370 lines
37 KiB
XML
370 lines
37 KiB
XML
<document id="AC2882206CFFDFDB06A14AFE0E0FF1B9" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFAE733032FAF8E5E293FBDD" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium tampolo Wesener 2009, new species" docType="treatment" docVersion="3" lastPageNumber="97" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="94" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="41107E6E9E93B35A5E104596CBC307CD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6815D536DFD5F6A7C182A01B03FC2156">
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<mods:title id="67230B1DE5DFE91D2E1C099EF8DD34FB">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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</mods:titleInfo>
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<mods:name id="C91162C2DE36D60D31867CDAAD5572CB" type="personal">
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<mods:role id="20BD0C0DC0170E9E72B925D003995F71">
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<mods:roleTerm id="752429CC65725E9B4E1EF531E570D221">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="53F69D50CD37E6BA9123CB8A179A96D6">Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EF6BFE13C1D8473EFCBDC05386C43AEE">text</mods:typeOfResource>
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<mods:titleInfo id="B56A1E48A2FBEADDD2BEC055D244F746">
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<mods:title id="E11EC175AD468746E377C21AC97174A5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8C8C086AFD3FFABE56BA063CA54CCBB6">
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<mods:date id="9BBD48F01F02426685F24CD3462E5E88">2009</mods:date>
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<mods:detail id="74D95D47B5C2CEAA8655336F4B275947" type="pubDate">
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<mods:number id="24085533E850B40F2C33DFFB7044C28B">2009-05-11</mods:number>
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</mods:detail>
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<mods:detail id="F5E5E41B55964BDBA57A598AF7283E83" type="volume">
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<mods:number id="169C0284372B59BCD016E77FC7CD993E">2097</mods:number>
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</mods:detail>
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<mods:detail id="BAE7FF6041539BA69C15B99B3452C075" type="issue">
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<mods:number id="F78D8C04A10941971CC70DEEF014B18B">1</mods:number>
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</mods:detail>
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<mods:extent id="B31223B331F42AF3673003B8D79B04BD" unit="page">
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<mods:start id="0CB98E0EA748EFADFF864CA79BFD2AA6">1</mods:start>
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<mods:end id="2B1621E7EC13AB6E6299F80A41C1259A">134</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="A3BFDC37706C39652B5BA5EDA881847A">
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<mods:url id="94DE553A72730F856FE98A28FDAEA900">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
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</mods:location>
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<mods:classification id="18BC1BF79337304134C0C9DC1246EFA6">journal article</mods:classification>
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<mods:identifier id="F27F06AD30DBBA27400DE7E7087423AF" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
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<mods:identifier id="32B094F1109274D82A3D2749B33FC5EF" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="1954967BDC4E5B43443A04854FE21150" type="Zenodo-Dep">5316536</mods:identifier>
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<mods:identifier id="AA96E7E0D9458B21CEFC2979F5685443" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
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</mods:mods>
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<treatment id="03FFBE1DFFAE733032FAF8E5E293FBDD" LSID="urn:lsid:plazi:treatment:03FFBE1DFFAE733032FAF8E5E293FBDD" httpUri="http://treatment.plazi.org/id/03FFBE1DFFAE733032FAF8E5E293FBDD" lastPageId="96" lastPageNumber="97" pageId="93" pageNumber="94">
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<subSubSection id="C34C5C80FFAE730D32FAF8E5E149F8A0" box="[151,590,1887,1913]" pageId="93" pageNumber="94" type="nomenclature">
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<paragraph id="8BE90F0BFFAE730D32FAF8E5E149F8A0" blockId="93.[151,590,1887,1913]" box="[151,590,1887,1913]" pageId="93" pageNumber="94">
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<heading id="D0A1B867FFAE730D32FAF8E5E149F8A0" bold="true" box="[151,590,1887,1913]" fontSize="11" level="1" pageId="93" pageNumber="94" reason="1">
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||
<emphasis id="B922D319FFAE730D32FAF8E5E149F8A0" bold="true" box="[151,590,1887,1913]" pageId="93" pageNumber="94">
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||
<emphasis id="B922D319FFAE730D32FAF8E5E2B1F8A0" bold="true" box="[151,438,1887,1913]" italics="true" pageId="93" pageNumber="94">
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<taxonomicName id="4C567488FFAE730D32FAF8E5E2B7F8A0" ID-CoL="5D7J8" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,432,1887,1913]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">Zoosphaerium tampolo</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A2116E62FFAE730D33D3F8E5E149F8A0" box="[446,590,1887,1913]" pageId="93" pageNumber="94" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C34C5C80FFAE730E32FAF812E260FE19" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94" type="etymology">
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<paragraph id="8BE90F0BFFAE730D32FAF812E69BF830" blockId="93.[151,1436,1960,2025]" pageId="93" pageNumber="94">
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<emphasis id="B922D319FFAE730D32FAF812E287F81B" bold="true" box="[151,384,1960,1986]" pageId="93" pageNumber="94">Derivatio nominis:</emphasis>
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<taxonomicName id="4C567488FFAE730D33E5F813E2EFF81B" authorityName="Wesener" authorityYear="2009" box="[392,488,1961,1986]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="tampolo">
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<emphasis id="B922D319FFAE730D33E5F813E2EFF81B" box="[392,488,1961,1986]" italics="true" pageId="93" pageNumber="94">tampolo</emphasis>
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</taxonomicName>
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, noun in apposition, refers to the type locality.
