990 lines
157 KiB
XML
990 lines
157 KiB
XML
<document id="3A27DE0A7E866FB5AC24A6C5DCB20506" ID-CLB-Dataset="28058" ID-DOI="10.11646/zootaxa.4508.1.4" ID-GBIF-Dataset="60d1bc89-5afc-4f3f-91c4-9b1c6da3f778" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606858" ID-ZooBank="DD50AB97-77F5-476F-9FFF-B33B96B43078" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585449037" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. & Morozov, Taras B." docDate="2018" docId="038C557E6804107EFF564BC5FD710211" docLanguage="en" docName="zootaxa.4508.1.4.pdf" docOrigin="Zootaxa 4508 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fatkullina imitata Grischenko & Gordon & Morozov 2018, n. sp." docType="treatment" docVersion="9" masterDocId="FFB52D0668061075FFC14E45FFF9076A" masterDocTitle="Fatkullina imitata n. sp., second species of a unique cheilostome bryozoan genus with reversed-polarity zooidal budding, and new family Fatkullinidae" updateTime="1698712868930" updateUser="ExternalLinkService">
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<mods:mods id="A933B1B98FF7B75575B8699FB372DF79" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="06F8BBB420E63DF728ACADBE6C3523C4">
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<mods:title id="29A79E554DF3E2AB845630C669D1549A">Fatkullina imitata n. sp., second species of a unique cheilostome bryozoan genus with reversed-polarity zooidal budding, and new family Fatkullinidae</mods:title>
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<mods:name id="CE6AC1A073B061DA182A13CBF9F5E6FA" type="personal">
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<mods:role id="3A8342D6525AB038CADB6390290571E3">
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<mods:roleTerm id="1F5832BF2DC177580DE5C0D011CDFDBB">Author</mods:roleTerm>
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<mods:namePart id="9E27D2FF0EF7E0197ABB3D2C305EF0B8">Grischenko, Andrei V.</mods:namePart>
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<mods:roleTerm id="85007C61F9C6267AB26A8E25610A7611">Author</mods:roleTerm>
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<mods:namePart id="95DE84C57986F9B2C9A90F741767A8C4">Gordon, Dennis P.</mods:namePart>
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<mods:name id="70893D488998C1F350D144125B5655D2" type="personal">
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<mods:roleTerm id="9F573411242ECF50F4E9EA56A95DD52A">Author</mods:roleTerm>
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<mods:namePart id="C2DE3EDB63FCB61F083C21EA31B1206C">Morozov, Taras B.</mods:namePart>
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<mods:typeOfResource id="C7DE75E5E1BB7C5945A54B2147AB2453">text</mods:typeOfResource>
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<mods:title id="E1E47E6F25551BD76CA1D447F97EF52B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="455F4CDE5FDCDCC03EF5C40E8F09AC54">
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<mods:date id="3BBEBC7CDB3901F6A6FEA95A559AE137">2018</mods:date>
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<mods:number id="F541044AE32C808E8BA0F5774425D2DD">2018-10-30</mods:number>
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<mods:detail id="6E0B0E047E73DCA51A95D455430322EC" type="volume">
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<mods:number id="7C610B60DBE6E5CCBC98258FC80E92E0">4508</mods:number>
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<mods:number id="3CC29F7464AFF9AAA527565C0AAF394A">1</mods:number>
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<mods:classification id="CAE4D7F5DE127979FDA650804F4B852E">journal volume</mods:classification>
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<mods:identifier id="3CC66E57603D7F4490D78E75DCB9C847" type="CLB-Dataset">28058</mods:identifier>
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<mods:identifier id="B9D3B27E32DEB9FEDEE8413A95010CB9" type="DOI">10.11646/zootaxa.4508.1.4</mods:identifier>
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<treatment id="038C557E6804107EFF564BC5FD710211" ID-DOI="http://doi.org/10.5281/zenodo.5975684" ID-GBIF-Taxon="154495680" ID-Zenodo-Dep="5975684" LSID="urn:lsid:plazi:treatment:038C557E6804107EFF564BC5FD710211" httpUri="http://treatment.plazi.org/id/038C557E6804107EFF564BC5FD710211" lastPageId="11" pageId="2">
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<subSubSection id="C33FB7E368041077FF564BC5FE4502F1" box="[151,444,1408,1435]" pageId="2" type="nomenclature">
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<paragraph id="8B9AE46868041077FF564BC5FE4502F1" blockId="2.[151,447,1408,1469]" box="[151,444,1408,1435]" pageId="2">
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<heading id="D0D2530468041077FF564BC5FE4502F1" bold="true" box="[151,444,1408,1435]" fontSize="11" level="1" pageId="2" reason="1">
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<emphasis id="B951387A68041077FF564BC5FE4502F1" bold="true" box="[151,444,1408,1435]" pageId="2">
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<taxonomicName id="4C259FEB68041077FF564BC5FE8A02F0" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[151,371,1408,1434]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
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<emphasis id="B951387A68041077FF564BC5FE8A02F0" bold="true" box="[151,371,1408,1434]" italics="true" pageId="2">Fatkullina imitata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A262850168041077FEBB4BC4FE4502F1" box="[378,444,1409,1435]" pageId="2" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FB7E368041077FF564BE1FD9E0168" pageId="2" type="description">
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<paragraph id="8B9AE46868041077FF564BE1FE4602D6" blockId="2.[151,447,1408,1469]" box="[151,447,1444,1469]" pageId="2">
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(
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<figureCitation id="131EF8ED68041077FF5E4BE1FF1902D6" box="[159,224,1444,1469]" captionStart="FIGURE 1" captionStartId="4.[151,250,1859,1881]" captionTargetBox="[195,1400,195,1840]" captionTargetId="figure@4.[195,1400,195,1840]" captionTargetPageId="4" captionText="FIGURE 1. General view of dried colonies of Fatkullina imitata n. sp. (A–D) and F. paradoxa Grischenko, Gordon & Taylor, 1998 (E). A, B, specimens R.V. Professor Kizevetter 2015, Stn 73; C, D, specimens paratype 8 NIWA 127751; E, specimen R.V. Professor Probatov 2013, Stn 5, 16 June 2013, 57.41694° N, 156.51806° E, 27 m, pebbles, sand, silt. A, B. Colonies encrusting pebbles. C, D. Colonies encrusting external (C), and internal (D) surface of shell fragment of Chlamys sp. E. Colony encrusting internal surface of bivalve shell fragment of Serripes sp., surrounded by other encrusting bryozoan species. Scale bars: 5 mm." httpUri="https://zenodo.org/record/2606860/files/figure.png" pageId="2">Figs 1</figureCitation>
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A–D, 2–6;
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<tableCitation id="C6A7D1D368041077FEA24BE1FE4A02D6" box="[355,435,1444,1469]" captionStart="TABLE 1" captionStartId="10.[151,239,440,463]" captionText="TABLE 1. Measurements (in mm, except number of marginal and frontal pores) of the holotype and paratype specimens of Fatkullina imitata n. sp. Abbreviations: ZL, zooid length; ZW, zooid width; ZD, zooid depth (height); OrL, orifice (without sinus) length; Or(s)L, orificial sinus length; OrW, orifice width; P(m)N, number of marginal pores (areolae); P(f)N, number of frontal pseudopores; AnL, ancestrula length; AnW, ancestrula width; AnOrL, ancestrular orifice (without sinus) length; AnOr(s)L, ancestrular orificial sinus length; AnOrW, ancestrular orifice width; N, number of measurements (unless otherwise indicated, n = 20)." pageId="2">Table 1</tableCitation>
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)
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||
</paragraph>
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<paragraph id="8B9AE46868041077FF564BA9FD9E0168" blockId="2.[151,615,1516,1538]" box="[151,615,1516,1538]" pageId="2">
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<treatmentCitationGroup id="AB35C34668041077FF564BA9FD9E0168" box="[151,615,1516,1538]" pageId="2">
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<taxonomicName id="4C259FEB68041077FF564BA9FEF80168" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[151,257,1516,1538]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
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<emphasis id="B951387A68041077FF564BA9FEF80168" box="[151,257,1516,1538]" italics="true" pageId="2">Fatkullina</emphasis>
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</taxonomicName>
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||
sp.:
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<treatmentCitation id="0A84C27968041077FEF04BA9FDFC0168" author="Grischenko, A. V." box="[305,517,1516,1538]" page="39" pageId="2" year="2015">
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<bibRefCitation id="EFB4999968041077FEF04BA9FE1B0168" author="Grischenko, A. V." box="[305,482,1516,1538]" pageId="2" pagination="37 - 42" refId="ref7717" refString="Grischenko, A. V. (2015) To the cheilostome bryozoan fauna from the continental slope of western Kamchatka. In: Tokranov, A. M. (Ed.), Conservation of Biodiversity of Kamchatka and Coastal Waters. Materials of 16 th international scientific conference. Kamchatpress, Petropavlovsk-Kamchatsky, pp. 37 - 42. [in Russian]" type="book chapter" year="2015">Grischenko 2015</bibRefCitation>
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||
: 39
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||
</treatmentCitation>
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||
(table 1).
|
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</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C33FB7E368041076FF56486BFE750572" lastPageId="3" pageId="2" type="materials_examined">
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<paragraph id="8B9AE46868041076FF56486BFE750572" blockId="2.[151,1436,1582,2003]" lastBlockId="3.[151,1436,151,2013]" lastPageId="3" pageId="2">
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<emphasis id="B951387A68041077FF56486BFE78012D" bold="true" box="[151,385,1582,1607]" pageId="2">Material examined.</emphasis>
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<materialsCitation id="3B4DEE3568041077FE46486BFB6901E5" ID-GBIF-Occurrence="2028138719" collectingDate="2008-06-16" collectionCode="ZIRAS" collectorName="Collection, R. V & S. G. Korostylyov." elevation="290" latitude="58.03833" location="Sea of Okhotsk" longLatPrecision="1" longitude="155.72028" municipality="Stn" pageId="2" specimenCode="ZIRAS 1" specimenCount="1" stateProvince="Kamchatka" typeStatus="holotype">
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<emphasis id="B951387A68041077FE46486BFE14012D" box="[391,493,1582,1607]" italics="true" pageId="2">
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<typeStatus id="549E5ACA68041077FE46486BFE14012D" box="[391,493,1582,1607]" pageId="2" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
:
|
||
<specimenCode id="DB834C1368041077FE3C486AFDA7012D" box="[509,606,1582,1607]" collectionCode="ZIRAS" pageId="2">ZIRAS 1</specimenCode>
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||
/50661, colony (18 ×
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<quantity id="4CDD498D68041077FC94486AFC5C012D" box="[853,933,1583,1607]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="2" unit="mm" value="17.0">17 mm</quantity>
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) encrusting flattened pebble; KamchatNIRO
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||
<collectorName id="26D081BE68041077FF564817FEB40100" box="[151,333,1618,1643]" pageId="2">Collection, R.V</collectorName>
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.
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<emphasis id="B951387A68041077FE9C4816FE6B0100" box="[349,402,1619,1642]" italics="true" pageId="2">Agat</emphasis>
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,
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<date id="FF9BC2A868041077FE674817FDB30101" box="[422,586,1618,1643]" pageId="2" value="2008-06-16">
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<collectingDate id="EFDF3B4068041077FE674817FDB30101" box="[422,586,1618,1643]" pageId="2" value="2008-06-16">16 June 2008</collectingDate>
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</date>
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,
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<collectingMunicipality id="6BFE7E1268041077FD9A4816FD790101" box="[603,640,1619,1643]" pageId="2">Stn</collectingMunicipality>
|
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2–S–2, continental slope of western
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||
<collectingRegion id="49E12A8A68041077FB834817FAC70101" box="[1090,1342,1618,1643]" country="Russia" name="Kamchatka" pageId="2">Kamchatka Peninsula</collectingRegion>
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,
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<location id="8EFAB2B368041077FA8E4816FF0001E5" LSID="urn:lsid:plazi:treatment:038C557E6804107EFF564BC5FD710211:8EFAB2B368041077FA8E4816FF0001E5" latitude="58.03833" longLatPrecision="1" longitude="155.72028" municipality="Stn" name="Sea of Okhotsk" pageId="2" stateProvince="Kamchatka">Sea of Okhotsk</location>
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,
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<geoCoordinate id="EE1182AF68041077FEC64833FE6401E4" box="[263,413,1654,1679]" direction="north" orientation="latitude" pageId="2" precision="1" value="58.03833">58.03833° N</geoCoordinate>
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,
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<geoCoordinate id="EE1182AF68041077FE6A4833FDB301E4" box="[427,586,1654,1679]" direction="east" orientation="longitude" pageId="2" precision="1" value="155.72028">155.72028° E</geoCoordinate>
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, depth
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<quantity id="4CDD498D68041077FD634832FD1601E4" box="[674,751,1655,1679]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="2" unit="m" value="290.0">
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<elevation id="0008035B68041077FD634832FD1601E4" box="[674,751,1655,1679]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="2" unit="m" value="290.0">290 m</elevation>
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</quantity>
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, on pebbles, coll.
