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<document id="9CB91EBFEC62DC238F797BA8F343F927" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFB8731F32FAF973E293FB34" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium trichordum Wesener 2009, new species" docType="treatment" docVersion="4" lastPageNumber="80" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="76" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="5CF708C348F4FDB35404A7EAFD651723" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0B0ACC0155A2EE625BC732B5BD76A0FD">
<mods:title id="70754446B8D4BDC927BFC244D999C549">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="D73629EC35FB173CCB7629DA8DEFEE1B" type="personal">
<mods:role id="710C13200B8FA63424DB35C9F8D9EDBB">
<mods:roleTerm id="8D5976D29DCFFCE98AC31B81C9CCBE0F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3298229EF342053C8DC89108CF8309F7">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="87B3BB1B5167343C7260D980FF6DAC00">text</mods:typeOfResource>
<mods:relatedItem id="F8CA8C6DF921B0DF3E62669815B18A31" type="host">
<mods:titleInfo id="2A3127029BC65EB18F5B090FDA29BAD2">
<mods:title id="9BAB02CFA2DA45A5F2670CA56C19AA65">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5531A982CDE1203A216B0624B39E0F51">
<mods:date id="15D2B676FAD3F1AB5CE58C593D04D3D7">2009</mods:date>
<mods:detail id="446D515F271C65D114B78F2C4F8DC35B" type="pubDate">
<mods:number id="5155A701D83164F9359A822A0655B425">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="4F27495A2423B5D0BC191A308F521B0F" type="volume">
<mods:number id="48634F2083BEC01DA14EF00F79AC72D7">2097</mods:number>
</mods:detail>
<mods:detail id="3E81807E71158309D0C2F45629E7A501" type="issue">
<mods:number id="CEA0DD5996999A049A1742951F92AE79">1</mods:number>
</mods:detail>
<mods:extent id="2F0B4FA1FC03BBA8FC698CCDB5820E72" unit="page">
<mods:start id="98266A712FC3761B3C9809DD6AC22A2B">1</mods:start>
<mods:end id="3286ACF9E4BE0CD372D78C1A8FC5B48A">134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="5346DE7CDD7943B81F342776688354C8">
<mods:url id="26DBBEDF470894115FFC98C771843B9A">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="E9F7758ED766BFBB183172267B151285">journal article</mods:classification>
<mods:identifier id="7743DEB41C03F7782F69EA7ABB19B55D" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="3FFA151166A7D18117D4108FCB337212" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="BCA95CE5B0091D94F2440C6DC64F1F42" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="02B0E2BF98A73C14FB6B0BB75DA83649" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFFB8731F32FAF973E293FB34" ID-DOI="http://doi.org/10.5281/zenodo.5325493" ID-Zenodo-Dep="5325493" LSID="urn:lsid:plazi:treatment:03FFBE1DFFB8731F32FAF973E293FB34" httpUri="http://treatment.plazi.org/id/03FFBE1DFFB8731F32FAF973E293FB34" lastPageId="79" lastPageNumber="80" pageId="75" pageNumber="76">
<subSubSection id="C34C5C80FFB8731B32FAF973E172F93A" box="[151,629,1737,1763]" pageId="75" pageNumber="76" type="nomenclature">
<paragraph id="8BE90F0BFFB8731B32FAF973E172F93A" blockId="75.[151,629,1737,1763]" box="[151,629,1737,1763]" pageId="75" pageNumber="76">
<heading id="D0A1B867FFB8731B32FAF973E172F93A" bold="true" box="[151,629,1737,1763]" fontSize="11" level="1" pageId="75" pageNumber="76" reason="1">
<emphasis id="B922D319FFB8731B32FAF973E172F93A" bold="true" box="[151,629,1737,1763]" pageId="75" pageNumber="76">
<emphasis id="B922D319FFB8731B32FAF973E2D9F93A" bold="true" box="[151,478,1737,1763]" italics="true" pageId="75" pageNumber="76">
<taxonomicName id="4C567488FFB8731B32FAF973E2DFF93A" ID-CoL="5D7JB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,472,1737,1763]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="trichordum" status="sp. nov.">Zoosphaerium trichordum</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2116E62FFB8731B3388F973E172F93A" box="[485,629,1737,1763]" pageId="75" pageNumber="76" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFB8731C32FAF8A8E749FAA7" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="etymology">
<paragraph id="8BE90F0BFFB8731B32FAF8A8E240F8A3" blockId="75.[151,1435,1810,1952]" pageId="75" pageNumber="76">
<emphasis id="B922D319FFB8731B32FAF8A8E285F8F5" bold="true" box="[151,386,1810,1836]" pageId="75" pageNumber="76">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FFB8731B33E6F8A9E109F8F5" authorityName="Wesener" authorityYear="2009" box="[395,526,1811,1836]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="trichordum">
<emphasis id="B922D319FFB8731B33E6F8A9E109F8F5" box="[395,526,1811,1836]" italics="true" pageId="75" pageNumber="76">trichordum</emphasis>
</taxonomicName>
, a composite noun in apposition, consisting of tri- for three, and chordum = chord, referring to the three stridulation ribs on the male harp, a unique feature for members of the genus
<taxonomicName id="4C567488FFB8731B32FAF8DAE247F8A0" authorityName="Pocock" authorityYear="1895" box="[151,320,1888,1913]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B922D319FFB8731B32FAF8DAE247F8A0" box="[151,320,1888,1913]" italics="true" pageId="75" pageNumber="76">Zoosphaerium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE90F0BFFB8731B32ABF83CE116F879" blockId="75.[151,1435,1810,1952]" box="[198,529,1926,1952]" pageId="75" pageNumber="76">
<emphasis id="B922D319FFB8731B32ABF83CE2A7F879" bold="true" box="[198,416,1926,1952]" pageId="75" pageNumber="76">Studied material:</emphasis>
2 M,
<specimenCount id="9D50C482FFB8731B338BF83CE116F879" box="[486,529,1926,1952]" pageId="75" pageNumber="76" type="generic">5 F.</specimenCount>
</paragraph>
<paragraph id="8BE90F0BFFBF731C32FAFB61E749FAA7" blockId="76.[151,1436,1243,1406]" pageId="76" pageNumber="77">
<emphasis id="B922D319FFBF731C32FAFB61E236FB2A" bold="true" box="[151,305,1243,1267]" pageId="76" pageNumber="77">FIGURE 051</emphasis>
.
<taxonomicName id="4C567488FFBF731C332FFB66E15AFB2A" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[322,605,1244,1267]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="trichordum" status="sp. nov.">
<emphasis id="B922D319FFBF731C332FFB66E15AFB2A" box="[322,605,1244,1267]" italics="true" pageId="76" pageNumber="77">Zoosphaerium trichordum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFBF731C300AFB61E1A0FB2A" bold="true" box="[615,679,1243,1267]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="A2116E62FFBF731C300AFB61E1A0FB2A" box="[615,679,1243,1267]" pageId="76" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A female
<typeStatus id="54EDB1A9FFBF731C3145FB66E080FB2A" box="[808,903,1244,1267]" pageId="76" pageNumber="77" type="paratype">paratype</typeStatus>
, B–E
<typeStatus id="54EDB1A9FFBF731C31A2FB66E728FB2A" box="[975,1071,1244,1267]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
,
<emphasis id="B922D319FFBF731C3651FB61E748FB2A" bold="true" box="[1084,1103,1243,1267]" pageId="76" pageNumber="77">A</emphasis>
: second left coxa with vulva, posterior view;
<emphasis id="B922D319FFBF731C3352FB44E256FACF" bold="true" box="[319,337,1278,1302]" pageId="76" pageNumber="77">B</emphasis>
: last two podomeres of anterior telopod, lateral view;
<emphasis id="B922D319FFBF731C31F7FB44E0AAFACF" bold="true" box="[922,941,1278,1302]" pageId="76" pageNumber="77">C</emphasis>
: right half of anterior telopods, anterior view;
<emphasis id="B922D319FFBF731C32FAFA9BE3ADFAE0" bold="true" box="[151,170,1313,1337]" pageId="76" pageNumber="77">D</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFBF731C3049FA9BE131FAE0" bold="true" box="[548,566,1313,1337]" pageId="76" pageNumber="77">E</emphasis>
: chela of posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FFBF731C36ACFA9BE7EFFAE0" bold="true" box="[1217,1256,1313,1337]" pageId="76" pageNumber="77">cav</emphasis>
= cavity;
<emphasis id="B922D319FFBF731C3736FA9BE684FAE0" bold="true" box="[1371,1411,1313,1337]" pageId="76" pageNumber="77">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FFBF731C3320FAF9E26AFA82" bold="true" box="[333,365,1347,1371]" pageId="76" pageNumber="77">Cx</emphasis>
= coxa;
<emphasis id="B922D319FFBF731C33A4FAF9E2ECFA82" bold="true" box="[457,491,1347,1371]" pageId="76" pageNumber="77">EP</emphasis>
= external plate of vulva;
<emphasis id="B922D319FFBF731C316CFAF9E027FA82" bold="true" box="[769,800,1347,1371]" pageId="76" pageNumber="77">IH</emphasis>
= inner horns;
<emphasis id="B922D319FFBF731C31ADFAF9E0DDFA82" bold="true" box="[960,986,1347,1371]" pageId="76" pageNumber="77">IP</emphasis>
= mesal plate of vulva;
<emphasis id="B922D319FFBF731C36B4FAF9E7F0FA82" bold="true" box="[1241,1271,1347,1371]" pageId="76" pageNumber="77">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFBF731C32BEFADCE3EFFAA7" bold="true" box="[211,232,1382,1406]" pageId="76" pageNumber="77">O</emphasis>
= operculum;
<emphasis id="B922D319FFBF731C33E9FADCE2B3FAA7" bold="true" box="[388,436,1382,1406]" pageId="76" pageNumber="77">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FFBF731C30E1FADCE1A9FAA7" bold="true" box="[652,686,1382,1406]" pageId="76" pageNumber="77">SR</emphasis>
= stridulation ribs. Scale bars =
<quantity id="4CAEA2EEFFBF731C3664FADDE74EFAA7" box="[1033,1097,1383,1406]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="76" pageNumber="77" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBF731C32AAFA16E7D3F95F" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="8BE90F0BFFBF731C32AAFA16E27DF9CD" blockId="76.[151,1436,1452,2017]" pageId="76" pageNumber="77">
<materialsCitation id="3B3E0556FFBF731C32AAFA16E27DF9CD" collectingDate="2003-12-10" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" county="Locus" elevation="175" latitude="-12.969999" location="Foret d'Ampondrabe" longLatPrecision="21" longitude="49.7" municipality="Province" pageId="76" pageNumber="77" specimenCode="BLF 9973" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="B922D319FFBF731C32AAFA16E245FA1F" bold="true" box="[199,322,1452,1478]" pageId="76" pageNumber="77">
<typeStatus id="54EDB1A9FFBF731C32AAFA16E23AFA1F" box="[199,317,1452,1478]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
:
</emphasis>
1 M (
<quantity id="4CAEA2EEFFBF731C33E1FA16E2E6FA1F" box="[396,481,1452,1478]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="76" pageNumber="77" unit="mm" value="36.0">36 mm</quantity>
long),
<emphasis id="B922D319FFBF731C305BFA16E1F3FA1F" bold="true" box="[566,756,1452,1478]" pageId="76" pageNumber="77">
<collectionCode id="ED4797CEFFBF731C305BFA16E177FA1F" box="[566,624,1452,1478]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="76" pageNumber="77" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFBF731C3015FA16E1F3FA1F" box="[632,756,1452,1478]" collectionCode="BLF" pageId="76" pageNumber="77">BLF 9973</specimenCode>
</emphasis>
,
<emphasis id="B922D319FFBF731C3169FA16E0A2FA1C" box="[772,933,1452,1477]" italics="true" pageId="76" pageNumber="77">
<collectingCounty id="62887787FFBF731C3169FA16E04DFA1C" box="[772,842,1452,1477]" pageId="76" pageNumber="77">Locus</collectingCounty>
typicus
</emphasis>
:
<collectingCountry id="F3414F9BFFBF731C31D9FA16E744FA1F" box="[948,1091,1452,1478]" name="Madagascar" pageId="76" pageNumber="77">Madagascar</collectingCountry>
,
<collectingMunicipality id="6B8D9571FFBF731C363DFA16E7BFFA1F" box="[1104,1208,1452,1478]" pageId="76" pageNumber="77">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FFBF731C36ACFA16E656FA1F" box="[1217,1361,1452,1478]" country="Madagascar" name="Antsiranana" pageId="76" pageNumber="77">Antsiranana</collectingRegion>
,
<location id="8E8959D0FFBF731C3733FA16E240FA34" LSID="urn:lsid:plazi:treatment:03FFBE1DFFB8731F32FAF973E293FB34:8E8959D0FFBF731C3733FA16E240FA34" country="Madagascar" county="Locus" latitude="-12.969999" longLatPrecision="21" longitude="49.7" municipality="Province" name="Foret d'Ampondrabe" pageId="76" pageNumber="77" stateProvince="Antsiranana">Forêt d'Ampondrabe</location>
,
<locationDeviation id="9A5B1629FFBF731C3339FA69E190FA34" box="[340,663,1491,1517]" location="Daraina" pageId="76" pageNumber="77">
<quantity id="4CAEA2EEFFBF731C3339FA69E2B1FA34" box="[340,438,1491,1517]" metricMagnitude="4" metricUnit="m" metricValue="2.63" pageId="76" pageNumber="77" unit="km" value="26.3">26.3 km</quantity>
10° NNE Daraina
</locationDeviation>
,
<geoCoordinate id="EE6269CCFFBF731C30C9FA69E02CFA34" box="[676,811,1491,1517]" degrees="12" direction="south" minutes="58" orientation="latitude" pageId="76" pageNumber="77" precision="15" seconds="12" value="-12.969999">12°58'12&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFBF731C3157FA69E0D6FA34" box="[826,977,1491,1517]" degrees="049" direction="east" minutes="42" orientation="longitude" pageId="76" pageNumber="77" precision="15" seconds="00" value="49.7">049°42'00&quot;E</geoCoordinate>
,
<quantity id="4CAEA2EEFFBF731C318DFA69E729FA34" box="[992,1070,1491,1517]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="76" pageNumber="77" unit="m" value="175.0">
<elevation id="007BE838FFBF731C318DFA69E729FA34" box="[992,1070,1491,1517]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="76" pageNumber="77" unit="m" value="175.0">175 m</elevation>
</quantity>
, tropical dry forest, coll.
<collectorName id="26A36ADDFFBF731C3733FA69E3E5F9CD" pageId="76" pageNumber="77">B. L. Fisher</collectorName>
,
<date id="FFE829CBFFBF731C3280FA40E271F9CD" box="[237,374,1530,1556]" pageId="76" pageNumber="77" value="2003-12-10">
<collectingDate id="EFACD023FFBF731C3280FA40E271F9CD" box="[237,374,1530,1556]" pageId="76" pageNumber="77" value="2003-12-10">10.xii.2003</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFBF731C32AAF99AE05AF9E3" blockId="76.[151,1436,1452,2017]" box="[199,861,1568,1594]" pageId="76" pageNumber="77">
<materialsCitation id="3B3E0556FFBF731C32AAF99AE05EF9E3" box="[199,857,1568,1594]" collectingDate="2003-12-10" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" county="Locus" elevation="175" latitude="-12.969999" location="Foret d'Ampondrabe" longLatPrecision="21" longitude="49.7" municipality="Province" pageId="76" pageNumber="77" specimenCode="BLF 9973" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<emphasis id="B922D319FFBF731C32AAF99AE24AF9E3" bold="true" box="[199,333,1568,1594]" pageId="76" pageNumber="77">
<typeStatus id="54EDB1A9FFBF731C32AAF99AE24EF9E3" box="[199,329,1568,1594]" pageId="76" pageNumber="77" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2 F,
<emphasis id="B922D319FFBF731C33E5F99AE143F9E3" bold="true" box="[392,580,1568,1594]" pageId="76" pageNumber="77">
<collectionCode id="ED4797CEFFBF731C33E5F99AE2C5F9E3" box="[392,450,1568,1594]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="76" pageNumber="77" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFBF731C33A7F99AE143F9E3" box="[458,580,1568,1594]" collectionCode="BLF" pageId="76" pageNumber="77">BLF 9973</specimenCode>
</emphasis>
, same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BE90F0BFFBF731C32FAF9FDE7D3F95F" blockId="76.[151,1436,1452,2017]" pageId="76" pageNumber="77">
Other material: 1 M, 3 F,
<materialsCitation id="3B3E0556FFBF731C33B1F9FCE7D7F95F" collectingDate="2003-12-07" collectionCode="CAS" collectorName="de Bekaraoka &amp; B. L. Fisher" elevation="150" latitude="-13.166667" location="Foret de Bekaraoka" longLatPrecision="21" longitude="49.71" municipality="Province" pageId="76" pageNumber="77" specimenCode="BLF 9871" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="B922D319FFBF731C33B1F9FCE1A5F9B9" bold="true" box="[476,674,1606,1632]" pageId="76" pageNumber="77">
<collectionCode id="ED4797CEFFBF731C33B1F9FCE111F9B9" box="[476,534,1606,1632]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="76" pageNumber="77" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFBF731C304EF9FCE1A5F9B9" box="[547,674,1606,1632]" collectionCode="BLF" pageId="76" pageNumber="77">BLF 9871</specimenCode>
</emphasis>
:
<collectingMunicipality id="6B8D9571FFBF731C30DBF9FDE019F9B8" box="[694,798,1607,1633]" pageId="76" pageNumber="77">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FFBF731C3147F9FDE0BDF9B8" box="[810,954,1607,1633]" country="Madagascar" name="Antsiranana" pageId="76" pageNumber="77">Antsiranana</collectingRegion>
,
<location id="8E8959D0FFBF731C31A6F9FDE7BBF9B8" LSID="urn:lsid:plazi:treatment:03FFBE1DFFB8731F32FAF973E293FB34:8E8959D0FFBF731C31A6F9FDE7BBF9B8" box="[971,1212,1607,1633]" latitude="-13.166667" longLatPrecision="21" longitude="49.71" municipality="Province" name="Foret de Bekaraoka" pageId="76" pageNumber="77" stateProvince="Antsiranana">
Forêt
<collectorName id="26A36ADDFFBF731C3679F9FDE7BBF9B8" box="[1044,1212,1607,1633]" pageId="76" pageNumber="77">de Bekaraoka</collectorName>
</location>
,
<locationDeviation id="9A5B1629FFBF731C36A1F9FDE3F0F95F" location="Daraina" pageId="76" pageNumber="77">
<quantity id="4CAEA2EEFFBF731C36A1F9FDE625F9B8" box="[1228,1314,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="6.8" pageId="76" pageNumber="77" unit="km" value="6.8">6.8 km</quantity>
60° ENE Daraina
</locationDeviation>
,
<geoCoordinate id="EE6269CCFFBF731C336FF9D6E28EF95F" box="[258,393,1644,1670]" degrees="13" direction="south" minutes="10" orientation="latitude" pageId="76" pageNumber="77" precision="15" seconds="00" value="-13.166667">13°10'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFBF731C33FBF9D6E12AF95F" box="[406,557,1644,1670]" degrees="049" direction="east" minutes="42" orientation="longitude" pageId="76" pageNumber="77" precision="15" seconds="36" value="49.71">049°42'36&quot;E</geoCoordinate>
,
<quantity id="4CAEA2EEFFBF731C3057F9D6E180F95F" box="[570,647,1644,1670]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="76" pageNumber="77" unit="m" value="150.0">
<elevation id="007BE838FFBF731C3057F9D6E180F95F" box="[570,647,1644,1670]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="76" pageNumber="77" unit="m" value="150.0">150 m</elevation>
</quantity>
, tropical dry forest, coll.
<collectorName id="26A36ADDFFBF731C31C0F9D6E73CF95F" box="[941,1083,1644,1670]" pageId="76" pageNumber="77">B. L. Fisher</collectorName>
,
<date id="FFE829CBFFBF731C362AF9D6E7D7F95F" box="[1095,1232,1644,1670]" pageId="76" pageNumber="77" value="2003-12-07">
<collectingDate id="EFACD023FFBF731C362AF9D6E7D7F95F" box="[1095,1232,1644,1670]" pageId="76" pageNumber="77" value="2003-12-07">07.xii.2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBF731C32ABF92BE29EF90B" pageId="76" pageNumber="77" type="distribution">
<paragraph id="8BE90F0BFFBF731C32ABF92BE29EF90B" blockId="76.[151,1436,1452,2017]" pageId="76" pageNumber="77">
<emphasis id="B922D319FFBF731C32ABF92BE2F5F972" bold="true" box="[198,498,1681,1707]" pageId="76" pageNumber="77">Distribution &amp; Ecology:</emphasis>
Recorded from the dry forests of Ampondrabe and Bekaraoka in North-Western
<collectingCountry id="F3414F9BFFBF731C32FAF902E223F90B" box="[151,292,1720,1746]" name="Madagascar" pageId="76" pageNumber="77">Madagascar</collectingCountry>
(Fig. 50).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBF731D32ABF964E265F833" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="diagnosis">
<paragraph id="8BE90F0BFFBF731D32ABF964E07AFE43" blockId="76.[151,1436,1452,2017]" lastBlockId="77.[151,1436,152,564]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<emphasis id="B922D319FFBF731C32ABF964E24FF921" bold="true" box="[198,328,1758,1784]" pageId="76" pageNumber="77">Diagnosis:</emphasis>
Up to
<quantity id="4CAEA2EEFFBF731C33F0F965E2F7F920" box="[413,496,1759,1785]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="76" pageNumber="77" unit="mm" value="50.0">50 mm</quantity>
long. Females slightly larger than males. Tergites and appendages olive-green, tergites with light brown posterior margin. Texture of tergites orange-peel like, with numerous minute impressions. Anal shield well-rounded, in males slightly bell-shaped, covered with numerous minute hairs. Hairs on anal shield and on anterior half of tergites standing in minute impressions. First antennomere with groove (Fig. 52A) and
<emphasis id="B922D319FFBF731C33D9F8C0E19FF84A" box="[436,664,1914,1939]" italics="true" pageId="76" pageNumber="77">sensilla basiconica</emphasis>
(Fig. 52C), disc with 35–45 apical cones (Fig. 52D). Movable finger of posterior telopods broad, on both sides completely covered with dense field of long hair, only on well-rounded tip glabrous (Figs 51D, E). Movable finger with triangular membranous lobe and three sclerotized spines (Fig. 51D), posterior side with
<emphasis id="B922D319FFBE731D308FFF22E01AFF68" box="[738,797,152,177]" italics="true" pageId="77" pageNumber="78">circa</emphasis>
8–10 small, sclerotized teeth (Fig. 51E). Fixed finger long and slender, basally with single small spine, distally with numerous sclerotized spots (Fig. 51D). Fixed finger almost as long as movable finger. Three stridulation ribs on male harp (Fig. 51C), posterior side of anterior telopods third podomere laterally with two sclerotized teeth (Fig. 51B). Anal shield with two locking carinae, first small, second 3 or 4 times longer than first, both carinae located towards laterotergites. Females with two or three stridulation ribs on each side of female washboard, operculum of vulva with two wellrounded, same sized tips not protruding above coxa (Fig. 51A).
</paragraph>
<paragraph id="8BE90F0BFFBE731D32ABFE1CE031FDED" blockId="77.[151,1436,152,564]" pageId="77" pageNumber="78">
<emphasis id="B922D319FFBE731D32ABFE1CE288FE19" bold="true" box="[198,399,422,448]" pageId="77" pageNumber="78">Similar species:</emphasis>
<taxonomicName id="4C567488FFBE731D33F6FE1DE071FE19" authority="Wesener &amp; Sierwald, 2005" authorityName="Wesener &amp; Sierwald" authorityYear="2005" box="[411,886,422,448]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="B922D319FFBE731D33F6FE1DE125FE19" box="[411,546,423,448]" italics="true" pageId="77" pageNumber="78">Z. villosum</emphasis>
Wesener &amp; Sierwald, 2005
</taxonomicName>
displays tergite texture and antenna features similar to
<emphasis id="B922D319FFBE731D3362FE77E2BEFE3F" box="[271,441,461,486]" italics="true" pageId="77" pageNumber="78">
<taxonomicName id="4C567488FFBE731D3362FE77E2B3FE3F" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[271,436,461,486]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="trichordum" status="sp. nov.">Z. trichordum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFBE731D33ACFE77E103FE3E" bold="true" box="[449,516,461,487]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="A2116E62FFBE731D33ACFE77E103FE3E" box="[449,516,461,487]" pageId="77" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. The latter is the only species in
<taxonomicName id="4C567488FFBE731D31E1FE77E732FE3F" authorityName="Pocock" authorityYear="1895" box="[908,1077,461,486]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B922D319FFBE731D31E1FE77E732FE3F" box="[908,1077,461,486]" italics="true" pageId="77" pageNumber="78">Zoosphaerium</emphasis>
</taxonomicName>
with three stridulation ribs on each male anterior telopod (Fig. 51C). The pubescence pattern of the posterior telopods third podomere (Figs 51D, E) is unique in the
<taxonomicName id="4C567488FFBE731D33DBFDA0E184FDEA" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[438,643,538,563]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="coquerelianum">
<emphasis id="B922D319FFBE731D33DBFDA0E184FDEA" box="[438,643,538,563]" italics="true" pageId="77" pageNumber="78">Z. coquerelianum</emphasis>
</taxonomicName>
species-group.
</paragraph>
<paragraph id="8BE90F0BFFBE731D32FAF837E265F833" blockId="77.[151,1436,1933,2026]" pageId="77" pageNumber="78">
<emphasis id="B922D319FFBE731D32FAF837E233F87C" bold="true" box="[151,308,1933,1957]" pageId="77" pageNumber="78">FIGURE 052</emphasis>
.
<taxonomicName id="4C567488FFBE731D3325F837E161F87D" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[328,614,1933,1956]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="trichordum" status="sp. nov.">
<emphasis id="B922D319FFBE731D3325F837E161F87D" box="[328,614,1933,1956]" italics="true" pageId="77" pageNumber="78">Zoosphaerium trichordum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFBE731D301EF837E1B1F87C" bold="true" box="[627,694,1933,1957]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="A2116E62FFBE731D301EF837E1B1F87C" box="[627,694,1933,1957]" pageId="77" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFBE731D30A7F837E02EF87D" box="[714,809,1933,1956]" pageId="77" pageNumber="78" type="paratype">paratype</typeStatus>
, SEM, antenna
<emphasis id="B922D319FFBE731D3188F837E0FFF87C" bold="true" box="[997,1016,1933,1957]" pageId="77" pageNumber="78">A</emphasis>
: antennomeres 1–3, lateral view;
<emphasis id="B922D319FFBE731D37EEF837E692F87C" bold="true" box="[1411,1429,1933,1957]" pageId="77" pageNumber="78">B</emphasis>
: antennomeres 4–6, lateral view
<emphasis id="B922D319FFBE731D3386F80AE2F9F811" bold="true" box="[491,510,1968,1992]" pageId="77" pageNumber="78">C</emphasis>
: 5th antennomere with
<emphasis id="B922D319FFBE731D3091F80AE0C0F81E" box="[764,967,1968,1991]" italics="true" pageId="77" pageNumber="78">sensilla basiconica</emphasis>
;
<emphasis id="B922D319FFBE731D31BBF80AE0EEF811" bold="true" box="[982,1001,1968,1992]" pageId="77" pageNumber="78">D</emphasis>
: apical view on disc. Abbreviation:
<emphasis id="B922D319FFBE731D3700F80AE681F811" bold="true" box="[1389,1414,1968,1992]" pageId="77" pageNumber="78">sb</emphasis>
=
<emphasis id="B922D319FFBE731D32FAF869E265F833" box="[151,354,2003,2026]" italics="true" pageId="77" pageNumber="78">sensilla basiconica</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBD731F32ABFF2DE293FB34" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79" type="description">
<paragraph id="8BE90F0BFFBD731E32ABFF2DE1CBFEFF" blockId="78.[151,1436,151,680]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32ABFF2DE267FF68" bold="true" box="[198,352,151,177]" pageId="78" pageNumber="79">Description:</emphasis>
<emphasis id="B922D319FFBD731E3304FF22E2F1FF68" box="[361,502,152,177]" italics="true" pageId="78" pageNumber="79">Body length</emphasis>
: Males (
<specimenCount id="9D50C482FFBD731E300DFF22E1F0FF6B" box="[608,759,152,178]" pageId="78" pageNumber="79" type="generic">2 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFBD731E31CCFF22E0FFFF6B" box="[929,1016,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="78" pageNumber="79" unit="mm" value="36.0">36 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFFBD731E3704FF22E3CFFF01" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="78" pageNumber="79" unit="mm" value="20.2">20.2 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFFBD731E302EFF04E1B7FF01" box="[579,688,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="78" pageNumber="79" unit="mm" value="10.5">10.5 mm</quantity>
. Females (
<specimenCount id="9D50C482FFBD731E315DFF04E0CFFF01" box="[816,968,190,216]" pageId="78" pageNumber="79" type="generic">5 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFBD731E3619FF04E7CDFF01" box="[1140,1226,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="78" pageNumber="79" unit="mm" value="49.0">49 mm</quantity>
, width up to
<quantity id="4CAEA2EEFFBD731E3705FF04E3CFFF26" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="78" pageNumber="79" unit="mm" value="25.4">25.4 mm</quantity>
, height up to
<quantity id="4CAEA2EEFFBD731E3303FF5FE2DCFF26" box="[366,475,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="78" pageNumber="79" unit="mm" value="14.4">14.4 mm</quantity>
. Too few specimens are known to prove statistically that females are larger than males. All known females are larger than males.
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAFE88E158FE43" blockId="78.[151,1436,151,680]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAFE88E256FE92" box="[199,337,306,331]" italics="true" pageId="78" pageNumber="79">Coloration:</emphasis>
Faded in alcohol. Head, collum, thoracic shield, tergites and anal shield olive green with light brown posterior margins. Antennae and legs dark green. In some specimens, all olive green is replaced by brown (because of alcohol influence?).
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAFE1DE116FE19" blockId="78.[151,1436,151,680]" box="[199,529,422,448]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAFE1DE208FE19" box="[199,271,423,448]" italics="true" pageId="78" pageNumber="79">Head:</emphasis>
Eyes with&gt;75 ocelli.
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAFE77E274FD82" blockId="78.[151,1436,151,680]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAFE77E238FE3F" box="[199,319,461,486]" italics="true" pageId="78" pageNumber="79">Antennae:</emphasis>
Length of antennomeres: 1&lt;2&gt;3&gt;4=5&lt;6 (Figs 52A, B), sixth antennomere longest, bearing disc with 35–45 sensory cones (Fig. 52D). First antennomere remarkably broader than the others (Fig. 52A), short, with groove and two rows of
<emphasis id="B922D319FFBD731E3052FDA0E027FDEA" box="[575,800,538,563]" italics="true" pageId="78" pageNumber="79">sensilla basiconica</emphasis>
(Fig. 52C). Sclerotized teeth at base of antennomeres 1–5 (Figs 52A, B).
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAFDD2E013FD71" blockId="78.[151,1436,151,680]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAFDD2E247FD58" box="[199,320,616,641]" italics="true" pageId="78" pageNumber="79">Mandible:</emphasis>
Five rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 53). Condylus with two strongly developed steps (Fig. 53).
</paragraph>
<paragraph id="8BE90F0BFFBD731E32FAF9E3E758F94A" blockId="78.[151,1435,1625,1683]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32FAF9E3E237F9A8" bold="true" box="[151,304,1625,1649]" pageId="78" pageNumber="79">FIGURE 053</emphasis>
.
<taxonomicName id="4C567488FFBD731E3352F9E3E15DF9A9" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[319,602,1625,1648]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="trichordum" status="sp. nov.">
<emphasis id="B922D319FFBD731E3352F9E3E15DF9A9" box="[319,602,1625,1648]" italics="true" pageId="78" pageNumber="79">Zoosphaerium trichordum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFBD731E300EF9E3E1A6F9A8" bold="true" box="[611,673,1625,1649]" pageId="78" pageNumber="79">
<taxonomicNameLabel id="A2116E62FFBD731E300EF9E3E1A6F9A8" box="[611,673,1625,1649]" pageId="78" pageNumber="79" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFBD731E30DFF9E3E017F9A9" box="[690,784,1625,1648]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
, SEM, right mandible, dorsal view. Abbreviations:
<emphasis id="B922D319FFBD731E372BF9E3E66AF9A8" bold="true" box="[1350,1389,1625,1649]" pageId="78" pageNumber="79">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFBD731E337BF9C1E231F94A" bold="true" box="[278,310,1659,1683]" pageId="78" pageNumber="79">Co</emphasis>
= condylus;
<emphasis id="B922D319FFBD731E33ACF9C1E2D8F94A" bold="true" box="[449,479,1659,1683]" pageId="78" pageNumber="79">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFBD731E30F0F9C1E1C5F94A" bold="true" box="[669,706,1659,1683]" pageId="78" pageNumber="79">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFBD731E3108F9C1E082F94A" bold="true" box="[869,901,1659,1683]" pageId="78" pageNumber="79">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAF97FE103F8DF" blockId="78.[151,1436,1733,2030]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAF97FE296F907" box="[199,401,1733,1758]" italics="true" pageId="78" pageNumber="79">Gnathochilarium</emphasis>
: Lateral of palpi four sensory cones, located together. Depression on posterior side of palpi with some sensory cones.
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAF8A8E761F8F5" blockId="78.[151,1436,1733,2030]" box="[199,1126,1810,1836]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAF8A8E220F8F2" box="[199,295,1810,1835]" italics="true" pageId="78" pageNumber="79">Collum:</emphasis>
Whole surface of collum covered with isolated hair of medium length.
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAF883E062F88A" blockId="78.[151,1436,1733,2030]" box="[199,869,1849,1875]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAF883E279F88B" box="[199,382,1849,1874]" italics="true" pageId="78" pageNumber="79">Thoracic shield</emphasis>
shaped like in other species of the genus.
</paragraph>
<paragraph id="8BE90F0BFFBD731E32AAF8DAE0DAF879" blockId="78.[151,1436,1733,2030]" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAF8DAE224F8A0" box="[199,291,1888,1913]" italics="true" pageId="78" pageNumber="79">Tergites</emphasis>
: Texture resembles those of an orange-peel. Anterior half of tergites with numerous hairs standing inside shallow pits. Paratergites projecting slightly posteriorly.
</paragraph>
<paragraph id="8BE90F0BFFBD731F32AAF817E2C0FF6B" blockId="78.[151,1436,1733,2030]" lastBlockId="79.[151,1436,152,1261]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<emphasis id="B922D319FFBD731E32AAF817E267F81F" box="[199,352,1965,1990]" italics="true" pageId="78" pageNumber="79">Endotergum:</emphasis>
Inner area with broad conical spines and long hair. Single row of large, elliptical cuticular impressions present, regularly distributed (Fig. 63B). Three rows of marginal bristles, slightly extending beyond tergite (Fig. 63B).
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAFF05E018FE95" blockId="79.[151,1436,152,1261]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32AAFF05E251FF01" box="[199,342,191,216]" italics="true" pageId="79" pageNumber="80">Anal shield:</emphasis>
Sexually dimorphic, in females well-rounded, neither bell-shaped nor tapered. Covered with numerous hairs standing in minute impressions. Underside carries two well-developed black locking carinae, anterior one small, but well developed. Posterior carina 3.5–4 times longer than first, straight. Carinae located closer to laterotergites than towards anal shield margin.
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAFEE3E3D0FE3E" blockId="79.[151,1436,152,1261]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32AAFEE3E20FFEAB" box="[199,264,345,370]" italics="true" pageId="79" pageNumber="80">Legs:</emphasis>
Tarsi of first pair of legs with five or six, second with seven or eight and third with 9 or 10 ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, 10–12 ventral spines and one apical spine. In 9th leg femur 2.0 times, tarsi 5.5 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAFE4EE04DFDD7" blockId="79.[151,1436,152,1261]" box="[199,842,500,526]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32AAFE4EE28BFDD4" box="[199,396,500,525]" italics="true" pageId="79" pageNumber="80">Stigmatic plates:</emphasis>
First lobe triangular, with pointed tip.
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAFDA0E69BFD71" blockId="79.[151,1436,152,1261]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32AAFDA0E2FFFDEA" box="[199,504,538,563]" italics="true" pageId="79" pageNumber="80">Female sexual characters</emphasis>
: Subanal plate with washboard, consisting of well-developed stridulation ribs, two ribs on each half. Stridulation ribs symmetrical and strong. Vulva operculum large, reaching up to apical edge of coxa, with two well-rounded, same sized lobes (Fig. 51A). Apical margin of operculum medially notched, basal margin straight. Mesal plate long and broad, protruding to half of operculum height (Fig. 51A).
</paragraph>
<paragraph id="8BE90F0BFFBC731F32A3FD0FE06EFD16" blockId="79.[151,1436,152,1261]" box="[206,873,693,719]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32A3FD0FE2E6FD17" box="[206,481,693,718]" italics="true" pageId="79" pageNumber="80">Male sexual characters</emphasis>
: Anal shield weakly bell-shaped.
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAFD66E772FC6E" blockId="79.[151,1436,152,1261]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32AAFD66E292FD2C" box="[199,405,732,757]" italics="true" pageId="79" pageNumber="80">Anterior telopods</emphasis>
(Figs 51B, C): Harp with three stridulation ribs, mesal rib part of inner margin, medial rib shortest, lateral rib longest. All three ribs straight and well-developed. Second podomere process lobe-like and little curved with rounded edges, protruding up to half of third podomeres height, apically with sclerotized spots and single spine. Third podomere long mesal margin of third podomere anteriorly located on lower level than apex of joint. Cavity mesally with numerous sclerotized spots and two sclerotized spines, laterally with two crenulated, small teeth. One sclerotized spot located closely to tip.
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAFC7EE293FB34" blockId="79.[151,1436,152,1261]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32AAFC7EE2A3FC04" box="[199,420,964,989]" italics="true" pageId="79" pageNumber="80">Posterior telopods</emphasis>
(Figs 51D, E): Movable finger of chela thick, 2.0 times longer than wide, weakly curved, apically slightly tapering. Hollowed-out inner margin with three sclerotized spines and one short lobe; posterior aspect with one row of
<emphasis id="B922D319FFBC731F3041FBABE160FBF3" box="[556,615,1041,1066]" italics="true" pageId="79" pageNumber="80">circa</emphasis>
8–10 small, sclerotized teeth. Fixed finger slenderer but as long as movable finger, tip of former weakly curved. Anterior side of fixed finger basally with one short sclerotized spine, apically with numerous sclerotized spots. Fixed finger on both sides covered with dense field of long hair. Third podomere on both sides covered completely with dense field of long hair, leaving only out small, glabrous area around tip. Inner horns of syncoxite apically curved. Tips pointed, whole horns covered with numerous short hairs.
</paragraph>
</subSubSection>
</treatment>
</document>
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