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<emphasis id="B922D319FFAE730D32ABF875E12FF830" bold="true" box="[198,552,1999,2025]" pageId="93" pageNumber="94">
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Studied material:
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<typeStatus id="54EDB1A9FFAE730D33C0F875E124F830" box="[429,547,1999,2025]" pageId="93" pageNumber="94" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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1 M (
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<quantity id="4CAEA2EEFFAE730D3017F875E1D6F830" box="[634,721,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="93" pageNumber="94" unit="mm" value="33.0">33 mm</quantity>
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long),
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<emphasis id="B922D319FFAE730D3147F875E0CAF830" bold="true" box="[810,973,1999,2025]" pageId="93" pageNumber="94">FMMC 3958</emphasis>
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,
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<emphasis id="B922D319FFAE730D31B2F86AE784F830" box="[991,1155,2000,2025]" italics="true" pageId="93" pageNumber="94">Locus typicus</emphasis>
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:
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<collectingCountry id="F3414F9BFFAE730D36F8F875E623F830" box="[1173,1316,1999,2025]" name="Madagascar" pageId="93" pageNumber="94">Madagascar</collectingCountry>
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, Province
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</paragraph>
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<paragraph id="8BE90F0BFFAD730E32FAFF22E230FF01" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
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<materialsCitation id="3B3E0556FFAD730E32FAFF22E234FF01" collectingDate="1997-04-03" collectingDateMax="1997-04-16" collectingDateMin="1997-04-03" collectorName="de Tampolo & S. Goodman" elevation="10" latitude="-17.286667" location="Forestiere de Tampolo" longLatPrecision="127" longitude="49.423332" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Toamasina">
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<collectingRegion id="4992C1E9FFAD730E32FAFF22E21FFF6B" box="[151,280,152,178]" country="Madagascar" name="Toamasina" pageId="94" pageNumber="95">Toamasina</collectingRegion>
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,
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<location id="8E8959D0FFAD730E335CFF22E153FF6B" LSID="urn:lsid:plazi:treatment:03FFBE1DFFAE733032FAF8E5E293FBDD:8E8959D0FFAD730E335CFF22E153FF6B" box="[305,596,152,178]" latitude="-17.286667" longLatPrecision="127" longitude="49.423332" name="Forestiere de Tampolo" pageId="94" pageNumber="95" stateProvince="Toamasina">
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Forestière
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<collectorName id="26A36ADDFFAD730E33D7FF22E153FF6B" box="[442,596,152,178]" pageId="94" pageNumber="95">de Tampolo</collectorName>
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</location>
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, littoral forest,
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<quantity id="4CAEA2EEFFAD730E3140FF22E07FFF6B" box="[813,888,152,178]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="10.0">
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||
<elevation id="007BE838FFAD730E3140FF22E07FFF6B" box="[813,888,152,178]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="10.0">10 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE6269CCFFAD730E31FCFF22E702FF6B" box="[913,1029,152,178]" degrees="17" direction="south" minutes="17.2" orientation="latitude" pageId="94" pageNumber="95" precision="92" value="-17.286667">17°17.2'S</geoCoordinate>
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,
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<geoCoordinate id="EE6269CCFFAD730E3673FF22E793FF6B" box="[1054,1172,152,178]" degrees="49" direction="east" minutes="25.4" orientation="longitude" pageId="94" pageNumber="95" precision="92" value="49.423332">49°25.4'E</geoCoordinate>
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||
, coll.
