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<document id="EEAD6914132E632ECD12CBE9FCFA9160" ID-CLB-Dataset="24671" ID-DOI="10.11646/zootaxa.4706.3.4" ID-GBIF-Dataset="47f9ebfa-4250-4218-9613-c1d1398ca33b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3568896" ID-ZooBank="29429537-8240-4E24-B926-6F23A7BD151E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575968595120" checkinUser="plazi" docAuthor="Farahi, Sara, Shishehbor, Parviz, Nemati, Alireza &amp; Witaliński, Wojciech" docDate="2019" docId="03C71F47FFD0FF9EDABCC2A7FA4AF91F" docLanguage="en" docName="zootaxa.4706.3.4.pdf" docOrigin="Zootaxa 4706 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trachygamasus Berlese 1906" docType="key" docVersion="6" lastPageNumber="448" masterDocId="FFFE673FFFD9FF97DA2BC332FFD6FFB0" masterDocTitle="Trachygamasus karuni sp. nov., a new mite species from Iran (Parasitiformes: Parasitidae)" masterLastPageNumber="450" masterPageNumber="439" pageNumber="448" updateTime="1698773684450" updateUser="ExternalLinkService">
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<mods:title id="CF41286DC90D6C5CC67A286571207164">Trachygamasus karuni sp. nov., a new mite species from Iran (Parasitiformes: Parasitidae)</mods:title>
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<mods:namePart id="6036849FC5DA7D64A002B62A02714F27">Farahi, Sara</mods:namePart>
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<mods:namePart id="1E792D07B1E99D9C397C17E8E39852DD">Shishehbor, Parviz</mods:namePart>
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<mods:namePart id="69BF71C2CBB845A53DB4FC4D587B252C">Nemati, Alireza</mods:namePart>
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<mods:namePart id="CBE82186A2906F0C1479E057D711AB14">Witaliński, Wojciech</mods:namePart>
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<mods:title id="9E1231B7D23837B67F5AE68493A73D7B">Zootaxa</mods:title>
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<mods:number id="42B7D77ADB97DE3B1B5C2F08FA30C9A8">2019-12-10</mods:number>
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<treatment id="03C71F47FFD0FF9EDABCC2A7FA4AF91F" ID-GBIF-Taxon="160687973" LSID="urn:lsid:plazi:treatment:03C71F47FFD0FF9EDABCC2A7FA4AF91F" httpUri="http://treatment.plazi.org/id/03C71F47FFD0FF9EDABCC2A7FA4AF91F" lastPageNumber="448" pageId="9" pageNumber="448">
<subSubSection id="C374FDDAFFD0FF9EDABCC2A7FF28FE47" pageId="9" pageNumber="448" type="nomenclature">
<paragraph id="8BD1AE51FFD0FF9EDABCC2A7FD66FE1F" blockId="9.[151,688,405,432]" box="[151,688,405,432]" pageId="9" pageNumber="448">
<heading id="D099193DFFD0FF9EDABCC2A7FD66FE1F" bold="true" box="[151,688,405,432]" fontSize="11" level="1" pageId="9" pageNumber="448" reason="1">
<emphasis id="B91A7243FFD0FF9EDABCC2A7FD66FE1F" bold="true" box="[151,688,405,432]" pageId="9" pageNumber="448">
Key to the adults of
<taxonomicName id="4C6ED5D2FFD0FF9EDBB8C2A7FD84FE00" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[403,594,405,432]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B91A7243FFD0FF9EDBB8C2A7FD84FE00" bold="true" box="[403,594,405,432]" italics="true" pageId="9" pageNumber="448">Trachygamasus</emphasis>
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species
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<paragraph id="8BD1AE51FFD0FF9EDABCC2EFFF28FE47" blockId="9.[151,254,477,503]" box="[151,254,477,503]" pageId="9" pageNumber="448">
<heading id="D099193DFFD0FF9EDABCC2EFFF28FE47" bold="true" box="[151,254,477,503]" fontSize="11" level="1" pageId="9" pageNumber="448" reason="1">
<emphasis id="B91A7243FFD0FF9EDABCC2EFFF28FE47" bold="true" box="[151,254,477,503]" pageId="9" pageNumber="448">Females</emphasis>
</heading>
</paragraph>
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<subSubSection id="C374FDDAFFD0FF9EDABCC110FA4AF91F" pageId="9" pageNumber="448" type="key">
<keyStep id="309AB6F4FFD0FF9EDABCC110FA4AFDE4" pageId="9" pageNumber="448">
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<keyLead id="309F0D64FFD0FF9EDABCC110FA4AFD87" box="[151,1436,545,567]" pageId="9" pageNumber="448">1. Central prong of gnathotectum distally widened and bifid or trifid apically........................................ 2</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC10DFA4AFDE4" blockId="9.[151,1437,545,1711]" box="[151,1436,575,597]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC10DFA4AFDE4" box="[151,1436,575,597]" pageId="9" pageNumber="448">- Central prong of gnathotectum acuminate, terminally pointed or rounded......................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC16EFA4AFD3F" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC16EFA4AFDC2" blockId="9.[151,1437,545,1711]" box="[151,1436,604,626]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC16EFA4AFDC2" box="[151,1436,604,626]" pageId="9" pageNumber="448">
2. All dorsal setae simple, seta
<emphasis id="B91A7243FFD0FF9EDBDFC16EFDD1FDC1" box="[500,519,604,625]" italics="true" pageId="9" pageNumber="448">al</emphasis>
on palpfemur bifid......................................
<taxonomicName id="4C6ED5D2FFD0FF9EDEBAC16FFA4AFDC2" authority="Costa, 1962" authorityName="Costa" authorityYear="1962" box="[1169,1436,604,626]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="macfarlanei">
<emphasis id="B91A7243FFD0FF9EDEBAC16FFB76FDC2" bold="true" box="[1169,1184,605,626]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE87C16EFAF2FDC2" bold="true" box="[1196,1316,604,626]" italics="true" pageId="9" pageNumber="448">macfarlanei</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDF01C16EFA4AFDC2" author="Costa, M." box="[1322,1436,604,626]" pageId="9" pageNumber="448" pagination="481 - 503" refId="ref7173" refString="Costa, M. (1962) Mites from the nests of the mole-rat (Spalax ehrenbergi) in Israel. Annals &amp; Magazine of Natural History, Series 13, 4, 481 - 503. https: // doi. org / 10.1080 / 00222936108651166" type="journal article" year="1962">Costa, 1962</bibRefCitation>
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<paragraph id="8BD1AE51FFD0FF9EDABCC148FA4AFD3F" blockId="9.[151,1437,545,1711]" box="[151,1436,633,655]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC148FA4AFD3F" box="[151,1436,633,655]" pageId="9" pageNumber="448">
- Some dorsal setae pilose, seta
<emphasis id="B91A7243FFD0FF9ED82DC148FDCFFD3F" box="[518,537,634,655]" italics="true" pageId="9" pageNumber="448">al</emphasis>
on palpfemur spatulate....................................................... 3
</keyLead>
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<keyStep id="309AB6F4FFD0FF9EDABCC1A5FA4AFD79" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC1A5FA4AFD1D" blockId="9.[151,1437,545,1711]" box="[151,1436,663,685]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC1A5FA4AFD1D" box="[151,1436,663,685]" pageId="9" pageNumber="448">
3. Dorsal setae
<emphasis id="B91A7243FFD0FF9EDB75C1A5FEA7FD1C" box="[350,369,663,684]" italics="true" pageId="9" pageNumber="448">j1</emphasis>
feathered...........................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE89C1A5FA4AFD1D" authority="Holzmann, 1969" authorityName="Holzmann" authorityYear="1969" box="[1186,1436,663,685]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B91A7243FFD0FF9EDE89C1A5FB67FD1C" bold="true" box="[1186,1201,663,684]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE96C1A5FB20FD1D" bold="true" box="[1213,1270,663,685]" italics="true" pageId="9" pageNumber="448">fallax</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDED7C1A5FA4AFD1D" author="Holzmann, C." box="[1276,1436,663,685]" pageId="9" pageNumber="448" pagination="3 - 24" refId="ref7332" refString="Holzmann, C. (1969) Die Familie Parasitidae Oudemans, 1901 (Eine systematische Studie aus dem Jahre 1955). Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 13, 3 - 24 + 25 - 55, 23 pls." type="book chapter" year="1969">Holzmann, 1969</bibRefCitation>
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</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC187FA4AFD79" blockId="9.[151,1437,545,1711]" box="[151,1436,692,714]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC187FA4AFD79" box="[151,1436,692,714]" pageId="9" pageNumber="448">
- Dorsal setae
<emphasis id="B91A7243FFD0FF9EDB75C186FEA7FD79" box="[350,369,692,713]" italics="true" pageId="9" pageNumber="448">j1</emphasis>
simple.................................................................................. 4
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</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC1E0FA4AFCB5" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC1E0FA4AFD57" blockId="9.[151,1437,545,1711]" box="[151,1436,721,743]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC1E0FA4AFD57" box="[151,1436,721,743]" pageId="9" pageNumber="448">
4. Opisthonotum with 21 pairs of setae..................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE8AC1E0FA4AFD57" authority="Karg, 1998" authorityName="Karg" authorityYear="1998" box="[1185,1436,721,743]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="biplumatus">
<emphasis id="B91A7243FFD0FF9EDE8AC1E0FB66FD57" bold="true" box="[1185,1200,722,743]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE97C1E3FAFDFD57" bold="true" box="[1212,1323,721,743]" italics="true" pageId="9" pageNumber="448">biplumatus</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDF1AC1E0FA4AFD57" author="Karg, W." box="[1329,1436,722,743]" pageId="9" pageNumber="448" pagination="185 - 214" refId="ref7693" refString="Karg, W. (1998) Zur Kenntnis der Eugamasides Karg mit neuen Arten aus den Regenwaldern von Ecuador (Acarina, Parasiti- formes). Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologische Reihe, 74, 185 - 214. https: // doi. org / 10.1002 / mmnz. 19980740204" type="journal article" year="1998">Karg, 1998</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC1DDFA4AFCB5" blockId="9.[151,1437,545,1711]" box="[151,1436,751,773]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC1DDFA4AFCB5" box="[151,1436,751,773]" pageId="9" pageNumber="448">- Opisthonotum with 12–18 pairs of setae................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC03EFA4AFC8F" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC03EFA4AFC91" blockId="9.[151,1437,545,1711]" box="[151,1436,780,802]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC03EFA4AFC91" box="[151,1436,780,802]" pageId="9" pageNumber="448">
5. Opisthonotum much narrow than podonotum, with 12 pairs of setae.................
<taxonomicName id="4C6ED5D2FFD0FF9EDE6AC03FFA4AFC91" authority="Yao, Jin &amp; Zhang, 2019" authorityName="Yao, Jin &amp; Zhang" authorityYear="2019" box="[1089,1436,780,802]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="hyalinus">
<emphasis id="B91A7243FFD0FF9EDE6AC03FFB86FC92" bold="true" box="[1089,1104,781,802]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE70C03EFB66FC92" bold="true" box="[1115,1200,780,802]" italics="true" pageId="9" pageNumber="448">hyalinus</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDE9CC03FFA4AFC91" author="Yao, M. &amp; Yi, T. - C. &amp; Guo, J. - J. &amp; Jin, D. - C. &amp; Zhang, R. - Z." box="[1207,1436,780,802]" pageId="9" pageNumber="448" pagination="1465 - 1489" refId="ref8231" refString="Yao, M., Yi, T. - C., Guo, J. - J., Jin, D. - C. &amp; Zhang, R. - Z. (2019) Three new species of Trachygamasus (Mesostigmata: Parasitidae) from China, with a key to world species of the genus. Systematic and Applied Acarology, 24, 1465 - 1489. https: // doi. org / 10.11158 / saa. 24.8.9" type="journal article" year="2019">Yao, Jin &amp; Zhang, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC018FA4AFC8F" blockId="9.[151,1437,545,1711]" box="[151,1436,809,831]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC018FA4AFC8F" box="[151,1436,809,831]" pageId="9" pageNumber="448">- Opisthonotal width similar to width of podonotum, with 14-18 pairs of setae...................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC075FA4AFC27" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC075FA4AFCC9" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC075FA4AFCC9" pageId="9" pageNumber="448">
6. Posterior margin of podonotum with convexity protruding into concavity in opisthonotal margin; opisthonotum with 14 pairs of setae................................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE1AC057FA4AFCC9" authority="Yao, Jin &amp; Zhang, 2019" authorityName="Yao, Jin &amp; Zhang" authorityYear="2019" box="[1073,1436,868,890]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="multisetus">
<emphasis id="B91A7243FFD0FF9EDE1AC057FB96FCCA" bold="true" box="[1073,1088,869,890]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE60C056FB66FCCA" bold="true" box="[1099,1200,868,890]" italics="true" pageId="9" pageNumber="448">multisetus</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDE9CC057FA4AFCC9" author="Yao, M. &amp; Yi, T. - C. &amp; Guo, J. - J. &amp; Jin, D. - C. &amp; Zhang, R. - Z." box="[1207,1436,868,890]" pageId="9" pageNumber="448" pagination="1465 - 1489" refId="ref8231" refString="Yao, M., Yi, T. - C., Guo, J. - J., Jin, D. - C. &amp; Zhang, R. - Z. (2019) Three new species of Trachygamasus (Mesostigmata: Parasitidae) from China, with a key to world species of the genus. Systematic and Applied Acarology, 24, 1465 - 1489. https: // doi. org / 10.11158 / saa. 24.8.9" type="journal article" year="2019">Yao, Jin &amp; Zhang, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC0B0FA4AFC27" blockId="9.[151,1437,545,1711]" box="[151,1436,897,919]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC0B0FA4AFC27" box="[151,1436,897,919]" pageId="9" pageNumber="448">- Posterior margin of podonotum straight or only sinuous; opisthonotum with 14–18 pairs of setae...................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC0ADFA4AFBBD" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC0ADFA4AFC62" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC0ADFA4AFC62" pageId="9" pageNumber="448">
7. Length of idiosoma less than 500, opisthonotum with 14 pairs of setae, endogynium mushroom-shaped in outline, comprised of two circular parts as in
<taxonomicName id="4C6ED5D2FFD0FF9EDBF7C08FFDA3FC61" baseAuthorityName="Willmann" baseAuthorityYear="1949" box="[476,629,956,978]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="ambulacralis">
<emphasis id="B91A7243FFD0FF9EDBF7C08FFE3FFC62" box="[476,489,957,978]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDBDDC08EFDA3FC61" box="[502,629,956,977]" italics="true" pageId="9" pageNumber="448">ambulacralis</emphasis>
</taxonomicName>
<emphasis id="B91A7243FFD0FF9ED8A8C08FFCDBFC62" box="[643,781,957,978]" italics="true" pageId="9" pageNumber="448">............</emphasis>
<taxonomicName id="4C6ED5D2FFD0FF9ED936C08FFBD5FC62" authority="(Karg, 1965)" authorityName="Karg" authorityYear="1965" baseAuthorityName="Karg" baseAuthorityYear="1965" box="[797,1027,956,978]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B91A7243FFD0FF9ED936C08FFCFAFC62" bold="true" box="[797,812,957,978]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9ED913C08EFC57FC62" bold="true" box="[824,897,956,978]" italics="true" pageId="9" pageNumber="448">gracilis</emphasis>
(
<bibRefCitation id="EFFFD3A0FFD0FF9ED9A5C08EFC2AFC62" author="Karg, W." box="[910,1020,956,978]" pageId="9" pageNumber="448" pagination="193 - 340" refId="ref7558" refString="Karg, W. (1965) Larvalsystematische und phylogenetische Untersuchungen sowie Revision des Systems der Gamasina Leach, 1815 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 41, 193 - 340. https: // doi. org / 10.1002 / mmnz. 19650410207" type="journal article" year="1965">Karg, 1965</bibRefCitation>
)
</taxonomicName>
(=
<taxonomicName id="4C6ED5D2FFD0FF9EDE0FC08FFAF6FC62" authority="Holzmann, 1969" authorityName="Holzmann" authorityYear="1969" box="[1060,1312,956,978]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B91A7243FFD0FF9EDE0FC08FFBE7FC62" box="[1060,1073,957,978]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE16C08FFBA0FC62" box="[1085,1142,957,978]" italics="true" pageId="9" pageNumber="448">minor</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDE56C08EFAF6FC62" author="Holzmann, C." box="[1149,1312,956,978]" pageId="9" pageNumber="448" pagination="3 - 24" refId="ref7332" refString="Holzmann, C. (1969) Die Familie Parasitidae Oudemans, 1901 (Eine systematische Studie aus dem Jahre 1955). Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 13, 3 - 24 + 25 - 55, 23 pls." type="book chapter" year="1969">Holzmann, 1969</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFFFD3A0FFD0FF9EDF02C08FFA43FC62" author="Karg, W." box="[1321,1429,956,978]" pageId="9" pageNumber="448" pagination="1 - 475" refId="ref7610" refString="Karg, W. (1971) Acari (Acarina), Milben Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasi- des), Raubmilben. Die Tierwelt Deutschlands, 59, 1 - 475." type="journal article" year="1971">Karg, 1971</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC0E8FA4AFBBD" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC0E8FA4AFBBD" pageId="9" pageNumber="448">
- Length of idiosoma exceeding 600, opisthonotum with 15–18 pairs of setae, endogynium ovate in outline, comprised of two circular parts, fused axially and forming the figure “8”.............................
<taxonomicName id="4C6ED5D2FFD0FF9EDE64C0C5FA4AFBBD" authority="(Willmann, 1949)" authorityName="Willmann" authorityYear="1949" baseAuthorityName="Willmann" baseAuthorityYear="1949" box="[1103,1436,1015,1037]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="ambulacralis">
<emphasis id="B91A7243FFD0FF9EDE64C0C5FB88FBBC" bold="true" box="[1103,1118,1015,1036]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE41C0C5FB3DFBBD" bold="true" box="[1130,1259,1015,1037]" italics="true" pageId="9" pageNumber="448">ambulacralis</emphasis>
(
<bibRefCitation id="EFFFD3A0FFD0FF9EDED3C0C5FA43FBBD" author="Willmann, C." box="[1272,1429,1015,1037]" pageId="9" pageNumber="448" pagination="136 - 142" refId="ref8139" refString="Willmann, C. (1949) Uber eine eigenartige Milbenfauna an faulender Kartoffeln. Veroffentlichungen aus dem Museum fur Natur - Volker - Handelskunde Bremen, A 1, 136 - 142." type="journal article" year="1949">Willmann, 1949</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC726FA4AFB31" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC726FA4AFBF7" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC726FA4AFBF7" pageId="9" pageNumber="448">8. Epigynium regular (equilateral) triangle with straight lateral margins; gnathotectum central prong narrow, similar to lateral ones................................................................................................ 9</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC77DFA4AFB31" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC77DFA4AFB31" pageId="9" pageNumber="448">- Epigynium bell-shaped, with sinuous lateral margins, central prong of gnathotectum broader and more solid than the lateral prongs............................................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC7B8FA4AFB47" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC7B8FA4AFB0C" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC7B8FA4AFB0C" pageId="9" pageNumber="448">
9. Sternal setae
<emphasis id="B91A7243FFD0FF9EDB4AC7B8FEA8FB2F" box="[353,382,1162,1183]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
simple and thin, distance between
<emphasis id="B91A7243FFD0FF9ED8E7C7B8FD3FFB2F" box="[716,745,1162,1183]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
greater than between
<emphasis id="B91A7243FFD0FF9ED990C7B8FC01FB2F" box="[955,983,1162,1183]" italics="true" pageId="9" pageNumber="448">st1</emphasis>
; opisthogaster with seven pairs of setae, length of idiosoma 670–700..........................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE5AC795FA4AFB0C" authority="Tichomirov, 1977" authorityName="Tichomirov" authorityYear="1977" box="[1137,1436,1191,1213]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B91A7243FFD0FF9EDE5AC795FB56FB0C" bold="true" box="[1137,1152,1191,1212]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDEA7C795FB3AFB0D" bold="true" box="[1164,1260,1191,1213]" italics="true" pageId="9" pageNumber="448">medianus</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDEDAC795FA4AFB0C" author="Tichomirov, S. I." box="[1265,1436,1191,1213]" pageId="9" pageNumber="448" pagination="55 - 107" refId="ref8052" refString="Tichomirov, S. I. (1977) The family Parasitidae. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil Inhabiting Mites, Mesostigmata. Nauka, Leningrad, pp. 55 - 107. [in Russian]" type="book chapter" year="1977">Tichomirov, 1977</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC7F7FA4AFB47" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC7F7FA4AFB47" pageId="9" pageNumber="448">
- Sternal setae
<emphasis id="B91A7243FFD0FF9EDB4BC7F6FEAAFB69" box="[352,380,1220,1241]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
stout, distance between
<emphasis id="B91A7243FFD0FF9ED848C7F6FDA9FB69" box="[611,639,1220,1241]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
smaller than between
<emphasis id="B91A7243FFD0FF9ED979C7F6FCB8FB69" box="[850,878,1220,1241]" italics="true" pageId="9" pageNumber="448">st1</emphasis>
; opisthogaster with eight pairs of setae, length of idiosoma
<emphasis id="B91A7243FFD0FF9EDAC9C7D0FF2FFB47" box="[226,249,1250,1271]" italics="true" pageId="9" pageNumber="448">ca</emphasis>
. 560..........................................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE82C7D0FA4AFB47" authority="Karg, 1978" authorityName="Karg" authorityYear="1978" box="[1193,1436,1249,1271]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B91A7243FFD0FF9EDE82C7D0FB6EFB47" bold="true" box="[1193,1208,1250,1271]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDEEFC7D3FAFDFB47" bold="true" box="[1220,1323,1249,1271]" italics="true" pageId="9" pageNumber="448">triangulus</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDF1AC7D0FA4AFB47" author="Karg, W." box="[1329,1436,1250,1271]" pageId="9" pageNumber="448" pagination="347 - 359" refId="ref7649" refString="Karg, W. (1978) Zur Kenntnis der Milbengattungen Parasitus Latreille, 1795 und Trachygamasus Berlese, 1906 (Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 105, 347 - 359." type="journal article" year="1978">Karg, 1978</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC7CDFA4AFADC" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC7CDFA4AFA82" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC7CDFA4AFA82" pageId="9" pageNumber="448">
10. Distance between sternal setae
<emphasis id="B91A7243FFD0FF9ED825C7CDFDFDFAA4" box="[526,555,1279,1300]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
evidently greater than between
<emphasis id="B91A7243FFD0FF9ED975C7CDFCACFAA4" box="[862,890,1279,1300]" italics="true" pageId="9" pageNumber="448">st3</emphasis>
; opisthogastric setae
<emphasis id="B91A7243FFD0FF9EDE62C7CDFBB9FAA4" box="[1097,1135,1279,1300]" italics="true" pageId="9" pageNumber="448">JV4</emphasis>
simple, similar to other ventral setae; epigynium width greater than length..........................
<taxonomicName id="4C6ED5D2FFD0FF9ED992C62FFA4AFA82" authority="(Berlese, 1892) sensu Holzmann, 1969" authorityName="sensu Holzmann" authorityYear="1969" baseAuthorityName="Berlese" baseAuthorityYear="1892" box="[953,1436,1308,1330]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B91A7243FFD0FF9ED992C62FFC1EFA82" bold="true" box="[953,968,1309,1330]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9ED9FFC62EFBF7FA82" bold="true" box="[980,1057,1308,1330]" italics="true" pageId="9" pageNumber="448">pusillus</emphasis>
(
<bibRefCitation id="EFFFD3A0FFD0FF9EDE05C62EFB62FA82" author="Berlese, A." box="[1070,1204,1308,1330]" pageId="9" pageNumber="448" pagination="1 - 10" refId="ref7131" refString="Berlese, A. (1892) Acari, Myriapoda et Scorpiones hucusque in Italia reperta, 69, 22 (text) pp., pls. 1 - 10. [Reprint by Junk, The Hague, 1979]" type="journal article" year="1892">Berlese, 1892</bibRefCitation>
)
<emphasis id="B91A7243FFD0FF9EDEEAC62FFB20FA82" box="[1217,1270,1309,1330]" italics="true" pageId="9" pageNumber="448">sensu</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDED7C62EFA4AFA82" author="Holzmann, C." box="[1276,1436,1308,1330]" pageId="9" pageNumber="448" pagination="3 - 24" refId="ref7332" refString="Holzmann, C. (1969) Die Familie Parasitidae Oudemans, 1901 (Eine systematische Studie aus dem Jahre 1955). Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 13, 3 - 24 + 25 - 55, 23 pls." type="book chapter" year="1969">Holzmann, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC608FA4AFADC" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC608FA4AFADC" pageId="9" pageNumber="448">
- Distance between sternal setae
<emphasis id="B91A7243FFD0FF9ED83BC608FDFAFAFF" box="[528,556,1338,1359]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
similar to that between
<emphasis id="B91A7243FFD0FF9ED93EC608FCE7FAFF" box="[789,817,1338,1359]" italics="true" pageId="9" pageNumber="448">st3</emphasis>
; opisthogastric setae
<emphasis id="B91A7243FFD0FF9EDE29C608FBF1FAFF" box="[1026,1063,1338,1359]" italics="true" pageId="9" pageNumber="448">JV4</emphasis>
enlarged and terminally pilose; epigynium width not greater than length....................................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC646FA4AFA17" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC646FA4AFA39" blockId="9.[151,1437,545,1711]" box="[151,1436,1396,1418]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC646FA4AFA39" box="[151,1436,1396,1418]" pageId="9" pageNumber="448">
11. Paragynal poroid
<emphasis id="B91A7243FFD0FF9EDBA7C646FE7CFA39" box="[396,426,1396,1417]" italics="true" pageId="9" pageNumber="448">
<date id="FFD08891FFD0FF9EDBA7C646FE7CFA39" box="[396,426,1396,1417]" pageId="9" pageNumber="448">iv3</date>
</emphasis>
circular rather than elongate................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE15C647FA4AFA39" authority="Farahi &amp; Witalinski, 1919" authorityName="Farahi &amp; Witalinski" authorityYear="1919" box="[1086,1436,1396,1418]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="karuni">
<emphasis id="B91A7243FFD0FF9EDE15C647FB9BFA3A" bold="true" box="[1086,1101,1397,1418]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE72C646FB4AFA3A" bold="true" box="[1113,1180,1396,1418]" italics="true" pageId="9" pageNumber="448">karuni</emphasis>
Farahi &amp; Witaliński, 1919
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC6A0FA4AFA17" blockId="9.[151,1437,545,1711]" box="[151,1436,1425,1447]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC6A0FA4AFA17" box="[151,1436,1425,1447]" pageId="9" pageNumber="448">
- Paragynal poroid
<emphasis id="B91A7243FFD0FF9EDBA7C6A0FE7CFA17" box="[396,426,1426,1447]" italics="true" pageId="9" pageNumber="448">
<date id="FFD08891FFD0FF9EDBA7C6A0FE7CFA17" box="[396,426,1426,1447]" pageId="9" pageNumber="448">iv3</date>
</emphasis>
elongate, similar in shape to poroids
<date id="FFD08891FFD0FF9ED939C6A0FC53FA17" box="[786,901,1425,1447]" pageId="9" pageNumber="448">
<emphasis id="B91A7243FFD0FF9ED939C6A0FCF9FA17" box="[786,815,1426,1447]" italics="true" pageId="9" pageNumber="448">iv1</emphasis>
and
<emphasis id="B91A7243FFD0FF9ED975C6A0FCADFA17" box="[862,891,1426,1447]" italics="true" pageId="9" pageNumber="448">iv2</emphasis>
.
</date>
.......................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC69DFA4AFA4F" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC69DFA4AFA52" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC69DFA4AFA52" pageId="9" pageNumber="448">
12. Endogynium narrow, tubular, with separate elliptic anterior part,
<emphasis id="B91A7243FFD0FF9ED967C682FCB5FA75" box="[844,867,1456,1477]" italics="true" pageId="9" pageNumber="448">ca</emphasis>
. 90 μm long, extending nearly along the entire length of the epigynium....................................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE50C6FFFA4AFA52" authority="Ma &amp; Wang, 1996" authorityName="Ma &amp; Wang" authorityYear="1996" box="[1147,1436,1484,1506]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B91A7243FFD0FF9EDE50C6FFFB5CFA52" bold="true" box="[1147,1162,1485,1506]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDEBDC6FEFB33FA52" bold="true" box="[1174,1253,1484,1506]" italics="true" pageId="9" pageNumber="448">borealis</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDEC0C6FFFA4AFA52" author="Ma, L. - M. &amp; Wang, S." box="[1259,1436,1484,1506]" pageId="9" pageNumber="448" pagination="81 - 88" refId="ref7836" refString="Ma, L. - M. &amp; Wang, S. (1996) Four new species of the family Parasitidae and discovery of the genus Trachygamasus in China. Acta Arachnologica Sinica, 5, 81 - 88. [in Chinese, with English abstract]" type="journal article" year="1996">Ma &amp; Wang, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC6D8FA4AFA4F" blockId="9.[151,1437,545,1711]" box="[151,1436,1513,1535]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC6D8FA4AFA4F" box="[151,1436,1513,1535]" pageId="9" pageNumber="448">- Endogynium cross-shaped, shorter than epigynium length.................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="309AB6F4FFD0FF9EDABCC535FA4AF91F" pageId="9" pageNumber="448">
<paragraph id="8BD1AE51FFD0FF9EDABCC535FA4AF9E7" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC535FA4AF9E7" pageId="9" pageNumber="448">
13. Podonotum and opisthonotum fused, setae
<emphasis id="B91A7243FFD0FF9ED8A8C535FD40F9AC" box="[643,662,1543,1564]" italics="true" pageId="9" pageNumber="448">j1</emphasis>
and
<emphasis id="B91A7243FFD0FF9ED8EDC535FD0DF9AC" box="[710,731,1543,1564]" italics="true" pageId="9" pageNumber="448">z1</emphasis>
on podonotal shield; sternal setae
<emphasis id="B91A7243FFD0FF9EDE07C535FB9EF9AC" box="[1068,1096,1543,1564]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
somewhat thicker and shorter than
<emphasis id="B91A7243FFD0FF9EDAC9C516FF28F989" box="[226,254,1572,1593]" italics="true" pageId="9" pageNumber="448">st1</emphasis>
; endogynium with open arms,
<emphasis id="B91A7243FFD0FF9ED803C517FDE9F98A" box="[552,575,1573,1594]" italics="true" pageId="9" pageNumber="448">ca</emphasis>
. 58 long, its length not exceeding half of the length of the epigynium................................................................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE81C570FA4AF9E7" authority="Witalinski, 1917" authorityName="Witalinski" authorityYear="1917" box="[1194,1436,1601,1623]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="gerdi">
<emphasis id="B91A7243FFD0FF9EDE81C570FB6FF9E7" bold="true" box="[1194,1209,1602,1623]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDEEEC573FB2EF9E7" bold="true" box="[1221,1272,1601,1623]" italics="true" pageId="9" pageNumber="448">gerdi</emphasis>
Witaliński, 1917
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1AE51FFD0FF9EDABCC56DFA4AF91F" blockId="9.[151,1437,545,1711]" pageId="9" pageNumber="448">
<keyLead id="309F0D64FFD0FF9EDABCC56DFA4AF91F" pageId="9" pageNumber="448">
- Podonotum and opisthonotum separate, setae
<emphasis id="B91A7243FFD0FF9ED88BC56DFD65F9C4" box="[672,691,1631,1652]" italics="true" pageId="9" pageNumber="448">j1</emphasis>
and
<emphasis id="B91A7243FFD0FF9ED8CFC56DFD2FF9C4" box="[740,761,1631,1652]" italics="true" pageId="9" pageNumber="448">
z
<quantity id="4C9603B4FFD0FF9ED8C4C56DFD2FF9C4" box="[751,761,1631,1652]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="448" unit="in" value="1.0">1</quantity>
</emphasis>
in front of podonotal shield; sternal setae
<emphasis id="B91A7243FFD0FF9EDEB2C56DFB63F9C4" box="[1177,1205,1631,1652]" italics="true" pageId="9" pageNumber="448">st2</emphasis>
clearly thicker and two times shorter than
<emphasis id="B91A7243FFD0FF9EDBBFC54EFE66F921" box="[404,432,1660,1681]" italics="true" pageId="9" pageNumber="448">st1</emphasis>
; endogynium with closed arms,
<emphasis id="B91A7243FFD0FF9ED8C2C54FFCD6F922" box="[745,768,1661,1682]" italics="true" pageId="9" pageNumber="448">ca</emphasis>
. 95 long, posteriorly ending at
<emphasis id="B91A7243FFD0FF9EDE06C54FFB9FF922" box="[1069,1097,1661,1682]" italics="true" pageId="9" pageNumber="448">st5</emphasis>
setae level.......................................................................................
<taxonomicName id="4C6ED5D2FFD0FF9EDE35C5A8FA4AF91F" authority="Yao, Jin &amp; Zhang, 2019" authorityName="Yao, Jin &amp; Zhang" authorityYear="2019" box="[1054,1436,1689,1711]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="pseudogerdi">
<emphasis id="B91A7243FFD0FF9EDE35C5A8FBFBF91F" bold="true" box="[1054,1069,1690,1711]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE12C5ABFB67F91F" bold="true" box="[1081,1201,1689,1711]" italics="true" pageId="9" pageNumber="448">pseudogerdi</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDE9CC5A8FA4AF91F" author="Yao, M. &amp; Yi, T. - C. &amp; Guo, J. - J. &amp; Jin, D. - C. &amp; Zhang, R. - Z." box="[1207,1436,1689,1711]" pageId="9" pageNumber="448" pagination="1465 - 1489" refId="ref8231" refString="Yao, M., Yi, T. - C., Guo, J. - J., Jin, D. - C. &amp; Zhang, R. - Z. (2019) Three new species of Trachygamasus (Mesostigmata: Parasitidae) from China, with a key to world species of the genus. Systematic and Applied Acarology, 24, 1465 - 1489. https: // doi. org / 10.11158 / saa. 24.8.9" type="journal article" year="2019">Yao, Jin &amp; Zhang, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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