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<document id="ACD85878FA7B80AA34C948E37EB3CEF8" ID-DOI="10.5281/zenodo.275861" ID-GBIF-Dataset="3498f3ee-e8eb-42cb-abc8-4492b21dc509" ID-ISSN="1175-5326" ID-Zenodo-Dep="275861" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460176750768" checkinUser="plazi" docAuthor="Scolaro, José Alejandro &amp; Pincheira-Donoso, Daniel" docDate="2010" docId="03C187F70D47FFCEFF32FDAEFE09FE48" docLanguage="en" docName="zt02393p032.pdf" docOrigin="Zootaxa 2393" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phymaturus castillensis Scolaro &amp; Pincheira-Donoso, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FFF8FF8F0D43FFC8FFA5FF83FFA9FFDC" masterDocTitle="Lizards at the end of the world: Two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina" masterLastPageNumber="32" masterPageNumber="17" pageNumber="21" updateTime="1698236093096" updateUser="plazi">
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<mods:title id="06345DCB879840F757A4B6B27607E485">Lizards at the end of the world: Two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina</mods:title>
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<mods:namePart id="F600522E5F052EA35BD0F850366547F8">Scolaro, José Alejandro</mods:namePart>
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<mods:namePart id="0615E44F06B4F5A2E1D565B1FD4EAE4C">Pincheira-Donoso, Daniel</mods:namePart>
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<mods:date id="6C693E1F4CA7C8EB8445DE0BE3CAE7D5">2010</mods:date>
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<treatment id="03C187F70D47FFCEFF32FDAEFE09FE48" ID-DOI="http://doi.org/10.5281/zenodo.5697590" ID-GBIF-Taxon="119383556" ID-Zenodo-Dep="5697590" LSID="urn:lsid:plazi:treatment:03C187F70D47FFCEFF32FDAEFE09FE48" httpUri="http://treatment.plazi.org/id/03C187F70D47FFCEFF32FDAEFE09FE48" lastPageId="6" lastPageNumber="23" pageId="4" pageNumber="21">
<subSubSection id="C372656A0D47FFCCFF32FDAEFEF4FDB6" pageId="4" pageNumber="21" type="nomenclature">
<paragraph id="8BD736E10D47FFCCFF32FDAEFDB2FD9B" blockId="4.[151,539,557,618]" box="[151,539,557,583]" pageId="4" pageNumber="21">
<heading id="D09F818D0D47FFCCFF32FDAEFDB2FD9B" bold="true" box="[151,539,557,583]" fontSize="11" level="1" pageId="4" pageNumber="21" reason="1">
<emphasis id="B91CEAF30D47FFCCFF32FDAEFDB2FD9B" bold="true" box="[151,539,557,583]" pageId="4" pageNumber="21">
<taxonomicName id="4C684D620D47FFCCFF32FDAEFE1FFD9B" box="[151,438,557,583]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="castillensis" status="sp. nov.">
<emphasis id="B91CEAF30D47FFCCFF32FDAEFE1FFD9B" bold="true" box="[151,438,557,583]" italics="true" pageId="4" pageNumber="21">Phymaturus castillensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57880D47FFCCFE18FDAEFDB2FD9B" box="[445,539,557,583]" pageId="4" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736E10D47FFCCFF32FDD3FEF4FDB6" blockId="4.[151,539,557,618]" box="[151,349,592,618]" pageId="4" pageNumber="21">
(
<figureCitation id="13532A640D47FFCCFF3AFDD3FEB9FDB6" box="[159,272,592,618]" captionStart="FIGURE 4" captionStartId="11.[151,255,1350,1374]" captionTargetBox="[151,1431,194,1326]" captionTargetId="figure@11.[151,1436,194,1329]" captionTargetPageId="11" captionText="FIGURE 4. Adult males and females of the two new species of Phymaturus of the patagonicus clade proposed in this study. Phymaturus castillensis holotype male (a) and paratype female (c) in dorsal view, and adult paratype male (e) in ventral view. All specimens from the type locality. Phymaturus videlai paratype male (b) and paratype female (d) in dorsal view, and female paratype (f) in ventral view. All from the type locality. (Photos: J. A. Scolaro, November 2008)." httpUri="https://zenodo.org/record/275865/files/figure.png" pageId="4" pageNumber="21">Figures 4</figureCitation>
a, c, e)
</paragraph>
</subSubSection>
<subSubSection id="C372656A0D47FFCCFF32FD19FD4BFD6D" box="[151,738,666,689]" pageId="4" pageNumber="21" type="description">
<paragraph id="8BD736E10D47FFCCFF32FD19FD4BFD6D" blockId="4.[151,738,666,689]" box="[151,738,666,689]" pageId="4" pageNumber="21">
<taxonomicName id="4C684D620D47FFCCFF32FD19FD88FD6D" authority="Cei" authorityName="Cei" box="[151,545,666,689]" class="Squamata" family="Liolaemidae" genus="Centrura" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="21" phylum="Chordata" rank="subSpecies" species="patagonica" subSpecies="patagonica">
<emphasis id="B91CEAF30D47FFCCFF32FD19FE5FFD6D" box="[151,502,666,689]" italics="true" pageId="4" pageNumber="21">Centrura patagonica patagonica</emphasis>
<bibRefCitation id="EFF94B100D47FFCCFE59FD19FD88FD6D" author="Cei" box="[508,545,666,689]" pageId="4" pageNumber="30" refString="Cei, J. M. (1986) Reptiles del centro, centro-oeste y sur de la Argentina. Herpetofauna de las zonas aridas y semiaridas. Museo Regionale di Scienze Naturali di Torino, Monografie, 4, 528 pp." type="book" year="1986">Cei</bibRefCitation>
</taxonomicName>
(1986: 182183).
</paragraph>
</subSubSection>
<subSubSection id="C372656A0D47FFCCFF32FD5CFB0EFC68" pageId="4" pageNumber="21" type="materials_examined">
<paragraph id="8BD736E10D47FFCCFF32FD5CFEF5FCB6" blockId="4.[151,1437,735,2031]" pageId="4" pageNumber="21">
<emphasis id="B91CEAF30D47FFCCFF32FD5CFEE1FD25" bold="true" box="[151,328,735,761]" pageId="4" pageNumber="21">
<typeStatus id="54D388430D47FFCCFF32FD5CFF7DFD25" box="[151,212,735,761]" pageId="4" pageNumber="21">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54D388430D47FFCCFEFFFD63FE66FD26" box="[346,463,736,762]" pageId="4" pageNumber="21" type="holotype">Holotype</typeStatus>
: MLP-R. 5441, adult male, collected in rocky outcrops (
<quantity id="4C909B040D47FFCCFBDEFD63FB60FD26" box="[1147,1225,736,762]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="21" unit="m" value="500.0">500 m</quantity>
asl) of Sierra del Castillo in La Juanita farm (
<geoCoordinate id="EE5C50260D47FFCCFE55FC86FDD4FCC3" box="[496,637,773,799]" direction="south" orientation="latitude" pageId="4" pageNumber="21" precision="15" value="-45.14167">45°08'30&quot;S</geoCoordinate>
,
<geoCoordinate id="EE5C50260D47FFCCFD29FC86FC8CFCC3" box="[652,805,773,799]" direction="west" orientation="longitude" pageId="4" pageNumber="21" precision="15" value="-69.17528">69°10'31&quot;W</geoCoordinate>
), adjacent to Provincial Road 24, at
<quantity id="4C909B040D47FFCCFB41FC86FA98FCC3" box="[1252,1329,773,799]" metricMagnitude="4" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="21" unit="km" value="58.0">58 km</quantity>
north of Sarmiento town, Chubut Province,
<collectingCountry id="F37F76710D47FFCCFDE3FCA9FD6AFC98" box="[582,707,810,836]" name="Argentina" pageId="4" pageNumber="21">Argentina</collectingCountry>
. Collected by J.A. Scolaro,
<collectingCountry id="F37F76710D47FFCCFBBCFCA9FB98FC98" box="[1049,1073,810,836]" name="Iceland" pageId="4" pageNumber="21">O</collectingCountry>
.F. Tappari and A. Marcus,
<date id="FFD610210D47FFCCFADBFCA9FEF1FCB6" pageId="4" pageNumber="21" value="2008-11-29">29 November 2008</date>
.
</paragraph>
<paragraph id="8BD736E10D47FFCCFF61FCF6FB0EFC68" blockId="4.[151,1437,735,2031]" pageId="4" pageNumber="21">
<typeStatus id="54D388430D47FFCCFF61FCF6FE97FC53" box="[196,318,885,911]" pageId="4" pageNumber="21" type="paratype">Paratypes</typeStatus>
: MLP-R 5442, adult male; MLP-R. 5443, adult female; MLP-R 5444 adult female; JAS-DC 1234 adult male; JAS-DC 1219 adult female. The same data as detailed for the
<typeStatus id="54D388430D47FFCCFB9CFC19FB0AFC68" box="[1081,1187,922,948]" pageId="4" pageNumber="21" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C372656A0D47FFCCFF61FC3CFB12FC06" box="[196,1211,959,986]" pageId="4" pageNumber="21" type="etymology">
<paragraph id="8BD736E10D47FFCCFF61FC3CFB12FC06" blockId="4.[151,1437,735,2031]" box="[196,1211,959,986]" pageId="4" pageNumber="21">
<emphasis id="B91CEAF30D47FFCCFF61FC3CFEE0FC05" bold="true" box="[196,329,959,985]" pageId="4" pageNumber="21">Etymology</emphasis>
. The species name refers to the terra typica where this species is restricted.
</paragraph>
</subSubSection>
<subSubSection id="C372656A0D47FFCCFF61FC66FABDFAA8" pageId="4" pageNumber="21" type="diagnosis">
<paragraph id="8BD736E10D47FFCCFF61FC66FABDFAA8" blockId="4.[151,1437,735,2031]" pageId="4" pageNumber="21">
<emphasis id="B91CEAF30D47FFCCFF61FC66FE95FC23" bold="true" box="[196,316,997,1023]" pageId="4" pageNumber="21">Diagnosis</emphasis>
.
<taxonomicName id="4C684D620D47FFCCFEEEFC66FDCAFC22" box="[331,611,997,1022]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D47FFCCFEEEFC66FDCAFC22" box="[331,611,997,1022]" italics="true" pageId="4" pageNumber="21">Phymaturus castillensis</emphasis>
</taxonomicName>
can be distinguished by a peculiar colour pattern similar to the colour patterns observed in females of some species of the
<taxonomicName id="4C684D620D47FFCCFCB8FB89FC1AFBFF" box="[797,947,1034,1059]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="patagonicus">
<emphasis id="B91CEAF30D47FFCCFCB8FB89FC1AFBFF" box="[797,947,1034,1059]" italics="true" pageId="4" pageNumber="21">patagonicus</emphasis>
</taxonomicName>
clade (
<figureCitation id="13532A640D47FFCCFBABFB89FBF0FBF8" box="[1038,1113,1034,1060]" captionStart="FIGURE 4" captionStartId="11.[151,255,1350,1374]" captionTargetBox="[151,1431,194,1326]" captionTargetId="figure@11.[151,1436,194,1329]" captionTargetPageId="11" captionText="FIGURE 4. Adult males and females of the two new species of Phymaturus of the patagonicus clade proposed in this study. Phymaturus castillensis holotype male (a) and paratype female (c) in dorsal view, and adult paratype male (e) in ventral view. All specimens from the type locality. Phymaturus videlai paratype male (b) and paratype female (d) in dorsal view, and female paratype (f) in ventral view. All from the type locality. (Photos: J. A. Scolaro, November 2008)." httpUri="https://zenodo.org/record/275865/files/figure.png" pageId="4" pageNumber="21">Fig. 4</figureCitation>
a, c). However, from most species, except
<emphasis id="B91CEAF30D47FFCCFEC1FBB3FE72FB95" box="[356,475,1072,1097]" italics="true" pageId="4" pageNumber="21">P. v i d e l a i</emphasis>
,
<taxonomicName id="4C684D620D47FFCCFE54FBB3FD33FB95" box="[497,666,1072,1097]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="indistictus">
<emphasis id="B91CEAF30D47FFCCFE54FBB3FD33FB95" box="[497,666,1072,1097]" italics="true" pageId="4" pageNumber="21">P. indistictus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D620D47FFCCFD7EFBB3FC33FB95" box="[731,922,1072,1097]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="patagonicus">
<emphasis id="B91CEAF30D47FFCCFD7EFBB3FC33FB95" box="[731,922,1072,1097]" italics="true" pageId="4" pageNumber="21">P. patagonicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D620D47FFCCFC09FBB3FBF6FB95" box="[940,1119,1072,1097]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D47FFCCFC09FBB3FBF6FB95" box="[940,1119,1072,1097]" italics="true" pageId="4" pageNumber="21">P. castillensis</emphasis>
</taxonomicName>
is considerably isolated geographically by hundreds of kilometers (
<figureCitation id="13532A640D47FFCCFD1FFBD6FCA1FBB3" box="[698,776,1109,1135]" captionStart="FIGURE 3" captionStartId="10.[151,255,1819,1843]" captionTargetBox="[215,1341,231,1768]" captionTargetId="figure@10.[211,1345,194,1798]" captionTargetPageId="10" captionText="FIGURE 3. Geographic distribution of all known members of the Phymaturus patagonicus clade. Note nonoverlapping geographical ranges of most species. Localities are indicated by numbers, and represent Phymaturus nevadoi (1), P. payuniae (2), P. zapalensis (3), P. tenebrosus (4), P. manuelae (5), P. spurcus (6), P. excelsus (7), P. spectabilis (8), P. agilis (9), P. ceii (10), P. somuncurensis (11), P. calcogaster (12), P. patagonicus (13), P. indistinctus (14), P. v i d e l a i sp. nov. (15), and P. castillensis sp. nov. (16). The wide and scattered geographical distribution shown for P. patagonicus has been considered as the result of several cryptic different species recognized until today as polymorphisms within this species, and many of them are currently under study in order to establish their status within Phymaturus." httpUri="https://zenodo.org/record/275864/files/figure.png" pageId="4" pageNumber="21">Fig. 3</figureCitation>
). From these three geographically closer species,
<taxonomicName id="4C684D620D47FFCCFA20FBD6FE88FB48" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D47FFCCFA20FBD6FE88FB48" italics="true" pageId="4" pageNumber="21">P. castillensis</emphasis>
</taxonomicName>
can be differentiated by its pattern of coloration (
<figureCitation id="13532A640D47FFCCFC2AFBF9FC4CFB48" box="[911,997,1146,1172]" captionStart="FIGURE 1" captionStartId="8.[151,256,1917,1941]" captionTargetBox="[292,1276,198,1906]" captionTargetId="figure@8.[291,1281,197,1909]" captionTargetPageId="8" captionText="FIGURE 1. Phymaturus species of the patagonicus clade, showing the remarkable diversity of patterns of coloration known within this lineage. Phymaturus agilis (a, male; b, female), P. calcogaster (c, male; d, female), P. ceii (e, male; f, female), P. excelsus (g, male; h, female), P. indistinctus (i, male; j, female), P. manuelae (k, male; l, female), and P. nevadoi (m, male; n, female). (Photos: all J. A. Scolaro, except P. nevadoi, by J. M. Cei)." httpUri="https://zenodo.org/record/275862/files/figure.png" pageId="4" pageNumber="21">Figs. 1</figureCitation>
,
<figureCitation id="13532A640D47FFCCFC53FBF9FBA1FB48" box="[1014,1032,1146,1172]" captionStart="FIGURE 2" captionStartId="9.[151,255,1914,1938]" captionTargetBox="[301,1285,195,1904]" captionTargetId="figure@9.[300,1287,194,1906]" captionTargetPageId="9" captionText="FIGURE 2. Phymaturus species of the patagonicus clade (continued from figure 1). Phymaturus patagonicus (a, male; b, female), P. payuniae (c, male; d, female), P. somuncurensis (e, male; f, female), P. spectabilis (g, male; h, female), P. spurcus (i, male; j, female), P. tenebrosus (k, male; l, female), and P. zapalensis (m, male; n, female). (Photos: J. A. Scolaro)." httpUri="https://zenodo.org/record/275863/files/figure.png" pageId="4" pageNumber="21">2</figureCitation>
,
<figureCitation id="13532A640D47FFCCFBB3FBF9FB8FFB48" box="[1046,1062,1146,1172]" captionStart="FIGURE 4" captionStartId="11.[151,255,1350,1374]" captionTargetBox="[151,1431,194,1326]" captionTargetId="figure@11.[151,1436,194,1329]" captionTargetPageId="11" captionText="FIGURE 4. Adult males and females of the two new species of Phymaturus of the patagonicus clade proposed in this study. Phymaturus castillensis holotype male (a) and paratype female (c) in dorsal view, and adult paratype male (e) in ventral view. All specimens from the type locality. Phymaturus videlai paratype male (b) and paratype female (d) in dorsal view, and female paratype (f) in ventral view. All from the type locality. (Photos: J. A. Scolaro, November 2008)." httpUri="https://zenodo.org/record/275865/files/figure.png" pageId="4" pageNumber="21">4</figureCitation>
) as well as the morphological differences detailed in the previous paragraph. In this new species there are no dichromatic differences between the sexes. The new species is a member of the
<taxonomicName id="4C684D620D47FFCCFC89FB46FC12FB02" box="[812,955,1221,1246]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="patagonicus">
<emphasis id="B91CEAF30D47FFCCFC89FB46FC12FB02" box="[812,955,1221,1246]" italics="true" pageId="4" pageNumber="21">patagonicus</emphasis>
</taxonomicName>
group, distinguished from the
<taxonomicName id="4C684D620D47FFCCFA8DFB46FA35FB02" box="[1320,1436,1221,1246]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="flagellifer">
<emphasis id="B91CEAF30D47FFCCFA8DFB46FA35FB02" box="[1320,1436,1221,1246]" italics="true" pageId="4" pageNumber="21">flagellifer</emphasis>
</taxonomicName>
group in having flat imbricate superciliaries rather than being rectangular and non-overlapping; slightly spiny and non-rugose caudal scales in verticilles (as seen among members of the
<taxonomicName id="4C684D620D47FFCCFBAEFA93FBD6FAF5" box="[1035,1151,1296,1321]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="flagellifer">
<emphasis id="B91CEAF30D47FFCCFBAEFA93FBD6FAF5" box="[1035,1151,1296,1321]" italics="true" pageId="4" pageNumber="21">flagellifer</emphasis>
</taxonomicName>
group). However, it has also the subocular scale not fragmented and separated from supralabials by two rows of lorilabials, as in most species of the
<taxonomicName id="4C684D620D47FFCCFE98FAD8FE65FAA8" box="[317,460,1371,1396]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="patagonicus">
<emphasis id="B91CEAF30D47FFCCFE98FAD8FE65FAA8" box="[317,460,1371,1396]" italics="true" pageId="4" pageNumber="21">patagonicus</emphasis>
</taxonomicName>
group, but not seen in the majority of members of the
<taxonomicName id="4C684D620D47FFCCFBE8FAD8FB68FAA8" box="[1101,1217,1371,1396]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="21" phylum="Chordata" rank="species" species="flagellifer">
<emphasis id="B91CEAF30D47FFCCFBE8FAD8FB68FAA8" box="[1101,1217,1371,1396]" italics="true" pageId="4" pageNumber="21">flagellifer</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C372656A0D47FFCEFF61FAFCFE09FE48" lastPageId="6" lastPageNumber="23" pageId="4" pageNumber="21" type="description">
<paragraph id="8BD736E10D47FFCDFF61FAFCFF43FD60" blockId="4.[151,1437,735,2031]" lastBlockId="5.[151,1437,152,2007]" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21">
<emphasis id="B91CEAF30D47FFCCFF61FAFCFDB3FA45" bold="true" box="[196,538,1407,1433]" pageId="4" pageNumber="21">
Description of the
<typeStatus id="54D388430D47FFCCFE0BFAFCFDB3FA45" box="[430,538,1407,1433]" pageId="4" pageNumber="21" type="holotype">holotype</typeStatus>
</emphasis>
. A medium-sized lizard; snout-vent length (SVL) 93.0 mm; tail
<quantity id="4C909B040D47FFCCFAB9FA03FA3EFA46" box="[1308,1431,1408,1434]" metricMagnitude="-1" metricUnit="m" metricValue="1.105" pageId="4" pageNumber="21" unit="mm" value="110.5">110.5 mm</quantity>
; head length
<quantity id="4C909B040D47FFCCFE82FA26FE3AFA63" box="[295,403,1445,1471]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="21" unit="mm" value="19.5">19.5 mm</quantity>
; head width
<quantity id="4C909B040D47FFCCFD8CFA26FD3CFA63" box="[553,661,1445,1471]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="4" pageNumber="21" unit="mm" value="17.3">17.3 mm</quantity>
; eye-nose distance 7.0 mm; forelimb length measured from to the insertion of the limb into the body wall to the end of the claw of the fourth finger,
<quantity id="4C909B040D47FFCCFBC4FA49FB64FA38" box="[1121,1229,1482,1508]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="4" pageNumber="21" unit="mm" value="33.3">33.3 mm</quantity>
; hind limb length measured from to the insertion of the limb into the body wall to the end of the claw of the fourth toe,
<quantity id="4C909B040D47FFCCFA89FA73FA3EF9D6" box="[1324,1431,1520,1546]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="21" unit="mm" value="50.8">50.8 mm</quantity>
; axilla-groin distance
<quantity id="4C909B040D47FFCCFE33F996FDA8F9F3" box="[406,513,1557,1583]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="4" pageNumber="21" unit="mm" value="46.5">46.5 mm</quantity>
(50.0 % of SVL); fourth finger length
<quantity id="4C909B040D47FFCCFC7EF996FBE0F9F3" box="[987,1097,1557,1583]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="4" pageNumber="21" unit="mm" value="11.2">11.2 mm</quantity>
; fourth toe length 17.0 mm; scales in dorsal head 19; scales around midbody 215; ventral scales between mental and precloacal pores 169; scales between rostral and frontal 14; supralabial scales 9-8; infralabial scales 9-8; subdigital lamellae on fourth finger 23; subdigital lamellae on fourth toe 28; precloacal pores 11; cephalic scales subpentagonals, smooth; supraorbital semicircles with large bulky scales, rounded, without azygous, incomplete posteriorly on both sides; no distinct rounded supraoculars; 78 imbricate and enlarged upper ciliaries; subocular scales rectangular, almost irregular but not fragmented, shorter than eye diameter, separated from supralabials by 2-2 irregular rows of lorilabials; preocular in contact with first lorilabial row; canthal separated from nasal by two scales; temporals smooth and rounded irregularly coniform, in 89 scales from auditive opening to the subocular; external auditory meatus enlarged, subellipsoidal longitudinally, with 56 very protruding or conically enlarged scales on its anterior border; diminute granular scales on posterior border; rostral undivided, wider than higher, separated by one row of medium scales from nasals; nasal large and surrounded by seven small scales; nasals separated by three small irregular scales; nostril rounded and large, over the centre of nasal scale; parietals irregular and rough with evident interparietal, surrounded by eight scales; nuchals strongly conical organized in 1214 irregular rows; post-auricular folds very developed with smooth conical scales; mental subpentagonal shorter than width, but higher than rostral, in contact with six irregular rectangular scales; two rows of 56 bilateral postmentals decreasing behind; dorsal scales smooth, conics, small and juxtaposed; mid-dorsal scales slightly rounded and smooth, decreasingly smaller and strongly conical toward the flanks; ventro-laterals and ventrals larger than dorsals, almost pentagonal, imbricate and smooth; two gular folds with rounded, small scales; 73 gulars between auditory meatus; caudal scales quadrangular and regularly imbricate in verticiles, proximally large, conical and smooth on dorsum, or slightly keeled, distally more rectangular and strongly keeled; scales on forelimbs subtriangular and smooth in the upper side, granular, rounded and subconical in the under side; scales in hind limbs strongly conical and slightly keeled in the dorsum but larger subpentagonal, imbricate and flat in the under side; in the femoral region, small granular scales in the lower side; infracarpals and infratarsals with round margins, becoming keeled to the base of fingers and toes. Subdigital lamellae of fingers keeled; fourth toe and finger claws very developed, almost
<quantity id="4C909B040D46FFCDFEDCFDFEFE7DFD4B" box="[377,468,637,663]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="22" unit="mm" value="2.5">2.5 mm</quantity>
of long. Eleven 11 orange-reddish precloacal glands on the scales of the cloacal region.
</paragraph>
<paragraph id="8BD736E10D46FFCDFF61FD44FDE0FC1E" blockId="5.[151,1437,152,2007]" pageId="5" pageNumber="22">
<emphasis id="B91CEAF30D46FFCDFF61FD44FEFEFD3D" bold="true" box="[196,343,711,737]" pageId="5" pageNumber="22">Coloration</emphasis>
. Colour pattern is similar in both sexes. The general pattern is characterized by two longitudinal series of pale-brown spots on a darker brownish dorsal background. These two parallel series of clear spots are conspicuous between the parietal area of the head and the base of the tail, where disappear gradually on the dorsal tail background. On the neck and fore-back, several black scales result in a partially blackish background. Similarly intense black spots are found on the pre- and post-humeral areas. On the ventral surface, the background colour varies from brick-red to intense orange. Colour pictures of males and females are shown in figures 4a, c, e.
</paragraph>
<paragraph id="8BD736E10D46FFCDFF61FC4EFCAFFA10" blockId="5.[151,1437,152,2007]" pageId="5" pageNumber="22">
<emphasis id="B91CEAF30D46FFCDFF61FC4EFDAAFC3B" bold="true" box="[196,515,973,999]" pageId="5" pageNumber="22">Morphological variation</emphasis>
. The sample analyzed comprised
<specimenCount id="9D6EFD680D46FFCDFC0DFC4EFBA5FC3B" box="[936,1036,973,999]" pageId="5" pageNumber="22" type="adult">16 adult</specimenCount>
males and
<specimenCount id="9D6EFD680D46FFCDFB3CFC4EFB57FC3B" box="[1177,1278,973,999]" pageId="5" pageNumber="22" type="adult">14 adult</specimenCount>
females (for means and SD see
<tableCitation id="C6EA035A0D46FFCDFED0FC71FE62FBD0" box="[373,459,1010,1036]" captionStart="TABLE 1" captionStartId="3.[151,240,300,324]" captionTargetBox="[151,1437,482,1551]" captionTargetPageId="3" captionText="TABLE 1. Patterns of morphological variation in morphometric and meristic traits among four geographically related" httpUri="http://table.plazi.org/id/DF1766690D40FFCBFF32FEAFFA35FE98" pageId="5" pageNumber="22" tableUuid="DF1766690D40FFCBFF32FEAFFA35FE98">Table 1</tableCitation>
). Analyses show slight size differences between the sexes, females being larger (in SVL) than males (mean SVL in females =
<quantity id="4C909B040D46FFCDFD3EFB9BFCA0FBEE" box="[667,777,1048,1074]" metricMagnitude="-2" metricUnit="m" metricValue="8.91" pageId="5" pageNumber="22" unit="mm" value="89.1">89.1 mm</quantity>
, range =
<quantity id="4C909B040D46FFCDFCDFFB9BFB83FBEE" box="[890,1066,1048,1074]" metricMagnitude="-2" metricUnit="m" metricValue="8.765" metricValueMax="9.42" metricValueMin="8.11" pageId="5" pageNumber="22" unit="mm" value="87.65" valueMax="94.2" valueMin="81.1">81.194.2 mm</quantity>
, SD = 4.0; mean SVL males = 88.0 mm, range = 78.493.0, SD = 2.5). Axilla-groin distance in females ranged
<quantity id="4C909B040D46FFCDFBEBFBBEFB50FB8B" box="[1102,1273,1085,1111]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" metricValueMax="5.25" metricValueMin="4.27" pageId="5" pageNumber="22" unit="mm" value="47.6" valueMax="52.5" valueMin="42.7">42.752.5 mm</quantity>
(mean =
<quantity id="4C909B040D46FFCDFACCFBBEFF61FBA0" metricMagnitude="-2" metricUnit="m" metricValue="4.86" pageId="5" pageNumber="22" unit="mm" value="48.6">48.6 mm</quantity>
, SD = 3.6, representing 50.855.7% of SVL); in males ranged =
<quantity id="4C909B040D46FFCDFC6CFBE1FBDCFBA0" box="[969,1141,1122,1148]" metricMagnitude="-2" metricUnit="m" metricValue="4.6049999999999995" metricValueMax="4.97" metricValueMin="4.24" pageId="5" pageNumber="22" unit="mm" value="46.05" valueMax="49.7" valueMin="42.4">42.449.7 mm</quantity>
(mean = 46.5; SD = 2.1, representing 49.656.4% of SVL). In both sexes, head length ranged
<quantity id="4C909B040D46FFCDFC1AFB0BFBC5FB7E" box="[959,1132,1160,1186]" metricMagnitude="-2" metricUnit="m" metricValue="1.755" metricValueMax="1.97" metricValueMin="1.54" pageId="5" pageNumber="22" unit="mm" value="17.55" valueMax="19.7" valueMin="15.4">15.419.7 mm</quantity>
, representing 18.321.4% of SVL. Head width ranged
<quantity id="4C909B040D46FFCDFE46FB2EFD38FB1B" box="[483,657,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.575" metricValueMax="1.73" metricValueMin="1.42" pageId="5" pageNumber="22" unit="mm" value="15.75" valueMax="17.3" valueMin="14.2">14.217.3 mm</quantity>
. Eye-nostril distance ranged 5.27.0 mm. Tail length ranged
<quantity id="4C909B040D46FFCDFAFFFB2EFEBBFB30" metricMagnitude="-1" metricUnit="m" metricValue="1.0645" metricValueMax="1.137" metricValueMin="0.992" pageId="5" pageNumber="22" unit="mm" value="106.45" valueMax="113.7" valueMin="99.2">99.2 113.7 mm</quantity>
, representing 1.181.30 times of SVL. Forelimb length ranged
<quantity id="4C909B040D46FFCDFBADFB51FB10FB30" box="[1032,1209,1234,1260]" metricMagnitude="-2" metricUnit="m" metricValue="3.3049999999999997" metricValueMax="3.6399999999999997" metricValueMin="2.9699999999999998" pageId="5" pageNumber="22" unit="mm" value="33.05" valueMax="36.4" valueMin="29.7">29.736.4 mm</quantity>
. Mean of hindlimb length in males was
<quantity id="4C909B040D46FFCDFE3FFB7BFDA5FACE" box="[410,524,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="4.96" pageId="5" pageNumber="22" unit="mm" value="49.6">49.6 mm</quantity>
, but in females ranged
<quantity id="4C909B040D46FFCDFC91FB7BFC40FACE" box="[820,1001,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="4.775" metricValueMax="5.1899999999999995" metricValueMin="4.36" pageId="5" pageNumber="22" unit="mm" value="47.75" valueMax="51.9" valueMin="43.6">43.651.9 mm</quantity>
(mean =
<quantity id="4C909B040D46FFCDFBC6FB7BFB7EFACE" box="[1123,1239,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="4.84" pageId="5" pageNumber="22" unit="mm" value="48.4">48.4 mm</quantity>
). Scales around midbody ranged
<quantity id="4C909B040D46FFCDFEC6FA9EFE42FAEB" box="[355,491,1309,1335]" metricMagnitude="0" metricUnit="m" metricValue="5.2705" metricValueMax="5.715" metricValueMin="4.826" pageId="5" pageNumber="22" unit="in" value="207.5" valueMax="225.0" valueMin="190.0">190225 in</quantity>
both sexes combined. Dorsal head scales ranged 1821. Ventrals ranged 152 180. Precloacal glands observed only in males, and ranged 811. Subocular scales fragmented in 12 parts. Two rows of lorilabials between suboculars and supralabials. Scales surrounding interparietal 69. Scales contacting mental 46. Scales between rostral-interparietal 1316. Fourth finger subdigital lamellae number 2225. Fourth toe subdigital lamellae number 2330.
</paragraph>
<paragraph id="8BD736E10D46FFCDFF61FA54FA99FA2E" blockId="5.[151,1437,152,2007]" box="[196,1328,1495,1522]" pageId="5" pageNumber="22">
<emphasis id="B91CEAF30D46FFCDFF61FA54FE5BFA2D" bold="true" box="[196,498,1495,1521]" pageId="5" pageNumber="22">Geographic distribution</emphasis>
.
<taxonomicName id="4C684D620D46FFCDFDA5FA5BFCB1FA2D" box="[512,792,1496,1521]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D46FFCDFDA5FA5BFCB1FA2D" box="[512,792,1496,1521]" italics="true" pageId="5" pageNumber="22">Phymaturus castillensis</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="54D388430D46FFCDFB92FA5BFBC3FA2E" box="[1079,1130,1496,1522]" pageId="5" pageNumber="22">type</typeStatus>
locality (
<figureCitation id="13532A640D46FFCDFB7CFA5BFA89FA2E" box="[1241,1312,1496,1522]" captionStart="FIGURE 3" captionStartId="10.[151,255,1819,1843]" captionTargetBox="[215,1341,231,1768]" captionTargetId="figure@10.[211,1345,194,1798]" captionTargetPageId="10" captionText="FIGURE 3. Geographic distribution of all known members of the Phymaturus patagonicus clade. Note nonoverlapping geographical ranges of most species. Localities are indicated by numbers, and represent Phymaturus nevadoi (1), P. payuniae (2), P. zapalensis (3), P. tenebrosus (4), P. manuelae (5), P. spurcus (6), P. excelsus (7), P. spectabilis (8), P. agilis (9), P. ceii (10), P. somuncurensis (11), P. calcogaster (12), P. patagonicus (13), P. indistinctus (14), P. v i d e l a i sp. nov. (15), and P. castillensis sp. nov. (16). The wide and scattered geographical distribution shown for P. patagonicus has been considered as the result of several cryptic different species recognized until today as polymorphisms within this species, and many of them are currently under study in order to establish their status within Phymaturus." httpUri="https://zenodo.org/record/275864/files/figure.png" pageId="5" pageNumber="22">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BD736E10D46FFCDFF61FA7EFCB8F89D" blockId="5.[151,1437,152,2007]" pageId="5" pageNumber="22">
<emphasis id="B91CEAF30D46FFCDFF61FA7EFE2AF9CB" bold="true" box="[196,387,1533,1559]" pageId="5" pageNumber="22">Natural history</emphasis>
. The biotope of
<taxonomicName id="4C684D620D46FFCDFDE3FA7EFD4CF9CA" box="[582,741,1533,1558]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D46FFCDFDE3FA7EFD4CF9CA" box="[582,741,1533,1558]" italics="true" pageId="5" pageNumber="22">P. castillensis</emphasis>
</taxonomicName>
is located in the arid Patagonic Phytogeographic Province, Central District, Erial subdistrict, in an ecotonal zone with the district of Subarbustive Steppe of the Argentinean Sierra (
<bibRefCitation id="EFF94B100D46FFCDFE22F9CBFDFBF9BE" author="Leon" box="[391,594,1608,1634]" pageId="5" pageNumber="31" refString="Leon, R. J. C., Bran, D., Collantes, M., Paruelo, J. M. &amp; Soriano, A. (1998) Grandes unidades de vegetacion de la Patagonia extra andina. Ecologia Austral, 8, 125 - 144." type="journal article" year="1998">
León
<emphasis id="B91CEAF30D46FFCDFE6BF9CBFDA3F9BD" box="[462,522,1608,1633]" italics="true" pageId="5" pageNumber="22">et al.</emphasis>
1998
</bibRefCitation>
). The predominant landscape is characterized by steppes only partially covered (&lt;50% of the surface) by small bushes and herbaceous coiron vegetation and graminea. In these Patagonian environments, the dominant bush species are
<taxonomicName id="4C684D620D46FFCDFC93F910FBA0F970" box="[822,1033,1683,1708]" class="Magnoliopsida" family="Asteraceae" genus="Nassauvia" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="ulicina">
<emphasis id="B91CEAF30D46FFCDFC93F910FBA0F970" box="[822,1033,1683,1708]" italics="true" pageId="5" pageNumber="22">Nassauvia ulicina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D620D46FFCDFBE7F910FAB3F970" box="[1090,1306,1683,1708]" class="Magnoliopsida" family="Asteraceae" genus="Chuquiraga" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="aurea">
<emphasis id="B91CEAF30D46FFCDFBE7F910FAB3F970" box="[1090,1306,1683,1708]" italics="true" pageId="5" pageNumber="22">Chuquiraga aurea</emphasis>
</taxonomicName>
, primarily. In the Sierra del Castillo and in the areas of escorial, the predominant vegetations consists primarily of higher bushes (~
<quantity id="4C909B040D46FFCDFEA9F95EFEF7F92B" box="[268,350,1757,1783]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="22" unit="m" value="1.7">1.70m</quantity>
) of the species
<taxonomicName id="4C684D620D46FFCDFD86F95EFCEEF92A" box="[547,839,1757,1782]" class="Reptilia" family="Liolaemidae" genus="Colliguaya" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="integerrima">
<emphasis id="B91CEAF30D46FFCDFD86F95EFCEEF92A" box="[547,839,1757,1782]" italics="true" pageId="5" pageNumber="22">Colliguaya integerrima</emphasis>
</taxonomicName>
, and others such as
<taxonomicName id="4C684D620D46FFCDFBE1F95EFA9BF92A" box="[1092,1330,1757,1782]" class="Magnoliopsida" family="Anacardiaceae" genus="Schinus" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="polygamus">
<emphasis id="B91CEAF30D46FFCDFBE1F95EFA9BF92A" box="[1092,1330,1757,1782]" italics="true" pageId="5" pageNumber="22">Schinus polygamus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFAE6F95EFF5EF8C0" class="Magnoliopsida" family="Solanaceae" genus="Lycium" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="chilense">
<emphasis id="B91CEAF30D46FFCDFAE6F95EFF5EF8C0" italics="true" pageId="5" pageNumber="22">Lycium chilense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFEA0F880FD60F8C0" authority="Nardophhyllum" authorityName="Nardophhyllum" box="[261,713,1794,1820]" class="Magnoliopsida" family="Berberidaceae" genus="Berberis" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis id="B91CEAF30D46FFCDFEA0F880FDADF8C0" box="[261,516,1795,1820]" italics="true" pageId="5" pageNumber="22">Berberis heterophylla</emphasis>
,
<emphasis id="B91CEAF30D46FFCDFDB7F880FD60F8C0" box="[530,713,1795,1820]" italics="true" pageId="5" pageNumber="22">Nardophhyllum</emphasis>
</taxonomicName>
obtusifolium,
<emphasis id="B91CEAF30D46FFCDFCD5F880FB9DF8C0" box="[880,1076,1795,1820]" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C684D620D46FFCDFCD5F880FB99F8C0" box="[880,1072,1795,1820]" class="Magnoliopsida" family="Asteraceae" genus="Chuquiraga" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="spp">Chuquiraga spp</taxonomicName>
.
</emphasis>
,
<taxonomicName id="4C684D620D46FFCDFBE7F880FAB2F8C0" box="[1090,1307,1795,1820]" class="Magnoliopsida" family="Verbenaceae" genus="Verbena" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="ligustrina">
<emphasis id="B91CEAF30D46FFCDFBE7F880FAB2F8C0" box="[1090,1307,1795,1820]" italics="true" pageId="5" pageNumber="22">Verbena ligustrina</emphasis>
</taxonomicName>
and scarce graminea grasses such as
<emphasis id="B91CEAF30D46FFCDFE61F8ABFD9EF89D" box="[452,567,1832,1857]" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C684D620D46FFCDFE61F8ABFD9BF89D" box="[452,562,1832,1857]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="spp">Stipa spp</taxonomicName>
.
</emphasis>
and
<emphasis id="B91CEAF30D46FFCDFDCAF8ABFCB8F89D" box="[623,785,1832,1857]" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C684D620D46FFCDFDCAF8ABFCA4F89D" box="[623,781,1832,1857]" class="Liliopsida" family="Poaceae" genus="Poa" kingdom="Plantae" order="Poales" pageId="5" pageNumber="22" phylum="Tracheophyta" rank="species" species="ligularis">Poa ligularis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD736E10D46FFCEFF63F8CEFABEFF20" blockId="5.[151,1437,152,2007]" lastBlockId="6.[151,1436,152,404]" lastPageId="6" lastPageNumber="23" pageId="5" pageNumber="22">
<taxonomicName id="4C684D620D46FFCDFF63F8CEFE4EF8BA" box="[198,487,1869,1894]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D46FFCDFF63F8CEFE4EF8BA" box="[198,487,1869,1894]" italics="true" pageId="5" pageNumber="22">Phymaturus castillensis</emphasis>
</taxonomicName>
selects rocky microhabitats in areas where other
<taxonomicName id="4C684D620D46FFCDFBD1F8CEFAAFF8BB" box="[1140,1286,1869,1895]" class="Reptilia" family="Liolaemidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="family">Liolaemidae</taxonomicName>
(
<emphasis id="B91CEAF30D46FFCDFA87F8CEFD0DF850" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C684D620D46FFCDFA87F8CEFEA6F850" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="elongatus">Liolaemus elongatus</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFEBEF8F0FE3FF850" box="[283,406,1907,1932]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="bibroni">L. bibroni</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFE06F8F0FDA3F850" box="[419,522,1907,1932]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="kingii">L. kingii</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFDBDF8F0FD0DF850" box="[536,676,1907,1932]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="fitzingeri">L. fitzingeri</taxonomicName>
</emphasis>
),
<taxonomicName id="4C684D620D46FFCDFD1BF8F1FCF0F850" box="[702,857,1906,1932]" class="Reptilia" family="Leiosauridae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="family">Leiosauridae</taxonomicName>
(
<emphasis id="B91CEAF30D46FFCDFCC9F8F0FD82F86D" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C684D620D46FFCDFCC9F8F0FBD9F850" box="[876,1136,1907,1932]" class="Reptilia" family="Leiosauridae" genus="Diplolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="bibronii">Diplolaemus bibronii</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFBD9F8F0FAA2F850" box="[1148,1291,1907,1932]" class="Reptilia" family="Leiosauridae" genus="Diplolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="darwinii">D. darwinii</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFAB2F8F0FF71F86D" class="Reptilia" family="Leiosauridae" genus="Leiosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="bellii">Leiosaurus bellii</taxonomicName>
,
<taxonomicName id="4C684D620D46FFCDFF41F81BFD82F86D" box="[228,555,1944,1969]" class="Reptilia" family="Leiosauridae" genus="Pristidactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="nigroiugulus">Pristidactylus nigroiugulus</taxonomicName>
</emphasis>
) and
<taxonomicName id="4C684D620D46FFCDFDD7F81BFC92F86E" box="[626,827,1944,1970]" class="Reptilia" family="Phyllodactylidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="family">Phyllodactylidae</taxonomicName>
(
<taxonomicName id="4C684D620D46FFCDFCE8F81BFB90F86D" box="[845,1081,1944,1969]" class="Reptilia" family="Phyllodactylidae" genus="Homonota" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B91CEAF30D46FFCDFCE8F81BFB90F86D" box="[845,1081,1944,1969]" italics="true" pageId="5" pageNumber="22">Homonota darwinii</emphasis>
</taxonomicName>
) species have been recorded. However, since
<taxonomicName id="4C684D620D46FFCDFEF6F83EFE5BF80A" box="[339,498,1981,2006]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="22" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D46FFCDFEF6F83EFE5BF80A" box="[339,498,1981,2006]" italics="true" pageId="5" pageNumber="22">P. castillensis</emphasis>
</taxonomicName>
is restricted to the rocky outcrops, few individuals of these other species can be found in direct coexistence with it. In addition, the colubrid snakes
<taxonomicName id="4C684D620D45FFCEFC10FF1BFB48FF6D" box="[949,1249,152,177]" class="Reptilia" family="Colubridae" genus="Philodryas" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="23" phylum="Chordata" rank="species" species="patagoniensis">
<emphasis id="B91CEAF30D45FFCEFC10FF1BFB48FF6D" box="[949,1249,152,177]" italics="true" pageId="6" pageNumber="23">Philodryas patagoniensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D620D45FFCEFABEFF1BFEA1FF0A" class="Reptilia" family="Colubridae" genus="Philodryas" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="23" phylum="Chordata" rank="species" species="trilineata">
<emphasis id="B91CEAF30D45FFCEFABEFF1BFEA1FF0A" italics="true" pageId="6" pageNumber="23">Philodryas trilineata</emphasis>
</taxonomicName>
, and the viperid
<taxonomicName id="4C684D620D45FFCEFE7CFF3EFD54FF0A" box="[473,765,189,214]" class="Reptilia" family="Viperidae" genus="Bothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="23" phylum="Chordata" rank="species" species="ammodytoides">
<emphasis id="B91CEAF30D45FFCEFE7CFF3EFD54FF0A" box="[473,765,189,214]" italics="true" pageId="6" pageNumber="23">Bothrops ammodytoides</emphasis>
</taxonomicName>
, are common elements of the reptile fauna at the same locality, and along with the Leiosaurids, they are likely to predate on the new
<taxonomicName id="4C684D620D45FFCEFB83FF61FB1DFF27" box="[1062,1204,226,251]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B91CEAF30D45FFCEFB83FF61FB1DFF27" box="[1062,1204,226,251]" italics="true" pageId="6" pageNumber="23">Phymaturus</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BD736E10D45FFCEFF61FE8BFE09FE48" blockId="6.[151,1436,152,404]" pageId="6" pageNumber="23">
Our field and lab observations reveal that
<taxonomicName id="4C684D620D45FFCEFD11FE8BFCFAFEFD" box="[692,851,264,289]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="23" phylum="Chordata" rank="species" species="castillensis">
<emphasis id="B91CEAF30D45FFCEFD11FE8BFCFAFEFD" box="[692,851,264,289]" italics="true" pageId="6" pageNumber="23">P. castillensis</emphasis>
</taxonomicName>
is viviparous, as observed in all the other species of the genus. In captivity, two females gave birth to one and two fully developed offspring early on
<date id="FFD610210D45FFCEFA97FEAEFF7AFEB3" pageId="6" pageNumber="23" value="2009-02">February 2009</date>
(7th9th), respectively. In the field, this species is often found eating plants, as also observed in most members of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>