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<collectorName id="26A36ADDFFAD730E3698FF22E69FFF6B" box="[1269,1432,152,178]" pageId="94" pageNumber="95">S. Goodman</collectorName>
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,
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||
<date id="FFE829CBFFAD730E32FAFF04E234FF01" box="[151,307,190,216]" pageId="94" pageNumber="95" value="1997-04-03" valueMax="1997-04-16" valueMin="1997-04-03">
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||
<collectingDate id="EFACD023FFAD730E32FAFF04E234FF01" box="[151,307,190,216]" pageId="94" pageNumber="95" value="1997-04-03" valueMax="1997-04-16" valueMin="1997-04-03">3–16.iv.1997</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8BE90F0BFFAD730E32AAFF5FE3D0FEAA" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
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||
<materialsCitation id="3B3E0556FFAD730E32AAFF5FE07FFF26" box="[199,888,229,255]" collectionCode="FMMC" pageId="94" pageNumber="95" specimenCode="FMMC 3958" specimenCount="1" typeStatus="paratype">
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||
<emphasis id="B922D319FFAD730E32AAFF5FE24AFF26" bold="true" box="[199,333,229,255]" pageId="94" pageNumber="95">
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||
<typeStatus id="54EDB1A9FFAD730E32AAFF5FE24EFF26" box="[199,329,229,255]" pageId="94" pageNumber="95" type="paratype">Paratypes</typeStatus>
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||
:
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||
</emphasis>
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||
2 M, 3 F,
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<emphasis id="B922D319FFAD730E33A8FF5FE163FF26" bold="true" box="[453,612,229,255]" pageId="94" pageNumber="95">
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||
<specimenCode id="DBF0A770FFAD730E33A8FF5FE163FF26" box="[453,612,229,255]" collectionCode="FMMC" pageId="94" pageNumber="95">FMMC 3958</specimenCode>
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||
</emphasis>
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||
, same data as holotype
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||
</materialsCitation>
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||
. 8 M, 2 F,
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<materialsCitation id="3B3E0556FFAD730E319EFF5FE69FFEFF" collectingDate="1997-04-04" collectionCode="FMMC" collectorName="S. F. Tampolo & B. L. Fisher" elevation="10" latitude="-17.283333" location="Fenoarivo Atn." longLatPrecision="1279" longitude="49.433334" pageId="94" pageNumber="95" specimenCode="FMMC 4030" specimenCount="1" typeStatus="paratype">
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||
<emphasis id="B922D319FFAD730E319EFF5FE79DFF26" bold="true" box="[1011,1178,229,255]" pageId="94" pageNumber="95">
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||
<specimenCode id="DBF0A770FFAD730E319EFF5FE793FF26" box="[1011,1172,229,255]" collectionCode="FMMC" pageId="94" pageNumber="95">FMMC 4030</specimenCode>
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,
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</emphasis>
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<collectorName id="26A36ADDFFAD730E36CCFF5FE640FF26" box="[1185,1351,229,255]" pageId="94" pageNumber="95">S. F. Tampolo</collectorName>
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||
,
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<quantity id="4CAEA2EEFFAD730E373FFF5FE69CFF26" box="[1362,1435,229,255]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="km" value="10.0">10 km</quantity>
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NNE
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<location id="8E8959D0FFAD730E32B6FEB6E297FEFF" LSID="urn:lsid:plazi:treatment:03FFBE1DFFAE733032FAF8E5E293FBDD:8E8959D0FFAD730E32B6FEB6E297FEFF" box="[219,400,268,294]" latitude="-17.283333" longLatPrecision="1279" longitude="49.433334" name="Fenoarivo Atn." pageId="94" pageNumber="95">Fenoarivo Atn.</location>
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, littoral forest,
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<quantity id="4CAEA2EEFFAD730E302BFEB6E183FEFF" box="[582,644,268,294]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="10.0">
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||
<elevation id="007BE838FFAD730E302BFEB6E183FEFF" box="[582,644,268,294]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="10.0">10 m</elevation>
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||
</quantity>
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,
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<geoCoordinate id="EE6269CCFFAD730E30FFFEB6E1F7FEFF" box="[658,752,268,294]" degrees="17" direction="south" minutes="17" orientation="latitude" pageId="94" pageNumber="95" precision="925" value="-17.283333">17°17'S</geoCoordinate>
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||
,
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<geoCoordinate id="EE6269CCFFAD730E3090FEB6E05AFEFF" box="[765,861,268,294]" degrees="49" direction="east" minutes="26" orientation="longitude" pageId="94" pageNumber="95" precision="925" value="49.433334">49°26'E</geoCoordinate>
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||
, coll.
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||
<collectorName id="26A36ADDFFAD730E31CBFEB6E732FEFF" box="[934,1077,268,294]" pageId="94" pageNumber="95">B. L. Fisher</collectorName>
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||
,
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||
<date id="FFE829CBFFAD730E362FFEB6E7C7FEFF" box="[1090,1216,268,294]" pageId="94" pageNumber="95" value="1997-04-04">
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||
<collectingDate id="EFACD023FFAD730E362FFEB6E7C7FEFF" box="[1090,1216,268,294]" pageId="94" pageNumber="95" value="1997-04-04">04.iv.1997</collectingDate>
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||
</date>
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||
,
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||
<collectingMethod id="5217771CFFAD730E36A0FEB6E65FFEFF" box="[1229,1368,268,294]" pageId="94" pageNumber="95">pitfall traps</collectingMethod>
|
||
. 3 M
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B3E0556FFAD730E32FAFE88E3D3FEAA" collectingDate="1997-04-10" collectionCode="FMMC" collectorName="B. L. Fisher" elevation="10" latitude="-17.283333" location="Tampolo" longLatPrecision="1279" longitude="49.416668" pageId="94" pageNumber="95" specimenCode="FMMC 4035" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B922D319FFAD730E32FAFE88E23EFE95" bold="true" box="[151,313,306,332]" pageId="94" pageNumber="95">
|
||
<specimenCode id="DBF0A770FFAD730E32FAFE88E23EFE95" box="[151,313,306,332]" collectionCode="FMMC" pageId="94" pageNumber="95">FMMC 4035</specimenCode>
|
||
</emphasis>
|
||
, S. F.
|
||
<location id="8E8959D0FFAD730E33E6FE88E2F1FE95" LSID="urn:lsid:plazi:treatment:03FFBE1DFFAE733032FAF8E5E293FBDD:8E8959D0FFAD730E33E6FE88E2F1FE95" box="[395,502,306,332]" latitude="-17.283333" longLatPrecision="1279" longitude="49.416668" name="Tampolo" pageId="94" pageNumber="95">Tampolo</location>
|
||
, littoral forest,
|
||
<quantity id="4CAEA2EEFFAD730E30C2FE88E1E8FE95" box="[687,751,306,332]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="10.0">
|
||
<elevation id="007BE838FFAD730E30C2FE88E1E8FE95" box="[687,751,306,332]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFAD730E3093FE88E062FE95" box="[766,869,306,332]" degrees="17" direction="south" minutes="17" orientation="latitude" pageId="94" pageNumber="95" precision="925" value="-17.283333">17°17' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFAD730E3119FE88E0DAFE95" box="[884,989,306,332]" degrees="49" direction="east" minutes="25" orientation="longitude" pageId="94" pageNumber="95" precision="925" value="49.416668">49°25' E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26A36ADDFFAD730E3644FE88E7BBFE95" box="[1065,1212,306,332]" pageId="94" pageNumber="95">B. L. Fisher</collectorName>
|
||
,
|
||
<date id="FFE829CBFFAD730E36A7FE88E640FE95" box="[1226,1351,306,332]" pageId="94" pageNumber="95" value="1997-04-10">
|
||
<collectingDate id="EFACD023FFAD730E36A7FE88E640FE95" box="[1226,1351,306,332]" pageId="94" pageNumber="95" value="1997-04-10">10.iv.1997</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="5217771CFFAD730E373BFE88E3D3FEAA" pageId="94" pageNumber="95">pitfall traps</collectingMethod>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAFEC5E260FE19" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAFEC5E283FE40" bold="true" box="[199,388,383,409]" pageId="94" pageNumber="95">Other material</emphasis>
|
||
: All from S. F. Tampolo: 4 M,
|
||
<emphasis id="B922D319FFAD730E309BFEC5E090FE40" bold="true" box="[758,919,383,409]" pageId="94" pageNumber="95">FMMC 7029</emphasis>
|
||
; 4 F,
|
||
<emphasis id="B922D319FFAD730E31B6FEC5E77BFE40" bold="true" box="[987,1148,383,409]" pageId="94" pageNumber="95">FMMC 7881</emphasis>
|
||
; 11 F,
|
||
<emphasis id="B922D319FFAD730E36A0FEC5E669FE40" bold="true" box="[1229,1390,383,409]" pageId="94" pageNumber="95">FMMC 7881</emphasis>
|
||
; 20 M,
|
||
<emphasis id="B922D319FFAD730E32D2FE1CE258FE19" bold="true" box="[191,351,422,448]" pageId="94" pageNumber="95">FMMC 7882</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFAD730E32ABFE77E61EFE3E" box="[198,1305,461,487]" pageId="94" pageNumber="95" type="distribution">
|
||
<paragraph id="8BE90F0BFFAD730E32ABFE77E61EFE3E" blockId="94.[151,1436,152,2032]" box="[198,1305,461,487]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32ABFE77E26EFE3E" bold="true" box="[198,361,461,487]" pageId="94" pageNumber="95">Distribution:</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54EDB1A9FFAD730E3018FE77E1A0FE3E" box="[629,679,461,487]" pageId="94" pageNumber="95">type</typeStatus>
|
||
locality, the Northern littoral forest of S. F. Tampolo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFAD730E32ABFE48E674FBAE" pageId="94" pageNumber="95" type="diagnosis">
|
||
<paragraph id="8BE90F0BFFAD730E32ABFE48E20EFC56" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32ABFE48E24FFDD5" bold="true" box="[198,328,498,524]" pageId="94" pageNumber="95">Diagnosis:</emphasis>
|
||
Up to
|
||
<quantity id="4CAEA2EEFFAD730E33CFFE48E2F0FDD5" box="[418,503,498,524]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="94" pageNumber="95" unit="mm" value="34.0">34 mm</quantity>
|
||
long. Color faded to olive-brownish in alcohol. Antenna, head and legs dark greenish. Texture of tergites glabrous. Anal shield well -rounded, covered with few, isolated hairs. First stigmatic plates greatly enlarged and well-rounded (Figs 64A, B). Disc of antenna with 20–30 apical cones (Fig. 65C). Movable finger of posterior telopods broad, with well-rounded tip and five sclerotized spines (Fig. 64F); posterior side with
|
||
<emphasis id="B922D319FFAD730E33A8FD37E107FD7F" box="[453,512,653,678]" italics="true" pageId="94" pageNumber="95">circa</emphasis>
|
||
10 small, sclerotized teeth (Fig. 64G). Fixed finger long and slender, curved, basally with one small spine, distally at inner margin with numerous sclerotized spots (Fig. 64F). Fixed finger slightly shorter than movable finger. Single stridulation rib on each male harp (Fig. 64D). Last podomere of anterior telopods laterally of cavity with 4 or 5 sclerotized teeth (Fig. 64E). Anal shield with two locking carinae, first small, second 2 times longer than first. Female with two stridulation ribs on each side of female washboard, operculum of vulva with two well-rounded, same sized tips not protruding above coxal margin (Fig. 4C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32ABFC21E674FBAE" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32ABFC21E28BFC6C" bold="true" box="[198,396,923,949]" pageId="94" pageNumber="95">Similar species:</emphasis>
|
||
<emphasis id="B922D319FFAD730E33FBFC26E119FC6C" box="[406,542,924,949]" italics="true" pageId="94" pageNumber="95">
|
||
<taxonomicName id="4C567488FFAD730E33FBFC26E11DFC6C" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[406,538,924,949]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">Z. tampolo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFAD730E3045FC21E16BFC6C" bold="true" box="[552,620,923,949]" pageId="94" pageNumber="95">
|
||
<taxonomicNameLabel id="A2116E62FFAD730E3045FC21E16BFC6C" box="[552,620,923,949]" pageId="94" pageNumber="95" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
occurs sympatrically with
|
||
<taxonomicName id="4C567488FFAD730E31DFFC26E690FC6F" authority="(Wesener & Sierwald, 2005)" baseAuthorityName="Wesener & Sierwald" baseAuthorityYear="2005" box="[946,1431,924,950]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="villosum">
|
||
<emphasis id="B922D319FFAD730E31DFFC26E731FC6C" box="[946,1078,924,949]" italics="true" pageId="94" pageNumber="95">Z. villosum</emphasis>
|
||
(Wesener & Sierwald, 2005)
|
||
</taxonomicName>
|
||
, from which it differs in the number of stridulation ribs on the male harp, the body size and the shape of the locking carinae on the anal shield.
|
||
<emphasis id="B922D319FFAD730E3055FC53E1C7FBDB" box="[568,704,1001,1026]" italics="true" pageId="94" pageNumber="95">
|
||
<taxonomicName id="4C567488FFAD730E3055FC53E1BBFBDB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[568,700,1001,1026]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">Z. tampolo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFAD730E30A4FC52E00AFBDB" bold="true" box="[713,781,1000,1026]" pageId="94" pageNumber="95">
|
||
<taxonomicNameLabel id="A2116E62FFAD730E30A4FC52E00AFBDB" box="[713,781,1000,1026]" pageId="94" pageNumber="95" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares some habitus characters with
|
||
<taxonomicName id="4C567488FFAD730E36A4FC53E2E7FBF3" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="platylabum">
|
||
<emphasis id="B922D319FFAD730E36A4FC53E669FBDB" box="[1225,1390,1001,1026]" italics="true" pageId="94" pageNumber="95">Z. platylabum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC772FAFFAD730E3712FC53E2DFFBF3" author="de Saussure, H. & Zehntner, L." pageId="94" pageNumber="95" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. & Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but differs from the latter in different shaped anterior and posterior telopods. The greatly enlarged first stigma plates and the presence of minute sclerotized teeth on the anterior side of the movable finger are a unique character of