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<collectorName id="26D081BE68041077FC074832FB6901E5" box="[966,1168,1654,1679]" pageId="2">S.G. Korostylyov.</collectorName>
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</materialsCitation>
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<materialsCitation id="3B4DEE3568041077FB5B4832FE6B01BD" ID-GBIF-Occurrence="2028138713" collectingDate="2008-06-16" collectionCode="ZIRAS" collectorName="Collection, R. V & S. G. Korostylyov." elevation="290" latitude="58.03833" location="Sea of Okhotsk" longLatPrecision="1" longitude="155.72028" municipality="Stn" pageId="2" specimenCode="ZIRAS 2" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
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<emphasis id="B951387A68041077FB5B4832FAF801E4" box="[1178,1281,1655,1678]" italics="true" pageId="2">
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<typeStatus id="549E5ACA68041077FB5B4832FAF801E4" box="[1178,1281,1655,1678]" pageId="2" type="paratype">Paratype</typeStatus>
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</emphasis>
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1:
|
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<specimenCode id="DB834C1368041077FAEA4832FA6A01E5" box="[1323,1427,1655,1679]" collectionCode="ZIRAS" pageId="2">ZIRAS 2</specimenCode>
|
||
/ 50662, two colonies (23 ×
|
||
<quantity id="4CDD498D68041077FE0D48DEFDE601D8" box="[460,543,1691,1715]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="2" unit="mm" value="21.0">21 mm</quantity>
|
||
, 15 ×
|
||
<quantity id="4CDD498D68041077FDAB48DEFD4401D9" box="[618,701,1691,1715]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" unit="mm" value="12.0">12 mm</quantity>
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) encrusting broken fragment of bivalve shell
|
||
<taxonomicName id="4C259FEB68041077FB0648DFFAD001D9" box="[1223,1321,1690,1715]" class="Bivalvia" family="Pectinidae" genus="Chlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="2" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A68041077FB0648DFFAD001D9" box="[1223,1321,1690,1715]" italics="true" pageId="2">Chlamys</emphasis>
|
||
</taxonomicName>
|
||
sp.; same data as for holotype
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4DEE3568041077FE6548FAFAF50074" ID-GBIF-Occurrence="2028138720" collectingDate="2015-07-03" collectionCode="ZIRAS" collectorName="Collection, R. V. & T. B. Morozov." elevation="145" latitude="58.13333" location="Sea of Okhotsk" longLatPrecision="1" longitude="156.01889" municipality="Stn" pageId="2" specimenCode="ZIRAS 3" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68041077FE6548FAFDF201BC" box="[420,523,1727,1750]" italics="true" pageId="2">
|
||
<typeStatus id="549E5ACA68041077FE6548FAFDF201BC" box="[420,523,1727,1750]" pageId="2" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
2:
|
||
<specimenCode id="DB834C1368041077FDFE48FAFD5201BD" box="[575,683,1726,1751]" collectionCode="ZIRAS" pageId="2">ZIRAS 3</specimenCode>
|
||
/50663, 12 fragments of single colony detached from pebble; KamchatNIRO
|
||
<collectorName id="26D081BE68041077FE8948A7FE070190" box="[328,510,1762,1787]" pageId="2">Collection, R.V.</collectorName>
|
||
<emphasis id="B951387A68041077FDC448A6FD100191" box="[517,745,1763,1787]" italics="true" pageId="2">Professor Kizevetter</emphasis>
|
||
,
|
||
<date id="FF9BC2A868041077FD3948A6FC860191" box="[760,895,1762,1787]" pageId="2" value="2015-07-03">
|
||
<collectingDate id="EFDF3B4068041077FD3948A6FC860191" box="[760,895,1762,1787]" pageId="2" value="2015-07-03">3 July 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMunicipality id="6BFE7E1268041077FC4D48A6FC480191" box="[908,945,1763,1787]" pageId="2">Stn</collectingMunicipality>
|
||
73, shelf of western
|
||
<collectingRegion id="49E12A8A68041077FB6048A7FA610191" box="[1185,1432,1762,1787]" country="Russia" name="Kamchatka" pageId="2">Kamchatka Peninsula</collectingRegion>
|
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,
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||
<location id="8EFAB2B368041077FF564942FEB40075" LSID="urn:lsid:plazi:treatment:038C557E6804107EFF564BC5FD710211:8EFAB2B368041077FF564942FEB40075" box="[151,333,1798,1823]" latitude="58.13333" longLatPrecision="1" longitude="156.01889" municipality="Stn" name="Sea of Okhotsk" pageId="2" stateProvince="Kamchatka">Sea of Okhotsk</location>
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,
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<geoCoordinate id="EE1182AF68041077FE9A4943FE090074" box="[347,496,1798,1823]" direction="north" orientation="latitude" pageId="2" precision="1" value="58.13333">58.13333° N</geoCoordinate>
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,
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<geoCoordinate id="EE1182AF68041077FE3F4943FD670074" box="[510,670,1798,1823]" direction="east" orientation="longitude" pageId="2" precision="1" value="156.01889">156.01889° E</geoCoordinate>
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, depth
|
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<quantity id="4CDD498D68041077FD344943FCB80074" box="[757,833,1798,1823]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="2" unit="m" value="145.0">
|
||
<elevation id="0008035B68041077FD344943FCB80074" box="[757,833,1798,1823]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="2" unit="m" value="145.0">145 m</elevation>
|
||
</quantity>
|
||
, on rock and shell, coll.
|
||
<collectorName id="26D081BE68041077FBA74942FAF50074" box="[1126,1292,1799,1822]" pageId="2">T.B. Morozov.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4DEE3568041077FAD44942FBE40029" ID-GBIF-Occurrence="2028138723" collectingDate="2015-07-03" collectionCode="ZIRAS" collectorName="Collection, R. V. & T. B. Morozov." elevation="145" latitude="58.13333" location="Sea of Okhotsk" longLatPrecision="1" longitude="156.01889" municipality="Stn" pageId="2" specimenCode="ZIRAS 4" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68041077FAD44942FA850074" box="[1301,1404,1799,1822]" italics="true" pageId="2">
|
||
<typeStatus id="549E5ACA68041077FAD44942FA850074" box="[1301,1404,1799,1822]" pageId="2" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
3:
|
||
<specimenCode id="DB834C1368041077FF56496EFEF80029" box="[151,257,1834,1859]" collectionCode="ZIRAS" pageId="2">ZIRAS 4</specimenCode>
|
||
/50664, seven intact ancestrular colonies; same data as for paratype
|
||
</materialsCitation>
|
||
2.
|
||
<materialsCitation id="3B4DEE3568041077FB884969FAF4000D" ID-GBIF-Occurrence="2028138716" collectingDate="2008-06-16" collectionCode="ZIRAS" collectorName="Collection, R. V & S. G. Korostylyov." elevation="290" latitude="58.03833" location="Sea of Okhotsk" longLatPrecision="1" longitude="155.72028" municipality="Stn" pageId="2" specimenCode="ZIRAS 5" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68041077FB884969FB490029" box="[1097,1200,1836,1859]" italics="true" pageId="2">
|
||
<typeStatus id="549E5ACA68041077FB884969FB490029" box="[1097,1200,1836,1859]" pageId="2" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
4:
|
||
<specimenCode id="DB834C1368041077FB1D496EFABC0029" box="[1244,1349,1834,1859]" collectionCode="ZIRAS" pageId="2">ZIRAS 5</specimenCode>
|
||
/50665, ancestrular colony (4 ×
|
||
<quantity id="4CDD498D68041077FE65490AFE1E000D" box="[420,487,1871,1895]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" unit="mm" value="3.0">3 mm</quantity>
|
||
) encrusting broken fragment of
|
||
<taxonomicName id="4C259FEB68041077FC94490BFC4E000D" box="[853,951,1870,1895]" class="Bivalvia" family="Pectinidae" genus="Chlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="2" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A68041077FC94490BFC4E000D" box="[853,951,1870,1895]" italics="true" pageId="2">Chlamys</emphasis>
|
||
</taxonomicName>
|
||
sp; same data as for holotype
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4DEE3568041077FAD9490AFD7E00B9" ID-GBIF-Occurrence="2028138714" collectingDate="2008-06-16" collectionCode="ZIRAS" collectorName="Collection, R. V & S. G. Korostylyov." elevation="285" latitude="58.01917" location="Sea of Okhotsk" longLatPrecision="1" longitude="155.71667" municipality="Stn" pageId="2" specimenCode="ZIRAS 6" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68041077FAD9490AFA86000C" box="[1304,1407,1871,1894]" italics="true" pageId="2">
|
||
<typeStatus id="549E5ACA68041077FAD9490AFA86000C" box="[1304,1407,1871,1894]" pageId="2" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
5:
|
||
<specimenCode id="DB834C1368041077FF564936FF0700E1" box="[151,254,1906,1931]" collectionCode="ZIRAS" pageId="2">ZIRAS 6</specimenCode>
|
||
/50666, colony (8 ×
|
||
<quantity id="4CDD498D68041077FE2B4936FDD600E1" box="[490,559,1907,1931]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" unit="mm" value="7.0">7 mm</quantity>
|
||
) detached from pebble; KamchatNIRO
|
||
<collectorName id="26D081BE68041077FC384937FB5200E0" box="[1017,1195,1906,1931]" pageId="2">Collection, R.V</collectorName>
|
||
.
|
||
<emphasis id="B951387A68041077FB764936FB1500E0" box="[1207,1260,1907,1930]" italics="true" pageId="2">Agat</emphasis>
|
||
,
|
||
<date id="FF9BC2A868041077FB3D4937FA6100E1" box="[1276,1432,1906,1931]" pageId="2" value="2008-06-16">
|
||
<collectingDate id="EFDF3B4068041077FB3D4937FA6100E1" box="[1276,1432,1906,1931]" pageId="2" value="2008-06-16">16 June 2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMunicipality id="6BFE7E1268041077FF5649D2FF4500C5" box="[151,188,1943,1967]" pageId="2">Stn</collectingMunicipality>
|
||
1–K–2, continental slope of western
|
||
<collectingRegion id="49E12A8A68041077FD9F49D3FCAD00C5" box="[606,852,1942,1967]" country="Russia" name="Kamchatka" pageId="2">Kamchatka Peninsula</collectingRegion>
|
||
,
|
||
<location id="8EFAB2B368041077FC9E49D2FBF700C5" LSID="urn:lsid:plazi:treatment:038C557E6804107EFF564BC5FD710211:8EFAB2B368041077FC9E49D2FBF700C5" box="[863,1038,1942,1967]" latitude="58.01917" longLatPrecision="1" longitude="155.71667" municipality="Stn" name="Sea of Okhotsk" pageId="2" stateProvince="Kamchatka">Sea of Okhotsk</location>
|
||
,
|
||
<geoCoordinate id="EE1182AF68041077FBD849D3FB5200C4" box="[1049,1195,1942,1967]" direction="north" orientation="latitude" pageId="2" precision="1" value="58.01917">58.01917° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1182AF68041077FB7749D3FAAB00C4" box="[1206,1362,1942,1967]" direction="east" orientation="longitude" pageId="2" precision="1" value="155.71667">155.71667° E</geoCoordinate>
|
||
, depth
|
||
<quantity id="4CDD498D68041077FF5649FFFF1A00B8" box="[151,227,1978,2003]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="2" unit="m" value="285.0">
|
||
<elevation id="0008035B68041077FF5649FFFF1A00B8" box="[151,227,1978,2003]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="2" unit="m" value="285.0">285 m</elevation>
|
||
</quantity>
|
||
, on pebbles, coll.
|
||
<collectorName id="26D081BE68041077FE7D49FEFD7E00B9" box="[444,647,1978,2003]" pageId="2">S.G. Korostylyov.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4DEE3568041076FD5149F9FF0407BF" ID-GBIF-Occurrence="2028138722" collectingDate="2008-06-16" collectionCode="NHMUK" collectorName="Collection, R. V & S. G. Korostylyov." elevation="290" lastPageId="3" latitude="58.03833" location="Sea of Okhotsk" longLatPrecision="1" longitude="155.72028" municipality="Stn" pageId="2" specimenCode="NHMUK 2017.7" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68041077FD5149F9FD0E00B9" box="[656,759,1980,2003]" italics="true" pageId="2">
|
||
<typeStatus id="549E5ACA68041077FD5149F9FD0E00B9" box="[656,759,1980,2003]" pageId="2" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
6:
|
||
<specimenCode id="DB834C1368041077FCE349FEFC1100B9" box="[802,1000,1979,2003]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" type="Museum">NHMUK 2017.7</specimenCode>
|
||
.11.7, extensive colony (35 ×
|
||
<quantity id="4CDD498D68041077FA8149FEFA6C00B9" box="[1344,1429,1979,2003]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" unit="mm" value="29.0">29 mm</quantity>
|
||
) encrusting cirripede fragment (with adjacent tiny colony—ancestrula and single daughter zooid); same data as for holotype
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4DEE3568051076FED04EF8FAF40676" ID-GBIF-Occurrence="2028138718" collectingDate="2013-08-11" collectionCode="NHMUK" collectorName="Collection, R. V. & T. B. Morozov." elevation="146" latitude="58.15" location="Sea of Okhotsk" longLatPrecision="1" longitude="156.05" municipality="Stn" pageId="3" specimenCode="NHMUK 2017.7" specimenCount="1" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68051076FED04EF8FE8107BE" box="[273,376,189,212]" italics="true" pageId="3">
|
||
<typeStatus id="549E5ACA68051076FED04EF8FE8107BE" box="[273,376,189,212]" pageId="3" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
7:
|
||
<specimenCode id="DB834C1368051076FE704EF8FD8607BE" box="[433,639,188,212]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" type="Museum">NHMUK 2017.7</specimenCode>
|
||
.11.8, colony encrusting fragment of
|
||
<taxonomicName id="4C259FEB68051076FB8D4EF9FB5707BF" box="[1100,1198,188,213]" class="Bivalvia" family="Pectinidae" genus="Chlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="3" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A68051076FB8D4EF9FB5707BF" box="[1100,1198,188,213]" italics="true" pageId="3">Chlamys</emphasis>
|
||
</taxonomicName>
|
||
sp; KamchatNIRO
|
||
<collectorName id="26D081BE68051076FF564E9AFEA9079D" box="[151,336,223,248]" pageId="3">Collection, R.V.</collectorName>
|
||
<emphasis id="B951387A68051076FE9D4EA5FDC00792" box="[348,569,223,248]" italics="true" pageId="3">Professor Probatov</emphasis>
|
||
,
|
||
<date id="FF9BC2A868051076FD8D4EA5FCF20792" box="[588,779,224,248]" pageId="3" value="2013-08-11">
|
||
<collectingDate id="EFDF3B4068051076FD8D4EA5FCF20792" box="[588,779,224,248]" pageId="3" value="2013-08-11">11 August 2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMunicipality id="6BFE7E1268051076FCDD4EA5FCB80792" box="[796,833,224,248]" pageId="3">Stn</collectingMunicipality>
|
||
82, shelf of western
|
||
<collectingRegion id="49E12A8A68051076FB844E9AFAB90792" box="[1093,1344,223,248]" country="Russia" name="Kamchatka" pageId="3">Kamchatka Peninsula</collectingRegion>
|
||
,
|
||
<location id="8EFAB2B368051076FA914EA5FF000677" LSID="urn:lsid:plazi:treatment:038C557E6804107EFF564BC5FD710211:8EFAB2B368051076FA914EA5FF000677" latitude="58.15" longLatPrecision="1" longitude="156.05" municipality="Stn" name="Sea of Okhotsk" pageId="3" stateProvince="Kamchatka">Sea of Okhotsk</location>
|
||
,
|
||
<geoCoordinate id="EE1182AF68051076FEC74F41FE620676" box="[262,411,260,285]" direction="north" orientation="latitude" pageId="3" precision="1" value="58.15">58.15000° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1182AF68051076FE684F41FDB10676" box="[425,584,260,285]" direction="east" orientation="longitude" pageId="3" precision="1" value="156.05">156.05000° E</geoCoordinate>
|
||
, depth
|
||
<elevation id="0008035B68051076FD5E4F41FD130676" box="[671,746,260,285]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="3" unit="m" value="146.0">
|
||
<quantity id="4CDD498D68051076FD5E4F41FD130676" box="[671,746,260,285]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="3" unit="m" value="146.0">146 m</quantity>
|
||
</elevation>
|
||
, on gravel from silty sand, coll.
