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<document id="D69FB619BE18629DC21D916E583AB14B" ID-CLB-Dataset="45663" ID-DOI="10.11646/zootaxa.3802.3.6" ID-GBIF-Dataset="c65cfca8-1f3c-4bd1-a77d-09d68b51c16d" ID-ISSN="1175-5326" ID-Zenodo-Dep="224836" ID-ZooBank="3B6166C5-4C54-447C-8459-FC689C637BAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460426964436" checkinUser="plazi" docAuthor="Dubatolov, Vladimir V., Kishida, Yasunori &amp; Wu, Chunsheng" docDate="2014" docId="D74587DEFFCEB83DFF051F511025FA8D" docLanguage="en" docName="zt03802p380.pdf" docOrigin="Zootaxa 3802 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraheliosia Dubatolov, Kishida &amp; Wu, 2014, gen. nov." docType="treatment" docVersion="9" lastPageNumber="374" masterDocId="2B7CFFA6FFCFB83CFF921D3D1165FFE9" masterDocTitle="Review of East Asian Heliosia (Ledidoptera: Erebidae: Arctiinae: Lithosiini) species, with description of a new genus" masterLastPageNumber="380" masterPageNumber="373" pageNumber="374" updateTime="1698330827806" updateUser="ExternalLinkService">
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<mods:title id="A0FE3782855A7A7A9E8893F3F0620E2F">Review of East Asian Heliosia (Ledidoptera: Erebidae: Arctiinae: Lithosiini) species, with description of a new genus</mods:title>
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<mods:namePart id="345CC826F9F051A754AD2BDF33A17017">Dubatolov, Vladimir V.</mods:namePart>
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<mods:namePart id="5672D6C82318C8828766C1248D9E7C18">Kishida, Yasunori</mods:namePart>
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<mods:namePart id="8E98469A7CD7C41A1F271879BE6462DF">Wu, Chunsheng</mods:namePart>
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<treatment id="D74587DEFFCEB83DFF051F511025FA8D" ID-DOI="http://doi.org/10.5281/zenodo.6137698" ID-GBIF-Taxon="119428835" ID-Zenodo-Dep="6137698" LSID="urn:lsid:plazi:treatment:D74587DEFFCEB83DFF051F511025FA8D" httpUri="http://treatment.plazi.org/id/D74587DEFFCEB83DFF051F511025FA8D" lastPageNumber="374" pageId="1" pageNumber="374">
<subSubSection id="17F66543FFCEB83DFF051F511046FD4E" pageId="1" pageNumber="374" type="nomenclature">
<paragraph id="5F5336C8FFCEB83DFF051F5110FCFD6F" blockId="1.[151,409,620,679]" box="[151,409,620,646]" pageId="1" pageNumber="374">
<heading id="041B81A4FFCEB83DFF051F5110FCFD6F" bold="true" box="[151,409,620,646]" fontSize="10" level="2" pageId="1" pageNumber="374" reason="2">
<emphasis id="6D98EADAFFCEB83DFF051F5110FCFD6F" bold="true" box="[151,409,620,646]" pageId="1" pageNumber="374">
<taxonomicName id="98EC4D4BFFCEB83DFF051F51104CFD6C" box="[151,297,620,645]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6D98EADAFFCEB83DFF051F51104CFD6C" bold="true" box="[151,297,620,645]" italics="true" pageId="1" pageNumber="374">Paraheliosia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="76AB57A1FFCEB83DFEA21F5010FCFD6F" box="[304,409,621,646]" pageId="1" pageNumber="374" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5F5336C8FFCEB83DFF051FB31046FD4E" blockId="1.[151,409,620,679]" box="[151,291,654,679]" pageId="1" pageNumber="374">
(
<figureCitation id="C7D72A4DFFCEB83DFF0D1FB31186FD4E" box="[159,227,654,679]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1345,1368]" captionTargetBox="[151,1436,195,1319]" captionTargetId="figure@2.[151,1436,195,1324]" captionTargetPageId="2" captionText="FIGURES 1 6. Heliosia types (1, 3, 5) and their labels (2, 4, 6). 1 2 — Heliosia punctinigra van Eecke, 1920, syntype, female, Indonesia, Java, Batavia (now—Jakarta); 3 4 — Heliosia punctata Fang, 1992, paratype, male, China, Sichuan, Emeishan; 5 6 — Paraheliosia elegans (Reich, 1937), holotype, male, China, Zhejiang, West Tien Mu Shan." httpUri="https://zenodo.org/record/224837/files/figure.png" pageId="1" pageNumber="374">Fig. 5</figureCitation>
,
<figureCitation id="C7D72A4DFFCEB83DFF7D1FB2107EFD4E" box="[239,283,655,679]" captionStart="FIGURES 7 14" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[151,1436,193,1834]" captionTargetId="figure@4.[151,1436,193,1839]" captionTargetPageId="4" captionText="FIGURES 7 14. Lithosiinae moths (7 9, 11), their labels (10) and male genitalia (12 14). 7 — Paraheliosia rufa (Leech, 1890), male, China, Gansu, Kang Xian; 8 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), male, Russia, Primorskiy Kray, 20 km E from Kirovskiy; 9 10 — Paraheliosia novirufa (Fang, 1992), holotype, male (9), China, Zhejiang, Zhoushan, and its labels (10); 11, 12 — Nudina artaxidia (Butler, 1881), male, Russia, Blagoveshchensk; 13 — Paraheliosia elegans (Reich, 1937), holotype, China, Zhejiang, West Tien Mu Shan; 14 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), Russia, Primorskiy Kray, 20 km E from Kirovskiy." httpUri="https://zenodo.org/record/224838/files/figure.png" pageId="1" pageNumber="374">79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="17F66543FFCEB83DFF051FEF13EAFD0E" box="[151,655,721,743]" pageId="1" pageNumber="374" type="materials_examined">
<paragraph id="5F5336C8FFCEB83DFF051FEF13EAFD0E" blockId="1.[151,655,721,743]" box="[151,655,721,743]" pageId="1" pageNumber="374">
<typeStatus id="8057886AFFCEB83DFF051FEF11ACFD0E" box="[151,201,722,743]" pageId="1" pageNumber="374">Type</typeStatus>
species:
<taxonomicName id="98EC4D4BFFCEB83DFEB41FEF1353FD0E" authority="Reich, 1937" authorityName="Reich" authorityYear="1937" box="[294,566,721,743]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="6D98EADAFFCEB83DFEB41FEF10D3FD0E" box="[294,438,721,743]" italics="true" pageId="1" pageNumber="374">Asura elegans</emphasis>
<bibRefCitation id="3B7D4B39FFCEB83DFE2E1FEC1353FD0E" author="Reich" box="[444,566,721,743]" pageId="1" pageNumber="374" refString="Reich, P. (1937) Die Arctiidae der Chinnaausbeute des Herrn Hermann Hone in Shanghai. Deutsche Entomologische Zeitschrift Iris 51: 113 - 130." type="journal article" year="1937">Reich, 1937</bibRefCitation>
</taxonomicName>
(
<figureCitation id="C7D72A4DFFCEB83DFDD61FEC13E4FD0E" box="[580,641,721,743]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1345,1368]" captionTargetBox="[151,1436,195,1319]" captionTargetId="figure@2.[151,1436,195,1324]" captionTargetPageId="2" captionText="FIGURES 1 6. Heliosia types (1, 3, 5) and their labels (2, 4, 6). 1 2 — Heliosia punctinigra van Eecke, 1920, syntype, female, Indonesia, Java, Batavia (now—Jakarta); 3 4 — Heliosia punctata Fang, 1992, paratype, male, China, Sichuan, Emeishan; 5 6 — Paraheliosia elegans (Reich, 1937), holotype, male, China, Zhejiang, West Tien Mu Shan." httpUri="https://zenodo.org/record/224837/files/figure.png" pageId="1" pageNumber="374">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="17F66543FFCEB83DFF051E2C1511FBF6" pageId="1" pageNumber="374" type="description">
<paragraph id="5F5336C8FFCEB83DFF051E2C13BBFCA3" blockId="1.[151,1437,785,1380]" pageId="1" pageNumber="374">
<emphasis id="6D98EADAFFCEB83DFF051E2C104CFCC3" bold="true" box="[151,297,785,810]" pageId="1" pageNumber="374">Description.</emphasis>
Antennae setose. Forewings yellowish, rose or red, with two nearly straight oblique lines or rows of small spots and sometimes various melanistic areas.
</paragraph>
<paragraph id="5F5336C8FFCEB83DFF551E691511FBF6" blockId="1.[151,1437,785,1380]" pageId="1" pageNumber="374">
<emphasis id="6D98EADAFFCEB83DFF551E691011FC84" bold="true" box="[199,372,852,877]" pageId="1" pageNumber="374">Male genitalia</emphasis>
(
<figureCitation id="C7D72A4DFFCEB83DFE141E691363FC84" box="[390,518,852,877]" captionStart-0="FIGURES 7 14" captionStart-1="FIGURES 15 16" captionStartId-0="4.[151,264,1860,1883]" captionStartId-1="5.[151,264,740,763]" captionTargetBox-0="[151,1436,193,1834]" captionTargetBox-1="[167,1428,193,717]" captionTargetId-0="figure@4.[151,1436,193,1839]" captionTargetId-1="figure@5.[151,1436,193,719]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 7 14. Lithosiinae moths (7 9, 11), their labels (10) and male genitalia (12 14). 7 — Paraheliosia rufa (Leech, 1890), male, China, Gansu, Kang Xian; 8 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), male, Russia, Primorskiy Kray, 20 km E from Kirovskiy; 9 10 — Paraheliosia novirufa (Fang, 1992), holotype, male (9), China, Zhejiang, Zhoushan, and its labels (10); 11, 12 — Nudina artaxidia (Butler, 1881), male, Russia, Blagoveshchensk; 13 — Paraheliosia elegans (Reich, 1937), holotype, China, Zhejiang, West Tien Mu Shan; 14 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), Russia, Primorskiy Kray, 20 km E from Kirovskiy." captionText-1="FIGURES 15 16. Male genitalia. 15 — Paraheliosia rufa (Leech, 1890), China, Gansu, Kang Xian; 16 — Heliosia punctata Fang, 1992, paratype, China, Sichuan, Emeishan." httpUri-0="https://zenodo.org/record/224838/files/figure.png" httpUri-1="https://zenodo.org/record/224839/files/figure.png" pageId="1" pageNumber="374">Fig. 1315</figureCitation>
,
<figureCitation id="C7D72A4DFFCEB83DFD871E691305FC85" box="[533,608,852,876]" captionStart="FIGURES 17 19" captionStartId="5.[151,264,1946,1969]" captionTargetBox="[175,1400,832,1909]" captionTargetId="figure@5.[168,1404,823,1919]" captionTargetPageId="5" captionText="FIGURES 17 19. Male genitalia of Paraheliosia gen. nov. (17 18) and Heliosia punctata Fang, 1992 (19). 17 — Paraheliosia novirufa (Fang, 1992) (from Fang, 1992); 18 — Paraheliosia novirufa (Fang, 1992) (from Fang, 2000); 19 — Heliosia punctata Fang, 1992 (from Fang, 2000)." httpUri="https://zenodo.org/record/224840/files/figure.png" pageId="1" pageNumber="374">1718</figureCitation>
): The genus is best marked my male genitalia structure: uncus straight, slightly curved at apex; valve base with short triangular processes; cucullus membranose; sacculus with a fingerlike apical process; juxta with two apical processes; saccus triangular; aedeagus short, with one or few strong apical spines; vesica with two groups of small spine-like cornuti. In general, uncus, valve, juxta and saccus structure, as well as two groups of small spine-like cornuti on vesica are very similar within the know species of the genus. The most remarkable distinguishing character is a number of strong spines at aedeagus apex.
</paragraph>
</subSubSection>
<subSubSection id="17F66543FFCEB83DFF5519111185FAF4" pageId="1" pageNumber="374" type="discussion">
<paragraph id="5F5336C8FFCEB83DFF5519111185FAF4" blockId="1.[151,1437,785,1380]" pageId="1" pageNumber="374">
<emphasis id="6D98EADAFFCEB83DFF551911105EFBAC" bold="true" box="[199,315,1068,1093]" pageId="1" pageNumber="374">Remarks.</emphasis>
By the wing pattern, the genus is similar to the monotypic
<taxonomicName id="98EC4D4BFFCEB83DFC4D1911159EFBAD" authority="Staudinger, 1887" authorityName="Staudinger" authorityYear="1887" box="[991,1275,1068,1093]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="6D98EADAFFCEB83DFC4D19111557FBAD" box="[991,1074,1068,1092]" italics="true" pageId="1" pageNumber="374">Nudina</emphasis>
<bibRefCitation id="3B7D4B39FFCEB83DFBA81911159EFBAD" author="Staudinger" box="[1082,1275,1068,1093]" pageId="1" pageNumber="374" refString="Staudinger, O. (1887) Neue Arten und Varietaten von Lepidopteren aus dem Amur-Gebiete. In: Romanoff, N. M. (Ed.), Memoires sur les lepidopteres. Vol. 3. M. M. Stassulewitch, St. - Petersbourg, pp. 126 - 232, pls. 6 - 12, 16 - 17." type="book chapter" year="1887">Staudinger, 1887</bibRefCitation>
</taxonomicName>
, with the
<typeStatus id="8057886AFFCEB83DFAFE191014F9FBAC" box="[1388,1436,1069,1093]" pageId="1" pageNumber="374">type</typeStatus>
species
<taxonomicName id="98EC4D4BFFCEB83DFF62196C1346FB81" authority="Butler, 1881" authorityName="Butler" authorityYear="1881" box="[240,547,1103,1128]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="artaxidia">
<emphasis id="6D98EADAFFCEB83DFF62196C101DFB81" box="[240,376,1104,1128]" italics="true" pageId="1" pageNumber="374">N. artaxidia</emphasis>
(
<bibRefCitation id="3B7D4B39FFCEB83DFE1A1972137FFB81" author="Butler" box="[392,538,1103,1128]" pageId="1" pageNumber="374" refString="Butler, A. G. (1881) Descriptions of new genera and species of Heterocerous Lepidoptera from Japan. Transactions of the Entomological Society of London, 1881, 1 - 23. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1873. tb 01519. x" type="journal article" year="1881">Butler, 1881</bibRefCitation>
)
</taxonomicName>
; the latter species also have a wide dark band at the fore- and hindwing outer margins (
<figureCitation id="C7D72A4DFFCEB83DFE9219491037FB65" box="[256,338,1140,1165]" captionStart="FIGURES 7 14" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[151,1436,193,1834]" captionTargetId="figure@4.[151,1436,193,1839]" captionTargetPageId="4" captionText="FIGURES 7 14. Lithosiinae moths (7 9, 11), their labels (10) and male genitalia (12 14). 7 — Paraheliosia rufa (Leech, 1890), male, China, Gansu, Kang Xian; 8 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), male, Russia, Primorskiy Kray, 20 km E from Kirovskiy; 9 10 — Paraheliosia novirufa (Fang, 1992), holotype, male (9), China, Zhejiang, Zhoushan, and its labels (10); 11, 12 — Nudina artaxidia (Butler, 1881), male, Russia, Blagoveshchensk; 13 — Paraheliosia elegans (Reich, 1937), holotype, China, Zhejiang, West Tien Mu Shan; 14 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), Russia, Primorskiy Kray, 20 km E from Kirovskiy." httpUri="https://zenodo.org/record/224838/files/figure.png" pageId="1" pageNumber="374">Fig. 11</figureCitation>
). However, the male genitalia of these genera are very different: in
<taxonomicName id="98EC4D4BFFCEB83DFBDD194915C7FB65" box="[1103,1186,1140,1164]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="6D98EADAFFCEB83DFBDD194915C7FB65" box="[1103,1186,1140,1164]" italics="true" pageId="1" pageNumber="374">Nudina</emphasis>
</taxonomicName>
(
<figureCitation id="C7D72A4DFFCEB83DFB2319491466FB65" box="[1201,1283,1140,1165]" captionStart="FIGURES 7 14" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[151,1436,193,1834]" captionTargetId="figure@4.[151,1436,193,1839]" captionTargetPageId="4" captionText="FIGURES 7 14. Lithosiinae moths (7 9, 11), their labels (10) and male genitalia (12 14). 7 — Paraheliosia rufa (Leech, 1890), male, China, Gansu, Kang Xian; 8 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), male, Russia, Primorskiy Kray, 20 km E from Kirovskiy; 9 10 — Paraheliosia novirufa (Fang, 1992), holotype, male (9), China, Zhejiang, Zhoushan, and its labels (10); 11, 12 — Nudina artaxidia (Butler, 1881), male, Russia, Blagoveshchensk; 13 — Paraheliosia elegans (Reich, 1937), holotype, China, Zhejiang, West Tien Mu Shan; 14 — Paraheliosia rufa ussuriensis (O. Bang-Haas, 1927), Russia, Primorskiy Kray, 20 km E from Kirovskiy." httpUri="https://zenodo.org/record/224838/files/figure.png" pageId="1" pageNumber="374">Fig. 12</figureCitation>
) the sacculus apex has two apical spines, while in the new one it is finger-like; the aedeagus apex in
<taxonomicName id="98EC4D4BFFCEB83DFBF219A515D6FB59" box="[1120,1203,1176,1200]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="6D98EADAFFCEB83DFBF219A515D6FB59" box="[1120,1203,1176,1200]" italics="true" pageId="1" pageNumber="374">Nudina</emphasis>
</taxonomicName>
with two sclerotized plates, but in
<taxonomicName id="98EC4D4BFFCEB83DFEBE198110D8FB3D" box="[300,445,1212,1236]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6D98EADAFFCEB83DFEBE198110D8FB3D" box="[300,445,1212,1236]" italics="true" pageId="1" pageNumber="374">Paraheliosia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="76AB57A1FFCEB83DFE511980134FFB3D" box="[451,554,1213,1237]" pageId="1" pageNumber="374" rank="genus">gen. nov.</taxonomicNameLabel>
it has one or few strong spines. Moreover, presence of two strong apical spines at the juxta apex and basal costal valve processes look to be well marked autapomorphic characters of the new genus.
</paragraph>
</subSubSection>
<subSubSection id="17F66543FFCEB83DFF55181A1025FA8D" pageId="1" pageNumber="374" type="etymology">
<paragraph id="5F5336C8FFCEB83DFF55181A1025FA8D" blockId="1.[151,1437,785,1380]" pageId="1" pageNumber="374">
<emphasis id="6D98EADAFFCEB83DFF55181A102BFAA9" bold="true" box="[199,334,1319,1344]" pageId="1" pageNumber="374">Etymology.</emphasis>
The generic name consists of two parts: the Greek prefix ‘παρα’ (meaning against), and the genus name
<taxonomicName id="98EC4D4BFFCEB83DFF481871105CFA8D" box="[218,313,1356,1380]" class="Insecta" family="Arctiidae" genus="Heliosia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="6D98EADAFFCEB83DFF481871105CFA8D" box="[218,313,1356,1380]" italics="true" pageId="1" pageNumber="374">Heliosia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>