|
||
<emphasis id="B922D319FFAD730E3014FBE7E1F8FBAF" box="[633,767,1117,1142]" italics="true" pageId="94" pageNumber="95">
|
||
<taxonomicName id="4C567488FFAD730E3014FBE7E1FCFBAF" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[633,763,1117,1142]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">Z. tampolo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFAD730E316AFBE7E04EFBAE" bold="true" box="[775,841,1117,1143]" pageId="94" pageNumber="95">
|
||
<taxonomicNameLabel id="A2116E62FFAD730E316AFBE7E04EFBAE" box="[775,841,1117,1143]" pageId="94" pageNumber="95" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
for the whole
|
||
<taxonomicName id="4C567488FFAD730E3199FBE7E7C5FBAF" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[1012,1218,1117,1142]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="coquerelianum">
|
||
<emphasis id="B922D319FFAD730E3199FBE7E7C5FBAF" box="[1012,1218,1117,1142]" italics="true" pageId="94" pageNumber="95">Z. coquerelianum</emphasis>
|
||
</taxonomicName>
|
||
species-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFAD733032ABFB39E293FBDD" lastPageId="96" lastPageNumber="97" pageId="94" pageNumber="95" type="description">
|
||
<paragraph id="8BE90F0BFFAD730E32ABFB39E004FB32" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32ABFB39E267FB44" bold="true" box="[198,352,1155,1181]" pageId="94" pageNumber="95">Description:</emphasis>
|
||
<emphasis id="B922D319FFAD730E3305FB3EE2F2FB44" box="[360,501,1156,1181]" italics="true" pageId="94" pageNumber="95">Body length</emphasis>
|
||
: Males (
|
||
<specimenCount id="9D50C482FFAD730E3031FB3EE005FB47" box="[604,770,1156,1182]" pageId="94" pageNumber="95" type="generic">15 specimens</specimenCount>
|
||
): length up to
|
||
<quantity id="4CAEA2EEFFAD730E31C5FB3EE0FAFB47" box="[936,1021,1156,1182]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="94" pageNumber="95" unit="mm" value="33.0">33 mm</quantity>
|
||
, width of thoracic shield up to 16.0 mm, height of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFAD730E3025FB11E1A0FB1C" box="[584,679,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="94" pageNumber="95" unit="mm" value="8.8">8.8 mm</quantity>
|
||
. Females (
|
||
<specimenCount id="9D50C482FFAD730E3145FB11E0C6FB1C" box="[808,961,1195,1221]" pageId="94" pageNumber="95" type="generic">4 specimens</specimenCount>
|
||
): length up to
|
||
<quantity id="4CAEA2EEFFAD730E361DFB11E7C0FB1C" box="[1136,1223,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="94" pageNumber="95" unit="mm" value="32.0">32 mm</quantity>
|
||
, width up to
|
||
<quantity id="4CAEA2EEFFAD730E3705FB11E3CFFB32" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="94" pageNumber="95" unit="mm" value="15.4">15.4 mm</quantity>
|
||
, height up to
|
||
<quantity id="4CAEA2EEFFAD730E330BFB6BE2C4FB32" box="[358,451,1233,1259]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="94" pageNumber="95" unit="mm" value="7.9">7.9 mm</quantity>
|
||
. Both sexes of similar size.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAFB42E09BFACB" blockId="94.[151,1436,152,2032]" box="[199,924,1272,1298]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAFB42E229FAC8" box="[199,302,1272,1297]" italics="true" pageId="94" pageNumber="95">Habitus:</emphasis>
|
||
medium sized, dark-colored, glabrous pill millipede.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAFAA5E21CFA86" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAFAA5E256FAE1" box="[199,337,1311,1336]" italics="true" pageId="94" pageNumber="95">Coloration:</emphasis>
|
||
Head, collum, thoracic shield, tergites and anal shield completely blackish, antennae and legs dark green.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAFAD6E116FA5F" blockId="94.[151,1436,152,2032]" box="[199,529,1388,1414]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAFAD6E217FA5C" box="[199,272,1388,1413]" italics="true" pageId="94" pageNumber="95">Head:</emphasis>
|
||
Eyes with>75 ocelli.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAFA29E74EF9F9" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAFA29E238FA75" box="[199,319,1427,1452]" italics="true" pageId="94" pageNumber="95">Antennae:</emphasis>
|
||
Length of antennomeres: 1>2=3>4=5<<6, sixth antennomere longest, as long as fourth and fifth combined (Fig. 65A). Sixth antennomere cylindrical, disc with 20–30 sensory cones (Fig. 65C). First antennomere remarkably broader than others (Fig. 65A), short, with groove and 1 or 2 rows of
|
||