|
||
<collectorName id="26D081BE68051076FBA64F40FAF40676" box="[1127,1293,261,284]" pageId="3">T.B. Morozov.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4DEE3568051076FAD74F40FCE106BA" ID-GBIF-Occurrence="2028138721" collectingDate="2013-08-11" collectionCode="NIWA" collectorName="Collection, R. V & Stn & R. V. Professor Probatov" elevation="146" latitude="58.15" location="Sea of Okhotsk" longLatPrecision="1" longitude="156.05" municipality="Stn" pageId="3" specimenCode="NIWA 127751" specimenCount="13" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68051076FAD74F40FA840676" box="[1302,1405,261,284]" italics="true" pageId="3">
|
||
<typeStatus id="549E5ACA68051076FAD74F40FA840676" box="[1302,1405,261,284]" pageId="3" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
8:
|
||
<specimenCode id="DB834C1368051076FF564F6DFEBC062A" box="[151,325,295,320]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="3" type="Other institutional type">NIWA 127751</specimenCode>
|
||
, colony (33 ×
|
||
<quantity id="4CDD498D68051076FDC54F6DFDA2062A" box="[516,603,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" unit="mm" value="30.0">30 mm</quantity>
|
||
) encrusting fragment of
|
||
<taxonomicName id="4C259FEB68051076FC4C4F62FC16062A" box="[909,1007,295,320]" class="Bivalvia" family="Pectinidae" genus="Chlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="3" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A68051076FC4C4F62FC16062A" box="[909,1007,295,320]" italics="true" pageId="3">Chlamys</emphasis>
|
||
</taxonomicName>
|
||
sp., with adjacent colony and tiny ancestrular colonies; same data as for paratype 7.
|
||
<emphasis id="B951387A68051076FD324F09FC2C060F" box="[755,981,332,357]" italics="true" pageId="3">Additional material</emphasis>
|
||
: KamchatNIRO
|
||
<collectorName id="26D081BE68051076FB614F09FAAE060E" box="[1184,1367,332,357]" pageId="3">Collection, R.V</collectorName>
|
||
.
|
||
<emphasis id="B951387A68051076FAA64F09FA650609" box="[1383,1436,332,355]" italics="true" pageId="3">Agat</emphasis>
|
||
(2008):
|
||
<collectorName id="26D081BE68051076FF374F35FEE206E2" box="[246,283,368,392]" pageId="3">Stn</collectorName>
|
||
2–S–2, one specimen;
|
||
<collectorName id="26D081BE68051076FDF94F35FDA406E2" box="[568,605,368,392]" pageId="3">Stn</collectorName>
|
||
3–W–1, two specimens;
|
||
<collectorName id="26D081BE68051076FC504F35FC4F06E2" box="[913,950,368,392]" pageId="3">Stn</collectorName>
|
||
3–W–2, six specimens;
|
||
<collectorName id="26D081BE68051076FB234F35FAFE06E2" box="[1250,1287,368,392]" pageId="3">Stn</collectorName>
|
||
4–N–2, one specimen;
|
||
<collectorName id="26D081BE68051076FED34FD1FECE06C6" box="[274,311,404,428]" pageId="3">Stn</collectorName>
|
||
5–E–1, one specimen;
|
||
<collectorName id="26D081BE68051076FD864FD1FD9506C6" box="[583,620,404,428]" pageId="3">Stn</collectorName>
|
||
5–E–2, three specimens.
|
||
<collectorName id="26D081BE68051076FC574FD0FB5406C7" box="[918,1197,404,429]" pageId="3">
|
||
R.V.
|
||
<emphasis id="B951387A68051076FC114FD1FB5406C7" box="[976,1197,404,429]" italics="true" pageId="3">Professor Probatov</emphasis>
|
||
</collectorName>
|
||
(2013):
|
||
<collectorName id="26D081BE68051076FAD34FD1FACE06C6" box="[1298,1335,404,428]" pageId="3">Stn</collectorName>
|
||
62, four specimens;
|
||
<collectorName id="26D081BE68051076FEDD4FFDFEB806BA" box="[284,321,440,464]" pageId="3">Stn</collectorName>
|
||
63, one specimen;
|
||
<collectorName id="26D081BE68051076FDE34FFDFDBE06BA" box="[546,583,440,464]" pageId="3">Stn</collectorName>
|
||
82,
|
||
<specimenCount id="9D232FE168051076FDBA4FFDFCE106BA" box="[635,792,439,464]" pageId="3" type="generic">13 specimens</specimenCount>
|
||
</materialsCitation>
|
||
. R.V.
|
||
<materialsCitation id="3B4DEE3568051076FC9E4FFDFB50069F" ID-GBIF-Occurrence="2028138715" collectionCode="TINRO" collectorName="Stn & R. V. Professor Kizevetter" pageId="3" specimenCount="96" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68051076FC9E4FFDFC4906A5" box="[863,944,440,463]" italics="true" pageId="3">
|
||
<collectionCode id="ED347CAD68051076FC9E4FFDFC4906A5" box="[863,944,440,463]" pageId="3">TINRO</collectionCode>
|
||
</emphasis>
|
||
(2014):
|
||
<collectorName id="26D081BE68051076FBD54FFDFBC006BA" box="[1044,1081,440,464]" pageId="3">Stn</collectorName>
|
||
63, one specimen;
|
||
<collectorName id="26D081BE68051076FADB4FFDFAC606BA" box="[1306,1343,440,464]" pageId="3">Stn</collectorName>
|
||
69, one specimen;
|
||
<collectorName id="26D081BE68051076FED04F99FECF069E" box="[273,310,476,500]" pageId="3">Stn</collectorName>
|
||
82, eight specimens.
|
||
<collectorName id="26D081BE68051076FDEE4F98FCA9069E" box="[559,848,476,500]" pageId="3">
|
||
R.V.
|
||
<emphasis id="B951387A68051076FDA84F99FCA9069E" box="[617,848,476,500]" italics="true" pageId="3">Professor Kizevetter</emphasis>
|
||
</collectorName>
|
||
(2015):
|
||
<collectorName id="26D081BE68051076FC724F99FC21069E" box="[947,984,476,500]" pageId="3">Stn</collectorName>
|
||
73,
|
||
<specimenCount id="9D232FE168051076FBCD4F99FB50069F" box="[1036,1193,476,501]" pageId="3" type="generic">96 specimens</specimenCount>
|
||
</materialsCitation>
|
||
. R.V.
|
||
<materialsCitation id="3B4DEE3568051076FB314F98FE710572" ID-GBIF-Occurrence="2028138717" collectionCode="TINRO" collectorName="Stn" pageId="3" specimenCount="43" stateProvince="Kamchatka" typeStatus="paratype">
|
||
<emphasis id="B951387A68051076FB314F98FAB8069E" box="[1264,1345,477,500]" italics="true" pageId="3">
|
||
<collectionCode id="ED347CAD68051076FB314F98FAB8069E" box="[1264,1345,477,500]" pageId="3">TINRO</collectionCode>
|
||
</emphasis>
|
||
(2016):
|
||
<collectorName id="26D081BE68051076FF564C45FF450572" box="[151,188,512,536]" pageId="3">Stn</collectorName>
|
||
74,
|
||
<specimenCount id="9D232FE168051076FF2D4C45FE710572" box="[236,392,511,536]" pageId="3" type="generic">43 specimens</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33FB7E368051076FF064C61FDE20535" pageId="3" type="etymology">
|
||
<paragraph id="8B9AE46868051076FF064C61FDE20535" blockId="3.[151,1436,151,2013]" pageId="3">
|
||
<emphasis id="B951387A68051076FF064C61FEB70557" bold="true" box="[199,334,548,573]" pageId="3">Etymology.</emphasis>
|
||
Latin adjective
|
||
<emphasis id="B951387A68051076FDC44C60FDA60556" box="[517,607,549,572]" italics="true" pageId="3">imitatus</emphasis>
|
||
(imitated, copied), alluding to the strong resemblance to the
|
||
<typeStatus id="549E5ACA68051076FAD24C60FABA0557" box="[1299,1347,549,573]" pageId="3">type</typeStatus>
|
||
species of the genus,
|
||
<taxonomicName id="4C259FEB68051076FEEC4C02FDED050A" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[301,532,583,608]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="3" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A68051076FEEC4C02FDED050A" box="[301,532,583,608]" italics="true" pageId="3">Fatkullina paradoxa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33FB7E36805107FFF064C29FB3006E2" lastPageId="10" pageId="3" type="description">
|
||
<paragraph id="8B9AE46868051076FF064C29FDC20452" blockId="3.[151,1436,151,2013]" pageId="3">
|
||
<emphasis id="B951387A68051076FF064C29FEA005EF" bold="true" box="[199,345,620,645]" pageId="3">Description.</emphasis>
|
||
Colony encrusting, multiserial, unilaminar, more or less circular, with undulating margin, attaining
|
||
<emphasis id="B951387A68051076FF3E4CD4FEF205C2" box="[255,267,657,680]" italics="true" pageId="3">c</emphasis>
|
||
.
|
||
<quantity id="4CDD498D68051076FEDB4CD5FE9105CD" box="[282,360,656,680]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="3" unit="mm" value="40.0">40 mm</quantity>
|
||
diameter. Colour (when dried) crimson (most typical), pink, dark red, deep purple to black (
|
||
<figureCitation id="131EF8ED68051076FAAE4CCAFF5905A6" captionStart="FIGURE 1" captionStartId="4.[151,250,1859,1881]" captionTargetBox="[195,1400,195,1840]" captionTargetId="figure@4.[195,1400,195,1840]" captionTargetPageId="4" captionText="FIGURE 1. General view of dried colonies of Fatkullina imitata n. sp. (A–D) and F. paradoxa Grischenko, Gordon & Taylor, 1998 (E). A, B, specimens R.V. Professor Kizevetter 2015, Stn 73; C, D, specimens paratype 8 NIWA 127751; E, specimen R.V. Professor Probatov 2013, Stn 5, 16 June 2013, 57.41694° N, 156.51806° E, 27 m, pebbles, sand, silt. A, B. Colonies encrusting pebbles. C, D. Colonies encrusting external (C), and internal (D) surface of shell fragment of Chlamys sp. E. Colony encrusting internal surface of bivalve shell fragment of Serripes sp., surrounded by other encrusting bryozoan species. Scale bars: 5 mm." httpUri="https://zenodo.org/record/2606860/files/figure.png" pageId="3">Fig. 1</figureCitation>
|
||
A–D); preserved polypides pink. Zooids relatively large, greatly variable in form (
|
||
<figureCitation id="131EF8ED68051076FB8F4CF1FB3605A6" box="[1102,1231,692,717]" captionStart="FIGURE 2" captionStartId="5.[151,250,1845,1867]" captionTargetBox="[151,1436,213,1824]" captionTargetId="figure@5.[151,1436,213,1824]" captionTargetPageId="5" captionText="FIGURE 2. Fatkullina imitata n. sp. A, Holotype ZIRAS 1/50661; B, paratype 5 ZIRAS 6/50666; C, paratype 4 ZIRAS 5/ 50665. A. General view of colony encrusting flattened pebble in boundary interactions with Celleporina ventricosa (Lorenz, 1886) (top left), Reussinella arctica (Osburn, 1950) (top right), Hippothoa arctica (Kluge, 1906), H. expansa Dawson, 1859 (bottom left) and Pachyegis princeps (Norman, 1903) (centrally to bottom right). B. General view of a medium-sized colony. C. General view of a young colony demonstrating a gradual shift in geometry of zooidal budding from spiral into reversedpolarity. Asterisks indicate zooids with a sideways (transverse) orientation. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2606862/files/figure.png" pageId="3">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="131EF8ED68051076FB1D4CF1FAD805A6" box="[1244,1313,692,716]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1846]" captionTargetBox="[218,1413,456,1798]" captionTargetId="figure@6.[218,1413,456,1798]" captionTargetPageId="6" captionText="FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="3">3C, D</figureCitation>
|
||
) including broadly hexagonal, oval, quadrangular, rhombic, pyriform, polygonal or irregular in outline, arranged quincuncially or less orderly, separated by fine undulating sutures between vertical walls; sutures occasionally concealed by secondary calcification.
|
||
</paragraph>
|
||
<paragraph id="8B9AE46868051076FF064D01FC9C0382" blockId="3.[151,1436,151,2013]" pageId="3">
|
||
Frontal shield (
|
||
<figureCitation id="131EF8ED68051076FEB34D01FE4E0436" box="[370,439,836,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1846]" captionTargetBox="[218,1413,456,1798]" captionTargetId="figure@6.[218,1413,456,1798]" captionTargetPageId="6" captionText="FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="3">Figs 3</figureCitation>
|
||
C–F, 4A–C, 5F, G) lepralioid, thick, convex, most elevated and inflated in distal third of zooid around secondary orifice, moderately convex to evenly flattened centrally and proximally; with finely granular-tubercular surface, uniformly perforated with conspicuous round to elongate oval areolar-septular pores along zooidal margins and numerous smaller, circular frontal pseudopores (
|
||
<figureCitation id="131EF8ED68051076FBC94DEAFB7F04AD" box="[1032,1158,943,968]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1846]" captionTargetBox="[218,1413,456,1798]" captionTargetId="figure@6.[218,1413,456,1798]" captionTargetPageId="6" captionText="FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="3">Figs 3E, F</figureCitation>
|
||
,
|
||
<figureCitation id="131EF8ED68051076FB544DEAFB0604A2" box="[1173,1279,943,968]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">5B, F, G</figureCitation>
|
||
); areolae and pseudopores becoming funnel-shaped with development of secondary calcification, with granulation extending deep inside; some pores occluded. Interior surface of frontal shield (
|
||
<figureCitation id="131EF8ED68051076FC0B4DB2FB8C037A" box="[970,1141,1015,1040]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">Fig. 5D, E, G</figureCitation>
|
||
) smooth, with numerous openings of pseudopores in center, and openings of tubular areolar pore channels around periphery. Some zooids, especially in elevated older parts of colony, with more-elongated area of calcification distal to orifice, with up to 3– 4 rows of pseudopores (
|
||
<figureCitation id="131EF8ED68051076FE744A21FE050316" box="[437,508,1124,1149]" captionStart="FIGURE 4" captionStartId="7.[151,250,1657,1679]" captionTargetBox="[151,1436,193,1636]" captionTargetId="figure@7.[151,1436,193,1636]" captionTargetPageId="7" captionText="FIGURE 4. Fatkullina imitata n. sp. A–C, E, F, paratype 1 ZIRAS 2/50662; D, holotype ZIRAS 1/50661. Intracolonial polymorphism. A. Strongly distended zooids in elevated colony center, with proportionally elongated and elevated distal (preoral) frontal wall terminating with conical and bulbous solid umbones (arrowheads). B. Two neighboring zooids, each bearing distal (preoral) conical knob (arrowheads), with lateral boundary occluded by secondary calcification, indicated by enlarged infundibular pseudopores; note minute shortened distal suture line separating preoral areas. C. Two oppositely orientated zooids with strongly swollen frontal shields and orifices shifted to the top of frontal surface. D. Irregularly hexagonal, flattened, uniformly perforated kenozooid with undulating margins, surrounded by six autozooids. E. Colony area comprising mixed kenozooids and autozooids; note irregular shape of flattened kenozooids possessing strongly reduced orificial areas remaining in form of larger frontal pseudopores (left to central kenozooids) or two minor pseudopores (right kenozooid). F. Colony area fully represented by flattened kenozooids with orifices constricted or closed by calcified cylindrical or mammiform projections with tiny circular to slit-like openings at their apices. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/2606870/files/figure.png" pageId="3">Fig. 4</figureCitation>
|
||
A–C). Distal margin of secondary orifice can bear a solid, elevated, conical or bulbous umbo (
|
||
<figureCitation id="131EF8ED68051076FE874AC2FE3803CA" box="[326,449,1159,1184]" captionStart="FIGURE 4" captionStartId="7.[151,250,1657,1679]" captionTargetBox="[151,1436,193,1636]" captionTargetId="figure@7.[151,1436,193,1636]" captionTargetPageId="7" captionText="FIGURE 4. Fatkullina imitata n. sp. A–C, E, F, paratype 1 ZIRAS 2/50662; D, holotype ZIRAS 1/50661. Intracolonial polymorphism. A. Strongly distended zooids in elevated colony center, with proportionally elongated and elevated distal (preoral) frontal wall terminating with conical and bulbous solid umbones (arrowheads). B. Two neighboring zooids, each bearing distal (preoral) conical knob (arrowheads), with lateral boundary occluded by secondary calcification, indicated by enlarged infundibular pseudopores; note minute shortened distal suture line separating preoral areas. C. Two oppositely orientated zooids with strongly swollen frontal shields and orifices shifted to the top of frontal surface. D. Irregularly hexagonal, flattened, uniformly perforated kenozooid with undulating margins, surrounded by six autozooids. E. Colony area comprising mixed kenozooids and autozooids; note irregular shape of flattened kenozooids possessing strongly reduced orificial areas remaining in form of larger frontal pseudopores (left to central kenozooids) or two minor pseudopores (right kenozooid). F. Colony area fully represented by flattened kenozooids with orifices constricted or closed by calcified cylindrical or mammiform projections with tiny circular to slit-like openings at their apices. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/2606870/files/figure.png" pageId="3">Fig. 4A, B</figureCitation>
|
||
, arrowheads), with orifice shifted proximally to a position about 1/4 to 1/3 zooid length from distal end. Two smaller, conical umbones can additionally flank sinus of secondary orifice, strongly reducing area of secondary orifice and conferring strictly triangular form.
|
||
</paragraph>
|
||
<paragraph id="8B9AE46868051076FF064AB1FF1F01F2" blockId="3.[151,1436,151,2013]" pageId="3">
|
||
Primary orifice (
|
||
<figureCitation id="131EF8ED68051076FE454AB1FE310267" box="[388,456,1268,1293]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">Fig. 5</figureCitation>
|
||
A–C, E) deeply sunken, wholly visible only at colony margin; transversely oval, with straight or weakly concave proximal margin and short, narrow U-shaped sinus; thin proximal margin lacking granulation. In fully completed zooids, primary orifice tilted proximally at angle of
|
||
<emphasis id="B951387A68051076FBAB4B78FB8F023E" box="[1130,1142,1341,1364]" italics="true" pageId="3">c</emphasis>
|
||
. 20–40° to frontal plane. Secondary orifice (
|
||
<figureCitation id="131EF8ED68051076FEAF4B1AFE300212" box="[366,457,1375,1400]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1846]" captionTargetBox="[218,1413,456,1798]" captionTargetId="figure@6.[218,1413,456,1798]" captionTargetPageId="6" captionText="FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="3">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation id="131EF8ED68051076FE174B1AFE0E0212" box="[470,503,1375,1400]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">5B</figureCitation>
|
||
) formed by thickening of frontal shield around primary orifice; roughly triangular to drop-shaped in outline, with finely granular tubercular margin; at highest elevation of frontal shield. Distal margin gently convex, arching and overhanging 1/3 to 2/3 of distal part of primary orifice and partially concealing its lateral margins (
|
||
<figureCitation id="131EF8ED68051076FE404B89FDFF028E" box="[385,518,1484,1509]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">Fig. 5B, C</figureCitation>
|
||
). Proximolateral margins sloping abruptly within, forming broad to narrowly triangular pseudosinus descending to smooth suboral rim of primary orifice. Lumen of secondary orifice (
|
||
<figureCitation id="131EF8ED68051076FAFC4BAAFF5E0146" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1846]" captionTargetBox="[218,1413,456,1798]" captionTargetId="figure@6.[218,1413,456,1798]" captionTargetPageId="6" captionText="FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="3">Figs 3E, F</figureCitation>
|
||
,
|
||
<figureCitation id="131EF8ED68051076FF724851FEEF0146" box="[179,278,1556,1581]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">5C, E, G</figureCitation>
|
||
) deep, tubular to infundibular, tilted proximally at angle about 30–45° to frontal plane. Some zooids in central colony region secondarily kenozooidal (
|
||
<figureCitation id="131EF8ED68051076FD0F4872FCE1013A" box="[718,792,1591,1616]" captionStart="FIGURE 4" captionStartId="7.[151,250,1657,1679]" captionTargetBox="[151,1436,193,1636]" captionTargetId="figure@7.[151,1436,193,1636]" captionTargetPageId="7" captionText="FIGURE 4. Fatkullina imitata n. sp. A–C, E, F, paratype 1 ZIRAS 2/50662; D, holotype ZIRAS 1/50661. Intracolonial polymorphism. A. Strongly distended zooids in elevated colony center, with proportionally elongated and elevated distal (preoral) frontal wall terminating with conical and bulbous solid umbones (arrowheads). B. Two neighboring zooids, each bearing distal (preoral) conical knob (arrowheads), with lateral boundary occluded by secondary calcification, indicated by enlarged infundibular pseudopores; note minute shortened distal suture line separating preoral areas. C. Two oppositely orientated zooids with strongly swollen frontal shields and orifices shifted to the top of frontal surface. D. Irregularly hexagonal, flattened, uniformly perforated kenozooid with undulating margins, surrounded by six autozooids. E. Colony area comprising mixed kenozooids and autozooids; note irregular shape of flattened kenozooids possessing strongly reduced orificial areas remaining in form of larger frontal pseudopores (left to central kenozooids) or two minor pseudopores (right kenozooid). F. Colony area fully represented by flattened kenozooids with orifices constricted or closed by calcified cylindrical or mammiform projections with tiny circular to slit-like openings at their apices. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/2606870/files/figure.png" pageId="3">Fig. 4</figureCitation>
|
||
D–F), with orifice constricted or closed, often with a cylindrical or mammiform projection over orifice. Operculum dark brown. Oral spines, avicularia and ooecia absent.
|
||
</paragraph>
|
||
<paragraph id="8B9AE46868051076FF0648E1FA610042" blockId="3.[151,1436,151,2013]" pageId="3">
|
||
Larval incubation presumably occurs in internal brood sac. Interzooidal communication via uniporous septula recessed between buttresses (
|
||
<figureCitation id="131EF8ED68051076FE2A4882FD94018A" box="[491,621,1735,1760]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1846]" captionTargetBox="[218,1413,456,1798]" captionTargetId="figure@6.[218,1413,456,1798]" captionTargetPageId="6" captionText="FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="3">Fig. 3A, B</figureCitation>
|
||
). Basal wall fully calcified (
|
||
<figureCitation id="131EF8ED68051076FC7F4882FBC5018A" box="[958,1084,1735,1760]" captionStart="FIGURE 5" captionStartId="8.[151,250,1494,1516]" captionTargetBox="[224,1376,193,1483]" captionTargetId="figure@8.[224,1376,193,1483]" captionTargetPageId="8" captionText="FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="3">Fig. 5F, G</figureCitation>
|
||
). Zooidal morphogenesis with reversed polarity, i.e. orifice situated at proximal end of zooid relative to colony orientation and directed towards ancestrula. Kenozooids rarely single, distributed among autozooids, more frequently multiple, united into clusters.
|
||
</paragraph>
|
||
<paragraph id="8B9AE46868051073FF064971FA8806E2" blockId="3.[151,1436,151,2013]" lastBlockId="6.[151,1436,151,392]" lastPageId="6" pageId="3">
|
||
Ancestrula (
|
||
<figureCitation id="131EF8ED68051076FE914971FE6B0027" box="[336,402,1844,1869]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="3">Fig. 6</figureCitation>
|
||
A–H) resembling autozooids in general appearance; roughly hexagonal, convex, with finely granular surface and a few marginal and frontal pores. Ancestrular orifice circular, with deep V-shaped sinus; cormidial, bounded proximally by distal part of frontal shield, distally and laterally by margins of vertical walls of distal and lateral periancestrular zooids; boundaries defined by fine sutures externally (
|
||
<figureCitation id="131EF8ED68051076FBA149E5FB4F00D3" box="[1120,1206,1952,1977]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="3">Fig. 6B</figureCitation>
|
||
) and internally (
|
||
<figureCitation id="131EF8ED68051076FAAE49E5FF4C00B7" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="3">Fig. 6F</figureCitation>
|
||
, arrowheads). Interior of ancestrular frontal shield (
|
||
<figureCitation id="131EF8ED68051076FD3E4981FCBB00B7" box="[767,834,1988,2013]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="3">Fig. 6</figureCitation>
|
||
D–F) with mixed (suboral umbonuloid and proximal lepralioid) components; suboral part unequally divided by four radial sutures (
|
||
<figureCitation id="131EF8ED68001073FBD74ED2FB9007DA" box="[1046,1129,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="6">Fig. 6F</figureCitation>
|
||
, arrowheads), two of them flanking the sinus, apparently corresponding externally (in part, at least) to sutures seen frontally between other components of ancestrular complex. Ring scar discrete (
|
||
<figureCitation id="131EF8ED68001073FCCC4E9AFC9B0792" box="[781,866,223,248]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="6">Fig. 6F</figureCitation>
|
||
), forming very regular boundary between exteriorwall microstructure (planar-spherulitic fabric) of umbonuloid part and extra-umbonuloid calcification. Umbo extensive, occupying about 58% of length of frontal shield. Proximal lepralioid part of shield with smooth interior surface containing scattered tiny, uniformly circular, pseudopores (
|
||
<figureCitation id="131EF8ED68001073FC584F09FC0A060F" box="[921,1011,332,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="6">Fig. 6E</figureCitation>
|
||
). Ancestrula unrecognizable in older region of large colonies owing to secondary calcification that conceals orifices and crosses zooidal boundaries.
|
||
</paragraph>
|
||
<caption id="DF5AB4E068021071FF564906FEE90098" httpUri="https://zenodo.org/record/2606860/files/figure.png" pageId="4" startId="4.[151,250,1859,1881]" targetBox="[195,1400,195,1840]" targetPageId="4">
|
||
<paragraph id="8B9AE46868021071FF564906FEE90098" blockId="4.[151,1436,1859,2034]" pageId="4">
|
||
<emphasis id="B951387A68021071FF564906FEEA0033" bold="true" box="[151,275,1859,1881]" pageId="4">FIGURE 1.</emphasis>
|
||
General view of dried colonies of
|
||
<taxonomicName id="4C259FEB68021071FDB24906FCD20033" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[627,811,1859,1881]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="4" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A68021071FDB24906FCD20033" box="[627,811,1859,1881]" italics="true" pageId="4">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A68021071FCF04906FC930033" bold="true" box="[817,874,1859,1881]" pageId="4">
|
||
<taxonomicNameLabel id="A262850168021071FCF04906FC930033" box="[817,874,1859,1881]" pageId="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A–D) and
|
||
<taxonomicName id="4C259FEB68021071FC254901FF330012" authority="Grischenko, Gordon & Taylor, 1998" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="4" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A68021071FC254901FC080033" box="[996,1009,1860,1881]" italics="true" pageId="4">F</emphasis>
|
||
.
|
||
<emphasis id="B951387A68021071FC3F4906FBA60033" box="[1022,1119,1859,1881]" italics="true" pageId="4">paradoxa</emphasis>
|
||
Grischenko, Gordon & Taylor, 1998
|
||
</taxonomicName>
|
||
(E). A, B, specimens R.V.
|
||
<emphasis id="B951387A68021071FE264927FD410012" box="[487,696,1890,1912]" italics="true" pageId="4">Professor Kizevetter</emphasis>
|
||
2015, Stn 73; C, D, specimens paratype 8 NIWA 127751; E, specimen R.V.
|
||
<emphasis id="B951387A68021071FF0D49C4FE6A00FD" box="[204,403,1921,1943]" italics="true" pageId="4">Professor Probatov</emphasis>
|
||
2013, Stn 5, 16 June 2013, 57.41694° N, 156.51806° E, 27 m, pebbles, sand, silt. A, B. Colonies encrusting pebbles. C, D. Colonies encrusting external (C), and internal (D) surface of shell fragment of
|
||
<taxonomicName id="4C259FEB68021071FB7549DAFAF400DF" box="[1204,1293,1951,1973]" class="Bivalvia" family="Pectinidae" genus="Chlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="4" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A68021071FB7549DAFAF400DF" box="[1204,1293,1951,1973]" italics="true" pageId="4">Chlamys</emphasis>
|
||
</taxonomicName>
|
||
sp. E. Colony encrusting internal surface of bivalve shell fragment of
|
||
<taxonomicName id="4C259FEB68021071FD1649FAFCD200BE" box="[727,811,1983,2004]" class="Bivalvia" family="Cardiidae" genus="Serripes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cardiida" pageId="4" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A68021071FD1649FAFCD200BE" box="[727,811,1983,2004]" italics="true" pageId="4">Serripes</emphasis>
|
||
</taxonomicName>
|
||
sp., surrounded by other encrusting bryozoan species. Scale bars: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5AB4E068031070FF564970FBB3009E" httpUri="https://zenodo.org/record/2606862/files/figure.png" pageId="5" startId="5.[151,250,1845,1867]" targetBox="[151,1436,213,1824]" targetPageId="5">
|
||
<paragraph id="8B9AE46868031070FF564970FBB3009E" blockId="5.[151,1437,1845,2036]" pageId="5">
|
||
<emphasis id="B951387A68031070FF564970FEED0021" bold="true" box="[151,276,1845,1867]" pageId="5">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C259FEB68031070FEDD4970FE2F0021" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[284,470,1845,1867]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="5" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A68031070FEDD4970FE2F0021" box="[284,470,1845,1867]" italics="true" pageId="5">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A68031070FE1F4970FDE00021" bold="true" box="[478,537,1845,1867]" pageId="5">
|
||
<taxonomicNameLabel id="A262850168031070FE1F4970FDE00021" box="[478,537,1845,1867]" pageId="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, Holotype ZIRAS 1/50661; B, paratype 5 ZIRAS 6/50666; C, paratype 4 ZIRAS 5/ 50665. A. General view of colony encrusting flattened pebble in boundary interactions with
|
||
<taxonomicName id="4C259FEB68031070FB924913FF2B00E4" authority="(Lorenz, 1886)" authorityName="Lorenz" authorityYear="1886" baseAuthorityName="Lorenz" baseAuthorityYear="1886" class="Gymnolaemata" family="Celleporidae" genus="Celleporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="5" phylum="Bryozoa" rank="species" species="ventricosa">
|
||
<emphasis id="B951387A68031070FB924913FAC70007" box="[1107,1342,1878,1901]" italics="true" pageId="5">Celleporina ventricosa</emphasis>
|
||
(Lorenz, 1886)
|
||
</taxonomicName>
|
||
(top left),
|
||
<taxonomicName id="4C259FEB68031070FE82493DFD5200E4" authority="(Osburn, 1950)" authorityName="Osburn" authorityYear="1950" baseAuthorityName="Osburn" baseAuthorityYear="1950" box="[323,683,1912,1934]" class="Gymnolaemata" family="Microporidae" genus="Reussinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="5" phylum="Bryozoa" rank="species" species="arctica">
|
||
<emphasis id="B951387A68031070FE82493DFDFE00E4" box="[323,519,1912,1934]" italics="true" pageId="5">Reussinella arctica</emphasis>
|
||
(Osburn, 1950)
|
||
</taxonomicName>
|
||
(top right),
|
||
<taxonomicName id="4C259FEB68031070FCEA493DFB7900E4" authority="(Kluge, 1906)" authorityName="Kluge" authorityYear="1906" baseAuthorityName="Kluge" baseAuthorityYear="1906" box="[811,1152,1912,1934]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="5" phylum="Bryozoa" rank="species" species="arctica">
|
||
<emphasis id="B951387A68031070FCEA493DFC1F00E4" box="[811,998,1912,1934]" italics="true" pageId="5">Hippothoa arctica</emphasis>
|
||
(Kluge, 1906)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C259FEB68031070FB4D493CFA6500E4" authority="Dawson, 1859" authorityName="Dawson" authorityYear="1859" box="[1164,1436,1912,1934]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="5" phylum="Bryozoa" rank="species" species="expansa">
|
||
<emphasis id="B951387A68031070FB4D493CFB6700E4" box="[1164,1182,1913,1934]" italics="true" pageId="5">H</emphasis>
|
||
.
|
||
<emphasis id="B951387A68031070FB6C493CFAF900E4" box="[1197,1280,1913,1934]" italics="true" pageId="5">expansa</emphasis>
|
||
Dawson, 1859
|
||
</taxonomicName>
|
||
(bottom left) and
|
||
<taxonomicName id="4C259FEB68031070FE8D49DCFD4400C5" authority="(Norman, 1903)" authorityName="Norman" authorityYear="1903" baseAuthorityName="Norman" baseAuthorityYear="1903" box="[332,701,1945,1967]" class="Gymnolaemata" family="Stomachetosellidae" genus="Pachyegis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="5" phylum="Bryozoa" rank="species" species="princeps">
|
||
<emphasis id="B951387A68031070FE8D49DCFDEB00C5" box="[332,530,1945,1967]" italics="true" pageId="5">Pachyegis princeps</emphasis>
|
||
(Norman, 1903)
|
||
</taxonomicName>
|
||
(centrally to bottom right). B. General view of a medium-sized colony. C. General view of a young colony demonstrating a gradual shift in geometry of zooidal budding from spiral into reversedpolarity. Asterisks indicate zooids with a sideways (transverse) orientation. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5AB4E068001073FF564965FF3F0084" httpUri="https://zenodo.org/record/2606866/files/figure.png" pageId="6" startId="6.[151,250,1824,1846]" targetBox="[218,1413,456,1798]" targetPageId="6">
|
||
<paragraph id="8B9AE46868001073FF564965FF3F0084" blockId="6.[151,1436,1824,2030]" pageId="6">
|
||
<emphasis id="B951387A68001073FF564965FEEA005C" bold="true" box="[151,275,1824,1846]" pageId="6">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C259FEB68001073FEDB4965FE2A005C" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[282,467,1824,1846]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="6" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A68001073FEDB4965FE2A005C" box="[282,467,1824,1846]" italics="true" pageId="6">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A68001073FE184965FDEA005C" bold="true" box="[473,531,1824,1846]" pageId="6">
|
||
<taxonomicNameLabel id="A262850168001073FE184965FDEA005C" box="[473,531,1824,1846]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5AB4E068011072FF56483CFB5800A8" httpUri="https://zenodo.org/record/2606870/files/figure.png" pageId="7" startId="7.[151,250,1657,1679]" targetBox="[151,1436,193,1636]" targetPageId="7">
|
||
<paragraph id="8B9AE46868011072FF56483CFB5800A8" blockId="7.[151,1436,1657,1986]" pageId="7">
|
||
<emphasis id="B951387A68011072FF56483CFEEE01E5" bold="true" box="[151,279,1657,1679]" pageId="7">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C259FEB68011072FEE0483CFE2401E5" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[289,477,1657,1679]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="7" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A68011072FEE0483CFE2401E5" box="[289,477,1657,1679]" italics="true" pageId="7">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A68011072FE26483CFDDD01E5" bold="true" box="[487,548,1657,1679]" pageId="7">
|
||
<taxonomicNameLabel id="A262850168011072FE26483CFDDD01E5" box="[487,548,1657,1679]" pageId="7" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–C, E, F, paratype 1 ZIRAS 2/50662; D, holotype ZIRAS 1/50661. Intracolonial polymorphism. A. Strongly distended zooids in elevated colony center, with proportionally elongated and elevated distal (preoral) frontal wall terminating with conical and bulbous solid umbones (arrowheads). B. Two neighboring zooids, each bearing distal (preoral) conical knob (arrowheads), with lateral boundary occluded by secondary calcification, indicated by enlarged infundibular pseudopores; note minute shortened distal suture line separating preoral areas. C. Two oppositely orientated zooids with strongly swollen frontal shields and orifices shifted to the top of frontal surface. D. Irregularly hexagonal, flattened, uniformly perforated kenozooid with undulating margins, surrounded by six autozooids. E. Colony area comprising mixed kenozooids and autozooids; note irregular shape of flattened kenozooids possessing strongly reduced orificial areas remaining in form of larger frontal pseudopores (left to central kenozooids) or two minor pseudopores (right kenozooid). F. Colony area fully represented by flattened kenozooids with orifices constricted or closed by calcified cylindrical or mammiform projections with tiny circular to slit-like openings at their apices. Scale bars: 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5AB4E0680E107DFF564B93FE8B009C" httpUri="https://zenodo.org/record/2606872/files/figure.png" pageId="8" startId="8.[151,250,1494,1516]" targetBox="[224,1376,193,1483]" targetPageId="8">
|
||
<paragraph id="8B9AE468680E107DFF564B93FE8B009C" blockId="8.[151,1437,1494,2038]" pageId="8">
|
||
<emphasis id="B951387A680E107DFF564B93FEEA0286" bold="true" box="[151,275,1494,1516]" pageId="8">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C259FEB680E107DFEDB4B93FE2A0287" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[282,467,1494,1517]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="8" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680E107DFEDB4B93FE2A0287" box="[282,467,1494,1517]" italics="true" pageId="8">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680E107DFE1B4B93FDEA0286" bold="true" box="[474,531,1494,1516]" pageId="8">
|
||
<taxonomicNameLabel id="A2628501680E107DFE1B4B93FDEA0286" box="[474,531,1494,1516]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5AB4E0680F107CFF564803FA800084" httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="9" startId="9.[151,250,1606,1628]" targetBox="[238,1356,183,1593]" targetPageId="9">
|
||
<paragraph id="8B9AE468680F107CFF564803FA800084" blockId="9.[151,1437,1606,2030]" pageId="9">
|
||
<emphasis id="B951387A680F107CFF564803FEEB0136" bold="true" box="[151,274,1606,1628]" pageId="9">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C259FEB680F107CFED94803FE290136" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[280,464,1606,1628]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="9" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680F107CFED94803FE290136" box="[280,464,1606,1628]" italics="true" pageId="9">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680F107CFE144803FDF70136" bold="true" box="[469,526,1606,1628]" pageId="9">
|
||
<taxonomicNameLabel id="A2628501680F107CFE144803FDF70136" box="[469,526,1606,1628]" pageId="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V.
|
||
<emphasis id="B951387A680F107CFBE44802FB970136" box="[1061,1134,1607,1628]" italics="true" pageId="9">TINRO</emphasis>
|
||
2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9AE468680C107FFF064ED2FB3006E2" blockId="10.[151,1436,151,392]" pageId="10">
|
||
Ancestrula buds triplet of somewhat spirally arranged periancestrular zooids, integrated into ancestrular complex. First daughter zooid of complex differentiating distally (
|
||
<figureCitation id="131EF8ED680C107FFCBA4EF9FC0E07BE" box="[891,1015,188,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="10">Fig. 6A, D</figureCitation>
|
||
), perpendicular to ancestrula, slightly curved along long axis, giving T-shaped appearance to initial stage of ancestrular complex (seen in a dozen such newly established colonies), and two periancestrular zooids then differentiating laterally on each side. Orifices of distolateral zooids can be orientated antiparallel to ancestrula (
|
||
<figureCitation id="131EF8ED680C107FFC994F62FC29062A" box="[856,976,295,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="10">Fig. 6B, C</figureCitation>
|
||
, right zooid;
|
||
<figureCitation id="131EF8ED680C107FFBA64F62FB45062A" box="[1127,1212,295,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="10">Fig. 6G</figureCitation>
|
||
, left zooid) or same direction as ancestrula (
|
||
<figureCitation id="131EF8ED680C107FFE604F09FE0E060F" box="[417,503,332,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1606,1628]" captionTargetBox="[238,1356,183,1593]" captionTargetId="figure@9.[238,1356,183,1593]" captionTargetPageId="9" captionText="FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." httpUri="https://zenodo.org/record/2606874/files/figure.png" pageId="10">Fig. 6G</figureCitation>
|
||
, right zooid). Weakly spiral budding pattern continued through zone of astogenetic change and visible throughout four to ten or more generations of zooids distant from ancestrula.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5AB4E0680C107FFF564FFDFD370505" pageId="10">
|
||
<paragraph id="8B9AE468680C107FFF564FFDFD370505" blockId="10.[151,1435,440,624]" pageId="10">
|
||
<emphasis id="B951387A680C107FFF564FFDFEF106A5" bold="true" box="[151,264,440,463]" pageId="10">TABLE 1.</emphasis>
|
||
Measurements (in mm, except number of marginal and frontal pores) of the holotype and paratype specimens of
|
||
<taxonomicName id="4C259FEB680C107FFF744F92FE800685" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[181,377,471,495]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680C107FFF744F92FE800685" box="[181,377,471,495]" italics="true" pageId="10">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680C107FFE404F9DFE390685" bold="true" box="[385,448,472,495]" pageId="10">
|
||
<taxonomicNameLabel id="A2628501680C107FFE404F9DFE390685" box="[385,448,472,495]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Abbreviations: ZL, zooid length; ZW, zooid width; ZD, zooid depth (height); OrL, orifice (without sinus) length; Or(s)L, orificial sinus length; OrW, orifice width; P(m)N, number of marginal pores (areolae); P(f)N, number of frontal pseudopores; AnL, ancestrula length; AnW, ancestrula width; AnOrL, ancestrular orifice (without sinus) length; AnOr(s)L, ancestrular orificial sinus length; AnOrW, ancestrular orifice width; N, number of measurements (unless otherwise indicated,
|
||
<emphasis id="B951387A680C107FFDB44C1CFD7A051A" box="[629,643,601,624]" italics="true" pageId="10">n</emphasis>
|
||
= 20).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9AE468680C107FFED74CC9FB3D0219" pageId="10">
|
||
<table id="F92516C8680CEF8AFF5E4CC9FA6D0219" box="[159,1428,652,1395]" gridcols="5" gridrows="14" pageId="10">
|
||
<tr id="3515E62A680CEF8AFF5E4CC9FA6D05C8" box="[159,1428,652,674]" gridrow="0" pageId="10" rowspan-0="1">
|
||
<th id="76C48F56680CEF8AFED74CC9FDE405C8" box="[278,541,652,674]" gridcol="1" gridrow="0" pageId="10">Holotype ZIRAS 1/50661</th>
|
||
<th id="76C48F56680CEF8AFDF64CC9FCB505C8" box="[567,844,652,674]" gridcol="2" gridrow="0" pageId="10">Paratype 1 ZIRAS 2/50662</th>
|
||
<th id="76C48F56680CEF8AFC9A4CC9FB8905C8" box="[859,1136,652,674]" gridcol="3" gridrow="0" pageId="10">Paratype 2 ZIRAS 3/50663</th>
|
||
<th id="76C48F56680CEF8AFB414CC9FA6D05C8" box="[1152,1428,652,674]" gridcol="4" gridrow="0" pageId="10">Paratype 3 ZIRAS 4/50664</th>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4CF0FA6D05A1" box="[159,1428,693,715]" gridrow="1" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4CF0FEFB05A1" box="[159,258,693,715]" gridcol="0" gridrow="1" pageId="10">ZL</th>
|
||
<td id="76C48F56680CEF8AFED74CF0FDE405A1" box="[278,541,693,715]" gridcol="1" gridrow="1" pageId="10">0.65–1.03 (0.84 ± 0.10)</td>
|
||
<td id="76C48F56680CEF8AFDF64CF0FCB505A1" box="[567,844,693,715]" gridcol="2" gridrow="1" pageId="10">0.65–1.05 (0.80 ± 0.09)</td>
|
||
<td id="76C48F56680CEF8AFC9A4CF0FB8905A1" box="[859,1136,693,715]" gridcol="3" gridrow="1" pageId="10">0.60–0.85 (0.75 ± 0.07)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4C9BFA6D059E" box="[159,1428,734,756]" gridrow="2" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4C9BFEFB059E" box="[159,258,734,756]" gridcol="0" gridrow="2" pageId="10">ZW</th>
|
||
<td id="76C48F56680CEF8AFED74C9BFDE4059E" box="[278,541,734,756]" gridcol="1" gridrow="2" pageId="10">0.48–0.80 (0.65 ± 0.09)</td>
|
||
<td id="76C48F56680CEF8AFDF64C9BFCB5059E" box="[567,844,734,756]" gridcol="2" gridrow="2" pageId="10">0.44–0.78 (0.62 ± 0.08)</td>
|
||
<td id="76C48F56680CEF8AFC9A4C9BFB89059E" box="[859,1136,734,756]" gridcol="3" gridrow="2" pageId="10">0.45–0.78 (0.61 ± 0.08)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4D4DFA6D0457" box="[159,1428,776,829]" gridrow="3" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4D4DFEFB0457" box="[159,258,776,829]" gridcol="0" gridrow="3" pageId="10">ZD</th>
|
||
<td id="76C48F56680CEF8AFED74D4DFDE40457" box="[278,541,776,829]" gridcol="1" gridrow="3" pageId="10">
|
||
0.53–0.68 (0.59 ± 0.06) (
|
||
<emphasis id="B951387A680C107FFEDF4D6DFED20457" box="[286,299,808,829]" italics="true" pageId="10">n</emphasis>
|
||
= 5)
|
||
</td>
|
||
<td id="76C48F56680CEF8AFDF64D4DFCB50457" box="[567,844,776,829]" gridcol="2" gridrow="3" pageId="10">
|
||
0.45–0.60 (0.51 ± 0.06) (
|
||
<emphasis id="B951387A680C107FFD814D6DFDB40457" box="[576,589,808,829]" italics="true" pageId="10">n</emphasis>
|
||
= 5)
|
||
</td>
|
||
<td id="76C48F56680CEF8AFC9A4D4DFB890457" box="[859,1136,776,829]" gridcol="3" gridrow="3" pageId="10">
|
||
0.45–0.65 (0.55 ± 0.09) (
|
||
<emphasis id="B951387A680C107FFCA54D6DFC880457" box="[868,881,808,829]" italics="true" pageId="10">n</emphasis>
|
||
= 5)
|
||
</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4D15FA6D040C" box="[159,1428,848,870]" gridrow="4" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4D15FEFB040C" box="[159,258,848,870]" gridcol="0" gridrow="4" pageId="10">OrL</th>
|
||
<td id="76C48F56680CEF8AFED74D15FDE4040C" box="[278,541,848,870]" gridcol="1" gridrow="4" pageId="10">0.10–0.15 (0.12 ± 0.02)</td>
|
||
<td id="76C48F56680CEF8AFDF64D15FCB5040C" box="[567,844,848,870]" gridcol="2" gridrow="4" pageId="10">0.10–0.15 (0.12 ± 0.01)</td>
|
||
<td id="76C48F56680CEF8AFC9A4D15FB89040C" box="[859,1136,848,870]" gridcol="3" gridrow="4" pageId="10">0.10–0.14 (0.12 ± 0.01)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4D3CFA6D04E5" box="[159,1428,889,911]" gridrow="5" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4D3CFEFB04E5" box="[159,258,889,911]" gridcol="0" gridrow="5" pageId="10">Or(s)L</th>
|
||
<td id="76C48F56680CEF8AFED74D3CFDE404E5" box="[278,541,889,911]" gridcol="1" gridrow="5" pageId="10">0.02–0.05 (0.03 ± 0.01)</td>
|
||
<td id="76C48F56680CEF8AFDF64D3CFCB504E5" box="[567,844,889,911]" gridcol="2" gridrow="5" pageId="10">0.02–0.05 (0.03 ± 0.01)</td>
|
||
<td id="76C48F56680CEF8AFC9A4D3CFB8904E5" box="[859,1136,889,911]" gridcol="3" gridrow="5" pageId="10">0.02–0.04 (0.03 ± 0.01)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4DE7FA6D04D2" box="[159,1428,930,952]" gridrow="6" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4DE7FEFB04D2" box="[159,258,930,952]" gridcol="0" gridrow="6" pageId="10">OrW</th>
|
||
<td id="76C48F56680CEF8AFED74DE7FDE404D2" box="[278,541,930,952]" gridcol="1" gridrow="6" pageId="10">0.16–0.23 (0.19 ± 0.02)</td>
|
||
<td id="76C48F56680CEF8AFDF64DE7FCB504D2" box="[567,844,930,952]" gridcol="2" gridrow="6" pageId="10">0.16–0.23 (0.19 ± 0.02)</td>
|
||
<td id="76C48F56680CEF8AFC9A4DE7FB8904D2" box="[859,1136,930,952]" gridcol="3" gridrow="6" pageId="10">0.16–0.21 (0.18 ± 0.01)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4D89FA6D0488" box="[159,1428,972,994]" gridrow="7" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4D89FEFB0488" box="[159,258,972,994]" gridcol="0" gridrow="7" pageId="10">P(m)N</th>
|
||
<td id="76C48F56680CEF8AFED74D89FDE40488" box="[278,541,972,994]" gridcol="1" gridrow="7" pageId="10">18–23 (21.05 ± 1.36)</td>
|
||
<td id="76C48F56680CEF8AFDF64D89FCB50488" box="[567,844,972,994]" gridcol="2" gridrow="7" pageId="10">19–25 (21.80 ± 1.70)</td>
|
||
<td id="76C48F56680CEF8AFC9A4D89FB890488" box="[859,1136,972,994]" gridcol="3" gridrow="7" pageId="10">17–23 (19.25 ± 1.68)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4DB0FA6D0361" box="[159,1428,1013,1035]" gridrow="8" pageId="10" rowspan-4="1">
|
||
<th id="76C48F56680CEF8AFF5E4DB0FEFB0361" box="[159,258,1013,1035]" gridcol="0" gridrow="8" pageId="10">P(f)N</th>
|
||
<td id="76C48F56680CEF8AFED74DB0FDE40361" box="[278,541,1013,1035]" gridcol="1" gridrow="8" pageId="10">16–32 (20.95 ± 3.38)</td>
|
||
<td id="76C48F56680CEF8AFDF64DB0FCB50361" box="[567,844,1013,1035]" gridcol="2" gridrow="8" pageId="10">16–36 (25.95 ± 5.75)</td>
|
||
<td id="76C48F56680CEF8AFC9A4DB0FB890361" box="[859,1136,1013,1035]" gridcol="3" gridrow="8" pageId="10">9–20 (14.10 ± 3.02)</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4A5BFA6D0339" box="[159,1428,1054,1107]" gridrow="9" pageId="10" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76C48F56680CEF8AFF5E4A5BFEFB0339" box="[159,258,1054,1107]" gridcol="0" gridrow="9" pageId="10">AnL</th>
|
||
<td id="76C48F56680CEF8AFB414A5BFA6D0339" box="[1152,1428,1054,1107]" gridcol="4" gridrow="9" pageId="10">
|
||
0.64–0.82 (0.70 ± 0.06) (
|
||
<emphasis id="B951387A680C107FFB494A7BFB6C0339" box="[1160,1173,1086,1107]" italics="true" pageId="10">n</emphasis>
|
||
= 7)
|
||
</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4A23FA6D03F1" box="[159,1428,1126,1179]" gridrow="10" pageId="10" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76C48F56680CEF8AFF5E4A23FEFB03F1" box="[159,258,1126,1179]" gridcol="0" gridrow="10" pageId="10">AnW</th>
|
||
<td id="76C48F56680CEF8AFB414A23FA6D03F1" box="[1152,1428,1126,1179]" gridcol="4" gridrow="10" pageId="10">
|
||
0.45–0.55 (0.50 ± 0.03) (
|
||
<emphasis id="B951387A680C107FFB494AC3FB6C03F1" box="[1160,1173,1158,1179]" italics="true" pageId="10">n</emphasis>
|
||
= 7)
|
||
</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4AEBFA6D0389" box="[159,1428,1198,1251]" gridrow="11" pageId="10" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76C48F56680CEF8AFF5E4AEBFEFB0389" box="[159,258,1198,1251]" gridcol="0" gridrow="11" pageId="10">AnOrL</th>
|
||
<td id="76C48F56680CEF8AFB414AEBFA6D0389" box="[1152,1428,1198,1251]" gridcol="4" gridrow="11" pageId="10">
|
||
0.12–0.14 (0.13 ± 0.01) (
|
||
<emphasis id="B951387A680C107FFB494A8BFB6C0389" box="[1160,1173,1230,1251]" italics="true" pageId="10">n</emphasis>
|
||
= 7)
|
||
</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4AB3FA6D0241" box="[159,1428,1270,1323]" gridrow="12" pageId="10" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76C48F56680CEF8AFF5E4AB3FEFB0241" box="[159,258,1270,1323]" gridcol="0" gridrow="12" pageId="10">AnOr(s)L</th>
|
||
<td id="76C48F56680CEF8AFB414AB3FA6D0241" box="[1152,1428,1270,1323]" gridcol="4" gridrow="12" pageId="10">
|
||
0.02–0.04 (0.03 ± 0.01) (
|
||
<emphasis id="B951387A680C107FFB494B53FB6C0241" box="[1160,1173,1302,1323]" italics="true" pageId="10">n</emphasis>
|
||
= 7)
|
||
</td>
|
||
</tr>
|
||
<tr id="3515E62A680CEF8AFF5E4B7BFA6D0219" box="[159,1428,1342,1395]" gridrow="13" pageId="10" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76C48F56680CEF8AFF5E4B7BFEFB0219" box="[159,258,1342,1395]" gridcol="0" gridrow="13" pageId="10">AnOrW</th>
|
||
<td id="76C48F56680CEF8AFB414B7BFA6D0219" box="[1152,1428,1342,1395]" gridcol="4" gridrow="13" pageId="10">
|
||
0.15–0.17 (0.16 ± 0.01) (
|
||
<emphasis id="B951387A680C107FFB494B1BFB6C0219" box="[1160,1173,1374,1395]" italics="true" pageId="10">n</emphasis>
|
||
= 7)
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C33FB7E3680C107EFF064BF2FC51047F" lastPageId="11" pageId="10" type="discussion">
|
||
<paragraph id="8B9AE468680C107FFF064BF2FC6D0172" blockId="10.[151,1436,1463,1560]" pageId="10">
|
||
<emphasis id="B951387A680C107FFF064BF2FEC202BA" bold="true" box="[199,315,1463,1488]" pageId="10">Remarks.</emphasis>
|
||
In having zooids with reversed polarity and occasionally possessing deep-purple to black colony colour,
|
||
<taxonomicName id="4C259FEB680C107FFF2A4B99FE4F029E" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[235,438,1500,1525]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680C107FFF2A4B99FE4F029E" box="[235,438,1500,1525]" italics="true" pageId="10">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680C107FFE7D4B99FE03029F" bold="true" box="[444,506,1500,1525]" pageId="10">
|
||
<taxonomicNameLabel id="A2628501680C107FFE7D4B99FE03029F" box="[444,506,1500,1525]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
strongly resembles the
|
||
<typeStatus id="549E5ACA680C107FFCC34B98FCCB029F" box="[770,818,1501,1525]" pageId="10">type</typeStatus>
|
||
species of the genus,
|
||
<taxonomicName id="4C259FEB680C107FFBE74B98FB55029F" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[1062,1196,1500,1525]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680C107FFBE74B98FB55029F" box="[1062,1196,1500,1525]" italics="true" pageId="10">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
, and can be confused with it. The two species differ, however, in the following characters:
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680C107FFF56480DFD8F01C2" blockId="10.[151,1437,1607,2029]" pageId="10">
|
||
1 Frontal shield convexity—maximally convex centrally to proximally, with a comparatively depressed orificial area in
|
||
<taxonomicName id="4C259FEB680C107FFEEB4828FE4F01EF" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[298,438,1644,1669]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680C107FFEEB4828FE4F01EF" box="[298,438,1644,1669]" italics="true" pageId="10">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; typically most elevated and inflated distally in circumorificial area, and moderately convex proximally in
|
||
<taxonomicName id="4C259FEB680C107FFE0448D4FDD601C2" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[453,559,1681,1704]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680C107FFE0448D4FDD601C2" box="[453,559,1681,1704]" italics="true" pageId="10">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680C107FFDF648CAFD8F01C2" bold="true" box="[567,630,1679,1704]" pageId="10">
|
||
<taxonomicNameLabel id="A2628501680C107FFDF648CAFD8F01C2" box="[567,630,1679,1704]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680C107FFF5648F1FE6500EA" blockId="10.[151,1437,1607,2029]" pageId="10">
|
||
2 Umbo(nes)—a prominent solid umbo or 2–3 smaller, scattered mamilliform umbones in the suboral/ subcentral part of the frontal shield in many zooids of
|
||
<taxonomicName id="4C259FEB680C107FFC81489CFC31019A" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[832,968,1751,1776]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680C107FFC81489CFC31019A" box="[832,968,1751,1776]" italics="true" pageId="10">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; a conical to bulbous umbo occasionally developing distal to the orifice in some zooids of
|
||
<taxonomicName id="4C259FEB680C107FFCDD48B8FC73007E" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[796,906,1789,1812]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680C107FFCDD48B8FC73007E" box="[796,906,1789,1812]" italics="true" pageId="10">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680C107FFC5448B9FC21007F" bold="true" box="[917,984,1788,1813]" pageId="10">
|
||
<taxonomicNameLabel id="A2628501680C107FFC5448B9FC21007F" box="[917,984,1788,1813]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, particularly in elevated older parts of colony. Further, 2–3 smaller umbones can occasionally replace the single prominent umbo randomly over the whole frontal shield in
|
||
<taxonomicName id="4C259FEB680C107FFE144900FDA70037" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[469,606,1860,1885]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680C107FFE144900FDA70037" box="[469,606,1860,1885]" italics="true" pageId="10">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
, whereas a pair of smaller umbones may flank the sinus proximolaterally in
|
||
<taxonomicName id="4C259FEB680C107FFF2D492CFEAF00EA" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[236,342,1897,1920]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680C107FFF2D492CFEAF00EA" box="[236,342,1897,1920]" italics="true" pageId="10">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680C107FFE9C4922FE6500EA" bold="true" box="[349,412,1895,1920]" pageId="10">
|
||
<taxonomicNameLabel id="A2628501680C107FFE9C4922FE6500EA" box="[349,412,1895,1920]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680C107FFF5649C9FAC80087" blockId="10.[151,1437,1607,2029]" pageId="10">
|
||
3 Shape of the primary orifice—more or less circular in outline (mean length and width 0.20 ×
|
||
<quantity id="4CDD498D680C107FFB2549C9FAB100CE" box="[1252,1352,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="10" unit="mm" value="0.21">0.21 mm</quantity>
|
||
in
|
||
<typeStatus id="549E5ACA680C107FFAAD49C8FA6500CF" box="[1388,1436,1933,1957]" pageId="10">type</typeStatus>
|
||
material) with a deep, widely V-shaped sinus in
|
||
<taxonomicName id="4C259FEB680C107FFD2B49F4FC8900A2" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[746,880,1967,1992]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680C107FFD2B49F4FC8900A2" box="[746,880,1967,1992]" italics="true" pageId="10">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; transversely oval (mean length and width 0.12 ×
|
||
<quantity id="4CDD498D680C107FFF0E4991FECD0086" box="[207,308,2004,2028]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="10" unit="mm" value="0.19">0.19 mm</quantity>
|
||
) with a straight proximal margin and narrowed, shallow U-shaped sinus in
|
||
<taxonomicName id="4C259FEB680C107FFB404990FB120086" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[1153,1259,2005,2028]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680C107FFB404990FB120086" box="[1153,1259,2005,2028]" italics="true" pageId="10">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680C107FFB334991FAC80087" bold="true" box="[1266,1329,2004,2029]" pageId="10">
|
||
<taxonomicNameLabel id="A2628501680C107FFB334991FAC80087" box="[1266,1329,2004,2029]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680D107EFF564EDDFBD90677" blockId="11.[151,1437,151,789]" pageId="11">
|
||
4 Depth of the primary orifice—little sunken, facing frontally and wholly visible in all zooids at all stages of astogeny in
|
||
<taxonomicName id="4C259FEB680D107EFEA34EF8FE1607BF" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[354,495,188,213]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFEA34EF8FE1607BF" box="[354,495,188,213]" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; deeply sunken, tilted proximally at about 20–40° to frontal plane, and partly concealed by the overhanging distal margin of the secondary orifice for 1/3 to 1/2 of its length, hence wholly visible only in developing zooids near the colony margin in
|
||
<taxonomicName id="4C259FEB680D107EFCB14F40FC230676" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[880,986,261,284]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFCB14F40FC230676" box="[880,986,261,284]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFC234F41FBD90677" bold="true" box="[994,1056,260,285]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFC234F41FBD90677" box="[994,1056,260,285]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680D107EFF564F62FD9F06BA" blockId="11.[151,1437,151,789]" pageId="11">
|
||
5 Relative proportions of the orifice—orifice width and sinus length are very similar and overlapping in
|
||
<taxonomicName id="4C259FEB680D107EFA464F6CFEC2060F" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFA464F6CFEC2060F" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C259FEB680D107EFEB24F08FE26060E" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[371,479,333,356]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFEB24F08FE26060E" box="[371,479,333,356]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFE264F09FDDE060F" bold="true" box="[487,551,332,357]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFE264F09FDDE060F" box="[487,551,332,357]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, respectively
|
||
<quantity id="4CDD498D680D107EFD064F09FC92060E" box="[711,875,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="11" unit="mm" value="0.215" valueMax="0.24" valueMin="0.19">0.19–0.24 mm</quantity>
|
||
(0.21 ±
|
||
<quantity id="4CDD498D680D107EFC0C4F09FBCD060E" box="[973,1076,332,356]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="11" unit="mm" value="0.02">0.02 mm</quantity>
|
||
) vs
|
||
<quantity id="4CDD498D680D107EFBA44F09FAF0060E" box="[1125,1289,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="11" unit="mm" value="0.195" valueMax="0.23" valueMin="0.16">0.16–0.23 mm</quantity>
|
||
(0.19 ±
|
||
<quantity id="4CDD498D680D107EFAAA4F09FF0606E2" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="11" unit="mm" value="0.02">0.02 mm</quantity>
|
||
), and
|
||
<quantity id="4CDD498D680D107EFE854F2AFE1106ED" box="[324,488,367,392]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="11" unit="mm" value="0.04" valueMax="0.05" valueMin="0.03">0.03–0.05 mm</quantity>
|
||
(0.04 ±
|
||
<quantity id="4CDD498D680D107EFD8B4F35FD4806E2" box="[586,689,368,392]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="11" unit="mm" value="0.01">0.01 mm</quantity>
|
||
) vs
|
||
<quantity id="4CDD498D680D107EFD234F2AFC7F06ED" box="[738,902,367,392]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="11" unit="mm" value="0.035" valueMax="0.05" valueMin="0.02">0.02–0.05 mm</quantity>
|
||
(0.03 ±
|
||
<quantity id="4CDD498D680D107EFC284F35FBAB06E2" box="[1001,1106,368,392]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="11" unit="mm" value="0.01">0.01 mm</quantity>
|
||
), whereas orifice length in
|
||
<taxonomicName id="4C259FEB680D107EFA464F34FEC206C7" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFA464F34FEC206C7" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
exceeds that in
|
||
<taxonomicName id="4C259FEB680D107EFE374FD0FD9A06C6" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[502,611,405,428]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFE374FD0FD9A06C6" box="[502,611,405,428]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFDAD4FD1FD5406C7" bold="true" box="[620,685,404,429]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFDAD4FD1FD5406C7" box="[620,685,404,429]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<quantity id="4CDD498D680D107EFD7C4FD1FC9D06C6" box="[701,868,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.7999999999999998" pageId="11" unit="mm" value="0.215" valueMax="0.25" valueMin="0.18">0.18–0.25 mm</quantity>
|
||
(0.20 ±
|
||
<quantity id="4CDD498D680D107EFC094FD1FBD606C6" box="[968,1071,404,428]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="11" unit="mm" value="0.02">0.02 mm</quantity>
|
||
) vs
|
||
<quantity id="4CDD498D680D107EFBA34FD1FAFE06C6" box="[1122,1287,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="11" unit="mm" value="0.125" valueMax="0.15" valueMin="0.1">0.10–0.15 mm</quantity>
|
||
(0.12 ±
|
||
<quantity id="4CDD498D680D107EFAAA4FD1FF0506BA" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="11" unit="mm" value="0.02">0.02 mm</quantity>
|
||
)) and the ranges do not overlap.
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680D107EFF564F99FE2A0572" blockId="11.[151,1437,151,789]" pageId="11">
|
||
6 Ratio of primary orifice length to zooid length—half as great in
|
||
<taxonomicName id="4C259FEB680D107EFC674F98FBF7069E" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[934,1038,477,500]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFC674F98FBF7069E" box="[934,1038,477,500]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFBD84F99FBA1069F" bold="true" box="[1049,1112,476,501]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFBD84F99FBA1069F" box="[1049,1112,476,501]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(about 14%) compared to
|
||
<taxonomicName id="4C259FEB680D107EFA464F98FEC20572" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFA464F98FEC20572" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
(about 29%).
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680D107EFF564C60FE0505EF" blockId="11.[151,1437,151,789]" pageId="11">
|
||
7 Secondary orifice—the primary and secondary orifices are nearly identical in form and roughly circular in
|
||
<taxonomicName id="4C259FEB680D107EFA464C60FEC3050A" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFA464C60FEC3050A" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; the primary orifice is transversely oval and the secondary orifice triangular to drop-shaped in outlines in
|
||
<taxonomicName id="4C259FEB680D107EFE8A4C28FE4F05EE" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[331,438,621,644]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFE8A4C28FE4F05EE" box="[331,438,621,644]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFE7C4C29FE0505EF" bold="true" box="[445,508,620,645]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFE7C4C29FE0505EF" box="[445,508,620,645]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680D107EFF564CD5FBA605A7" blockId="11.[151,1437,151,789]" pageId="11">
|
||
8 The cormidial nature of the primary orifice—cormidial in several generations of periancestrular zooids in
|
||
<taxonomicName id="4C259FEB680D107EFA464CD4FEC305A7" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFA464CD4FEC305A7" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; cormidial only in zooids of the ancestrular complex in
|
||
<taxonomicName id="4C259FEB680D107EFC6F4CF0FBE005A6" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[942,1049,693,716]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFC6F4CF0FBE005A6" box="[942,1049,693,716]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFBE14CF1FBA605A7" bold="true" box="[1056,1119,692,717]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFBE14CF1FBA605A7" box="[1056,1119,692,717]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9AE468680D107EFF564C9DFC51047F" blockId="11.[151,1437,151,789]" pageId="11">
|
||
9 Colour of dried colonies (colour not seen in living colonies)—black in
|
||
<taxonomicName id="4C259FEB680D107EFC334C9CFB80059A" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[1010,1145,727,752]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="paradoxa">
|
||
<emphasis id="B951387A680D107EFC334C9CFB80059A" box="[1010,1145,727,752]" italics="true" pageId="11">F. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; mostly dark red/crimson in
|
||
<taxonomicName id="4C259FEB680D107EFF2D4CB8FEAF047E" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[236,342,765,788]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFF2D4CB8FEAF047E" box="[236,342,765,788]" italics="true" pageId="11">F. imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFE9C4CB9FE65047F" bold="true" box="[349,412,764,789]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFE9C4CB9FE65047F" box="[349,412,764,789]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(sometimes pink, rarely deep purple to black).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33FB7E3680D107EFF064D01FD870381" pageId="11" type="biology_ecology">
|
||
<paragraph id="8B9AE468680D107EFF064D01FD870381" blockId="11.[151,1436,836,1403]" pageId="11">
|
||
<emphasis id="B951387A680D107EFF064D01FED20437" bold="true" box="[199,299,836,861]" pageId="11">Ecology.</emphasis>
|
||
The new species was documented in areas of mixed hard and soft bottoms, including pebbles and rocks with shell, gravel and sand/silt admixtures. Of the 197 colonies examined, 79.69% were found on pebbles, 10.15% and 1.02% respectively on shell fragments of the bivalves
|
||
<taxonomicName id="4C259FEB680D107EFCBE4DC9FC1804CF" box="[895,993,908,933]" class="Bivalvia" family="Pectinidae" genus="Chlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="11" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951387A680D107EFCBE4DC9FC1804CF" box="[895,993,908,933]" italics="true" pageId="11">Chlamys</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName id="4C259FEB680D107EFBFF4DC8FADA04CF" baseAuthorityName="Deshayes" baseAuthorityYear="1839" box="[1086,1315,908,933]" class="Bivalvia" family="Anomiidae" genus="Monia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="11" phylum="Mollusca" rank="species" species="macrochisma">
|
||
<emphasis id="B951387A680D107EFBFF4DC8FADA04CF" box="[1086,1315,908,933]" italics="true" pageId="11">Monia macrochisma</emphasis>
|
||
</taxonomicName>
|
||
, 8.12% on barnacles, 0.51% on tubes of serpulid polychaetes and 0.51% on sponges. Co-occurring invertebrates mostly included the echinoderms
|
||
<taxonomicName id="4C259FEB680D107EFE1D4D91FCE10487" box="[476,792,980,1005]" class="Echinoidea" family="Strongylocentrotidae" genus="Strongylocentrotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echinoida" pageId="11" phylum="Echinodermata" rank="species" species="pallidus">
|
||
<emphasis id="B951387A680D107EFE1D4D91FCE10487" box="[476,792,980,1005]" italics="true" pageId="11">Strongylocentrotus pallidus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C259FEB680D107EFCA54D91FBA90487" authorityName="H. L. Clark" authorityYear="1911" box="[868,1104,980,1005]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="11" phylum="Echinodermata" rank="species" species="leptoctenia">
|
||
<emphasis id="B951387A680D107EFCA54D91FBA90487" box="[868,1104,980,1005]" italics="true" pageId="11">Ophiura leptoctenia</emphasis>
|
||
</taxonomicName>
|
||
, the sedentary polychaetes
|
||
<taxonomicName id="4C259FEB680D107EFF564DB2FE9F037A" baseAuthorityName="Müller" baseAuthorityYear="1776" box="[151,358,1015,1040]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orbiniida" pageId="11" phylum="Annelida" rank="species" species="armiger">
|
||
<emphasis id="B951387A680D107EFF564DB2FE9F037A" box="[151,358,1015,1040]" italics="true" pageId="11">Scoloplos armiger</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C259FEB680D107EFE614DB2FD65037A" baseAuthorityName="Zachs" baseAuthorityYear="1923" box="[416,668,1015,1040]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="11" phylum="Annelida" rank="species" species="oculata">
|
||
<emphasis id="B951387A680D107EFE614DB2FD65037A" box="[416,668,1015,1040]" italics="true" pageId="11">Galathowenia oculata</emphasis>
|
||
</taxonomicName>
|
||
, the gammaridean amphipod
|
||
<emphasis id="B951387A680D107EFC314DB2FB6C037A" box="[1008,1173,1015,1040]" italics="true" pageId="11">Melita dentata</emphasis>
|
||
and the bivalve
|
||
<taxonomicName id="4C259FEB680D107EFA944DB2FF19035F" baseAuthorityName="Couthouy" baseAuthorityYear="1838" class="Bivalvia" family="Yoldiidae" genus="Yoldia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nuculanida" pageId="11" phylum="Mollusca" rank="species" species="myalis">
|
||
<emphasis id="B951387A680D107EFA944DB2FF19035F" italics="true" pageId="11">Yoldia myalis</emphasis>
|
||
</taxonomicName>
|
||
, as well as hydroids, barnacles, sponges and about 70–90 other bryozoan species, the most abundant being
|
||
<taxonomicName id="4C259FEB680D107EFF564A7AFDED0332" authority="(Kluge, 1906)" authorityName="Kluge" authorityYear="1906" baseAuthorityName="Kluge" baseAuthorityYear="1906" box="[151,532,1087,1112]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="arctica">
|
||
<emphasis id="B951387A680D107EFF564A7AFE9E0332" box="[151,359,1087,1112]" italics="true" pageId="11">Hippothoa arctica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB49999680D107EFEB84A7AFDF50332" author="Kluge, H. & G. A" box="[377,524,1087,1112]" pageId="11" pagination="31 - 55" refId="ref8373" refString="Kluge, H. [G. A.] (1906) Erganzungsbericht uber die van der " Olga " - Expedition gesammelten Bryozoen. Wissenschaftliche Meeresuntersuchungen, Neue Folge, 8, 31 - 55." type="journal article" year="1906">Kluge, 1906</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C259FEB680D107EFDE24A04FCAE0332" authority="Dawson, 1859" authorityName="Dawson" authorityYear="1859" box="[547,855,1087,1112]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="expansa">
|
||
<emphasis id="B951387A680D107EFDE24A04FDCE0332" box="[547,567,1089,1112]" italics="true" pageId="11">H</emphasis>
|
||
.
|
||
<emphasis id="B951387A680D107EFD884A04FD5C0332" box="[585,677,1089,1112]" italics="true" pageId="11">expansa</emphasis>
|
||
<bibRefCitation id="EFB49999680D107EFD6E4A05FCAE0332" author="Dawson, J. W." box="[687,855,1087,1112]" pageId="11" pagination="255 - 257" refId="ref7139" refString="Dawson, J. W. (1859) Polyzoa. Geological Survey of Canada Report of Progress for the Year 1858, 255 - 257." type="journal article" year="1859">Dawson, 1859</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C259FEB680D107EFCA74A7AFAE10332" authority="(Hassall, 1842)" authorityName="Hassall" authorityYear="1842" baseAuthorityName="Hassall" baseAuthorityYear="1842" box="[870,1304,1087,1112]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="ventricosa">
|
||
<emphasis id="B951387A680D107EFCA74A7AFBA40332" box="[870,1117,1087,1112]" italics="true" pageId="11">Escharella ventricosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB49999680D107EFBAE4A7AFAE90332" author="Hassall, A. H." box="[1135,1296,1087,1112]" pageId="11" pagination="407 - 414" refId="ref8230" refString="Hassall, A. H. (1842) Remarks on the genus Lepralia of Dr. Johnston, with descriptions of six undescribed species, and notices of two other Zoophytes. Annals and Magazine of Natural History, Series 1, 9 (59), 407 - 414. https: // doi. org / 10.1080 / 03745484209445355" type="journal article" year="1842">Hassall, 1842</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C259FEB680D107EFAE64A7AFE700315" authority="(Busk, 1856)" authorityName="Busk" authorityYear="1856" baseAuthorityName="Busk" baseAuthorityYear="1856" class="Gymnolaemata" family="Stomachetosellidae" genus="Ragionula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomata" pageId="11" phylum="Bryozoa" rank="species" species="rosacea">
|
||
<emphasis id="B951387A680D107EFAE64A7AFF170315" italics="true" pageId="11">Ragionula rosacea</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB49999680D107EFF3F4A23FE790315" author="Busk, G." box="[254,384,1126,1151]" pageId="11" pagination="32 - 36" refId="ref6699" refString="Busk, G. (1856) Polyzoa collected by Mr. M'Andrew on the coast of Norway and Finmark in 1856. Annals and Magazine of Natural History, Series 2, 18, 32 - 36, pl. 1." type="journal article" year="1856">Busk, 1856</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C259FEB680D107EFE014A23FC3E0315" authority="(Androsova, 1958)" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" baseAuthorityName="Androsova" baseAuthorityYear="1958" box="[448,967,1126,1151]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="tuberculata">
|
||
<emphasis id="B951387A680D107EFE014A23FD110315" box="[448,744,1126,1151]" italics="true" pageId="11">Stomacrustula tuberculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB49999680D107EFD364A23FC460315" author="Androsova, E. I." box="[759,959,1126,1151]" pageId="11" pagination="90 - 204" refId="ref6426" refString="Androsova, E. I. (1958) Bryozoa of the order Cheilostomata of the northern part of the Sea of Japan. Issledovaniya Dal'nevostochnikh Morei SSSR, 5, 90 - 204. [in Russian]" type="journal article" year="1958">Androsova, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The number of
|
||
<taxonomicName id="4C259FEB680D107EFB434A2DFB170315" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[1154,1262,1128,1151]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata">
|
||
<emphasis id="B951387A680D107EFB434A2DFB680315" box="[1154,1169,1128,1151]" italics="true" pageId="11">F</emphasis>
|
||
.
|
||
<emphasis id="B951387A680D107EFB5E4A2DFB170315" box="[1183,1262,1128,1151]" italics="true" pageId="11">imitata</emphasis>
|
||
</taxonomicName>
|
||
colonies/m
|
||
<superScript id="7C504920680D107EFAB04A26FA80031B" attach="left" box="[1393,1401,1123,1137]" fontSize="6" pageId="11">2</superScript>
|
||
by collection year was 4–28 (2008), 4–40 (2013), 4–32 (2014), 384 (2015) and 172 (2016). Colonies of
|
||
<taxonomicName id="4C259FEB680D107EFAEC4AC9FA6203C9" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[1325,1435,1163,1187]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata">
|
||
<emphasis id="B951387A680D107EFAEC4AC9FAC503C9" box="[1325,1340,1164,1187]" italics="true" pageId="11">F</emphasis>
|
||
.
|
||
<emphasis id="B951387A680D107EFA8D4AC9FA6203C9" box="[1356,1435,1164,1187]" italics="true" pageId="11">imitata</emphasis>
|
||
</taxonomicName>
|
||
occasionally cover a considerable area of any particular substratum, appearing as the ‘background’ species at certain sites where shingly bottoms prevail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33FB7E3680D107EFF064AB3FD710211" pageId="11" type="distribution">
|
||
<paragraph id="8B9AE468680D107EFF064AB3FD710211" blockId="11.[151,1436,836,1403]" pageId="11">
|
||
<emphasis id="B951387A680D107EFF064AB3FE990265" bold="true" box="[199,352,1270,1295]" pageId="11">Distribution.</emphasis>
|
||
<taxonomicName id="4C259FEB680D107EFEA94AB3FDCD0265" authority="Grischenko & Gordon & Morozov, 2018" authorityName="Grischenko & Gordon & Morozov" authorityYear="2018" box="[360,564,1270,1295]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" phylum="Bryozoa" rank="species" species="imitata" status="sp. nov.">
|
||
<emphasis id="B951387A680D107EFEA94AB3FDCD0265" box="[360,564,1270,1295]" italics="true" pageId="11">Fatkullina imitata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951387A680D107EFDFD4AB3FD840265" bold="true" box="[572,637,1270,1295]" pageId="11">
|
||
<taxonomicNameLabel id="A2628501680D107EFDFD4AB3FD840265" box="[572,637,1270,1295]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is currently known from 16 stations in the depth range
|
||
<quantity id="4CDD498D680D107EFB364AB3FA800265" box="[1271,1401,1270,1295]" metricMagnitude="2" metricUnit="m" metricValue="2.305" metricValueMax="3.61" metricValueMin="1.0" pageId="11" unit="m" value="230.5" valueMax="361.0" valueMin="100.0">100–361 m</quantity>
|
||
, in an area bounded by the coordinates 56.98389–
|
||
<geoCoordinate id="EE1182AF680D107EFD154B5FFC900258" box="[724,873,1306,1331]" direction="north" orientation="latitude" pageId="11" precision="1" value="58.18528">58.18528° N</geoCoordinate>
|
||
and 154.93389–
|
||
<geoCoordinate id="EE1182AF680D107EFBF74B5FFB230258" box="[1078,1242,1306,1331]" direction="east" orientation="longitude" pageId="11" precision="1" value="156.05">156.05000° E</geoCoordinate>
|
||
. It can thus be categorized as a high-Boreal, Pacific, Asian, sublittoral to upper bathyal species, endemic to the western
|
||
<collectingRegion id="49E12A8A680D107EFF564B27FEEE0211" box="[151,279,1378,1403]" country="Russia" name="Kamchatka" pageId="11">Kamchatka</collectingRegion>
|
||
shelf and slope, Sea of Okhotsk.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |