" specimenCode="W017" specimenCount="68" specimenCount-female="21" stateProvince="Toliara" typeStatus="holotype">
14 ♂,
<specimenCount box="[466,528,624,651]" pageId="82" pageNumber="83" type="female">21 ♀</specimenCount>
,
<specimenCount box="[546,648,624,650]" pageId="82" pageNumber="83" type="generic">47 imm.</specimenCount>
<emphasis box="[659,762,624,650]" italics="true" pageId="82" pageNumber="83">
<typeStatus box="[659,756,624,650]" pageId="82" pageNumber="83">Holotype</typeStatus>
:
</emphasis>
1 ♂ (
<quantity box="[840,928,624,651]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="82" pageNumber="83" unit="mm" value="42.0">42 mm</quantity>
long),
<collectionCode box="[1018,1108,624,650]" pageId="82" pageNumber="83">FMMC</collectionCode>
<specimenCode box="[140,216,660,686]" collectionCode="FMMC" pageId="82" pageNumber="83">W017</specimenCode>
D,
<collectingCountry box="[249,383,659,685]" name="Madagascar" pageId="82" pageNumber="83">Madagascar</collectingCountry>
,
<collectingCounty box="[395,493,659,685]" pageId="82" pageNumber="83">Province</collectingCounty>
<collectingRegion box="[498,580,659,685]" country="Madagascar" name="Toliara" pageId="82" pageNumber="83">Toliara</collectingRegion>
, RNI
<collectingMunicipality box="[649,788,659,685]" pageId="82" pageNumber="83">Andohahela</collectingMunicipality>
, parcel II,
<location LSID="urn:lsid:plazi:treatment:1772122E7855FF92FF013D72ACC0EB7B:9A04F5E37855FF94FC073DE7ACA7E8FC" box="[906,1022,659,685]" country="Madagascar" county="Province" latitude="-24.954933" longLatPrecision="1" longitude="46.62023" municipality="Andohahela" name="Tsimelahy" pageId="82" pageNumber="83" stateProvince="Toliara">Tsimelahy</location>
, gallery forest, in alluvial debris,
<geoCoordinate box="[414,564,694,721]" degrees="24" direction="south" minutes="57.296" orientation="latitude" pageId="82" pageNumber="83" precision="1" value="-24.954933">24°57.296 S</geoCoordinate>
,
<geoCoordinate box="[575,727,695,721]" degrees="46" direction="east" minutes="37.214" orientation="longitude" pageId="82" pageNumber="83" precision="1" value="46.62023">46°37.214 E</geoCoordinate>
, leg.
<collectorName box="[783,907,695,721]" pageId="82" pageNumber="83">T. Wesener</collectorName>
et al.,
<date box="[981,1104,695,721]" pageId="82" pageNumber="83" value="2007-05-24">
<collectingDate box="[981,1104,695,721]" pageId="82" pageNumber="83" value="2007-05-24">24.V.2007</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2608818201" box="[140,953,730,756]" collectingDate="2007-05-24" collectionCode="FMMC" collectorName="T. Wesener" country="Madagascar" county="Province" latitude="-24.954933" location="Tsimelahy" longLatPrecision="1" longitude="46.62023" municipality="Andohahela" pageId="82" pageNumber="83" specimenCode="W017" specimenCount="12" specimenCount-female="6" stateProvince="Toliara" typeStatus="paratype">
<emphasis box="[140,247,731,756]" italics="true" pageId="82" pageNumber="83">
<typeStatus box="[140,242,731,756]" pageId="82" pageNumber="83">Paratypes</typeStatus>
:
</emphasis>
4 ♂,
<specimenCount box="[320,365,730,756]" pageId="82" pageNumber="83" type="female">6 ♀</specimenCount>
,
<specimenCount box="[382,471,730,756]" pageId="82" pageNumber="83" type="generic">6 imm.</specimenCount>
,
<collectionCode box="[485,575,730,756]" pageId="82" pageNumber="83">FMMC</collectionCode>
<specimenCode box="[584,660,730,756]" collectionCode="FMMC" pageId="82" pageNumber="83">W017</specimenCode>
D, same data as holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818285" collectingDate="2007-05-24" collectionCode="CAS" collectorName="T. Wesener" country="Madagascar" county="Province" latitude="-24.954933" location="Tsimelahy" longLatPrecision="1" longitude="46.62023" municipality="Andohahela" pageId="82" pageNumber="83" specimenCode="W017" specimenCount="3" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
1 ♂,
<specimenCount box="[1030,1075,730,756]" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
,
<specimenCount pageId="82" pageNumber="83" type="generic">2 imm.</specimenCount>
,
<collectionCode box="[218,271,765,791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="82" pageNumber="83" type="Museum">CAS</collectionCode>
<specimenCode box="[280,356,765,791]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="82" pageNumber="83" type="Museum">W017</specimenCode>
D, same data as holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818310" box="[661,979,765,792]" collectionCode="ZMUC" country="Madagascar" pageId="82" pageNumber="83" specimenCode="W017" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
1 ♂,
<specimenCount box="[722,768,765,792]" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
,
<collectionCode box="[784,872,765,791]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="82" pageNumber="83">ZMUC</collectionCode>
<specimenCode box="[881,957,765,791]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="82" pageNumber="83">W017</specimenCode>
D
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818237" collectingDate="2007-05-24" collectionCode="ZMH" collectorName="T. Wesener" country="Madagascar" county="Province" latitude="-24.954933" location="Tsimelahy" longLatPrecision="1" longitude="46.62023" municipality="Andohahela" pageId="82" pageNumber="83" specimenCode="W017" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
1 ♂,
<specimenCount box="[1055,1100,765,792]" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
,
<collectionCode box="[140,209,801,827]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="82" pageNumber="83" type="Museum">ZMH</collectionCode>
<specimenCode box="[216,292,800,826]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="82" pageNumber="83" type="Museum">W017</specimenCode>
D, same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<materialsCitation ID-GBIF-Occurrence="2608818256" location="University" pageId="82" pageNumber="83" specimenCount="14" specimenCount-female="4" stateProvince="Antananarivo">
<emphasis bold="true" box="[185,488,835,861]" pageId="82" pageNumber="83">
<typeStatus box="[185,483,835,861]" isOtherTypeStatus="true" pageId="82" pageNumber="83">
<emphasis box="[185,257,835,861]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Other</emphasis>
material examined
</typeStatus>
:
</emphasis>
4 ♂,
<specimenCount box="[546,587,836,862]" pageId="82" pageNumber="83" type="female">4 ♀</specimenCount>
,
<specimenCount box="[600,698,835,862]" pageId="82" pageNumber="83" type="generic">10 imm.</specimenCount>
, W017D, same data as type material, but stored as vouchers at
<location LSID="urn:lsid:plazi:treatment:1772122E7855FF92FF013D72ACC0EB7B:9A04F5E37855FF94FE2E3C13AD4EE9D0" box="[419,535,871,897]" name="University" pageId="82" pageNumber="83" stateProvince="Antananarivo">University</location>
<collectingRegion box="[542,698,871,897]" country="Madagascar" name="Antananarivo" pageId="82" pageNumber="83">Antananarivo</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818347" collectingDate="2007-06-15" collectionCode="FMMC" collectorName="Ebosika &amp; T. Wesener" latitude="-24.9459" location="Andohahela mountain" longLatPrecision="1" longitude="46.675514" pageId="82" pageNumber="83" specimenCode="W016" specimenCount="13" specimenCount-female="4" stateProvince="Antananarivo">
1 ♂,
<specimenCount box="[764,806,871,897]" pageId="82" pageNumber="83" type="female">4 ♀</specimenCount>
,
<specimenCount box="[819,904,871,897]" pageId="82" pageNumber="83" type="generic">9 imm.</specimenCount>
,
<collectionCode box="[914,1004,871,897]" pageId="82" pageNumber="83">FMMC</collectionCode>
<specimenCode box="[1010,1082,871,897]" collectionCode="FMMC" pageId="82" pageNumber="83">W016</specimenCode>
A,
<location LSID="urn:lsid:plazi:treatment:1772122E7855FF92FF013D72ACC0EB7B:9A04F5E37855FF94FF013CFEAED6E9F5" box="[140,399,906,932]" latitude="-24.9459" longLatPrecision="1" longitude="46.675514" name="Andohahela mountain" pageId="82" pageNumber="83" stateProvince="Antananarivo">Andohahela mountain</location>
chain,
<collectorName box="[490,582,906,932]" pageId="82" pageNumber="83">Ebosika</collectorName>
, rainforest,
<geoCoordinate box="[723,903,906,933]" degrees="24" direction="south" minutes="56" orientation="latitude" pageId="82" pageNumber="83" precision="1" seconds="45.24" value="-24.9459">24°5645.24” S</geoCoordinate>
,
<geoCoordinate box="[919,1103,906,932]" degrees="46" direction="east" minutes="40" orientation="longitude" pageId="82" pageNumber="83" precision="1" seconds="31.84" value="46.675514">46°4031.84” E</geoCoordinate>
, leg.
<collectorName box="[186,311,942,968]" pageId="82" pageNumber="83">T. Wesener</collectorName>
et al.,
<date box="[389,524,941,968]" pageId="82" pageNumber="83" value="2007-06-15">
<collectingDate box="[389,524,941,968]" pageId="82" pageNumber="83" value="2007-06-15">15.VI.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818295" collectingDate="2007-06-15" collectionCode="FMMC" collectorName="Ebosika &amp; T. Wesener" latitude="-24.9459" location="Andohahela mountain" longLatPrecision="1" longitude="46.675514" pageId="82" pageNumber="83" specimenCount="13" specimenCount-female="4" stateProvince="Antananarivo">
1 ♂,
<specimenCount box="[595,638,941,968]" pageId="82" pageNumber="83" type="female">4 ♀</specimenCount>
,
<specimenCount box="[653,740,941,967]" pageId="82" pageNumber="83" type="generic">9 imm.</specimenCount>
, W016A, same data as previous, University
<collectingRegion box="[264,421,976,1002]" country="Madagascar" name="Antananarivo" pageId="82" pageNumber="83">Antananarivo</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="82" pageNumber="83" type="diagnosis">
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis bold="true" box="[185,454,1011,1037]" pageId="82" pageNumber="83">
<emphasis box="[185,324,1011,1037]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Differential</emphasis>
diagnosis:
</emphasis>
see
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[508,653,1011,1038]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species">
<emphasis box="[508,653,1011,1038]" italics="true" pageId="82" pageNumber="83">A. laticlavius</emphasis>
</taxonomicName>
for a differentiation of both taxa. It is possible that
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[289,478,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[289,478,1047,1073]" italics="true" pageId="82" pageNumber="83">Ostinobolus rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[485,546,1047,1073]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
can also be found in the Ebosika rainforest. Both species live in the same habitat and are of similar size and colour.
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[878,969,1082,1108]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species">
<emphasis box="[878,969,1082,1108]" italics="true" pageId="82" pageNumber="83">O. rufus</emphasis>
</taxonomicName>
is, however, entirely red (
<figureCitation box="[288,392,1117,1143]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, GJ male holotype; C, E, F male paratype. A head, lateral view B legs 17, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="82" pageNumber="83">Fig. 48C</figureCitation>
) while the mesozonites are dark olive greenish in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species">
<emphasis italics="true" pageId="82" pageNumber="83">Alluviobolus tsimelahy</emphasis>
</taxonomicName>
(
<figureCitation box="[256,360,1152,1178]" captionStart="Figure 42" captionStartId="77.[139,211,599,621]" captionTargetBox="[139,1107,165,580]" captionTargetId="figure@77.[139,1108,165,580]" captionTargetPageId="77" captionText="Figure 42. Alluviobolus tsimelahy sp. n., A living male from Tsimelahy B male and female specimen from Ebosika." figureDoi="http://doi.org/10.5281/zenodo.3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="82" pageNumber="83">Fig. 42A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="description">
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis bold="true" box="[185,335,1187,1213]" pageId="82" pageNumber="83">
<emphasis box="[185,330,1187,1213]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[343,494,1188,1213]" italics="true" pageId="82" pageNumber="83">Measurements</emphasis>
: males with 4749 body rings, circa
<quantity box="[915,1042,1188,1214]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="82" pageNumber="83" unit="mm" value="42.5" valueMax="44.0" valueMin="41.0">4144 mm</quantity>
long,
<quantity box="[140,279,1223,1249]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="82" pageNumber="83" unit="mm" value="3.95" valueMax="4.3" valueMin="3.6">3.64.3 mm</quantity>
wide. Females with 4749 body rings,
<quantity box="[710,834,1222,1249]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="6.0" metricValueMin="5.2" pageId="82" pageNumber="83" unit="mm" value="56.0" valueMax="60.0" valueMin="52.0">5260 mm</quantity>
long, 5.56.0 mm wide.
</paragraph>
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis box="[185,300,1258,1284]" italics="true" pageId="82" pageNumber="83">Coloration</emphasis>
on mesozonites dark olive green, metazonites red (
<figureCitation box="[867,970,1258,1284]" captionStart="Figure 42" captionStartId="77.[139,211,599,621]" captionTargetBox="[139,1107,165,580]" captionTargetId="figure@77.[139,1108,165,580]" captionTargetPageId="77" captionText="Figure 42. Alluviobolus tsimelahy sp. n., A living male from Tsimelahy B male and female specimen from Ebosika." figureDoi="http://doi.org/10.5281/zenodo.3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="82" pageNumber="83">Fig. 42A</figureCitation>
). Rings dorsally with a slender orange stripe. Head, legs, antennae and telson red (
<figureCitation box="[953,1037,1293,1320]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Figs 45</figureCitation>
AC).
<emphasis box="[140,183,1329,1354]" italics="true" pageId="82" pageNumber="83">Eyes</emphasis>
with 2224 ocelli arranged in 4 or 5 rows (
<figureCitation box="[687,791,1328,1355]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45A</figureCitation>
).
<emphasis box="[812,915,1329,1355]" italics="true" pageId="82" pageNumber="83">Antennae</emphasis>
protruding back to ring 5 (
<figureCitation box="[254,357,1363,1390]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45A</figureCitation>
).
</paragraph>
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis box="[185,403,1399,1425]" italics="true" pageId="82" pageNumber="83">Male coxal processes</emphasis>
on coxae 35 (
<figureCitation box="[586,690,1398,1425]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45B</figureCitation>
), 3 and 4 large, swollen, process on coxa 5 smaller (
<figureCitation box="[315,418,1434,1461]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, AG, IM male, H female. A head, lateral view B legs 17, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="82" pageNumber="83">Fig. 46B</figureCitation>
).
<emphasis box="[438,598,1434,1460]" italics="true" pageId="82" pageNumber="83">Preanal process</emphasis>
absent (
<figureCitation box="[692,799,1434,1461]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Figs 45C</figureCitation>
,
<figureCitation box="[811,865,1434,1461]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, AG, IM male, H female. A head, lateral view B legs 17, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="82" pageNumber="83">46C</figureCitation>
).
</paragraph>
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis box="[185,376,1469,1495]" italics="true" pageId="82" pageNumber="83">Anterior gonopod</emphasis>
sternite elevated into a wide lobe, apically tapering. Tip wellrounded (
<figureCitation box="[251,352,1504,1531]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45F</figureCitation>
). Coxite and telopodite inconspicuous (
<figureCitation box="[802,911,1504,1531]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45H</figureCitation>
).
</paragraph>
<paragraph blockId="82.[140,1108,624,1672]" pageId="82" pageNumber="83">
<emphasis box="[185,378,1539,1565]" italics="true" pageId="82" pageNumber="83">Posterior gonopods</emphasis>
: telopodite with a membranous fringe (
<figureCitation box="[816,908,1539,1566]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45I</figureCitation>
). Apical sclerite (
<emphasis box="[1095,1107,1540,1565]" italics="true" pageId="82" pageNumber="83">x</emphasis>
in
<figureCitation box="[169,258,1574,1601]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45I</figureCitation>
) slender, triangular, regularly protruding into a slender, acute tip (
<figureCitation box="[990,1095,1574,1601]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="82" pageNumber="83">Fig. 45G</figureCitation>
).
</paragraph>
<paragraph blockId="82.[140,1108,624,1672]" lastBlockId="83.[140,1108,1328,1672]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<emphasis bold="true" box="[185,459,1609,1635]" pageId="82" pageNumber="83">
<emphasis box="[185,333,1609,1635]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Intraspecific</emphasis>
variation:
</emphasis>
specimens from the rainforest of Ebosika show minute differences in the posterior gonopods towards those of the
<typeStatus box="[820,868,1646,1672]" pageId="82" pageNumber="83">type</typeStatus>
series (compare
<figureCitation captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. AC, FI male holotype, D, E female paratype. A head, lateral view B legs 17, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">Figs 45G, I</figureCitation>
with
<figureCitation box="[287,460,1328,1355]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, AG, IM male, H female. A head, lateral view B legs 17, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="83" pageNumber="84">Figs 46J, L, M</figureCitation>
). Ebosika specimens possess apically on the telopodite a short protruding process towards the membranous fringe (
<figureCitation box="[811,912,1363,1389]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, AG, IM male, H female. A head, lateral view B legs 17, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="83" pageNumber="84">Fig. 46L</figureCitation>
, marked with an arrow). However, size, number of body rings, colour pattern and the shape of male coxal processes (
<figureCitation box="[322,405,1434,1461]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, AG, IM male, H female. A head, lateral view B legs 17, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="83" pageNumber="84">Figs 46</figureCitation>
AC) are identical in specimens from both populations.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767334" ID-Zenodo-Dep="3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="83" pageNumber="84" startId="83.[140,212,1014,1036]" targetBox="[140,1108,165,990]" targetPageId="83">
<paragraph blockId="83.[140,1108,1014,1260]" pageId="83" pageNumber="84">
<emphasis bold="true" box="[140,252,1014,1036]" pageId="83" pageNumber="84">Figure 46.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[260,459,1014,1035]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[260,459,1014,1035]" italics="true" pageId="83" pageNumber="84">Alluviobolus tsimelahy</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[466,518,1015,1036]" pageId="83" pageNumber="84" rank="species">sp. n.</taxonomicNameLabel>
specimens from Ebosika, AG, IM male, H female.
<emphasis bold="true" box="[1031,1050,1014,1036]" pageId="83" pageNumber="84">A</emphasis>
head, lateral view
<emphasis bold="true" box="[250,267,1046,1068]" pageId="83" pageNumber="84">B</emphasis>
legs 17, ventral view
<emphasis bold="true" box="[478,497,1046,1068]" pageId="83" pageNumber="84">C</emphasis>
telson
<emphasis bold="true" box="[564,583,1046,1068]" pageId="83" pageNumber="84">D</emphasis>
gnathochilarium, ventral view
<emphasis bold="true" box="[872,887,1046,1068]" pageId="83" pageNumber="84">E</emphasis>
gnathochilarium, mentum
<emphasis bold="true" box="[186,200,1078,1100]" pageId="83" pageNumber="84">F</emphasis>
gnathochilarium, dorsal view on endochilarium
<emphasis bold="true" box="[659,710,1078,1100]" pageId="83" pageNumber="84">G, H</emphasis>
left midbody legs
<emphasis bold="true" box="[886,894,1078,1100]" pageId="83" pageNumber="84">I</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[245,253,1110,1132]" pageId="83" pageNumber="84">Į</emphasis>
right posterior gonopod, anterior view
<emphasis bold="true" box="[620,637,1110,1132]" pageId="83" pageNumber="84">K</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[954,969,1110,1132]" pageId="83" pageNumber="84">L</emphasis>
right posterior gonopod, telopodite, posterior view
<emphasis bold="true" box="[477,498,1142,1164]" pageId="83" pageNumber="84">M</emphasis>
right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="distribution">
<paragraph blockId="83.[140,1108,1328,1672]" lastBlockId="84.[140,1108,167,299]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
<emphasis bold="true" box="[185,483,1469,1495]" pageId="83" pageNumber="84">
<emphasis box="[185,334,1469,1495]" inLineHeading="true" pageId="83" pageNumber="84" reason="1">Distribution</emphasis>
and ecology
</emphasis>
:
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[497,627,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species">
<emphasis box="[497,627,1469,1495]" italics="true" pageId="83" pageNumber="84">A. tsimelahy</emphasis>
</taxonomicName>
is until now the only known Malagasy species of
<taxonomicName authorityName="Cook" authorityYear="1895" box="[217,344,1504,1530]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
which occurs in the spiny forest and rainforest ecosystem, more precisely in the spiny forest at Tsimelahy and just
<quantity box="[645,703,1539,1566]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="83" pageNumber="84" unit="km" value="5.0">5 km</quantity>
east in the rainforest of Ebosika. The record from the spiny forest of Tsimelahy, however, is tainted, because the only place where
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[212,342,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis box="[212,342,1610,1636]" italics="true" pageId="83" pageNumber="84">Alluviobolus</emphasis>
</taxonomicName>
specimens were found was very close to the river, inside wet debris. It is currently unknown if the
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[454,585,1645,1671]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species">
<emphasis box="[454,585,1645,1671]" italics="true" pageId="83" pageNumber="84">A. tsimelahy</emphasis>
</taxonomicName>
specimens from the spiny forest area represent a stable population or whether specimens just floated out of the rainforest and into the spiny forest during the wet season. It is perhaps not a coincident that the specimens from Ebosika were collected close to the same river as those in Tsimelahy (
<figureCitation box="[893,975,237,263]" captionStart="Figure 43" captionStartId="77.[139,211,1099,1121]" captionTargetBox="[139,1111,687,1080]" captionTargetId="figure@77.[139,1108,688,1080]" captionTargetPageId="77" captionText="Figure 43. Distribution map of the genus Alluviobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820825" httpUri="https://zenodo.org/record/3820825/files/figure.png" pageId="84" pageNumber="85">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[185,921,272,299]" pageId="84" pageNumber="85" type="etymology">
<paragraph blockId="84.[140,1108,167,299]" box="[185,921,272,299]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[185,323,272,298]" pageId="84" pageNumber="85">
<emphasis box="[185,318,272,298]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[330,431,272,298]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species">
<emphasis box="[330,431,272,298]" italics="true" pageId="84" pageNumber="85">tsimelahy</emphasis>
</taxonomicName>
, noun in apposition, after the
<typeStatus box="[778,826,273,299]" pageId="84" pageNumber="85">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>