<emphasis id="B922D319FFAD730E372FFA5AE211F9F9" italics="true" pageId="94" pageNumber="95">sensilla basiconica</emphasis>
|
||
(Fig. 65B). Sclerotized teeth at base of antennomeres 1–5 (Fig. 65A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF997E02EF9B7" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF997E247F99F" box="[199,320,1581,1606]" italics="true" pageId="94" pageNumber="95">Mandible:</emphasis>
|
||
5 or 6 rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 65D). Condylus with two strongly developed steps (Fig. 65D).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF9C1E103F962" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF9C1E296F94D" box="[199,401,1659,1684]" italics="true" pageId="94" pageNumber="95">Gnathochilarium</emphasis>
|
||
: Laterally of palpi four sensory cones, located together. Depression on posterior side of palpi with some sensory cones.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF972E1AEF93B" blockId="94.[151,1436,152,2032]" box="[199,681,1736,1762]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF972E220F938" box="[199,295,1736,1761]" italics="true" pageId="94" pageNumber="95">Collum:</emphasis>
|
||
Median part of collum glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF955E1E8F8D1" blockId="94.[151,1436,152,2032]" box="[199,751,1774,1800]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF955E279F8D1" box="[199,382,1775,1800]" italics="true" pageId="94" pageNumber="95">Thoracic shield</emphasis>
|
||
surface structured like tergites.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF8AFE021F8F6" blockId="94.[151,1436,152,2032]" box="[199,806,1813,1839]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF8AFE224F8F7" box="[199,291,1813,1838]" italics="true" pageId="94" pageNumber="95">Tergites</emphasis>
|
||
: Paratergites projecting slightly posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF886E69BF8A4" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF886E267F88C" box="[199,352,1852,1877]" italics="true" pageId="94" pageNumber="95">Endotergum:</emphasis>
|
||
Inner area with thin conical spines and long hair. Single row of small, circular cuticular impression, regularly distributed (Fig. 74A). Two rows of marginal bristles, reaching up to margin (Fig. 74A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAD730E32AAF833E140F829" blockId="94.[151,1436,152,2032]" pageId="94" pageNumber="95">
|
||
<emphasis id="B922D319FFAD730E32AAF833E25FF87B" box="[199,344,1929,1954]" italics="true" pageId="94" pageNumber="95">Anal shield:</emphasis>
|
||
Well-rounded, neither bell-shaped nor tapered. Covered with few isolated hairs. Underside carries two well-developed black locking carinae, anterior one small, but well-developed. Posterior carina 1.5–2 times longer than anterior one.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFAC730F32FAF88BE0A5F857" blockId="95.[151,1435,1841,1934]" pageId="95" pageNumber="96">
|
||
<emphasis id="B922D319FFAC730F32FAF88BE22AF890" bold="true" box="[151,301,1841,1865]" pageId="95" pageNumber="96">FIGURE 065</emphasis>
|
||
.
|
||
<taxonomicName id="4C567488FFAC730F3357F888E136F890" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,561,1842,1865]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">
|
||
<emphasis id="B922D319FFAC730F3357F888E136F890" box="[314,561,1842,1865]" italics="true" pageId="95" pageNumber="96">Zoosphaerium tampolo</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFAC730F3055F88BE174F890" bold="true" box="[568,627,1841,1865]" pageId="95" pageNumber="96">
|
||
<taxonomicNameLabel id="A2116E62FFAC730F3055F88BE174F890" box="[568,627,1841,1865]" pageId="95" pageNumber="96" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="54EDB1A9FFAC730F30EDF88BE1D8F891" box="[640,735,1841,1864]" pageId="95" pageNumber="96" type="paratype">paratype</typeStatus>
|
||
, SEM,
|
||
<emphasis id="B922D319FFAC730F3140F88BE047F890" bold="true" box="[813,832,1841,1865]" pageId="95" pageNumber="96">A</emphasis>
|
||
: antenna, lateral view;
|
||
<emphasis id="B922D319FFAC730F3654F88BE74CF890" bold="true" box="[1081,1099,1841,1865]" pageId="95" pageNumber="96">B</emphasis>
|
||
: antenna, first antennomere;
|
||
<emphasis id="B922D319FFAC730F37ECF88BE693F890" bold="true" box="[1409,1428,1841,1865]" pageId="95" pageNumber="96">C</emphasis>
|
||
: apical view on sixth antennomere with disc;
|
||
<emphasis id="B922D319FFAC730F30EFF8E9E192F8B2" bold="true" box="[642,661,1875,1899]" pageId="95" pageNumber="96">D</emphasis>
|
||
: right mandible, mesal view. Abbreviations:
|
||
<emphasis id="B922D319FFAC730F36EEF8E9E7AEF8B2" bold="true" box="[1155,1193,1875,1899]" pageId="95" pageNumber="96">3iT</emphasis>
|
||
= 3 inner teeth;
|
||
<emphasis id="B922D319FFAC730F3709F8E9E683F8B2" bold="true" box="[1380,1412,1875,1899]" pageId="95" pageNumber="96">Co</emphasis>
|
||
= condylus;
|
||
<emphasis id="B922D319FFAC730F3368F8CCE224F857" bold="true" box="[261,291,1910,1934]" pageId="95" pageNumber="96">eT</emphasis>
|
||
= external tooth;
|
||
<emphasis id="B922D319FFAC730F338DF8CCE102F857" bold="true" box="[480,517,1910,1934]" pageId="95" pageNumber="96">mp</emphasis>
|
||
= molar plate;
|
||
<emphasis id="B922D319FFAC730F30C4F8CCE1CEF857" bold="true" box="[681,713,1910,1934]" pageId="95" pageNumber="96">pL</emphasis>
|
||
= pectinate lamella.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFF93733032AAFF22E216FEFF" blockId="96.[151,1437,152,1028]" pageId="96" pageNumber="97">
|
||
<emphasis id="B922D319FF93733032AAFF22E20FFF68" box="[199,264,152,177]" italics="true" pageId="96" pageNumber="97">Legs:</emphasis>
|
||
Tarsi of first pair of legs with four or five, second with five to six and third pair with seven to eight ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, 12–14 ventral spines and one apical spine. In 9th leg femur 2.4 times, tarsi 4.8 times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFF93733032AAFE88E7CDFE95" blockId="96.[151,1437,152,1028]" box="[199,1226,306,332]" pageId="96" pageNumber="97">
|
||
<emphasis id="B922D319FF93733032AAFE88E28BFE92" box="[199,396,306,331]" italics="true" pageId="96" pageNumber="97">Stigmatic plates:</emphasis>
|
||
First lobe long and broad, reaching as high as prefemur (Figs 64A, B).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFF93733032AAFEE3E0A8FE3E" blockId="96.[151,1437,152,1028]" pageId="96" pageNumber="97">
|
||
<emphasis id="B922D319FF93733032AAFEE3E2FFFEAB" box="[199,504,345,370]" italics="true" pageId="96" pageNumber="97">Female sexual characters</emphasis>
|
||
: Subanal plate with washboard, consisting of well-developed stridulation ribs, two ribs on each half. Stridulation ribs symmetrical and strong. Vulva large, covering more than 3/4 of coxa. Operculum reaching up to apical coxal edge, with two well-rounded, same sized lobes (Fig. 64C). Mesal plate long and broad, reaching up to half of operculum height (Fig. 64C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFF93733032AAFE4EE08AFDD7" blockId="96.[151,1437,152,1028]" box="[199,909,500,526]" pageId="96" pageNumber="97">
|
||
<emphasis id="B922D319FF93733032AAFE4EE2DDFDD4" box="[199,474,500,525]" italics="true" pageId="96" pageNumber="97">Male sexual characters</emphasis>
|
||
: Anal shield not sexually dimorphic.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFF93733032AAFDA0E3BAFD16" blockId="96.[151,1437,152,1028]" pageId="96" pageNumber="97">
|
||
<emphasis id="B922D319FF93733032AAFDA0E293FDEA" box="[199,404,538,563]" italics="true" pageId="96" pageNumber="97">Anterior telopods</emphasis>
|
||
(Figs 64D, E): Harp with single stridulation rib, rib straight and well-developed. Second podomere process lobe-like and little curved with rounded edges, reaching up to 1/3 of third podomere height, apically with sclerotized spots. Third podomere cavity mesally with numerous sclerotized spots and three sclerotized spines, laterally with row of 4 or 5 crenulated, large teeth. One sclerotized spot located closely to tip.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFF93733032AAFD66E293FBDD" blockId="96.[151,1437,152,1028]" pageId="96" pageNumber="97">
|
||
<emphasis id="B922D319FF93733032AAFD66E2A4FD2C" box="[199,419,732,757]" italics="true" pageId="96" pageNumber="97">Posterior telopods</emphasis>
|
||
(Figs 64E, F): Movable finger of chela thick, 1.85 times longer than wide, weakly curved, apically slightly tapering. Hollowed-out inner margin with five sclerotized spines, posterior aspect with
|
||
<emphasis id="B922D319FF93733032B9FC93E208FC9B" box="[212,271,809,834]" italics="true" pageId="96" pageNumber="97">circa</emphasis>
|
||
10 small, sclerotized teeth, anterior aspect apically with four minute, non-sclerotized teeth. Fixed finger shorter and more slender than movable finger. Fixed finger tip weakly curved, anterior side basally with single, short, sclerotized spine, apically with numerous sclerotized spots. Fixed finger on both sides covered with few long hairs. Movable finger on posterior side medially with few hairs, remaining parts of posterior telopods glabrous. Inner horns of syncoxite apically curved posteriorly. Tips sharp, whole horns covered with numerous short hairs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |