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<mods:title id="F17D1F588307C44978528864709BB62C">On the identities of the sesarmid crabs, Sesarma villosum A. Milne-Edwards 1869, and Clistocoeloma suvaense Edmondson, 1951, with the description of a new species from the West Pacific (Decapoda, Brachyura, Thoracotremata)</mods:title>
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<mods:namePart id="5A867EFEBABD1AD3B9A58CE25DB06D75">Ng, Bee Yan Lee Ngan Kee</mods:namePart>
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<mods:namePart id="5286C66B609FB2EB48E94A6AB4CFB552">Ng, Peter K. L.</mods:namePart>
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<emphasis id="7E957179FFFEFFDDFF30FF29FE44E9FF" bold="true" box="[151,405,151,178]" italics="true" pageId="9" pageNumber="226">Clistocoeloma nobile</emphasis>
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<paragraph id="4C5EAD6BFFFEFFDDFF30FF05FD13E998" blockId="9.[151,706,151,214]" box="[151,706,187,214]" pageId="9" pageNumber="226">(Figs, 2B, C, 5E, F, 6E–K, 7A–C, 8C, D, 11A–D)</paragraph>
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<emphasis id="7E957179FFFEFFDDFF30FEBEFF3FE85A" box="[151,238,256,279]" italics="true" pageId="9" pageNumber="226">Sesarma</emphasis>
(
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<emphasis id="7E957179FFFEFFDDFECFFEBEFE7DE855" box="[360,428,256,280]" italics="true" pageId="9" pageNumber="226">villosa</emphasis>
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— 
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<bibRefCitation id="2870D09AFFFEFFDDFE62FEBEFD6CE85A" author="De Man, J. G." box="[453,701,255,280]" pageId="9" pageNumber="226" pagination="75 - 218" refId="ref10460" refString="De Man, J. G. (1897) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 9 (1), 75 - 218. [in German]" type="journal article" year="1897">De Man 1897: 153–161</bibRefCitation>
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(in part).— 
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<bibRefCitation id="2870D09AFFFEFFDDFC9EFEBEFC2EE855" author="De Man, J. G." box="[825,1023,256,280]" pageId="9" pageNumber="226" pagination="677 - 708" refId="ref10528" refString="De Man, J. G. (1898) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Sechster (Schluss-) Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (4), 677 - 708, pls. 28 - 38." type="journal article" year="1898">De Man 1898: 702</bibRefCitation>
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(list) (in part), pl. 29 fig. 30a, 30e [not 
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<emphasis id="7E957179FFFEFFDDFF60FEA1FEAAE87B" box="[199,379,286,310]" italics="true" pageId="9" pageNumber="226">Sesarma villosum</emphasis>
A. 
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<emphasis id="7E957179FFFEFFDDFF30FE83FF3FE819" box="[151,238,317,340]" italics="true" pageId="9" pageNumber="226">Sesarma</emphasis>
(
<emphasis id="7E957179FFFEFFDDFF5AFE83FE55E818" box="[253,388,317,341]" italics="true" pageId="9" pageNumber="226">Holometopus</emphasis>
) 
<emphasis id="7E957179FFFEFFDDFE34FE83FE34E818" box="[403,485,317,341]" italics="true" pageId="9" pageNumber="226">villosum</emphasis>
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<emphasis id="7E957179FFFEFFDDFE42FE83FE2EE818" box="[485,511,317,341]" italics="true" pageId="9" pageNumber="226">—</emphasis>
<treatmentCitation id="CD408B7AFFFEFFDDFE58FE83FD62E818" author="McNeill, F. A." box="[511,691,317,341]" page="79" pageId="9" pageNumber="226" year="1968">
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(in part).— 
<treatmentCitation id="CD408B7AFFFEFFDDFC8DFE83FC35E818" author="Rathbun, M. J." box="[810,996,317,341]" page="35" pageId="9" pageNumber="226" year="1907">
<bibRefCitation id="2870D09AFFFEFFDDFC8DFE83FC35E818" author="Rathbun, M. J." box="[810,996,317,341]" pageId="9" pageNumber="226" pagination="25 - 74" refId="ref11841" refString="Rathbun, M. J. (1907) Reports on the scientific results of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. IX. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. X: The Brachyura. Memoirs of the Museum of Comparative Zoology at Harvard College, 35 (2), 25 - 74." type="journal article" year="1907">Rathbun 1907: 35</bibRefCitation>
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.
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</paragraph>
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<emphasis id="7E957179FFFEFFDDFF30FEE2FF3FE83E" box="[151,238,348,371]" italics="true" pageId="9" pageNumber="226">Sesarma</emphasis>
(
<emphasis id="7E957179FFFEFFDDFF58FEE2FE57E839" box="[255,390,348,372]" italics="true" pageId="9" pageNumber="226">Holometopus</emphasis>
) 
<emphasis id="7E957179FFFEFFDDFE30FEE2FE0AE839" box="[407,475,348,372]" italics="true" pageId="9" pageNumber="226">villosa</emphasis>
</taxonomicName>
— 
<treatmentCitation id="CD408B7AFFFEFFDDFE54FEE2FD00E839" author="Tesch, J. J." box="[499,721,348,372]" page="208" pageId="9" pageNumber="226" year="1917">
<bibRefCitation id="2870D09AFFFEFFDDFE54FEE2FD00E839" author="Tesch, J. J." box="[499,721,348,372]" pageId="9" pageNumber="226" pagination="127 - 260" refId="ref12609" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, Leiden, 3 (2 - 3), 127 - 260, pls. 15 - 17. https: // doi. org / 10.5962 / bhl. title. 10640" type="journal article" year="1917">Tesch 1917: 208–210</bibRefCitation>
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(in part), 231 (list), 237 (key), 239 (footnote), 260, pl. 17 fig. 2 [not 
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<emphasis id="7E957179FFFEFFDDFF60FEC5FEAAE8DF" box="[199,379,378,402]" italics="true" pageId="9" pageNumber="226">Sesarma villosum</emphasis>
A. 
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<treatmentCitationGroup id="6CF18A45FFFEFFDDFF30FE27FA4DE8FC" box="[151,1436,409,433]" pageId="9" pageNumber="226">
<emphasis id="7E957179FFFEFFDDFF30FE27FE5AE8FC" box="[151,395,409,433]" italics="true" pageId="9" pageNumber="226">
<taxonomicName id="8BE1D6E8FFFEFFDDFF30FE27FEA1E8FC" ID-CoL="84GRJ" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[151,368,409,433]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="villosum">Chiromantes villosum</taxonomicName>
—
</emphasis>
<treatmentCitation id="CD408B7AFFFEFFDDFE2CFE27FDB6E8FC" author="Nomoto, A. &amp; Yodo, S. &amp; Kimura, S. &amp; Kishino, T. &amp; Sakano, M. &amp; Wada, K." box="[395,615,409,433]" page="9" pageId="9" pageNumber="226" year="1999">
<bibRefCitation id="2870D09AFFFEFFDDFE2CFE27FDB6E8FC" author="Nomoto, A. &amp; Yodo, S. &amp; Kimura, S. &amp; Kishino, T. &amp; Sakano, M. &amp; Wada, K." box="[395,615,409,433]" pageId="9" pageNumber="226" pagination="5 - 9" refId="ref11720" refString="Nomoto, A., Yodo, S., Kimura, S., Kishino, T., Sakano, M. &amp; Wada, K. (1999) Six rare brachyuran species of the family Grapsidae, recorded from the Kinokawa River Estuary, Wakayama Prefecture. Nanki Seibutu, 41 (1), 5 - 9 [in Japanese]" type="journal article" year="1999">
Nomoto 
<emphasis id="7E957179FFFEFFDDFE44FE24FDDEE8FC" box="[483,527,409,433]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 1999: 9
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, pl. 1 fig. 6.— 
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<bibRefCitation id="2870D09AFFFEFFDDFD53FE27FBDEE8FC" author="Kishino, T. &amp; Yonegawa, T. &amp; Nomoto, A. &amp; Kimura, S. &amp; Wada, K." box="[756,1039,409,433]" pageId="9" pageNumber="226" pagination="15 - 22" refId="ref10767" refString="Kishino, T., Yonegawa, T., Nomoto, A., Kimura, S. &amp; Wada, K. (2001 a) Twelve rare species of brachyuran crabs recorded in the brackish waters of Amami-Oshima, Kagoshima Prefecture, Japan. Nanki Seibutsu, 43 (1), 15 - 22. [in Japanese]" type="journal article" year="2001">
Kishino 
<emphasis id="7E957179FFFEFFDDFCEBFE24FCA9E8FC" box="[844,888,409,433]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2001a: 17–18
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, pl. 2 fig. 2.— 
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<bibRefCitation id="2870D09AFFFEFFDDFB3BFE27FA4DE8FC" author="Kishino, T. &amp; Yonegawa, T. &amp; Nomoto, A. &amp; Kimura, S. &amp; Wada, K." box="[1180,1436,409,433]" pageId="9" pageNumber="226" pagination="125 - 131" refId="ref10830" refString="Kishino, T., Yonegawa, T., Nomoto, A., Kimura, S. &amp; Wada, K. (2001 b) Brachyuran crab species recorded in the brackish waters of Amami-Oshima, Kagoshima Prefecture, Japan. Nanki Seibutsu, 43 (2), 125 - 131. [in Japanese]" type="journal article" year="2001">
Kishino 
<emphasis id="7E957179FFFEFFDDFB53FE24FACEE8FC" box="[1268,1311,409,433]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2001b: 127
</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
(list), table 2.— 
<bibRefCitation id="2870D09AFFFEFFDDFEC2FE06FD9CE89D" author="Shokita, S. &amp; Nagai, T. &amp; Fujita, Y. &amp; Naruse, T. &amp; Nagamatsu, T. &amp; Yamazaki, T. &amp; Shinjo, M. &amp; Nagata, Y." box="[357,589,440,464]" pageId="9" pageNumber="226" pagination="73 - 86" refId="ref12340" refString="Shokita, S., Nagai, T., Fujita, Y., Naruse, T., lto, A., Nagamatsu, T., Yamazaki, T., Shinjo, M. &amp; Nagata, Y. (2002) [= Distribution and abundance of crustaceans in the Ohura River mangrove swamp of Okinawa Island, Japan]. In: A Comprehensive Study for Mangrove Ecosystem in Okinawa. Research Institute for Subtropics, Naha, Okinawa, pp. 73 - 86. [in Japanese with an English summary]" type="book chapter" year="2002">
Shokita 
<emphasis id="7E957179FFFEFFDDFE1CFE06FE39E89D" box="[443,488,440,464]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2002: 78
</bibRefCitation>
(list), 85, photo 4A–1 fig. A [not 
<taxonomicName id="8BE1D6E8FFFEFFDDFC04FE06FA8FE882" ID-CoL="4WZ2R" authority="A. Milne-Edwards, 1869" authorityName="A. Milne-Edwards" authorityYear="1869" box="[931,1374,439,464]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFFEFFDDFC04FE06FB86E89D" box="[931,1111,440,464]" italics="true" pageId="9" pageNumber="226">Sesarma villosum</emphasis>
A. 
<bibRefCitation id="2870D09AFFFEFFDDFBDBFE06FA8FE882" author="Milne-Edwards, A." box="[1148,1374,439,464]" pageId="9" pageNumber="226" pagination="25 - 31" refId="ref11446" refString="Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]" type="journal article" year="1869">Milne-Edwards, 1869</bibRefCitation>
</taxonomicName>
]. 
<taxonomicName id="8BE1D6E8FFFEFFDDFF30FE68FE18E8A3" authorityName="Edmondson" authorityYear="1951" box="[151,457,470,494]" class="Malacostraca" family="Sesarmidae" genus="Cleistocoeloma" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFEFFDDFF30FE68FEE4E8A3" box="[151,309,470,494]" italics="true" pageId="9" pageNumber="226">Cleistocoeloma</emphasis>
(sic) 
<emphasis id="7E957179FFFEFFDDFEC9FE69FE18E8A3" box="[366,457,471,494]" italics="true" pageId="9" pageNumber="226">suvaense</emphasis>
</taxonomicName>
— 
<bibRefCitation id="2870D09AFFFEFFDDFE44FE69FD79E8A3" author="Tan, C. G. S. &amp; Ng, P. K. L." box="[483,680,470,495]" pageId="9" pageNumber="226" pagination="75 - 84" refId="ref12558" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84. https: // doi. org / 10.1007 / BF 00005655" type="journal article" year="1994">Tan &amp; Ng 1994: 82</bibRefCitation>
(list) [not 
<taxonomicName id="8BE1D6E8FFFEFFDDFCB3FE68FB1AE8A3" ID-CoL="5YZJP" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[788,1227,470,494]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFEFFDDFCB3FE68FBD6E8A3" box="[788,1031,470,494]" italics="true" pageId="9" pageNumber="226">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFFEFFDDFBA9FE68FB1AE8A3" author="Edmondson, C. H." box="[1038,1227,470,494]" pageId="9" pageNumber="226" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="4C5EAD6BFFFEFFDDFF30FE4BFB7BEB40" blockId="9.[151,1437,255,678]" box="[151,1194,501,525]" pageId="9" pageNumber="226">
<taxonomicName id="8BE1D6E8FFFEFFDDFF30FE4BFE40EB40" authorityName="Edmondson" authorityYear="1951" box="[151,401,501,525]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="suvaensis">
<emphasis id="7E957179FFFEFFDDFF30FE4BFE40EB40" box="[151,401,501,525]" italics="true" pageId="9" pageNumber="226">Clistocoeloma suvaensis</emphasis>
</taxonomicName>
—Wee &amp; Ng 1996: 82 (list) [not 
<taxonomicName id="8BE1D6E8FFFEFFDDFD4FFE4BFB4EEB40" ID-CoL="5YZJP" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[744,1183,501,525]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFEFFDDFD4FFE4BFC0AEB40" box="[744,987,501,525]" italics="true" pageId="9" pageNumber="226">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFFEFFDDFC45FE4BFB4EEB40" author="Edmondson, C. H." box="[994,1183,501,525]" pageId="9" pageNumber="226" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="4C5EAD6BFFFEFFDDFF30FDAAFA4CEB07" blockId="9.[151,1437,255,678]" pageId="9" pageNumber="226">
<taxonomicName id="8BE1D6E8FFFEFFDDFF30FDAAFE57EB61" ID-CoL="5YZK2" authorityName="A. Milne-Edwards" authorityYear="1869" box="[151,390,532,556]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFFEFFDDFF30FDAAFE57EB61" box="[151,390,532,556]" italics="true" pageId="9" pageNumber="226">Clistocoeloma villosum</emphasis>
</taxonomicName>
— 
<bibRefCitation id="2870D09AFFFEFFDDFE07FDAAFD96EB61" author="Davie, P. J. F." box="[416,583,532,556]" pageId="9" pageNumber="226" pagination="1 - 641" refId="ref10217" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda - Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. K. (Eds.), Zoological Catalogues of Australia. Vol. 19.3 B. CSIRO Publishing, Melbourne, pp. i - xiv + 1 - 641." type="book chapter" year="2002">Davie 2002: 221</bibRefCitation>
(in part).— 
<bibRefCitation id="2870D09AFFFEFFDDFD19FDAAFC4FEB61" author="Komai, T. &amp; Nagai, T. &amp; Yogi, A. &amp; Naruse, T. &amp; Fujita, Y. &amp; Shokita, S." box="[702,926,532,556]" pageId="9" pageNumber="226" pagination="33 - 63" refId="ref10699" refString="Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. &amp; Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63." type="journal article" year="2004">
Komai 
<emphasis id="7E957179FFFEFFDDFCACFDAAFCE6EB61" box="[779,823,532,556]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2004: 38
</bibRefCitation>
, 40–42, fig. 1B, 3.— 
<bibRefCitation id="2870D09AFFFEFFDDFBD7FDAAFAA2EB66" author="Karasawa, T. &amp; Kimura, S. &amp; Kuroda, M. &amp; Nomoto, A." box="[1136,1395,531,556]" pageId="9" pageNumber="226" pagination="60 - 62" refId="ref10640" refString="Karasawa, T., Kimura, S., Kuroda, M. &amp; Nomoto, A. (2006) Record of Clistocoeloma villosum (Crustacea, Brachyura, Sesarmidae) from the Waka River Estuary in Wakayama Prefecture, Japan. Nanki-Seibutu, 48 (1), 60 - 62. [in Japanese]" type="journal article" year="2006">
Karasawa 
<emphasis id="7E957179FFFEFFDDFB79FDAAFAC0EB61" box="[1246,1297,532,556]" italics="true" pageId="9" pageNumber="226">et al.</emphasis>
2006: 60
</bibRefCitation>
, 61, fig. 1. 
<emphasis id="7E957179FFFEFFDDFEA4FD8DFECBEB07" box="[259,282,563,586]" italics="true" pageId="9" pageNumber="226">—</emphasis>
<bibRefCitation id="2870D09AFFFEFFDDFEBEFD8CFE2EEB07" author="Schubart, C. D. &amp; Liu, H. - C. &amp; Ng, P. K. L." box="[281,511,562,586]" pageId="9" pageNumber="226" pagination="1 - 29" refId="ref12174" refString="Schubart, C. D., Liu, H. - C. &amp; Ng, P. K. L. (2009) Revision of Selatium Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), with description of a new genus and two new species. Zootaxa, 2154 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2154.1.1" type="journal article" year="2009">
Schubart 
<emphasis id="7E957179FFFEFFDDFEDEFD8DFE77EB07" box="[377,422,562,586]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2009: 3
</bibRefCitation>
(list), fig. 18 (in part).— 
<bibRefCitation id="2870D09AFFFEFFDDFD5EFD8CFC28EB07" author="Lee, S. - K. &amp; Jung, J. &amp; Kim, W." box="[761,1017,562,586]" pageId="9" pageNumber="226" pagination="179 - 181" refId="ref11086" refString="Lee, S. - K., Jung, J. &amp; Kim, W. (2010) A new report on sesarmid crab Clistocoeloma villosum (Crustacea: Decapoda: Brachyura) from Korea. Korean Journal of Systematic Zoology, 26 (2), 179 - 181. https: // doi. org / 10.5635 / KJSZ. 2010.26.2.179" type="journal article" year="2010">
Lee 
<emphasis id="7E957179FFFEFFDDFC81FD8DFC83EB07" box="[806,850,562,586]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2010: 179–181
</bibRefCitation>
, fig. 1 A–H.—Maenosono &amp; Sakai 2016:
</paragraph>
<paragraph id="4C5EAD6BFFFEFFDDFF60FDEFFB88EBEB" blockId="9.[151,1437,255,678]" pageId="9" pageNumber="226">
6, fig. 2D.— 
<bibRefCitation id="2870D09AFFFEFFDDFEE0FDEFFDDFEB24" author="Yuhara, T. &amp; Yokooka, H. &amp; Taru, M." box="[327,526,593,617]" pageId="9" pageNumber="226" pagination="19" refId="ref12778" refString="Yuhara, T., Yokooka, H. &amp; Taru, M. (2017) Range extension of the sesarmid crab Clistocoeloma villosum along the eastern Pacific coast of the Izu Peninsula, Japan. Marine Biodiversity Records, 10, 19. https: // doi. org / 10.1186 / s 41200 - 017 - 0122 - 1" type="journal article" year="2017">
Yuhara 
<emphasis id="7E957179FFFEFFDDFE3CFDECFE18EB24" box="[411,457,593,617]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2017
</bibRefCitation>
: [1–4], fig. 2.— 
<bibRefCitation id="2870D09AFFFEFFDDFD16FDEFFC27EB24" author="Yuhara, T. &amp; Yokooka, H." box="[689,1014,593,617]" pageId="9" pageNumber="226" pagination="8 - 13" refId="ref12718" refString="Yuhara, T. &amp; Yokooka, H. (2019) Life history of the sesarmid crab, Clistocoeloma villosum, on the southern coast of the Izu Peninsula, central Japan. Plankton &amp; Benthos Research, 14 (1), 8 - 13. https: // doi. org / 10.3800 / pbr. 14.8" type="journal article" year="2019">Yuhara &amp; Yokooka 2019: 8–11</bibRefCitation>
, figs. 2C, 3.— 
<bibRefCitation id="2870D09AFFFEFFDDFB29FDEFFA8FEB24" author="Li, J. - J. &amp; Hsu, J. - W. &amp; Ng, N. K. &amp; Shih, H. - T." box="[1166,1374,593,617]" pageId="9" pageNumber="226" pagination="1207 - 1230" refId="ref10922" refString="Li, J. - J., Hsu, J. - W., Ng, N. K. &amp; Shih, H. - T. (2019) Eight new records of crabs (Decapoda, Brachyura: Sesarmidae, Varunidae) from the coasts of Taiwan. Crustaceana, 92 (10), 1207 - 1230. https: // doi. org / 10.1163 / 15685403 - 00003935" type="journal article" year="2019">
Li 
<emphasis id="7E957179FFFEFFDDFB09FDECFB0DEB24" box="[1198,1244,593,617]" italics="true" pageId="9" pageNumber="226">et al</emphasis>
. 2019: 1208
</bibRefCitation>
(list), 1211–1214, figs. 1B, 3. – 
<bibRefCitation id="2870D09AFFFEFFDDFE69FDCEFCC8EBCA" author="Takakura, H. &amp; Komai, T." box="[462,793,623,648]" pageId="9" pageNumber="226" pagination="25 - 31" refId="ref12498" refString="Takakura, H. &amp; Komai, T. (2019) New records of a semiterrestrial sesarmid crab Clistocoeloma villosum (A. Milne-Edwards, 1869) (Crustacea: Decapoda: Brachyura: Grapsoidea) from Izu and Boso Peninsulas, Japan. Cancer, 28, 25 - 31. [in Japanese with English abstract]" type="journal article" year="2019">Takakura &amp; Komai 2019: 25–30</bibRefCitation>
, figs. 1–5 [not 
<taxonomicName id="8BE1D6E8FFFEFFDDFC16FDCEFABDEBCA" ID-CoL="4WZ2R" authority="A. Milne-Edwards, 1869" authorityName="A. Milne-Edwards" authorityYear="1869" box="[945,1388,623,648]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFFEFFDDFC16FDCEFBB4EBC5" box="[945,1125,624,648]" italics="true" pageId="9" pageNumber="226">Sesarma villosum</emphasis>
A. 
<bibRefCitation id="2870D09AFFFEFFDDFB2DFDCEFABDEBCA" author="Milne-Edwards, A." box="[1162,1388,623,648]" pageId="9" pageNumber="226" pagination="25 - 31" refId="ref11446" refString="Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]" type="journal article" year="1869">Milne-Edwards, 1869</bibRefCitation>
</taxonomicName>
]. 
<taxonomicName id="8BE1D6E8FFFEFFDDFF30FD30FE79EBEB" ID-CoL="5YZJP" authorityName="Edmondson" authorityYear="1951" box="[151,424,654,678]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFEFFDDFF30FD30FEF8EBEB" box="[151,297,654,678]" italics="true" pageId="9" pageNumber="226">Clistocoeloma</emphasis>
cf. 
<emphasis id="7E957179FFFEFFDDFEF7FD31FE79EBEB" box="[336,424,655,678]" italics="true" pageId="9" pageNumber="226">suvaense</emphasis>
</taxonomicName>
<emphasis id="7E957179FFFEFFDDFE0FFD31FE13EBEB" box="[424,450,655,678]" italics="true" pageId="9" pageNumber="226">—</emphasis>
<bibRefCitation id="2870D09AFFFEFFDDFE65FD30FDC7EBEB" author="Li, J. - J." box="[450,534,654,678]" pageId="9" pageNumber="226" pagination="84 - 91" refId="ref10890" refString="Li, J. - J. (2015) Land crabs of Christmas Island and Taiwan. Taiwan Natural Science, 126, 84 - 91. [in Chinese]" type="journal article" year="2015">Li 2015</bibRefCitation>
: fig. 15 [not 
<taxonomicName id="8BE1D6E8FFFEFFDDFD30FD30FB9FEBEB" ID-CoL="5YZJP" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[663,1102,654,678]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="226" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFEFFDDFD30FD30FC5BEBEB" box="[663,906,654,678]" italics="true" pageId="9" pageNumber="226">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFFEFFDDFC36FD30FB9FEBEB" author="Edmondson, C. H." box="[913,1102,654,678]" pageId="9" pageNumber="226" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFFEFFDEFF30FD6BFE14E8C4" lastPageId="10" lastPageNumber="227" pageId="9" pageNumber="226" type="materials_examined">
<paragraph id="4C5EAD6BFFFEFFDDFF30FD6BFE3EEAA6" blockId="9.[151,1437,724,2011]" pageId="9" pageNumber="226">
<emphasis id="7E957179FFFEFFDDFF30FD6BFE95EBA3" bold="true" box="[151,324,724,751]" pageId="9" pageNumber="226">Type material.</emphasis>
<materialsCitation id="FC89A736FFFEFFDDFEEEFD6BFB25EA5E" ID-GBIF-Occurrence="4161964336" collectingDate="2013-05-20" collectionCode="ZRC" collectorName="II Expedition" country="Singapore" latitude="1.2153333" location="St. John's Island" longLatPrecision="13" longitude="103.8515" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0970" specimenCount="1" typeStatus="holotype">
<emphasis id="7E957179FFFEFFDDFEEEFD6BFE6EEBA2" bold="true" box="[329,447,725,751]" pageId="9" pageNumber="226">
<typeStatus id="935A13C9FFFEFFDDFEEEFD6BFE6BEBA2" box="[329,442,725,751]" pageId="9" pageNumber="226" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (15.3 × 
<quantity id="8B19008EFFFEFFDDFDF1FD6BFD6BEBA2" box="[598,698,725,751]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="226" unit="mm" value="12.7">12.7 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFD68FD6BFC58EBA2" box="[719,905,725,751]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0970</specimenCode>
), stn SW7 (SS-00221), pontoon at south lagoon, 
<location id="493EFBB0FFFEFFDDFF30FD47FE8BEA5F" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFF30FD47FE8BEA5F" box="[151,346,760,787]" country="Singapore" latitude="1.2153333" longLatPrecision="13" longitude="103.8515" name="St. John's Island" pageId="9" pageNumber="226">St. John’s Island</location>
, 
<collectingCountry id="34F6EDFBFFFEFFDDFECEFD46FE0CEA5F" box="[361,477,760,786]" name="Singapore" pageId="9" pageNumber="226">Singapore</collectingCountry>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFE4AFD47FDB4EA5E" box="[493,613,761,787]" degrees="1" direction="north" minutes="12.92" orientation="latitude" pageId="9" pageNumber="226" precision="9" value="1.2153333">1°12.92’N</geoCoordinate>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFDD3FD46FCD4EA5E" box="[628,773,760,787]" degrees="103" direction="east" minutes="51.09" orientation="longitude" pageId="9" pageNumber="226" precision="9" value="103.8515">103°51.09’E</geoCoordinate>
, coll. CMBS 
<collectorName id="E114C8BDFFFEFFDDFC0EFD47FB94EA5F" box="[937,1093,760,787]" pageId="9" pageNumber="226">II Expedition</collectorName>
, 
<date id="385F8BABFFFEFFDDFBF4FD47FB25EA5E" box="[1107,1268,761,787]" pageId="9" pageNumber="226" value="2013-05-20">
<collectingDate id="281B7243FFFEFFDDFBF4FD47FB25EA5E" box="[1107,1268,761,787]" pageId="9" pageNumber="226" value="2013-05-20">20 May 2013</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFAA5FD47FB07EA7B" ID-GBIF-Occurrence="4161964343" collectingDate="2013-05-20" collectingDateMax="2018-09-30" collectingDateMin="1999-08-16" collectionCode="ZRC" collectorName="II Expedition" country="Singapore" latitude="1.2153333" location="St. John's Island" longLatPrecision="13" longitude="103.8515" municipality="Lazarus Island" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0971" specimenCount="1" specimenCount-female="1" specimenCount-male="10" typeStatus="paratype">
<emphasis id="7E957179FFFEFFDDFAA5FD47FA52EA5E" bold="true" box="[1282,1411,761,787]" pageId="9" pageNumber="226">
<typeStatus id="935A13C9FFFEFFDDFAA5FD47FAAFEA5E" box="[1282,1406,761,787]" pageId="9" pageNumber="226" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFA29FD47FF32EA7B" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(18.7 × 
<quantity id="8B19008EFFFEFFDDFE98FCA2FE72EA7A" box="[319,419,796,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="9" pageNumber="226" unit="mm" value="15.2">15.2 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFE1FFCA3FDA0EA7A" box="[440,625,797,823]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0971</specimenCode>
), same locality and collection information as Holotype.
</materialsCitation>
— 
<materialsCitation id="FC89A736FFFEFFDDFB53FCA2FDF9EA32" ID-GBIF-Occurrence="4161964323" collectingDate="1999-08-16" collectingDateMax="2018-09-30" collectingDateMin="1999-08-16" collectionCode="ZRC" collectorName="C. D. Schubart" country="Singapore" location="Kusu Island" municipality="Lazarus Island" pageId="9" pageNumber="226" specimenCode="ZRC 2002.0154, ZRC 2002.0155, ZRC 2022.0972" specimenCount="13" specimenCount-female="3" specimenCount-male="10" typeStatus="holotype">
<specimenCount id="5AE766E2FFFEFFDDFB53FCA2FA97EA7B" box="[1268,1350,796,822]" pageId="9" pageNumber="226" type="male">3 males</specimenCount>
(16.2 × 
<quantity id="8B19008EFFFEFFDDFF30FCFFFF2CEA16" box="[151,253,833,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="226" unit="mm" value="13.8">13.8 mm</quantity>
, 15.9 × 
<quantity id="8B19008EFFFEFFDDFEF2FCFFFE6AEA16" box="[341,443,833,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="226" unit="mm" value="14.2">14.2 mm</quantity>
, 12.6 × 11.0 mm), 
<specimenCount id="5AE766E2FFFEFFDDFD2BFCFFFD3CEA17" box="[652,749,832,859]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(13.2 × 
<quantity id="8B19008EFFFEFFDDFCECFCFFFC7FEA16" box="[843,942,833,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="9" pageNumber="226" unit="mm" value="11.2">11.2 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFC63FCFFFBAFEA16" box="[964,1150,832,859]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2002.0154</specimenCode>
), 
<location id="493EFBB0FFFEFFDDFB37FCFFFAC8EA17" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFB37FCFFFAC8EA17" box="[1168,1305,832,859]" country="Singapore" municipality="Lazarus Island" name="Kusu Island" pageId="9" pageNumber="226">Kusu Island</location>
, 
<collectingCountry id="34F6EDFBFFFEFFDDFA83FCFEFA49EA17" box="[1316,1432,832,858]" name="Singapore" pageId="9" pageNumber="226">Singapore</collectingCountry>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFF76FCDBFEA5EA33" box="[209,372,868,895]" pageId="9" pageNumber="226">C.D. Schubart</collectorName>
, 
<date id="385F8BABFFFEFFDDFE26FCDAFDF5EA32" box="[385,548,868,895]" pageId="9" pageNumber="226" value="2000-04-15">
<collectingDate id="281B7243FFFEFFDDFE26FCDAFDF5EA32" box="[385,548,868,895]" pageId="9" pageNumber="226" value="2000-04-15">15 April 2000</collectingDate>
</date>
.
</materialsCitation>
— 
<materialsCitation id="FC89A736FFFEFFDDFDE0FCDBFD79EAEF" ID-GBIF-Occurrence="4161964342" collectingDate="1999-08-16" collectingDateMax="2018-09-30" collectingDateMin="1999-08-16" collectionCode="ZRC" collectorName="C. D. Schubart &amp; N. Sivasothi" country="Singapore" location="Kusu Island" municipality="Lazarus Island" pageId="9" pageNumber="226" specimenCode="ZRC 2002.0154, ZRC 2002.0155, ZRC 2022.0972" specimenCount="13" specimenCount-female="3" specimenCount-male="10" typeStatus="holotype">
<specimenCount id="5AE766E2FFFEFFDDFDE0FCDBFD5FEA33" box="[583,654,868,895]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(15.0 × 13.0 mm) (
<specimenCode id="1C470510FFFEFFDDFCD7FCDBFBFCEA32" box="[880,1069,868,895]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2002.0155</specimenCode>
), Sungei Buloh, 
<collectingCountry id="34F6EDFBFFFEFFDDFB4DFCDAFA8FEA33" box="[1258,1374,868,894]" name="Singapore" pageId="9" pageNumber="226">Singapore</collectingCountry>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFF30FC37FEE6EAEF" box="[151,311,904,931]" pageId="9" pageNumber="226">C.D. Schubart</collectorName>
&amp; 
<collectorName id="E114C8BDFFFEFFDDFEFDFC37FE36EAEF" box="[346,487,904,931]" pageId="9" pageNumber="226">N. Sivasothi</collectorName>
, 
<date id="385F8BABFFFEFFDDFE55FC36FD75EAEF" box="[498,676,904,931]" pageId="9" pageNumber="226" value="1999-08-16">
<collectingDate id="281B7243FFFEFFDDFE55FC36FD75EAEF" box="[498,676,904,931]" pageId="9" pageNumber="226" value="1999-08-16">16 August 1999</collectingDate>
</date>
.
</materialsCitation>
— 
<materialsCitation id="FC89A736FFFEFFDDFD60FC36FE3DEAA6" ID-GBIF-Occurrence="4161964310" collectingDate="1999-08-16" collectingDateMax="2018-09-30" collectingDateMin="1999-08-16" collectionCode="ZRC" collectorName="B. Y. Lee" country="Singapore" location="Kusu Island" municipality="Lazarus Island" pageId="9" pageNumber="226" specimenCode="ZRC 2002.0154, ZRC 2002.0155, ZRC 2022.0972" specimenCount="13" specimenCount-female="3" specimenCount-male="10" typeStatus="holotype">
<specimenCount id="5AE766E2FFFEFFDDFD60FC36FCCDEAEF" box="[711,796,904,930]" pageId="9" pageNumber="226" type="male">6 males</specimenCount>
(16.9 × 
<quantity id="8B19008EFFFEFFDDFCDCFC37FC30EAEE" box="[891,993,905,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="9" pageNumber="226" unit="mm" value="14.3">14.3 mm</quantity>
, 15.8 × 
<quantity id="8B19008EFFFEFFDDFB9DFC36FB71EAEE" box="[1082,1184,904,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="226" unit="mm" value="13.5">13.5 mm</quantity>
, 15.2 × 
<quantity id="8B19008EFFFEFFDDFB5DFC37FAB1EAEE" box="[1274,1376,905,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="9" pageNumber="226" unit="mm" value="13.1">13.1 mm</quantity>
, 15.1 × 13.0 mm), 
<specimenCount id="5AE766E2FFFEFFDDFE8EFC13FE47EA8B" box="[297,406,940,967]" pageId="9" pageNumber="226" type="female">2 females</specimenCount>
(15.1 × 
<quantity id="8B19008EFFFEFFDDFE52FC13FD8DEA8A" box="[501,604,941,967]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="9" pageNumber="226" unit="mm" value="12.8">12.8 mm</quantity>
, 12.7 × 
<quantity id="8B19008EFFFEFFDDFD10FC13FCCDEA8A" box="[695,796,941,967]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="9" pageNumber="226" unit="mm" value="10.3">10.3 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFC95FC13FC3FEA8A" box="[818,1006,941,967]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0972</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFBA7FC13FB76EA8B" box="[1024,1191,940,967]" pageId="9" pageNumber="226">Lazarus Island</collectingMunicipality>
, 
<collectingCountry id="34F6EDFBFFFEFFDDFB14FC12FAF6EA8B" box="[1203,1319,940,966]" name="Singapore" pageId="9" pageNumber="226">Singapore</collectingCountry>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFACCFC13FF10EAA6" pageId="9" pageNumber="226">B.Y. Lee</collectorName>
<emphasis id="7E957179FFFEFFDDFF6FFC6FFED1EAA7" box="[200,256,976,1002]" italics="true" pageId="9" pageNumber="226">et al.</emphasis>
, 
<date id="385F8BABFFFEFFDDFEA9FC6FFE3DEAA6" box="[270,492,976,1003]" pageId="9" pageNumber="226" value="2018-09-30">
<collectingDate id="281B7243FFFEFFDDFEA9FC6FFE3DEAA6" box="[270,492,976,1003]" pageId="9" pageNumber="226" value="2018-09-30">30 September 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="4C5EAD6BFFFEFFDEFF60FC4AFE14E8C4" blockId="9.[151,1437,724,2011]" lastBlockId="10.[151,1437,150,393]" lastPageId="10" lastPageNumber="227" pageId="9" pageNumber="226">
<emphasis id="7E957179FFFEFFDDFF60FC4AFDD1ED43" bold="true" box="[199,512,1012,1038]" pageId="9" pageNumber="226">Other material examined.</emphasis>
<materialsCitation id="FC89A736FFFEFFDDFDAEFC4AFE86ED7E" ID-GBIF-Occurrence="4161964309" collectingDate="2003-05" collectorName="J. Jung" country="South Korea" location="Island" municipality="Gosan" pageId="9" pageNumber="226" specimenCount="1" specimenCount-female="1" stateProvince="Jeju">
<emphasis id="7E957179FFFEFFDDFDAEFC4AFD7DED42" bold="true" box="[521,684,1012,1039]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFDAEFC4AFD77ED42" box="[521,678,1012,1039]" name="South Korea" pageId="9" pageNumber="226">South Korea</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFD12FC4BFCC9ED43" box="[693,792,1012,1039]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(12.8 × 
<quantity id="8B19008EFFFEFFDDFCD9FC4BFC34ED42" box="[894,997,1013,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="9" pageNumber="226" unit="mm" value="10.8">10.8 mm</quantity>
) (WS900), 
<collectingMunicipality id="AC3A3711FFFEFFDDFBCAFC4BFB66ED42" box="[1133,1207,1013,1039]" pageId="9" pageNumber="226">Gosan</collectingMunicipality>
, 
<collectingRegion id="8E256389FFFEFFDDFB62FC4AFB23ED43" box="[1221,1266,1012,1038]" country="South Korea" name="Jeju" pageId="9" pageNumber="226">Jeju</collectingRegion>
<location id="493EFBB0FFFEFFDDFB5CFC4AFA93ED43" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFB5CFC4AFA93ED43" box="[1275,1346,1012,1038]" country="South Korea" municipality="Gosan" name="Island" pageId="9" pageNumber="226" stateProvince="Jeju">Island</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFA2DFC4BFF1EED7E" pageId="9" pageNumber="226">J. Jung</collectorName>
, 
<date id="385F8BABFFFEFFDDFF7AFBA7FE86ED7E" box="[221,343,1049,1075]" pageId="9" pageNumber="226" value="2003-05">
<collectingDate id="281B7243FFFEFFDDFF7AFBA7FE86ED7E" box="[221,343,1049,1075]" pageId="9" pageNumber="226" value="2003-05">May 2003</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFEDDFBA7FAE1ED7E" ID-GBIF-Occurrence="4161964308" box="[378,1328,1048,1075]" collectingDate="2003-05" collectorName="J. Jung" country="South Korea" location="Island" municipality="Gosan" pageId="9" pageNumber="226" specimenCount="1" specimenCount-female="1" stateProvince="Jeju">
<specimenCount id="5AE766E2FFFEFFDDFEDDFBA7FE08ED7F" box="[378,473,1048,1075]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(14.5 × 
<quantity id="8B19008EFFFEFFDDFDE5FBA7FD7BED7E" box="[578,682,1049,1075]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="9" pageNumber="226" unit="mm" value="12.1">12.1 mm</quantity>
) (WS899), 
<collectingMunicipality id="AC3A3711FFFEFFDDFC94FBA7FCACED7E" box="[819,893,1049,1075]" pageId="9" pageNumber="226">Gosan</collectingMunicipality>
, 
<collectingRegion id="8E256389FFFEFFDDFC2BFBA6FC68ED7F" box="[908,953,1048,1074]" country="South Korea" name="Jeju" pageId="9" pageNumber="226">Jeju</collectingRegion>
<location id="493EFBB0FFFEFFDDFC64FBA6FBDBED7F" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFC64FBA6FBDBED7F" box="[963,1034,1048,1074]" country="South Korea" municipality="Gosan" name="Island" pageId="9" pageNumber="226" stateProvince="Jeju">Island</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFBF3FBA7FB76ED7E" box="[1108,1191,1049,1075]" pageId="9" pageNumber="226">J. Jung</collectorName>
, 
<date id="385F8BABFFFEFFDDFB12FBA7FAE1ED7E" box="[1205,1328,1049,1075]" pageId="9" pageNumber="226" value="2003-05">
<collectingDate id="281B7243FFFEFFDDFB12FBA7FAE1ED7E" box="[1205,1328,1049,1075]" pageId="9" pageNumber="226" value="2003-05">May 2003</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFAF4FBA7FC14ED1A" ID-GBIF-Occurrence="4161964341" collectingDate="2003-05" collectorName="J. Jung" country="South Korea" location="Island" municipality="Gosan" pageId="9" pageNumber="226" specimenCount="1" specimenCount-male="1" stateProvince="Jeju">
<specimenCount id="5AE766E2FFFEFFDDFAF4FBA7FA4DED7F" box="[1363,1436,1048,1075]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(16.2 × 
<quantity id="8B19008EFFFEFFDDFF57FB83FE87ED1A" box="[240,342,1085,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="226" unit="mm" value="14.2">14.2 mm</quantity>
) (WS898), 
<collectingMunicipality id="AC3A3711FFFEFFDDFE7CFB83FDF4ED1A" box="[475,549,1085,1111]" pageId="9" pageNumber="226">Gosan</collectingMunicipality>
, 
<collectingRegion id="8E256389FFFEFFDDFD96FB82FD8FED1B" box="[561,606,1084,1110]" country="South Korea" name="Jeju" pageId="9" pageNumber="226">Jeju</collectingRegion>
<location id="493EFBB0FFFEFFDDFDC2FB82FD7DED1B" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFDC2FB82FD7DED1B" box="[613,684,1084,1110]" country="South Korea" municipality="Gosan" name="Island" pageId="9" pageNumber="226" stateProvince="Jeju">Island</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFD56FB83FC93ED1A" box="[753,834,1085,1111]" pageId="9" pageNumber="226">J. Jung</collectorName>
, 
<date id="385F8BABFFFEFFDDFCE9FB83FC14ED1A" box="[846,965,1085,1111]" pageId="9" pageNumber="226" value="2003-05">
<collectingDate id="281B7243FFFEFFDDFCE9FB83FC14ED1A" box="[846,965,1085,1111]" pageId="9" pageNumber="226" value="2003-05">May 2003</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFC68FB83FE7BED36" ID-GBIF-Occurrence="4161964303" country="Japan" location="Japan" pageId="9" pageNumber="226" specimenCount="3" specimenCount-male="3">
<emphasis id="7E957179FFFEFFDDFC68FB83FBF3ED1A" bold="true" box="[975,1058,1085,1111]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFC68FB83FBCDED1A" box="[975,1052,1085,1111]" name="Japan" pageId="9" pageNumber="226">Japan</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFB8EFB83FB51ED1B" box="[1065,1152,1084,1111]" pageId="9" pageNumber="226" type="male">3 males</specimenCount>
(18.2 × 15.0 mm, 12.7 × 11.0 mm, 9.4 × 
<quantity id="8B19008EFFFEFFDDFEEFFBDFFE73ED36" box="[328,418,1121,1147]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="9" pageNumber="226" unit="mm" value="8.1">8.1 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="FC89A736FFFEFFDDFE12FBDFFF05EDD3" ID-GBIF-Occurrence="4161964305" collectingDate="2016-07-18" collectionCode="ZRC" collectorName="J. J. Li" country="Japan" location="Yagaji-jima" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0990" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa">
<specimenCount id="5AE766E2FFFEFFDDFE12FBDFFDC7ED37" box="[437,534,1120,1147]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(19.0 × 
<quantity id="8B19008EFFFEFFDDFDD2FBDEFD0BED36" box="[629,730,1120,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="9" pageNumber="226" unit="mm" value="15.8">15.8 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFD56FBDFFC7DED36" box="[753,940,1121,1147]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0990</specimenCode>
), 
<location id="493EFBB0FFFEFFDDFC1AFBDEFB92ED37" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFC1AFBDEFB92ED37" box="[957,1091,1120,1146]" country="Japan" name="Yagaji-jima" pageId="9" pageNumber="226" stateProvince="Okinawa">Yagaji-jima</location>
, 
<collectingRegion id="8E256389FFFEFFDDFBE9FBDEFB67ED37" box="[1102,1206,1120,1146]" country="Japan" name="Okinawa" pageId="9" pageNumber="226">Okinawa</collectingRegion>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFB5EFBDFFAEEED37" box="[1273,1343,1120,1147]" pageId="9" pageNumber="226">J.J. Li</collectorName>
, 
<date id="385F8BABFFFEFFDDFAECFBDFFF05EDD3" pageId="9" pageNumber="226" value="2016-07-18">
<collectingDate id="281B7243FFFEFFDDFAECFBDFFF05EDD3" pageId="9" pageNumber="226" value="2016-07-18">18 July 2016</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFF50FB3AFABEEDD2" ID-GBIF-Occurrence="4161964337" box="[247,1391,1156,1183]" collectingDate="1978-05-23" collectionCode="ZRC" country="Japan" location="Yagaji Island" municipality="Nago City" pageId="9" pageNumber="226" specimenCode="ZRC 1999.0254" specimenCount="1">
1 damaged female (11.9 × 
<quantity id="8B19008EFFFEFFDDFD82FB3BFD5AEDD2" box="[549,651,1157,1183]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="226" unit="mm" value="10.1">10.1 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFD04FB3BFC8EEDD2" box="[675,863,1156,1183]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 1999.0254</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFCD3FB3BFC3BEDD3" box="[884,1002,1156,1183]" pageId="9" pageNumber="226">Nago City</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFC52FB3AFB5CEDD3" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFC52FB3AFB5CEDD3" box="[1013,1165,1156,1182]" country="Japan" municipality="Nago City" name="Yagaji Island" pageId="9" pageNumber="226">Yagaji Island</location>
, coll. 
<date id="385F8BABFFFEFFDDFB74FB3BFABEEDD2" box="[1235,1391,1157,1183]" pageId="9" pageNumber="226" value="1978-05-23">
<collectingDate id="281B7243FFFEFFDDFB74FB3BFABEEDD2" box="[1235,1391,1157,1183]" pageId="9" pageNumber="226" value="1978-05-23">23 May 1978</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFA34FB3BFB0CED8E" ID-GBIF-Occurrence="4161964302" collectingDate="2009-11-16" collectionCode="ZRC" collectorName="T. Komai" country="Japan" location="Kumeijima" pageId="9" pageNumber="226" specimenCode="ZRC 2011.1037" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="5AE766E2FFFEFFDDFA34FB3BFF1CED8F" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(9.4 × 8.0 mm) (
<specimenCode id="1C470510FFFEFFDDFE2DFB17FD92ED8E" box="[394,579,1193,1219]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2011.1037</specimenCode>
), 
<location id="493EFBB0FFFEFFDDFDF2FB16FD07ED8F" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFDF2FB16FD07ED8F" box="[597,726,1192,1218]" country="Japan" name="Kumeijima" pageId="9" pageNumber="226" stateProvince="Okinawa">Kumeijima</location>
, 
<collectingRegion id="8E256389FFFEFFDDFD46FB16FC98ED8F" box="[737,841,1192,1218]" country="Japan" name="Okinawa" pageId="9" pageNumber="226">Okinawa</collectingRegion>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFC2DFB17FC27ED8F" box="[906,1014,1192,1219]" pageId="9" pageNumber="226">T. Komai</collectorName>
, 
<date id="385F8BABFFFEFFDDFBA7FB16FB0CED8E" box="[1024,1245,1192,1219]" pageId="9" pageNumber="226" value="2009-11-16">
<collectingDate id="281B7243FFFEFFDDFBA7FB16FB0CED8E" box="[1024,1245,1192,1219]" pageId="9" pageNumber="226" value="2009-11-16">16 November 2009</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFAA7FB17FEA4EC46" ID-GBIF-Occurrence="4161964328" collectingDate="2009-11-18" collectionCode="ZRC" collectorName="Kumeijima &amp; Expedition" country="Japan" latitude="26.34939" location="Kumejima" longLatPrecision="1" longitude="126.72608" municipality="Ohara" pageId="9" pageNumber="226" specimenCode="ZRC 2011.1036" specimenCount="1" specimenCount-female="1">
<specimenCount id="5AE766E2FFFEFFDDFAA7FB17FA8AED8F" box="[1280,1371,1192,1219]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(10.3 × 
<quantity id="8B19008EFFFEFFDDFF09FB73FED7EDAA" box="[174,262,1229,1255]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="9" pageNumber="226" unit="mm" value="8.8">8.8 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFEBBFB73FE06EDAB" box="[284,471,1228,1255]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2011.1036</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFE4EFB72FDE1EDAB" box="[489,560,1228,1254]" pageId="9" pageNumber="226">Ohara</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFD9BFB72FD64EDAB" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFD9BFB72FD64EDAB" box="[572,693,1228,1254]" country="Japan" latitude="26.34939" longLatPrecision="1" longitude="126.72608" municipality="Ohara" name="Kumejima" pageId="9" pageNumber="226">Kumejima</location>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFD66FB72FC8EEDAA" box="[705,863,1228,1255]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="9" pageNumber="226" precision="1" seconds="57.8" value="26.34939">26°20’57.8”N</geoCoordinate>
<geoCoordinate id="29D5CBACFFFEFFDDFCC1FB72FBC3EDAB" box="[870,1042,1228,1254]" degrees="126" direction="east" minutes="43" orientation="longitude" pageId="9" pageNumber="226" precision="1" seconds="33.9" value="126.72608">126°43’33.9”E</geoCoordinate>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFBF1FB72FB05EDAB" box="[1110,1236,1228,1254]" pageId="9" pageNumber="226">Kumeijima</collectorName>
2009 
<collectorName id="E114C8BDFFFEFFDDFABDFB72FA49EDAB" box="[1306,1432,1228,1254]" pageId="9" pageNumber="226">Expedition</collectorName>
, 
<date id="385F8BABFFFEFFDDFF30FB4FFEA4EC46" box="[151,373,1264,1291]" pageId="9" pageNumber="226" value="2009-11-18">
<collectingDate id="281B7243FFFEFFDDFF30FB4FFEA4EC46" box="[151,373,1264,1291]" pageId="9" pageNumber="226" value="2009-11-18">18 November 2009</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFED8FB4EFE9DEC62" ID-GBIF-Occurrence="4161964326" collectingDate="2014-08-25" collectionCode="ZRC" collectorName="J. J. Li" country="Taiwan" location="Xiangjiaowan" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0991" specimenCount="1">
<emphasis id="7E957179FFFEFFDDFED8FB4EFE31EC47" bold="true" box="[383,480,1264,1290]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFED8FB4EFE0AEC47" box="[383,475,1264,1290]" name="Taiwan" pageId="9" pageNumber="226">Taiwan</collectingCountry>
:
</emphasis>
1 ovigerous female (15.8 × 
<quantity id="8B19008EFFFEFFDDFC86FB4FFC56EC46" box="[801,903,1265,1291]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="9" pageNumber="226" unit="mm" value="13.4">13.4 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFC39FB4FFB88EC46" box="[926,1113,1265,1291]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0991</specimenCode>
), 
<location id="493EFBB0FFFEFFDDFBCBFB4EFADDEC47" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFBCBFB4EFADDEC47" box="[1132,1292,1264,1290]" country="Taiwan" name="Xiangjiaowan" pageId="9" pageNumber="226">Xiangjiaowan</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFAF6FB4FFA46EC47" box="[1361,1431,1264,1291]" pageId="9" pageNumber="226">J.J. Li</collectorName>
, 
<date id="385F8BABFFFEFFDDFF30FAAAFE9DEC62" box="[151,332,1300,1327]" pageId="9" pageNumber="226" value="2014-08-25">
<collectingDate id="281B7243FFFEFFDDFF30FAAAFE9DEC62" box="[151,332,1300,1327]" pageId="9" pageNumber="226" value="2014-08-25">25 August 2014</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFEC8FAABFD88EC1E" ID-GBIF-Occurrence="4161964318" collectingDate="2017-09-03" collectionCode="ZRC" collectorName="Z. C. Ji" country="Taiwan" county="Pintung County" location="Hengchun Township" municipality="Dintanzi" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0992" specimenCount="1">
1 ovigerous female (16.0 × 
<quantity id="8B19008EFFFEFFDDFD3AFAABFCD0EC62" box="[669,769,1301,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="9" pageNumber="226" unit="mm" value="13.2">13.2 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFCB0FAABFC00EC62" box="[791,977,1301,1327]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0992</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFC44FAAAFB97EC63" box="[995,1094,1300,1326]" pageId="9" pageNumber="226">Dintanzi</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFBF6FAAAFAEAEC63" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFBF6FAAAFAEAEC63" box="[1105,1339,1300,1326]" country="Taiwan" county="Pintung County" municipality="Dintanzi" name="Hengchun Township" pageId="9" pageNumber="226">Hengchun Township</location>
, 
<collectingCounty id="A53FD5E7FFFEFFDDFAE2FAAAFF3AEC1E" pageId="9" pageNumber="226">Pintung County</collectingCounty>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFE88FA87FEACEC1F" box="[303,381,1336,1363]" pageId="9" pageNumber="226">Z.C. Ji</collectorName>
, 
<date id="385F8BABFFFEFFDDFE2EFA87FD88EC1E" box="[393,601,1336,1363]" pageId="9" pageNumber="226" value="2017-09-03">
<collectingDate id="281B7243FFFEFFDDFE2EFA87FD88EC1E" box="[393,601,1336,1363]" pageId="9" pageNumber="226" value="2017-09-03">3 September 2017</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFDC3FA87FBCBEC1E" ID-GBIF-Occurrence="4161964313" box="[612,1050,1336,1363]" country="Hong Kong" location="Hong Kong" pageId="9" pageNumber="226" specimenCount="1" specimenCount-male="1">
<emphasis id="7E957179FFFEFFDDFDC3FA87FD26EC1E" bold="true" box="[612,759,1337,1363]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFDC3FA87FD23EC1E" box="[612,754,1337,1363]" name="Hong Kong" pageId="9" pageNumber="226">Hong Kong</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFD59FA87FC9BEC1F" box="[766,842,1336,1363]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(12.4 × 
<quantity id="8B19008EFFFEFFDDFC0DFA87FBC2EC1E" box="[938,1043,1337,1363]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="9" pageNumber="226" unit="mm" value="10.7">10.7 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="FC89A736FFFEFFDDFB81FA87FC59EC3B" ID-GBIF-Occurrence="4161964306" collectingDate="2016-05-09" collectionCode="ZRC" collectorName="K. Wong" country="Hong Kong" location="Sai Kung" municipality="To Kwa Peng" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0993" specimenCount="1" specimenCount-female="1">
<specimenCount id="5AE766E2FFFEFFDDFB81FA87FB56EC1F" box="[1062,1159,1336,1363]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(12.5 × 
<quantity id="8B19008EFFFEFFDDFB4FFA86FA9CEC1E" box="[1256,1357,1336,1363]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="226" unit="mm" value="10.5">10.5 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFAC3FA87FEC2EC3A" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0993</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFE81FAE3FE15EC3A" box="[294,452,1373,1399]" pageId="9" pageNumber="226">To Kwa Peng</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFE76FAE2FDEFEC3A" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFE76FAE2FDEFEC3A" box="[465,574,1372,1399]" country="Hong Kong" municipality="To Kwa Peng" name="Sai Kung" pageId="9" pageNumber="226">Sai Kung</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFD23FAE3FD3DEC3A" box="[644,748,1373,1399]" pageId="9" pageNumber="226">K. Wong</collectorName>
, 
<date id="385F8BABFFFEFFDDFD5EFAE3FC59EC3B" box="[761,904,1372,1399]" pageId="9" pageNumber="226" value="2016-05-09">
<collectingDate id="281B7243FFFEFFDDFD5EFAE3FC59EC3B" box="[761,904,1372,1399]" pageId="9" pageNumber="226" value="2016-05-09">9 May 2016</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFC0CFAE2FBF4ECD6" ID-GBIF-Occurrence="4161964314" collectingDate="2017-09-18" collectionCode="ZRC" collectorName="C. Cheng &amp; A. Quadros" country="Hong Kong" location="Northeast Hong Kong" municipality="Kak Po" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0994" specimenCount="1" specimenCount-male="1">
<specimenCount id="5AE766E2FFFEFFDDFC0CFAE2FC23EC3B" box="[939,1010,1372,1398]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(17.3 × 15.0 mm) (
<specimenCode id="1C470510FFFEFFDDFB73FAE3FA40EC3A" box="[1236,1425,1373,1399]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0994</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFF30FA3EFF3DECD6" box="[151,236,1408,1435]" pageId="9" pageNumber="226">Kak Po</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFF50FA3EFE21ECD6" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFF50FA3EFE21ECD6" box="[247,496,1408,1435]" country="Hong Kong" municipality="Kak Po" name="Northeast Hong Kong" pageId="9" pageNumber="226">Northeast Hong Kong</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFD96FA3FFD4BECD7" box="[561,666,1408,1435]" pageId="9" pageNumber="226">C. Cheng</collectorName>
&amp; 
<collectorName id="E114C8BDFFFEFFDDFD1CFA3FFCEEECD7" box="[699,831,1408,1435]" pageId="9" pageNumber="226">A. Quadros</collectorName>
, 
<date id="385F8BABFFFEFFDDFCEEFA3FFBF4ECD6" box="[841,1061,1408,1435]" pageId="9" pageNumber="226" value="2017-09-18">
<collectingDate id="281B7243FFFEFFDDFCEEFA3FFBF4ECD6" box="[841,1061,1408,1435]" pageId="9" pageNumber="226" value="2017-09-18">18 September 2017</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFB89FA3EFF1EECF2" ID-GBIF-Occurrence="4161964339" country="Philippines" location="Philippines" pageId="9" pageNumber="226" specimenCount="1" specimenCount-male="1">
<emphasis id="7E957179FFFEFFDDFB89FA3EFB6FECD7" bold="true" box="[1070,1214,1408,1434]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFB89FA3EFB69ECD7" box="[1070,1208,1408,1434]" name="Philippines" pageId="9" pageNumber="226">Philippines</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFB63FA3FFADFECD7" box="[1220,1294,1408,1435]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(17.4 × 
<quantity id="8B19008EFFFEFFDDFACCFA3FFF16ECF2" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="9" pageNumber="226" unit="mm" value="14.9">14.9 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="FC89A736FFFEFFDDFF7CFA1BFCDCECAE" ID-GBIF-Occurrence="4161964321" collectingDate="2004-06-30" collectionCode="ZRC" collectorName="Panglao Marine Biodiversity Project" country="Philippines" location="Panglao Island" municipality="Danao" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0973" specimenCount="1" specimenCount-female="1">
<specimenCount id="5AE766E2FFFEFFDDFF7CFA1BFEECECF3" box="[219,317,1444,1471]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(14.9 × 
<quantity id="8B19008EFFFEFFDDFE38FA1BFDD4ECF2" box="[415,517,1445,1471]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="226" unit="mm" value="12.7">12.7 mm</quantity>
) [photographed] (
<specimenCode id="1C470510FFFEFFDDFD77FA1BFC5CECF2" box="[720,909,1445,1471]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0973</specimenCode>
), stn M52, 
<collectingMunicipality id="AC3A3711FFFEFFDDFBABFA1BFB84ECF2" box="[1036,1109,1445,1471]" pageId="9" pageNumber="226">Danao</collectingMunicipality>
embayment, 
<location id="493EFBB0FFFEFFDDFB48FA1AFA49ECF3" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFB48FA1AFA49ECF3" box="[1263,1432,1444,1470]" country="Philippines" municipality="Danao" name="Panglao Island" pageId="9" pageNumber="226">Panglao Island</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFF68FA76FDB8ECAF" box="[207,617,1480,1506]" pageId="9" pageNumber="226">Panglao Marine Biodiversity Project</collectorName>
, 
<date id="385F8BABFFFEFFDDFDD3FA77FCDCECAE" box="[628,781,1481,1507]" pageId="9" pageNumber="226" value="2004-06-30">
<collectingDate id="281B7243FFFEFFDDFDD3FA77FCDCECAE" box="[628,781,1481,1507]" pageId="9" pageNumber="226" value="2004-06-30">30 June 2004</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFC97FA77FDA2EF4A" ID-GBIF-Occurrence="4161964312" collectingDate="2001-11-25" collectionCode="ZRC" collectorName="H. - C. Liu" country="Philippines" location="Kawasan" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0974" specimenCount="1" specimenCount-female="1" stateProvince="Cebu">
<specimenCount id="5AE766E2FFFEFFDDFC97FA77FC5AECAF" box="[816,907,1480,1507]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(15.9 × 
<quantity id="8B19008EFFFEFFDDFC4FFA77FB9CECAE" box="[1000,1101,1481,1507]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="9" pageNumber="226" unit="mm" value="13.4">13.4 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFBC5FA77FACCECAE" box="[1122,1309,1481,1507]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0974</specimenCode>
), 
<location id="493EFBB0FFFEFFDDFA88FA77FA49ECAE" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFA88FA77FA49ECAE" box="[1327,1432,1481,1507]" country="Philippines" name="Kawasan" pageId="9" pageNumber="226" stateProvince="Cebu">Kawasan</location>
, 
<collectingRegion id="8E256389FFFEFFDDFF30FA52FF04EF4B" box="[151,213,1516,1542]" country="Philippines" name="Cebu" pageId="9" pageNumber="226">Cebu</collectingRegion>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFEBDFA53FE5BEF4B" box="[282,394,1516,1543]" pageId="9" pageNumber="226">H.-C. Liu</collectorName>
, 
<date id="385F8BABFFFEFFDDFE31FA52FDA2EF4A" box="[406,627,1516,1543]" pageId="9" pageNumber="226" value="2001-11-25">
<collectingDate id="281B7243FFFEFFDDFE31FA52FDA2EF4A" box="[406,627,1516,1543]" pageId="9" pageNumber="226" value="2001-11-25">25 November 2001</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFDD8FA52FBB5EF66" ID-GBIF-Occurrence="4161964331" collectingDate="2010-04-14" collectionCode="ZRC" country="Vietnam" latitude="8.66875" location="Bay Canh Island" longLatPrecision="1" longitude="106.68222" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0975" specimenCount="1" specimenCount-male="1">
<emphasis id="7E957179FFFEFFDDFDD8FA52FD3FEF4B" bold="true" box="[639,750,1516,1542]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFDD8FA52FD38EF4B" box="[639,745,1516,1542]" name="Vietnam" pageId="9" pageNumber="226">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFD52FA53FC90EF4B" box="[757,833,1516,1543]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
[photographed] (17.4 × 
<quantity id="8B19008EFFFEFFDDFBFFFA52FB6FEF4A" box="[1112,1214,1516,1543]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="9" pageNumber="226" unit="mm" value="15.2">15.2 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFB72FA53FA40EF4A" box="[1237,1425,1516,1543]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0975</specimenCode>
), stn CD-I-08 (22), 
<location id="493EFBB0FFFEFFDDFECDF9AFFDFDEF67" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFECDF9AFFDFDEF67" box="[362,556,1552,1579]" country="Vietnam" latitude="8.66875" longLatPrecision="1" longitude="106.68222" name="Bay Canh Island" pageId="9" pageNumber="226">Bay Canh Island</location>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFD9DF9AEFD1DEF66" box="[570,716,1552,1579]" degrees="08" direction="north" minutes="40.125" orientation="latitude" pageId="9" pageNumber="226" precision="1" value="8.66875">08°40.125’N</geoCoordinate>
<geoCoordinate id="29D5CBACFFFEFFDDFD71F9AEFCA4EF67" box="[726,885,1552,1578]" degrees="106" direction="east" minutes="40.933" orientation="longitude" pageId="9" pageNumber="226" precision="1" value="106.68222">106°40.933’E</geoCoordinate>
, coll. 
<date id="385F8BABFFFEFFDDFC19F9AFFBB5EF66" box="[958,1124,1552,1579]" pageId="9" pageNumber="226" value="2010-04-14">
<collectingDate id="281B7243FFFEFFDDFC19F9AFFBB5EF66" box="[958,1124,1552,1579]" pageId="9" pageNumber="226" value="2010-04-14">14 April 2010</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFB20F9AFFECBEF3E" ID-GBIF-Occurrence="4161964316" collectingDate="2010-04-14" collectionCode="ZRC" country="Vietnam" latitude="8.66875" location="Bay Canh Island" longLatPrecision="1" longitude="106.68222" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0976" specimenCount="1" specimenCount-female="1">
<specimenCount id="5AE766E2FFFEFFDDFB20F9AFFB37EF67" box="[1159,1254,1552,1579]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
[photographed] (14.1 × 12.0 mm) (
<specimenCode id="1C470510FFFEFFDDFEDDF98BFDE8EF03" box="[378,569,1588,1615]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0976</specimenCode>
), stn CD-I-08 (24), 
<location id="493EFBB0FFFEFFDDFC82F98BFC3BEF03" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFC82F98BFC3BEF03" box="[805,1002,1588,1615]" country="Vietnam" latitude="8.66875" longLatPrecision="1" longitude="106.68222" name="Bay Canh Island" pageId="9" pageNumber="226">Bay Canh Island</location>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFC5EF98AFB5AEF02" box="[1017,1163,1588,1615]" degrees="08" direction="north" minutes="40.125" orientation="latitude" pageId="9" pageNumber="226" precision="1" value="8.66875">08°40.125’N</geoCoordinate>
<geoCoordinate id="29D5CBACFFFEFFDDFB31F98AFAE4EF03" box="[1174,1333,1588,1614]" degrees="106" direction="east" minutes="40.933" orientation="longitude" pageId="9" pageNumber="226" precision="1" value="106.68222">106°40.933’E</geoCoordinate>
, coll. 
<date id="385F8BABFFFEFFDDFA27F98BFECBEF3E" pageId="9" pageNumber="226" value="2010-04-14">
<collectingDate id="281B7243FFFEFFDDFA27F98BFECBEF3E" pageId="9" pageNumber="226" value="2010-04-14">14 April 2010</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFE9AF9E7FD62EFDA" ID-GBIF-Occurrence="4161964324" collectingDate="2010-04-14" collectionCode="ZRC" country="Vietnam" latitude="8.66875" location="Bay Canh Island" longLatPrecision="1" longitude="106.68222" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0977" specimenCount="1" specimenCount-male="1">
<specimenCount id="5AE766E2FFFEFFDDFE9AF9E7FE57EF3F" box="[317,390,1624,1651]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
[photographed] (16.0 × 
<quantity id="8B19008EFFFEFFDDFD0FF9E6FCC1EF3E" box="[680,784,1624,1651]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="226" unit="mm" value="13.5">13.5 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFC8DF9E7FC38EF3E" box="[810,1001,1625,1651]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0977</specimenCode>
), stn CD-I-08 (23), 
<location id="493EFBB0FFFEFFDDFB73F9E7FA49EF3F" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFB73F9E7FA49EF3F" box="[1236,1432,1624,1651]" country="Vietnam" latitude="8.66875" longLatPrecision="1" longitude="106.68222" name="Bay Canh Island" pageId="9" pageNumber="226">Bay Canh Island</location>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFF30F9C2FEF9EFDA" box="[151,296,1660,1687]" degrees="08" direction="north" minutes="40.125" orientation="latitude" pageId="9" pageNumber="226" precision="1" value="8.66875">08°40.125’N</geoCoordinate>
<geoCoordinate id="29D5CBACFFFEFFDDFE97F9C2FE1EEFDB" box="[304,463,1660,1686]" degrees="106" direction="east" minutes="40.933" orientation="longitude" pageId="9" pageNumber="226" precision="1" value="106.68222">106°40.933’E</geoCoordinate>
, coll. 
<date id="385F8BABFFFEFFDDFDB4F9C3FD62EFDA" box="[531,691,1660,1687]" pageId="9" pageNumber="226" value="2010-04-14">
<collectingDate id="281B7243FFFEFFDDFDB4F9C3FD62EFDA" box="[531,691,1660,1687]" pageId="9" pageNumber="226" value="2010-04-14">14 April 2010</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="5AE766E2FFFEFFDDFD72F9C3FCF8EFDB" box="[725,809,1660,1687]" pageId="9" pageNumber="226" type="male">8 males</specimenCount>
(16.5 × 
<quantity id="8B19008EFFFEFFDDFC2EF9C3FC3EEFDA" box="[905,1007,1661,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="226" unit="mm" value="14.2">14.2 mm</quantity>
, 14.0 × 12.0 mm, 7.6 × 
<quantity id="8B19008EFFFEFFDDFB59F9C2FA87EFDA" box="[1278,1366,1660,1687]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="9" pageNumber="226" unit="mm" value="6.3">6.3 mm</quantity>
, 6.1 × 
<quantity id="8B19008EFFFEFFDDFF30F91EFF20EFF6" box="[151,241,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="9" pageNumber="226" unit="mm" value="5.1">5.1 mm</quantity>
), 
<materialsCitation id="FC89A736FFFEFFDDFEA3F91FFC7EEF92" ID-GBIF-Occurrence="4161964320" collectingDate="2010-04-14" collectionCode="ZRC" country="Vietnam" latitude="8.66875" location="Bay Canh Island" longLatPrecision="1" longitude="106.68222" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0978" specimenCount="17" specimenCount-female="17">
<specimenCount id="5AE766E2FFFEFFDDFEA3F91FFEAFEFF7" box="[260,382,1696,1723]" pageId="9" pageNumber="226" type="female">17 females</specimenCount>
(16.2 × 
<quantity id="8B19008EFFFEFFDDFE79F91FFD95EFF6" box="[478,580,1697,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="9" pageNumber="226" unit="mm" value="13.1">13.1 mm</quantity>
, 14.9 × 
<quantity id="8B19008EFFFEFFDDFD38F91EFCD4EFF6" box="[671,773,1696,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="9" pageNumber="226" unit="mm" value="12.6">12.6 mm</quantity>
, 7.6 × 
<quantity id="8B19008EFFFEFFDDFCF6F91EFC7BEFF6" box="[849,938,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="226" unit="mm" value="6.5">6.5 mm</quantity>
, 5.0 × 
<quantity id="8B19008EFFFEFFDDFC51F91FFB9CEFF6" box="[1014,1101,1697,1723]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="226" unit="mm" value="4.3">4.3 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFBC3F91FFACEEFF6" box="[1124,1311,1697,1723]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0978</specimenCode>
), stn CD-I-08, 
<location id="493EFBB0FFFEFFDDFF65F97BFE53EF93" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFF65F97BFE53EF93" box="[194,386,1732,1759]" country="Vietnam" latitude="8.66875" longLatPrecision="1" longitude="106.68222" name="Bay Canh Island" pageId="9" pageNumber="226">Bay Canh Island</location>
, 
<geoCoordinate id="29D5CBACFFFEFFDDFE29F97AFDF1EF92" box="[398,544,1732,1759]" degrees="08" direction="north" minutes="40.125" orientation="latitude" pageId="9" pageNumber="226" precision="1" value="8.66875">08°40.125’N</geoCoordinate>
<geoCoordinate id="29D5CBACFFFEFFDDFD8FF97AFD16EF93" box="[552,711,1732,1758]" degrees="106" direction="east" minutes="40.933" orientation="longitude" pageId="9" pageNumber="226" precision="1" value="106.68222">106°40.933’E</geoCoordinate>
, coll. 
<date id="385F8BABFFFEFFDDFCAAF97BFC7EEF92" box="[781,943,1732,1759]" pageId="9" pageNumber="226" value="2010-04-14">
<collectingDate id="281B7243FFFEFFDDFCAAF97BFC7EEF92" box="[781,943,1732,1759]" pageId="9" pageNumber="226" value="2010-04-14">14 April 2010</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFEFFDDFC1DF97AFB61EE4E" ID-GBIF-Occurrence="4161964344" collectingDate="2014-07-18" collectionCode="ZRC" collectorName="Pulau Tioman" country="Malaysia" location="Sungei Baharu" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0979" specimenCount="1" specimenCount-male="1">
<emphasis id="7E957179FFFEFFDDFC1DF97AFBE1EF93" bold="true" box="[954,1072,1732,1758]" pageId="9" pageNumber="226">
<collectingCountry id="34F6EDFBFFFEFFDDFC1DF97AFBFAEF93" box="[954,1067,1732,1758]" name="Malaysia" pageId="9" pageNumber="226">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFEFFDDFB9FF97BFB55EF93" box="[1080,1156,1732,1759]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(14.0 × 
<quantity id="8B19008EFFFEFFDDFB40F97BFA9DEF92" box="[1255,1356,1733,1759]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="9" pageNumber="226" unit="mm" value="11.8">11.8 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFAC3F97BFEC2EE4E" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0979</specimenCode>
), 
<location id="493EFBB0FFFEFFDDFE81F956FE1DEE4F" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFE81F956FE1DEE4F" box="[294,460,1768,1794]" country="Malaysia" name="Sungei Baharu" pageId="9" pageNumber="226">Sungei Baharu</location>
mangrove, 
<collectorName id="E114C8BDFFFEFFDDFDF5F956FD23EE4F" box="[594,754,1768,1794]" pageId="9" pageNumber="226">Pulau Tioman</collectorName>
, coll. LSM4263 students, 
<date id="385F8BABFFFEFFDDFBBEF957FB61EE4E" box="[1049,1200,1768,1795]" pageId="9" pageNumber="226" value="2014-07-18">
<collectingDate id="281B7243FFFEFFDDFBBEF957FB61EE4E" box="[1049,1200,1768,1795]" pageId="9" pageNumber="226" value="2014-07-18">18 July 2014</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFB74F957FF05EE07" ID-GBIF-Occurrence="4161964322" collectingDate="2016-07-16" collectionCode="ZRC" collectorName="H. H. Tan &amp; M. A. H. Chua" country="Malaysia" county="Telok Bakau" location="Pulau Tioman" municipality="Pulau Tulai" pageId="9" pageNumber="226" specimenCode="ZRC 2016.0140" specimenCount="1" specimenCount-male="1" stateProvince="Pahang">
<specimenCount id="5AE766E2FFFEFFDDFB74F957FAC8EE4F" box="[1235,1305,1768,1795]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(11.6 × 
<quantity id="8B19008EFFFEFFDDFADEF957FF15EE6A" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="9" pageNumber="226" unit="mm" value="9.8">9.8 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFF7CF8B3FE47EE6B" box="[219,406,1804,1831]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2016.0140</specimenCode>
), 
<collectingCounty id="A53FD5E7FFFEFFDDFE0EF8B2FDE8EE6B" box="[425,569,1804,1830]" pageId="9" pageNumber="226">Telok Bakau</collectingCounty>
, 
<collectingMunicipality id="AC3A3711FFFEFFDDFDE2F8B2FD18EE6B" box="[581,713,1804,1830]" pageId="9" pageNumber="226">Pulau Tulai</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFD73F8B2FCA5EE6B" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFD73F8B2FCA5EE6B" box="[724,884,1804,1830]" country="Malaysia" county="Telok Bakau" municipality="Pulau Tulai" name="Pulau Tioman" pageId="9" pageNumber="226" stateProvince="Pahang">Pulau Tioman</location>
, 
<collectingRegion id="8E256389FFFEFFDDFCD8F8B2FC05EE6B" box="[895,980,1804,1830]" country="Malaysia" name="Pahang" pageId="9" pageNumber="226">Pahang</collectingRegion>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFBBFF8B3FBAEEE6A" box="[1048,1151,1805,1831]" pageId="9" pageNumber="226">H.H. Tan</collectorName>
&amp; 
<collectorName id="E114C8BDFFFEFFDDFB04F8B3FAEEEE6B" box="[1187,1343,1804,1831]" pageId="9" pageNumber="226">M.A.H. Chua</collectorName>
, 
<date id="385F8BABFFFEFFDDFAEDF8B2FF05EE07" pageId="9" pageNumber="226" value="2016-07-16">
<collectingDate id="281B7243FFFEFFDDFAEDF8B2FF05EE07" pageId="9" pageNumber="226" value="2016-07-16">16 July 2016</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFF50F88EFF00EE22" ID-GBIF-Occurrence="4161964330" collectingDate="2014-07-15" collectionCode="ZRC" collectorName="B. Y. Lee" country="Malaysia" location="Pulau Tioman" municipality="Sungei Baharu" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0980" specimenCount="1" specimenCount-male="1">
<specimenCount id="5AE766E2FFFEFFDDFF50F88EFEEDEE07" box="[247,316,1840,1866]" pageId="9" pageNumber="226" type="male">1 male</specimenCount>
(12.8 × 11.0 mm) (
<specimenCode id="1C470510FFFEFFDDFDA8F88FFD18EE06" box="[527,713,1841,1867]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0980</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFD7DF88EFCAFEE07" box="[730,894,1840,1866]" pageId="9" pageNumber="226">Sungei Baharu</collectingMunicipality>
mangrove, 
<location id="493EFBB0FFFEFFDDFBA7F88EFB4FEE07" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFBA7F88EFB4FEE07" box="[1024,1182,1840,1866]" country="Malaysia" municipality="Sungei Baharu" name="Pulau Tioman" pageId="9" pageNumber="226">Pulau Tioman</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFB79F88FFA93EE06" box="[1246,1346,1841,1867]" pageId="9" pageNumber="226">B.Y. Lee</collectorName>
, 
<date id="385F8BABFFFEFFDDFAEBF88EFF00EE22" pageId="9" pageNumber="226" value="2014-07-15">
<collectingDate id="281B7243FFFEFFDDFAEBF88EFF00EE22" pageId="9" pageNumber="226" value="2014-07-15">15 July 2014</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFF54F8EBFE75EEDE" ID-GBIF-Occurrence="4161964315" collectingDate="2014-07-15" collectionCode="ZRC" collectorName="B. Y. Lee" country="Malaysia" elevation="8" location="Pulau Tioman" municipality="Sungei Baharu" pageId="9" pageNumber="226" specimenCode="ZRC 2022.0981" specimenCount="2" specimenCount-male="2">
<specimenCount id="5AE766E2FFFEFFDDFF54F8EBFE9BEE23" box="[243,330,1876,1903]" pageId="9" pageNumber="226" type="male">2 males</specimenCount>
(12.5 × 11.0 mm, 8.2 × 
<quantity id="8B19008EFFFEFFDDFDC3F8EBFD76EE22" box="[612,679,1877,1903]" metricMagnitude="0" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="226" unit="m" value="7.8">
<elevation id="C7CC4A58FFFEFFDDFDC3F8EBFD76EE22" box="[612,679,1877,1903]" metricMagnitude="0" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="226" unit="m" value="7.8">7.8 m</elevation>
</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFD18F8EBFCABEE22" box="[703,890,1877,1903]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2022.0981</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFC28F8EAFBE7EE23" box="[911,1078,1876,1902]" pageId="9" pageNumber="226">Sungei Baharu</collectingMunicipality>
mangrove, 
<location id="493EFBB0FFFEFFDDFB19F8EAFA8EEE23" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFB19F8EAFA8EEE23" box="[1214,1375,1876,1902]" country="Malaysia" municipality="Sungei Baharu" name="Pulau Tioman" pageId="9" pageNumber="226">Pulau Tioman</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFF30F8C7FF2FEEDE" box="[151,254,1913,1939]" pageId="9" pageNumber="226">B.Y. Lee</collectorName>
, 
<date id="385F8BABFFFEFFDDFEACF8C6FE75EEDE" box="[267,420,1912,1939]" pageId="9" pageNumber="226" value="2014-07-15">
<collectingDate id="281B7243FFFEFFDDFEACF8C6FE75EEDE" box="[267,420,1912,1939]" pageId="9" pageNumber="226" value="2014-07-15">15 July 2014</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFEFFDDFE60F8C7FDC8EEFA" ID-GBIF-Occurrence="4161964329" collectingDate="2000-06-22" collectionCode="ZRC" collectorName="C. D. Schubart" country="Malaysia" location="Kota Kinabalu" municipality="Pulau Manukan" pageId="9" pageNumber="226" specimenCode="ZRC 2000.1655" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="5AE766E2FFFEFFDDFE60F8C7FDF4EEDF" box="[455,549,1912,1939]" pageId="9" pageNumber="226" type="female">1 female</specimenCount>
(17.6 × 
<quantity id="8B19008EFFFEFFDDFD2EF8C6FD21EEDE" box="[649,752,1912,1939]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="9" pageNumber="226" unit="mm" value="14.6">14.6 mm</quantity>
) (
<specimenCode id="1C470510FFFEFFDDFCAFF8C7FC14EEDE" box="[776,965,1912,1939]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC 2000.1655</specimenCode>
), 
<collectingMunicipality id="AC3A3711FFFEFFDDFC7EF8C6FB5EEEDF" box="[985,1167,1912,1938]" pageId="9" pageNumber="226">Pulau Manukan</collectingMunicipality>
, 
<location id="493EFBB0FFFEFFDDFB3CF8C7FA95EEDF" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFB3CF8C7FA95EEDF" box="[1179,1348,1912,1939]" country="Malaysia" municipality="Pulau Manukan" name="Kota Kinabalu" pageId="9" pageNumber="226" stateProvince="Sabah">Kota Kinabalu</location>
, 
<collectingRegion id="8E256389FFFEFFDDFAF6F8C6FA49EEDF" box="[1361,1432,1912,1938]" country="Malaysia" name="Sabah" pageId="9" pageNumber="226">Sabah</collectingRegion>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFF77F823FEA2EEFB" box="[208,371,1948,1975]" pageId="9" pageNumber="226">C.D. Schubart</collectorName>
, 
<date id="385F8BABFFFEFFDDFE27F823FDC8EEFA" box="[384,537,1949,1975]" pageId="9" pageNumber="226" value="2000-06-22">
<collectingDate id="281B7243FFFEFFDDFE27F823FDC8EEFA" box="[384,537,1949,1975]" pageId="9" pageNumber="226" value="2000-06-22">22 June 2000</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="5AE766E2FFFEFFDDFD9CF823FD79EEFB" box="[571,680,1948,1975]" pageId="9" pageNumber="226" type="female">2 females</specimenCount>
(12.8 × 
<quantity id="8B19008EFFFEFFDDFCADF822FCA0EEFA" box="[778,881,1948,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="226" unit="mm" value="10.5">10.5 mm</quantity>
, 9.0 × 
<quantity id="8B19008EFFFEFFDDFC66F823FBCDEEFA" box="[961,1052,1949,1975]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="9" pageNumber="226" unit="mm" value="7.7">7.7 mm</quantity>
), 1 ovigerous female (12.5 × 
<quantity id="8B19008EFFFEFFDDFACCF823FF16EE96" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="226" unit="mm" value="10.1">10.1 mm</quantity>
), 
<materialsCitation id="FC89A736FFFEFFDDFF7BF87FFABCEE97" ID-GBIF-Occurrence="4161964304" box="[220,1389,1984,2011]" collectingDate="1969-10-05" collectionCode="ZRC" collectorName="C. L. Soh" country="Malaysia" location="Straits" pageId="9" pageNumber="226" specimenCount="1" specimenCount-juvenile="1" stateProvince="Johor">
<specimenCount id="5AE766E2FFFEFFDDFF7BF87FFE9FEE97" box="[220,334,1984,2011]" pageId="9" pageNumber="226" type="juvenile">1 juvenile</specimenCount>
female (5.5 × 
<quantity id="8B19008EFFFEFFDDFE5CF87FFD84EE96" box="[507,597,1985,2011]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="226" unit="mm" value="4.9">4.9 mm</quantity>
) (
<collectionCode id="2AF035AEFFFEFFDDFDCAF87FFD74EE96" box="[621,677,1985,2011]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="226">ZRC</collectionCode>
1969.10.13.3), 
<collectingRegion id="8E256389FFFEFFDDFCFCF87EFC77EE97" box="[859,934,1984,2010]" country="Malaysia" name="Johor" pageId="9" pageNumber="226">Johore</collectingRegion>
<location id="493EFBB0FFFEFFDDFC08F87EFC28EE97" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFEFFDDFC08F87EFC28EE97" box="[943,1017,1984,2010]" country="Malaysia" name="Straits" pageId="9" pageNumber="226" stateProvince="Johor">Straits</location>
, coll. 
<collectorName id="E114C8BDFFFEFFDDFBE6F87FFB7AEE97" box="[1089,1195,1984,2011]" pageId="9" pageNumber="226">C.L. Soh</collectorName>
, 
<date id="385F8BABFFFEFFDDFB1FF87EFABCEE97" box="[1208,1389,1984,2011]" pageId="9" pageNumber="226" value="1969-10-05">
<collectingDate id="281B7243FFFEFFDDFB1FF87EFABCEE97" box="[1208,1389,1984,2011]" pageId="9" pageNumber="226" value="1969-10-05">5 October 1969</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="5AE766E2FFFEFFDEFA37F87EFF09E9FC" lastPageId="10" lastPageNumber="227" pageId="9" pageNumber="226" type="male">3 males</specimenCount>
(12.8 × 11.0 mm, 9.8 × 
<quantity id="8B19008EFFFDFFDEFE47FF29FDE6E9FC" box="[480,567,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="10" pageNumber="227" unit="mm" value="8.8">8.8 mm</quantity>
, 9.7 × 
<quantity id="8B19008EFFFDFFDEFD27FF29FD09E9FC" box="[640,728,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="227" unit="mm" value="8.6">8.6 mm</quantity>
), 
<materialsCitation id="FC89A736FFFDFFDEFD4DFF29FABCE998" ID-GBIF-Occurrence="4161964325" collectingDate="2006-11-20" collectionCode="ZRC" collectorName="A. D. Tran" country="Malaysia" location="Langkawi Island" municipality="Sungei Lengara" pageId="10" pageNumber="227" specimenCode="ZRC 2022.0982" specimenCount="4" specimenCount-female="4">
<specimenCount id="5AE766E2FFFDFFDEFD4DFF29FC84E9FC" box="[746,853,151,177]" pageId="10" pageNumber="227" type="female">4 females</specimenCount>
(15.8 × 
<quantity id="8B19008EFFFDFFDEFC17FF29FBC4E9FC" box="[944,1045,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="10" pageNumber="227" unit="mm" value="13.8">13.8 mm</quantity>
, 13.4 × 
<quantity id="8B19008EFFFDFFDEFBCBFF29FB01E9FC" box="[1132,1232,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="10" pageNumber="227" unit="mm" value="11.3">11.3 mm</quantity>
, 9.3 × 8.0 mm, 7.7 × 
<quantity id="8B19008EFFFDFFDEFF0AFF05FED5E998" box="[173,260,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="227" unit="mm" value="6.6">6.6 mm</quantity>
) (
<specimenCode id="1C470510FFFDFFDEFEBDFF05FE05E998" box="[282,468,187,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="227">ZRC 2022.0982</specimenCode>
), near 
<collectingMunicipality id="AC3A3711FFFDFFDEFDBAFF05FD01E998" box="[541,720,187,213]" pageId="10" pageNumber="227">Sungei Lengara</collectingMunicipality>
, 
<location id="493EFBB0FFFDFFDEFD7DFF05FC49E998" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFDFFDEFD7DFF05FC49E998" box="[730,920,187,213]" country="Malaysia" municipality="Sungei Lengara" name="Langkawi Island" pageId="10" pageNumber="227">Langkawi Island</location>
, coll. 
<collectorName id="E114C8BDFFFDFFDEFC7FFF05FB98E998" box="[984,1097,187,213]" pageId="10" pageNumber="227">A.D. Tran</collectorName>
<emphasis id="7E957179FFFDFFDEFBE8FF05FB56E998" box="[1103,1159,187,213]" italics="true" pageId="10" pageNumber="227">et al.</emphasis>
, 
<date id="385F8BABFFFDFFDEFB33FF05FABCE998" box="[1172,1389,187,213]" pageId="10" pageNumber="227" value="2006-11-20">
<collectingDate id="281B7243FFFDFFDEFB33FF05FABCE998" box="[1172,1389,187,213]" pageId="10" pageNumber="227" value="2006-11-20">20 November 2006</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFFDFFDEFA28FF05FE1EE850" ID-GBIF-Occurrence="4161964335" collectingDate="2006-11-20" collectionCode="ZRC" collectorName="T. Naruse" country="Malaysia" location="Langkawi" municipality="Sungei Temurun" pageId="10" pageNumber="227" specimenCode="ZRC 2022.0983" specimenCount="2" specimenCount-male="2">
<specimenCount id="5AE766E2FFFDFFDEFA28FF05FF09E9B4" pageId="10" pageNumber="227" type="male">2 males</specimenCount>
(12.5 × 11.0 mm, 9.6 × 
<quantity id="8B19008EFFFDFFDEFDA6FF61FD8AE9B4" box="[513,603,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="10" pageNumber="227" unit="mm" value="8.3">8.3 mm</quantity>
) (
<specimenCode id="1C470510FFFDFFDEFDD2FF61FCE5E9B4" box="[629,820,223,249]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="227">ZRC 2022.0983</specimenCode>
), river mouth of 
<collectingMunicipality id="AC3A3711FFFDFFDEFC5BFF61FB6DE9B4" box="[1020,1212,223,249]" pageId="10" pageNumber="227">Sungei Temurun</collectingMunicipality>
, 
<location id="493EFBB0FFFDFFDEFB6DFF61FAECE9B4" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFDFFDEFB6DFF61FAECE9B4" box="[1226,1341,223,249]" country="Malaysia" municipality="Sungei Temurun" name="Langkawi" pageId="10" pageNumber="227">Langkawi</location>
, coll. 
<collectorName id="E114C8BDFFFDFFDEFA21FF61FF38E850" pageId="10" pageNumber="227">T. Naruse</collectorName>
, 
<date id="385F8BABFFFDFFDEFF53FEBDFE1EE850" box="[244,463,259,285]" pageId="10" pageNumber="227" value="2006-11-20">
<collectingDate id="281B7243FFFDFFDEFF53FEBDFE1EE850" box="[244,463,259,285]" pageId="10" pageNumber="227" value="2006-11-20">20 November 2006</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="5AE766E2FFFDFFDEFE56FEBDFD82E850" box="[497,595,259,285]" pageId="10" pageNumber="227" type="male">10 males</specimenCount>
(13.3 × 
<quantity id="8B19008EFFFDFFDEFD15FEBDFCC7E850" box="[690,790,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="10" pageNumber="227" unit="mm" value="11.4">11.4 mm</quantity>
, 13.2 × 11.0 mm, 8.7 × 
<quantity id="8B19008EFFFDFFDEFB87FEBDFBA9E850" box="[1056,1144,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="227" unit="mm" value="7.3">7.3 mm</quantity>
, 7.4 × 
<quantity id="8B19008EFFFDFFDEFB63FEBDFACFE850" box="[1220,1310,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="227" unit="mm" value="6.6">6.6 mm</quantity>
), 
<materialsCitation id="FC89A736FFFDFFDEFA97FEBDFF03E828" ID-GBIF-Occurrence="4161964338" collectingDate="1969-10-12" collectionCode="ZRC" collectorName="C. L. Soh" country="Malaysia" location="Straits" pageId="10" pageNumber="227" specimenCount="3" specimenCount-female="3" stateProvince="Johor">
<specimenCount id="5AE766E2FFFDFFDEFA97FEBDFA4DE850" box="[1328,1436,259,285]" pageId="10" pageNumber="227" type="female">3 females</specimenCount>
(11.5 × 
<quantity id="8B19008EFFFDFFDEFF57FE99FE98E80C" box="[240,329,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="227" unit="mm" value="9.7">9.7 mm</quantity>
, 6.7 × 
<quantity id="8B19008EFFFDFFDEFE3EFE98FE23E80C" box="[409,498,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="10" pageNumber="227" unit="mm" value="5.7">5.7 mm</quantity>
, 6.7 × 
<quantity id="8B19008EFFFDFFDEFDE5FE98FD4BE80C" box="[578,666,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="227" unit="mm" value="5.5">5.5 mm</quantity>
) (
<collectionCode id="2AF035AEFFFDFFDEFD16FE99FD38E80C" box="[689,745,295,321]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="227">ZRC</collectionCode>
1969.10.14.1–10), 
<collectingRegion id="8E256389FFFDFFDEFC60FE99FBC3E80C" box="[967,1042,295,321]" country="Malaysia" name="Johor" pageId="10" pageNumber="227">Johore</collectingRegion>
<location id="493EFBB0FFFDFFDEFBBDFE99FBB5E80C" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFDFFDEFBBDFE99FBB5E80C" box="[1050,1124,295,321]" country="Malaysia" name="Straits" pageId="10" pageNumber="227" stateProvince="Johor">Straits</location>
, coll. 
<collectorName id="E114C8BDFFFDFFDEFB0EFE99FAC0E80C" box="[1193,1297,295,321]" pageId="10" pageNumber="227">C.L. Soh</collectorName>
, 
<date id="385F8BABFFFDFFDEFAB9FE99FF03E828" pageId="10" pageNumber="227" value="1969-10-12">
<collectingDate id="281B7243FFFDFFDEFAB9FE99FF03E828" pageId="10" pageNumber="227" value="1969-10-12">12 October 1969</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFFDFFDEFF79FEF4FE13E8C4" ID-GBIF-Occurrence="4161964319" collectingDate="2018-12" collectorName="H. H. Tan" country="Indonesia" county="Mawali Bay" location="Manado" municipality="Pulau Lembeh" pageId="10" pageNumber="227" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
<emphasis id="7E957179FFFDFFDEFF79FEF4FE8DE829" bold="true" box="[222,348,330,356]" pageId="10" pageNumber="227">
<collectingCountry id="34F6EDFBFFFDFFDEFF79FEF4FE86E829" box="[222,343,330,356]" name="Indonesia" pageId="10" pageNumber="227">Indonesia</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFFDFFDEFEC4FEF5FE7FE828" box="[355,430,331,357]" pageId="10" pageNumber="227" type="male">1 male</specimenCount>
(11.2 × 
<quantity id="8B19008EFFFDFFDEFDA9FEF5FDB8E828" box="[526,617,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="227" unit="mm" value="9.7">9.7 mm</quantity>
), near NAD resort, 
<collectingCounty id="A53FD5E7FFFDFFDEFCE1FEF5FC00E828" box="[838,977,331,357]" pageId="10" pageNumber="227">Mawali Bay</collectingCounty>
, 
<collectingMunicipality id="AC3A3711FFFDFFDEFC7BFEF5FB53E828" box="[988,1154,331,357]" pageId="10" pageNumber="227">Pulau Lembeh</collectingMunicipality>
, 
<location id="493EFBB0FFFDFFDEFB29FEF5FB3CE828" LSID="urn:lsid:plazi:treatment:C4481C7DFFFEFFC5FF30FF29FBF7EB2C:493EFBB0FFFDFFDEFB29FEF5FB3CE828" box="[1166,1261,331,357]" country="Indonesia" county="Mawali Bay" municipality="Pulau Lembeh" name="Manado" pageId="10" pageNumber="227" stateProvince="Sulawesi Utara">Manado</location>
, 
<collectingRegion id="8E256389FFFDFFDEFB5FFEF5FAB1E828" box="[1272,1376,331,357]" country="Indonesia" name="Sulawesi Utara" pageId="10" pageNumber="227">Sulawesi</collectingRegion>
, coll. 
<collectorName id="E114C8BDFFFDFFDEFF30FED1FED1E8C4" box="[151,256,367,393]" pageId="10" pageNumber="227">H.H. Tan</collectorName>
, 
<date id="385F8BABFFFDFFDEFEABFED1FE13E8C4" box="[268,450,367,393]" pageId="10" pageNumber="227" value="2018-12">
<collectingDate id="281B7243FFFDFFDEFEABFED1FE13E8C4" box="[268,450,367,393]" pageId="10" pageNumber="227" value="2018-12">December 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="189EFDE3FFFDFFDEFF30F900FAD2EE53" ID-DOI="http://doi.org/10.5281/zenodo.8162500" ID-Zenodo-Dep="8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="10" pageNumber="227" startId="10.[151,250,1726,1750]" targetBox="[151,1435,417,1702]" targetPageId="10" targetType="figure">
<paragraph id="4C5EAD6BFFFDFFDEFF30F900FAD2EE53" blockId="10.[151,1436,1726,1822]" pageId="10" pageNumber="227">
<emphasis id="7E957179FFFDFFDEFF30F900FEC3EF9B" bold="true" box="[151,274,1726,1750]" pageId="10" pageNumber="227">FIGURE 5.</emphasis>
A, B, 
<taxonomicName id="8BE1D6E8FFFDFFDEFEF3F900FC84EF9B" authority="(A. Milne-Edwards, 1869)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[340,853,1726,1750]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="227" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFFDFFDEFEF3F900FD93EF9B" box="[340,578,1726,1750]" italics="true" pageId="10" pageNumber="227">Clistocoeloma villosum</emphasis>
(A. 
<bibRefCitation id="2870D09AFFFDFFDEFDC8F900FC9FEF9B" author="Milne-Edwards, A." box="[623,846,1726,1750]" pageId="10" pageNumber="227" pagination="25 - 31" refId="ref11446" refString="Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]" type="journal article" year="1869">Milne-Edwards, 1869</bibRefCitation>
)
</taxonomicName>
, lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, 
<taxonomicName id="8BE1D6E8FFFDFFDEFF74F95DFDDAEFB7" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[211,523,1762,1786]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="227" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFDFFDEFF74F95DFF35EFB7" box="[211,228,1763,1786]" italics="true" pageId="10" pageNumber="227">C</emphasis>
. 
<emphasis id="7E957179FFFDFFDEFF57F95DFE9AEFB7" box="[240,331,1763,1786]" italics="true" pageId="10" pageNumber="227">suvaense</emphasis>
<bibRefCitation id="2870D09AFFFDFFDEFEF6F95CFDDAEFB7" author="Edmondson, C. H." box="[337,523,1762,1786]" pageId="10" pageNumber="227" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, 
<taxonomicName id="8BE1D6E8FFFDFFDEFB82F95DFB50EFB7" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[1061,1153,1762,1786]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="227" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFFDFFDEFB82F95DFBE7EFB7" box="[1061,1078,1763,1786]" italics="true" pageId="10" pageNumber="227">C</emphasis>
. 
<emphasis id="7E957179FFFDFFDEFBE5F95CFB50EFB7" box="[1090,1153,1762,1786]" italics="true" pageId="10" pageNumber="227">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFFDFFDEFB20F95CFB11EFB7" bold="true" box="[1159,1216,1762,1786]" pageId="10" pageNumber="227">
<taxonomicNameLabel id="65A6CC02FFFDFFDEFB20F95CFB11EFB7" box="[1159,1216,1762,1786]" pageId="10" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore. A, C, E, right cheliped top view; B, D, F, right cheliped ventral view.
</paragraph>
</caption>
<caption id="189EFDE3FFFCFFDFFF30FBE4FAF5EC68" ID-DOI="http://doi.org/10.5281/zenodo.8162504" ID-Zenodo-Dep="8162504" httpUri="https://zenodo.org/record/8162504/files/figure.png" pageId="11" pageNumber="228" startId="11.[151,250,1114,1138]" targetBox="[151,1435,181,1089]" targetPageId="11" targetType="figure">
<paragraph id="4C5EAD6BFFFCFFDFFF30FBE4FAF5EC68" blockId="11.[151,1437,1114,1318]" pageId="11" pageNumber="228">
<emphasis id="7E957179FFFCFFDFFF30FBE4FEC4ED3F" bold="true" box="[151,277,1114,1138]" pageId="11" pageNumber="228">FIGURE 6.</emphasis>
P5 dorsal view. A–C, 
<taxonomicName id="8BE1D6E8FFFCFFDFFDA2FBE4FC12ED3F" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[517,963,1114,1138]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="228" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFCFFDFFDA2FBE4FD2AED3F" box="[517,763,1114,1138]" italics="true" pageId="11" pageNumber="228">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFFCFFDFFCA2FBE4FC12ED3F" author="Edmondson, C. H." box="[773,963,1114,1138]" pageId="11" pageNumber="228" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
; A, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji; B, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; C, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa; D, 
<taxonomicName id="8BE1D6E8FFFCFFDFFA22FBC0FE2EEDF7" authority="(A. Milne-Edwards, 1869)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="228" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFFCFFDFFA22FBC0FA47EDD8" box="[1413,1430,1150,1173]" italics="true" pageId="11" pageNumber="228">C</emphasis>
. 
<emphasis id="7E957179FFFCFFDFFF30FB1CFF3CEDF7" box="[151,237,1186,1210]" italics="true" pageId="11" pageNumber="228">villosum</emphasis>
(A. 
<bibRefCitation id="2870D09AFFFCFFDFFEBEFB1CFE29EDF7" author="Milne-Edwards, A." box="[281,504,1186,1210]" pageId="11" pageNumber="228" pagination="25 - 31" refId="ref11446" refString="Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]" type="journal article" year="1869">Milne-Edwards, 1869</bibRefCitation>
)
</taxonomicName>
, lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–K, 
<taxonomicName id="8BE1D6E8FFFCFFDFFB25FB1CFB0FEDF7" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[1154,1246,1186,1210]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="228" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFFCFFDFFB25FB1CFB42EDF4" box="[1154,1171,1186,1209]" italics="true" pageId="11" pageNumber="228">C</emphasis>
. 
<emphasis id="7E957179FFFCFFDFFB38FB1CFB0FEDF7" box="[1183,1246,1186,1210]" italics="true" pageId="11" pageNumber="228">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFFCFFDFFB43FB1CFACDEDF7" bold="true" box="[1252,1308,1186,1210]" pageId="11" pageNumber="228">
<taxonomicNameLabel id="65A6CC02FFFCFFDFFB43FB1CFACDEDF7" box="[1252,1308,1186,1210]" pageId="11" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; E, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; F, paratype male (15.0 × 13.0 mm) (ZRC 2002.0155) Singapore; G, male (16.0 × 13.5 mm) (ZRC 2022.0977), Vietnam; H, male (17.4 × 14.9 mm) (ZRC 2022.0973), Philippines; I, male (17.3 × 15.0 mm) (ZRC 2022.0994), Hong Kong; J, male (18.2 × 15.0 mm), Japan; K, male (16.2 × 14.2 mm) (WS898), Korea.
</paragraph>
</caption>
<subSubSection id="04FBFEE0FFFCFFDFFF60FAEAFD7BEFB7" pageId="11" pageNumber="228" type="diagnosis">
<paragraph id="4C5EAD6BFFFCFFDFFF60FAEAFD7BEFB7" blockId="11.[151,1437,1364,2003]" pageId="11" pageNumber="228">
<emphasis id="7E957179FFFCFFDFFF60FAEAFE93EC23" bold="true" box="[199,322,1364,1390]" pageId="11" pageNumber="228">Diagnosis.</emphasis>
Carapace slightly wider than long,almost square; postfrontal lobes weak, divided by weak longitudinal grooves; median pair slightly prominent, nearly equal in proportions. Inner orbital angle present, rounded, distinct, slight gap between inner orbital angle and frontal margin (
<figureCitation id="D4DAB1EEFFFCFFDFFC8DFA22FC52ECFB" box="[810,899,1436,1462]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="11" pageNumber="228">Fig. 7C</figureCitation>
). Anterolateral margin of carapace with distinct external orbital angle, second tooth blunt, weak or absent; posterolateral margin slightly convex (
<figureCitation id="D4DAB1EEFFFCFFDFFB7DFA7EFA8FEC97" box="[1242,1374,1472,1498]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="11" pageNumber="228">Figs. 2B, C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFFCFFDFFACDFA7EFA5EEC97" box="[1386,1423,1472,1498]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="11" pageNumber="228">7A</figureCitation>
). Chelipeds dorsal surface of dactylus of chela with between 20–31 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (
<figureCitation id="D4DAB1EEFFFCFFDFFCF1F9B6FC7FEF6E" box="[854,942,1544,1571]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="11" pageNumber="228">Fig. 5E</figureCitation>
); presence of single longitudinal pectinated ridge with between 28–45 chitinous comb-like tubercles on dorsal surface of chela; row of 10–20 granules clustered behind pectinated ridge on chela; inner surface of palm granulated, vertical rows of 6–10 granules in adult male (
<figureCitation id="D4DAB1EEFFFCFFDFFF38F9CAFF26EFC2" box="[159,247,1652,1679]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="11" pageNumber="228">Fig. 5F</figureCitation>
). P5 merus relatively short, wide, length approximately 0.36–0.43 times width (
<figureCitation id="D4DAB1EEFFFCFFDFFB2BF9CAFAD9EFC3" box="[1164,1288,1652,1678]" captionStart="FIGURE 6" captionStartId="11.[151,250,1114,1138]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-18@11.[151,1435,181,1089]" captionTargetPageId="11" captionText="FIGURE 6. P5 dorsal view. A–C, Clistocoeloma suvaense Edmondson, 1951; A, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji; B, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; C, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa; D, C. villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–K, C. nobile n. sp.; E, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; F, paratype male (15.0 × 13.0 mm) (ZRC 2002.0155) Singapore; G, male (16.0 × 13.5 mm) (ZRC 2022.0977), Vietnam; H, male (17.4 × 14.9 mm) (ZRC 2022.0973), Philippines; I, male (17.3 × 15.0 mm) (ZRC 2022.0994), Hong Kong; J, male (18.2 × 15.0 mm), Japan; K, male (16.2 × 14.2 mm) (WS898), Korea." figureDoi="http://doi.org/10.5281/zenodo.8162504" httpUri="https://zenodo.org/record/8162504/files/figure.png" pageId="11" pageNumber="228">Fig. 6E–K</figureCitation>
). G1 slightly curved, distal region slightly wide; chitinous tip narrow, short; tip with dense tufts of setae, single row of plumose setae on exterior lateral margin; chitinous tip visible when denuded (
<figureCitation id="D4DAB1EEFFFCFFDFFC1AF902FB9AEF9B" box="[957,1099,1724,1750]" captionStart="FIGURE 11" captionStartId="16.[151,250,1172,1196]" captionTargetBox="[153,1434,183,1146]" captionTargetId="figure-32@16.[151,1436,181,1148]" captionTargetPageId="16" captionText="FIGURE 11. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore, left G1. A, dorsal view; B, left lateral view; C, ventral view; D, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162516" httpUri="https://zenodo.org/record/8162516/files/figure.png" pageId="11" pageNumber="228">Fig. 11A–D</figureCitation>
). Female vulvae with central operculum, round, slightly protruded (
<figureCitation id="D4DAB1EEFFFCFFDFFDE4F95EFD4FEFB7" box="[579,670,1760,1786]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="11" pageNumber="228">Fig. 8D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFFCFFD9FF60F8BAFDF6EEA1" lastPageId="13" lastPageNumber="230" pageId="11" pageNumber="228" type="description">
<paragraph id="4C5EAD6BFFFCFFDFFF60F8BAFC8EEE9F" blockId="11.[151,1437,1364,2003]" pageId="11" pageNumber="228">
<emphasis id="7E957179FFFCFFDFFF60F8BAFE88EE53" bold="true" box="[199,345,1796,1822]" pageId="11" pageNumber="228">Description.</emphasis>
Carapace slightly wider than long, almost square; covered with dense coat of setae, including small tufts of setae resembling tubercles; carapace margin fringed with dense, short setae; carapace smooth when denuded. Postfrontal lobes weak, divided by weak longitudinal grooves; median pair slightly prominent, nearly equal in proportions. Inner orbital angle present, rounded, distinct, slight gap between inner orbital angle and frontal margin (
<figureCitation id="D4DAB1EEFFFCFFDFFF52F82AFE9FEEE3" box="[245,334,1940,1966]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="11" pageNumber="228">Fig. 7C</figureCitation>
). Anterolateral margin of carapace with distinct external orbital angle, second tooth blunt, weak or absent; posterolateral margin slightly convex (
<figureCitation id="D4DAB1EEFFFCFFDFFD39F806FCF0EE9F" box="[670,801,1976,2002]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="11" pageNumber="228">Figs. 2B, C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFFCFFDFFC8BF806FC83EE9F" box="[812,850,1976,2002]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="11" pageNumber="228">7A</figureCitation>
).
</paragraph>
<caption id="189EFDE3FFFBFFD8FF30F829FD76EE9E" ID-DOI="http://doi.org/10.5281/zenodo.8162506" ID-Zenodo-Dep="8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="12" pageNumber="229" startId="12.[151,250,1943,1967]" targetBox="[228,1359,181,1920]" targetPageId="12" targetType="figure">
<paragraph id="4C5EAD6BFFFBFFD8FF30F829FD76EE9E" blockId="12.[151,1436,1943,2004]" pageId="12" pageNumber="229">
<emphasis id="7E957179FFFBFFD8FF30F829FEC3EEFD" bold="true" box="[151,274,1943,1968]" pageId="12" pageNumber="229">FIGURE 7.</emphasis>
<taxonomicName id="8BE1D6E8FFFBFFD8FEB0F826FE3FEEFD" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[279,494,1944,1968]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="229" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFFBFFD8FEB0F826FE3FEEFD" box="[279,494,1944,1968]" italics="true" pageId="12" pageNumber="229">Clistocoeloma nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFFBFFD8FE53F826FDFDEEFD" bold="true" box="[500,556,1944,1968]" pageId="12" pageNumber="229">
<taxonomicNameLabel id="65A6CC02FFFBFFD8FE53F826FDFDEEFD" box="[500,556,1944,1968]" pageId="12" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace.
</paragraph>
</caption>
<caption id="189EFDE3FFFAFFD9FF30FB95FCB2EDC1" ID-DOI="http://doi.org/10.5281/zenodo.8162510" ID-Zenodo-Dep="8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="13" pageNumber="230" startId="13.[151,250,1067,1091]" targetBox="[151,1435,181,1044]" targetPageId="13" targetType="figure">
<paragraph id="4C5EAD6BFFFAFFD9FF30FB95FCB2EDC1" blockId="13.[151,1437,1067,1164]" pageId="13" pageNumber="230">
<emphasis id="7E957179FFFAFFD9FF30FB95FEC3ED09" bold="true" box="[151,274,1067,1092]" pageId="13" pageNumber="230">FIGURE 8.</emphasis>
Female pleon, thoracic sternums and vulvae. A, B, 
<taxonomicName id="8BE1D6E8FFFAFFD9FCBEFB92FB1CED09" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[793,1229,1068,1092]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="230" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFFAFFD9FCBEFB92FBDCED0E" box="[793,1037,1068,1092]" italics="true" pageId="13" pageNumber="230">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFFAFFD9FBB4FB92FB1CED09" author="Edmondson, C. H." box="[1043,1229,1068,1092]" pageId="13" pageNumber="230" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, 
<taxonomicName id="8BE1D6E8FFFAFFD9FE53FBEEFD1BED25" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[500,714,1104,1128]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="230" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFFAFFD9FE53FBEEFD1BED25" box="[500,714,1104,1128]" italics="true" pageId="13" pageNumber="230">Clistocoeloma nobile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65A6CC02FFFAFFD9FD77FBEEFCD6ED2A" box="[720,775,1104,1127]" pageId="13" pageNumber="230" rank="species">n. sp.</taxonomicNameLabel>
, paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae.
</paragraph>
</caption>
<paragraph id="4C5EAD6BFFFAFFD9FF60FB04FDF3EDB5" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
Antenna short. Antennules folding transversely. Third maxilliped with merus relatively rounded. Epistomial margin relatively narrow (
<figureCitation id="D4DAB1EEFFFAFFD9FE1BFB60FDC4EDB5" box="[444,533,1246,1272]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="13" pageNumber="230">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFFAFFD9FF60FABCFED3ECB8" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
Chelipeds subequal in size; large, robust in male; outer surface of palm smooth when denuded; dorsal surface of carpus covered with small tufts of setae resembling tubercles, smooth when denuded.; dorsal surface of dactylus of chela with between 20–31 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (
<figureCitation id="D4DAB1EEFFFAFFD9FEE2FAD0FE4BECC4" box="[325,410,1390,1417]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="13" pageNumber="230">Fig. 5E</figureCitation>
); absent or weak granules in female and juveniles; presence of single longitudinal pectinated ridge with between 28–45 chitinous comb-like tubercles on dorsal surface of chela; row of 10–20 granules clustered behind pectinated ridge on chela; inner surface of palm granulated, vertical rows of 6–10 granules in adult male (
<figureCitation id="D4DAB1EEFFFAFFD9FF38FA64FF24ECB8" box="[159,245,1498,1525]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="13" pageNumber="230">Fig. 5F</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFFAFFD9FF60FA40FD01EF2D" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
P2–P5 covered with dense coat of setae, with small tufts of setae resembling tubercles, longer setae on dorsal and ventral margins except distal portion of dactylus (
<figureCitation id="D4DAB1EEFFFAFFD9FCAEF99CFC43EF71" box="[777,914,1570,1596]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="13" pageNumber="230">Figs. 2B, C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFFAFFD9FC07F99CFC17EF71" box="[928,966,1570,1596]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="13" pageNumber="230">7A</figureCitation>
); P5 merus relatively short, wide, length approximately 0.36–0.43 times width (
<figureCitation id="D4DAB1EEFFFAFFD9FDEDF9F8FD15EF2D" box="[586,708,1606,1632]" captionStart="FIGURE 6" captionStartId="11.[151,250,1114,1138]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-18@11.[151,1435,181,1089]" captionTargetPageId="11" captionText="FIGURE 6. P5 dorsal view. A–C, Clistocoeloma suvaense Edmondson, 1951; A, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji; B, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; C, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa; D, C. villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–K, C. nobile n. sp.; E, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; F, paratype male (15.0 × 13.0 mm) (ZRC 2002.0155) Singapore; G, male (16.0 × 13.5 mm) (ZRC 2022.0977), Vietnam; H, male (17.4 × 14.9 mm) (ZRC 2022.0973), Philippines; I, male (17.3 × 15.0 mm) (ZRC 2022.0994), Hong Kong; J, male (18.2 × 15.0 mm), Japan; K, male (16.2 × 14.2 mm) (WS898), Korea." figureDoi="http://doi.org/10.5281/zenodo.8162504" httpUri="https://zenodo.org/record/8162504/files/figure.png" pageId="13" pageNumber="230">Fig. 6E–K</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFFAFFD9FF60F9D4FBAEEFBC" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
Male thoracic sternum sparsely covered with short setae, smooth when denuded; sternites 1 and 2 fused, with suture between fused sternites 3 and 4; sternopleonal cavity extends nearly to edge of sternites 1 and 2. Male pleon wide and stout, long setae fringed pleon margin; somite 3 widest; somite 6 with curved edge; telson wide, dome-shaped (
<figureCitation id="D4DAB1EEFFFAFFD9FF53F968FE9DEFBD" box="[244,332,1750,1776]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="13" pageNumber="230">Fig. 7B</figureCitation>
); male pleonal locking mechanism absent, without tubercle on sternite 5.
</paragraph>
<paragraph id="4C5EAD6BFFFAFFD9FF60F944FDD4EE11" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
G1 slightly curved, distal region slightly wide; chitinous tip narrow, short; tip with dense tufts of setae, single row of plumose setae on exterior lateral margin; chitinous tip visible when denuded (
<figureCitation id="D4DAB1EEFFFAFFD9FBF7F8A0FB0AEE75" box="[1104,1243,1822,1848]" captionStart="FIGURE 11" captionStartId="16.[151,250,1172,1196]" captionTargetBox="[153,1434,183,1146]" captionTargetId="figure-32@16.[151,1436,181,1148]" captionTargetPageId="16" captionText="FIGURE 11. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore, left G1. A, dorsal view; B, left lateral view; C, ventral view; D, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162516" httpUri="https://zenodo.org/record/8162516/files/figure.png" pageId="13" pageNumber="230">Fig. 11A–D</figureCitation>
). G2 shorter than G1, slightly curved, tip rounded.
</paragraph>
<paragraph id="4C5EAD6BFFFAFFD9FF60F8D8FBAEEEE9" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
Female pleon rounded, relatively wider than long; telson dome-shaped (
<figureCitation id="D4DAB1EEFFFAFFD9FC7DF8D8FBE1EECD" box="[986,1072,1894,1920]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="13" pageNumber="230">Fig. 8C</figureCitation>
). Vulvae with central operculum, round, slightly protruded; sternal vulvar cover slightly raised above operculum (
<figureCitation id="D4DAB1EEFFFAFFD9FBBFF834FBA2EEE9" box="[1048,1139,1930,1956]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="13" pageNumber="230">Fig. 8D</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFFAFFD9FF60F810FDF6EEA1" blockId="13.[151,1437,1210,2028]" pageId="13" pageNumber="230">
<emphasis id="7E957179FFFAFFD9FF60F810FE8EEE85" bold="true" box="[199,351,1966,1992]" pageId="13" pageNumber="230">Colouration.</emphasis>
Chelipeds with fingers yellowish white and purple to purplish red palm (
<figureCitation id="D4DAB1EEFFFAFFD9FB23F810FB2AEE85" box="[1156,1275,1966,1992]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="13" pageNumber="230">Fig. 2B, C</figureCitation>
). In life, entire crab covered in a thin layer of mud.
</paragraph>
</subSubSection>
<caption id="189EFDE3FFF9FFDAFF30F829FC42EEBA" ID-DOI="http://doi.org/10.5281/zenodo.8162512" ID-Zenodo-Dep="8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="14" pageNumber="231" startId="14.[151,250,1943,1967]" targetBox="[212,1375,183,1917]" targetPageId="14" targetType="figure">
<paragraph id="4C5EAD6BFFF9FFDAFF30F829FC42EEBA" blockId="14.[151,1436,1943,2040]" pageId="14" pageNumber="231">
<emphasis id="7E957179FFF9FFDAFF30F829FEC3EEE2" bold="true" box="[151,274,1943,1967]" pageId="14" pageNumber="231">FIGURE 9.</emphasis>
Left G1. A–D, 
<taxonomicName id="8BE1D6E8FFF9FFDAFE0AF826FC7DEEE2" authority="(A. Milne-Edwards, 1869)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[429,940,1943,1968]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="231" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF9FFDAFE0AF826FD4BEEFD" box="[429,666,1944,1968]" italics="true" pageId="14" pageNumber="231">Clistocoeloma villosum</emphasis>
(A. 
<bibRefCitation id="2870D09AFFF9FFDAFD61F826FC74EEE2" author="Milne-Edwards, A." box="[710,933,1943,1968]" pageId="14" pageNumber="231" pagination="25 - 31" refId="ref11446" refString="Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]" type="journal article" year="1869">Milne-Edwards, 1869</bibRefCitation>
)
</taxonomicName>
, lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, 
<taxonomicName id="8BE1D6E8FFF9FFDAFE82F802FD8EEE9E" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[293,607,1979,2004]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="231" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF9FFDAFE82F802FEE7EE9E" box="[293,310,1980,2003]" italics="true" pageId="14" pageNumber="231">C</emphasis>
. 
<emphasis id="7E957179FFF9FFDAFEE4F802FE4FEE9E" box="[323,414,1980,2003]" italics="true" pageId="14" pageNumber="231">suvaense</emphasis>
<bibRefCitation id="2870D09AFFF9FFDAFE03F802FD8EEE9E" author="Edmondson, C. H." box="[420,607,1979,2004]" pageId="14" pageNumber="231" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="189EFDE3FFF8FFDBFF30F829FCDEEEBA" ID-DOI="http://doi.org/10.5281/zenodo.8162514" ID-Zenodo-Dep="8162514" httpUri="https://zenodo.org/record/8162514/files/figure.png" pageId="15" pageNumber="232" startId="15.[151,250,1943,1967]" targetBox="[216,1372,183,1917]" targetPageId="15" targetType="figure">
<paragraph id="4C5EAD6BFFF8FFDBFF30F829FCDEEEBA" blockId="15.[151,1436,1943,2040]" pageId="15" pageNumber="232">
<emphasis id="7E957179FFF8FFDBFF30F829FEF1EEE2" bold="true" box="[151,288,1943,1967]" pageId="15" pageNumber="232">FIGURE 10.</emphasis>
Left G1. A–D, 
<taxonomicName id="8BE1D6E8FFF8FFDBFE66F826FCA9EEE2" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[449,888,1943,1968]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF8FFDBFE66F826FD64EEE2" box="[449,693,1944,1968]" italics="true" pageId="15" pageNumber="232">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFF8FFDBFD1BF826FCA9EEE2" author="Edmondson, C. H." box="[700,888,1943,1968]" pageId="15" pageNumber="232" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E–H, 
<taxonomicName id="8BE1D6E8FFF8FFDBFF30F802FE00EE9E" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[151,465,1979,2004]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF8FFDBFF30F802FF79EE9E" box="[151,168,1980,2003]" italics="true" pageId="15" pageNumber="232">C</emphasis>
. 
<emphasis id="7E957179FFF8FFDBFF12F802FEC1EE9E" box="[181,272,1980,2003]" italics="true" pageId="15" pageNumber="232">suvaense</emphasis>
<bibRefCitation id="2870D09AFFF8FFDBFEB0F802FE00EE9E" author="Edmondson, C. H." box="[279,465,1979,2004]" pageId="15" pageNumber="232" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa, A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="189EFDE3FFE7FFC4FF30FB2AFC0BED82" ID-DOI="http://doi.org/10.5281/zenodo.8162516" ID-Zenodo-Dep="8162516" httpUri="https://zenodo.org/record/8162516/files/figure.png" pageId="16" pageNumber="233" startId="16.[151,250,1172,1196]" targetBox="[153,1434,183,1146]" targetPageId="16" targetType="figure">
<paragraph id="4C5EAD6BFFE7FFC4FF30FB2AFC0BED82" blockId="16.[151,1436,1172,1232]" pageId="16" pageNumber="233">
<emphasis id="7E957179FFE7FFC4FF30FB2AFEF1EDE1" bold="true" box="[151,288,1172,1196]" pageId="16" pageNumber="233">FIGURE 11.</emphasis>
<taxonomicName id="8BE1D6E8FFE7FFC4FE8FFB2AFDD0EDE1" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[296,513,1172,1196]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FE8FFB2AFDD0EDE1" box="[296,513,1172,1196]" italics="true" pageId="16" pageNumber="233">Clistocoeloma nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FDAEFB2AFD95EDE1" bold="true" box="[521,580,1172,1196]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FDAEFB2AFD95EDE1" box="[521,580,1172,1196]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore, left G1. A, dorsal view; B, left lateral view; C, ventral view; D, right lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="04FBFEE0FFE7FFC4FF60FB40FDDAEC71" pageId="16" pageNumber="233" type="etymology">
<paragraph id="4C5EAD6BFFE7FFC4FF60FB40FDDAEC71" blockId="16.[151,1437,1278,2025]" pageId="16" pageNumber="233">
<emphasis id="7E957179FFE7FFC4FF60FB40FE9FEC55" bold="true" box="[199,334,1278,1304]" pageId="16" pageNumber="233">Etymology.</emphasis>
The term, “ 
<taxonomicName id="8BE1D6E8FFE7FFC4FE76FB40FDC6EC55" authorityName="Ng &amp; Ng" authorityYear="2023" box="[465,535,1278,1304]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile">
<emphasis id="7E957179FFE7FFC4FE76FB40FDC6EC55" box="[465,535,1278,1304]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
” alludes to the distinctive colour of the chela of the species; purple being a colour normally associated with royalty.
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFE7FFC5FF60FAF8FCE9E8B8" lastPageId="17" lastPageNumber="234" pageId="16" pageNumber="233" type="discussion">
<paragraph id="4C5EAD6BFFE7FFC4FF60FAF8FEB6EE79" blockId="16.[151,1437,1278,2025]" pageId="16" pageNumber="233">
<emphasis id="7E957179FFE7FFC4FF60FAF8FEEAEC2D" bold="true" box="[199,315,1350,1376]" pageId="16" pageNumber="233">Remarks.</emphasis>
<taxonomicName id="8BE1D6E8FFE7FFC4FEE6FAF8FDFEEC2D" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[321,559,1350,1376]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FEE6FAF8FDFEEC2D" box="[321,559,1350,1376]" italics="true" pageId="16" pageNumber="233">Clistocoeloma nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FD91FAF8FDA5EC2D" bold="true" box="[566,628,1350,1376]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FD91FAF8FDA5EC2D" box="[566,628,1350,1376]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a mix of diagnostic characters of 
<taxonomicName id="8BE1D6E8FFE7FFC4FBB1FAF9FB44EC2D" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[1046,1173,1350,1377]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE7FFC4FBB1FAF9FBF8EC2C" box="[1046,1065,1351,1377]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FB91FAF8FB44EC2D" box="[1078,1173,1350,1376]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BE1D6E8FFE7FFC4FB6DFAF9FA9EEC2D" authorityName="Edmondson" authorityYear="1951" box="[1226,1359,1351,1377]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FB6DFAF9FB0CEC2C" box="[1226,1245,1351,1377]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FB4DFAF9FA9EEC2D" box="[1258,1359,1351,1376]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
. In the form of the carapace, 
<taxonomicName id="8BE1D6E8FFE7FFC4FE20FAD5FE3CECC9" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[391,493,1386,1413]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FE20FAD5FE4BECC8" box="[391,410,1387,1413]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FE00FAD4FE3CECC9" box="[423,493,1386,1412]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FE54FAD4FDE1ECC9" bold="true" box="[499,560,1386,1412]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FE54FAD4FDE1ECC9" box="[499,560,1386,1412]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a weak external orbital angle on the anterolateral margin of carapace (
<figureCitation id="D4DAB1EEFFE7FFC4FAC3FAD4FF0DECE5" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="16" pageNumber="233">Figs. 2B, C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFE7FFC4FF4FFA30FEDCECE5" box="[232,269,1422,1448]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[228,1359,181,1920]" captionTargetId="figure-32@12.[228,1359,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162506" httpUri="https://zenodo.org/record/8162506/files/figure.png" pageId="16" pageNumber="233">7A</figureCitation>
), and this is intermediate in form between 
<taxonomicName id="8BE1D6E8FFE7FFC4FD56FA31FCA0ECE5" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[753,881,1422,1449]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE7FFC4FD56FA31FCD5ECE4" box="[753,772,1423,1449]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FCB5FA30FCA0ECE5" box="[786,881,1422,1448]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
(which has no external orbital angle visible; 
<figureCitation id="D4DAB1EEFFE7FFC4FAD7FA30FF68EC81" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[164,1423,181,1773]" captionTargetId="figure-32@2.[164,1423,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162488" httpUri="https://zenodo.org/record/8162488/files/figure.png" pageId="16" pageNumber="233">Fig. 1A</figureCitation>
) and 
<taxonomicName id="8BE1D6E8FFE7FFC4FF50FA0DFEADEC81" authorityName="Edmondson" authorityYear="1951" box="[247,380,1459,1485]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FF50FA0DFEDBEC80" box="[247,266,1459,1485]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FEB0FA0DFEADEC81" box="[279,380,1459,1484]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
(which has a distinct external orbital angle; 
<figureCitation id="D4DAB1EEFFE7FFC4FCC3FA0CFC17EC81" box="[868,966,1458,1484]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="16" pageNumber="233">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFE7FFC4FC76FA0CFC24EC81" box="[977,1013,1458,1484]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="16" pageNumber="233">3A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFE7FFC4FBA7FA0CFBF7EC81" box="[1024,1062,1458,1484]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="16" pageNumber="233">4A</figureCitation>
). This character is very consistent in the large series of specimens of 
<taxonomicName id="8BE1D6E8FFE7FFC4FDBAFA69FD55ECBD" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[541,644,1494,1521]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FDBAFA69FDE1ECBC" box="[541,560,1495,1521]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FD99FA68FD55ECBD" box="[574,644,1494,1520]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FD2CFA68FD1BECBD" bold="true" box="[651,714,1494,1520]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FD2CFA68FD1BECBD" box="[651,714,1494,1520]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
examined. With regards to the armature on the dorsal margin of the dactylus of the male chela, 
<taxonomicName id="8BE1D6E8FFE7FFC4FE55FA45FD88EF59" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[498,601,1530,1557]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FE55FA45FDD4EF58" box="[498,517,1531,1557]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FDB4FA44FD88EF59" box="[531,601,1530,1556]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FDC7FA44FD4EEF59" bold="true" box="[608,671,1530,1556]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FDC7FA44FD4EEF59" box="[608,671,1530,1556]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has 20–31 granules (
<figureCitation id="D4DAB1EEFFE7FFC4FC2AFA44FC32EF58" box="[909,995,1530,1557]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="16" pageNumber="233">Fig. 5E</figureCitation>
), in contrast to only16 granules present in 
<taxonomicName id="8BE1D6E8FFE7FFC4FF11F9A1FEE9EF75" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[182,312,1566,1593]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE7FFC4FF11F9A1FF18EF74" box="[182,201,1567,1593]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FF7EF9A0FEE9EF75" box="[217,312,1566,1592]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
(
<figureCitation id="D4DAB1EEFFE7FFC4FEEDF9A0FE77EF74" box="[330,422,1566,1593]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="16" pageNumber="233">Fig. 5A</figureCitation>
). The number of granules in 
<taxonomicName id="8BE1D6E8FFE7FFC4FD50F9A1FCAEEF75" authorityName="Edmondson" authorityYear="1951" box="[759,895,1567,1593]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FD50F9A1FCDBEF74" box="[759,778,1567,1593]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FCBDF9A1FCAEEF75" box="[794,895,1567,1592]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
(22–27; 
<figureCitation id="D4DAB1EEFFE7FFC4FC4EF9A0FB9BEF75" box="[1001,1098,1566,1592]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="16" pageNumber="233">Figs. 3E</figureCitation>
, 
<figureCitation id="D4DAB1EEFFE7FFC4FBFFF9A0FBADEF74" box="[1112,1148,1566,1593]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="16" pageNumber="233">5C</figureCitation>
), however, overlaps with that of 
<taxonomicName id="8BE1D6E8FFE7FFC4FF4EF9FDFE80EF11" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[233,337,1602,1629]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FF4EF9FDFF2DEF10" box="[233,252,1603,1629]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FEACF9FCFE80EF11" box="[267,337,1602,1628]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FEFEF9FCFE48EF11" bold="true" box="[345,409,1602,1628]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FEFEF9FCFE48EF11" box="[345,409,1602,1628]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The G1of 
<taxonomicName id="8BE1D6E8FFE7FFC4FDB1F9FDFD4CEF11" authorityName="Edmondson" authorityYear="1951" box="[534,669,1603,1629]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FDB1F9FDFDF8EF10" box="[534,553,1603,1629]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FD9FF9FDFD4CEF11" box="[568,669,1603,1628]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
is also very similar to that of 
<taxonomicName id="8BE1D6E8FFE7FFC4FC53F9FDFB8DEF11" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[1012,1116,1602,1629]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FC53F9FDFBD6EF10" box="[1012,1031,1603,1629]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FBB1F9FCFB8DEF11" box="[1046,1116,1602,1628]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FBC3F9FCFB75EF11" bold="true" box="[1124,1188,1602,1628]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FBC3F9FCFB75EF11" box="[1124,1188,1602,1628]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, with the lateral view appearing stout with a distinct angle before chitinous tip in 
<taxonomicName id="8BE1D6E8FFE7FFC4FCE6F9D9FC18EFCD" authorityName="Edmondson" authorityYear="1951" box="[833,969,1639,1665]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FCE6F9D9FC85EFCC" box="[833,852,1639,1665]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FCC3F9D9FC18EFCD" box="[868,969,1639,1664]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
(
<figureCitation id="D4DAB1EEFFE7FFC4FC7DF9D8FBE8EFCD" box="[986,1081,1638,1664]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="16" pageNumber="233">Figs. 9F</figureCitation>
, 
<figureCitation id="D4DAB1EEFFE7FFC4FBE1F9D8FB44EFCD" box="[1094,1173,1638,1664]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[216,1372,183,1917]" captionTargetId="figure-18@15.[214,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 10. Left G1. A–D, Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E–H, C. suvaense Edmondson, 1951, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa, A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162514" httpUri="https://zenodo.org/record/8162514/files/figure.png" pageId="16" pageNumber="233">10B, F</figureCitation>
) while the lateral view appearing narrower with a weak angle visibile before the chitinous tip in 
<taxonomicName id="8BE1D6E8FFE7FFC4FC1BF935FBF0EFE9" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[956,1057,1674,1701]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FC1BF935FC1EEFE8" box="[956,975,1675,1701]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FC7CF934FBF0EFE9" box="[987,1057,1674,1700]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FB80F934FBB5EFE9" bold="true" box="[1063,1124,1674,1700]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FB80F934FBB5EFE9" box="[1063,1124,1674,1700]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D4DAB1EEFFE7FFC4FBD4F934FB06EFE9" box="[1139,1239,1674,1700]" captionStart="FIGURE 11" captionStartId="16.[151,250,1172,1196]" captionTargetBox="[153,1434,183,1146]" captionTargetId="figure-32@16.[151,1436,181,1148]" captionTargetPageId="16" captionText="FIGURE 11. Clistocoeloma nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore, left G1. A, dorsal view; B, left lateral view; C, ventral view; D, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162516" httpUri="https://zenodo.org/record/8162516/files/figure.png" pageId="16" pageNumber="233">Fig. 11B</figureCitation>
). The lateral view of the G1 of 
<taxonomicName id="8BE1D6E8FFE7FFC4FE8AF911FE7FEF85" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[301,430,1710,1737]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE7FFC4FE8AF911FE91EF84" box="[301,320,1711,1737]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FEE8F910FE7FEF85" box="[335,430,1710,1736]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
, on the other hand, appears narrow with no clear angle before chitinous tip (
<figureCitation id="D4DAB1EEFFE7FFC4FABBF910FAA1EF85" box="[1308,1392,1710,1736]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="16" pageNumber="233">Fig 9B</figureCitation>
). In addition, 
<taxonomicName id="8BE1D6E8FFE7FFC4FEA6F96DFEB6EFA1" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[257,359,1746,1773]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FEA6F96DFEC5EFA0" box="[257,276,1747,1773]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FE86F96CFEB6EFA1" box="[289,359,1746,1772]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FEC9F96CFE7CEFA1" bold="true" box="[366,429,1746,1772]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FEC9F96CFE7CEFA1" box="[366,429,1746,1772]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can also be separated from 
<taxonomicName id="8BE1D6E8FFE7FFC4FD42F96DFCBBEFA1" authorityName="Edmondson" authorityYear="1951" box="[741,874,1747,1773]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FD42F96DFD29EFA0" box="[741,760,1747,1773]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FCA2F96DFCBBEFA1" box="[773,874,1747,1772]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
in having a rounded and slightly protruded vulvar cover (
<figureCitation id="D4DAB1EEFFE7FFC4FF42F948FEEEEE5D" box="[229,319,1782,1808]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="16" pageNumber="233">Fig. 8D</figureCitation>
); which is ovate and less protruded vulvar cover in 
<taxonomicName id="8BE1D6E8FFE7FFC4FC27F949FBD7EE5D" authorityName="Edmondson" authorityYear="1951" box="[896,1030,1783,1809]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FC27F949FC42EE5C" box="[896,915,1783,1809]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FC06F949FBD7EE5D" box="[929,1030,1783,1808]" italics="true" pageId="16" pageNumber="233">suvaense</emphasis>
</taxonomicName>
(
<figureCitation id="D4DAB1EEFFE7FFC4FBB2F948FBBCEE5D" box="[1045,1133,1782,1808]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="16" pageNumber="233">Fig. 8B</figureCitation>
). Females of 
<taxonomicName id="8BE1D6E8FFE7FFC4FAA3F949FA52EE5D" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[1284,1411,1782,1809]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" sensu="stricto" species="villosum">
<emphasis id="7E957179FFE7FFC4FAA3F949FAC6EE5C" box="[1284,1303,1783,1809]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FA83F948FA52EE5D" box="[1316,1411,1782,1808]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65A6CC02FFE7FFC4FA2DF949FF68EE78" pageId="16" pageNumber="233" sensu="stricto">s. str.</taxonomicNameLabel>
are not known.
</paragraph>
<paragraph id="4C5EAD6BFFE7FFC5FF60F880FC0FE9B4" blockId="16.[151,1437,1278,2025]" lastBlockId="17.[151,1437,151,609]" lastPageId="17" lastPageNumber="234" pageId="16" pageNumber="233">
On the basis of these characters, it is clear that most of the records of “ 
<taxonomicName id="8BE1D6E8FFE7FFC4FC53F881FBA4EE15" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[1012,1141,1854,1881]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE7FFC4FC53F881FBD6EE14" box="[1012,1031,1855,1881]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FBB1F880FBA4EE15" box="[1046,1141,1854,1880]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
” from the IWP, including 
<collectingCountry id="34F6EDFBFFE7FFC4FF30F8DCFEDAEE31" box="[151,267,1890,1916]" name="Singapore" pageId="16" pageNumber="233">Singapore</collectingCountry>
, belong to 
<taxonomicName id="8BE1D6E8FFE7FFC4FE22F8DDFE3DEE31" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[389,492,1890,1917]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FE22F8DDFE49EE30" box="[389,408,1891,1917]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FE01F8DCFE3DEE31" box="[422,492,1890,1916]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FE55F8DCFDE1EE31" bold="true" box="[498,560,1890,1916]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FE55F8DCFDE1EE31" box="[498,560,1890,1916]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
instead. For the record by 
<bibRefCitation id="2870D09AFFE7FFC4FCFDF8DCFB87EE31" author="Tesch, J. J." box="[858,1110,1890,1916]" pageId="16" pageNumber="233" pagination="127 - 260" refId="ref12609" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, Leiden, 3 (2 - 3), 127 - 260, pls. 15 - 17. https: // doi. org / 10.5962 / bhl. title. 10640" type="journal article" year="1917">Tesch (1917: 208–210</bibRefCitation>
, pl. 17 fig. 2), his description and figures of the female specimen match 
<taxonomicName id="8BE1D6E8FFE7FFC4FDD5F839FD09EEED" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[626,728,1926,1953]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE7FFC4FDD5F839FD54EEEC" box="[626,645,1927,1953]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FD35F838FD09EEED" box="[658,728,1926,1952]" italics="true" pageId="16" pageNumber="233">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE7FFC4FD78F838FCCFEEED" bold="true" box="[735,798,1926,1952]" pageId="16" pageNumber="233">
<taxonomicNameLabel id="65A6CC02FFE7FFC4FD78F838FCCFEEED" box="[735,798,1926,1952]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. 
<bibRefCitation id="2870D09AFFE7FFC4FC8BF838FC6FEEED" author="Tesch, J. J." box="[812,958,1926,1952]" pageId="16" pageNumber="233" pagination="127 - 260" refId="ref12609" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, Leiden, 3 (2 - 3), 127 - 260, pls. 15 - 17. https: // doi. org / 10.5962 / bhl. title. 10640" type="journal article" year="1917">Tesch (1917)</bibRefCitation>
had noted that the New 
<collectingCountry id="34F6EDFBFFE7FFC4FB77F838FAF0EEED" box="[1232,1313,1926,1952]" name="Guinea-Bissau" pageId="16" pageNumber="233">Guinea</collectingCountry>
specimens has a distinct external orbital angle, in contrast to the 
<typeStatus id="935A13C9FFE7FFC4FD53F815FCF5EE88" box="[756,804,1963,1989]" pageId="16" pageNumber="233">type</typeStatus>
of 
<taxonomicName id="8BE1D6E8FFE7FFC4FCECF815FC1AEE89" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[843,971,1962,1989]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE7FFC4FCECF815FC8FEE88" box="[843,862,1963,1989]" italics="true" pageId="16" pageNumber="233">C</emphasis>
. 
<emphasis id="7E957179FFE7FFC4FCCBF814FC1AEE89" box="[876,971,1962,1988]" italics="true" pageId="16" pageNumber="233">villosum</emphasis>
</taxonomicName>
described by De Man (1987: 644) which has an indistinct one. The older records of “ 
<taxonomicName id="8BE1D6E8FFE7FFC4FDDEF870FC43EEA5" authorityName="Edmondson" authorityYear="1951" box="[633,914,1998,2024]" class="Malacostraca" family="Sesarmidae" genus="Cleistocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="233" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFE7FFC4FDDEF870FC43EEA5" box="[633,914,1998,2024]" italics="true" pageId="16" pageNumber="233">Cleistocoeloma suvaense</emphasis>
</taxonomicName>
” from 
<collectingCountry id="34F6EDFBFFE7FFC4FC46F870FB82EEA5" box="[993,1107,1998,2024]" name="Singapore" pageId="16" pageNumber="233">Singapore</collectingCountry>
by 
<bibRefCitation id="2870D09AFFE7FFC4FBDEF871FAE6EEA5" author="Tan, C. G. S. &amp; Ng, P. K. L." box="[1145,1335,1998,2025]" pageId="16" pageNumber="233" pagination="75 - 84" refId="ref12558" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84. https: // doi. org / 10.1007 / BF 00005655" type="journal article" year="1994">Tan &amp; Ng (1994)</bibRefCitation>
and 
<bibRefCitation id="2870D09AFFE7FFC5FACCF871FEFBE9FC" author="Wee, Y. C. &amp; Ng, P. K. L." lastPageId="17" lastPageNumber="234" pageId="16" pageNumber="233" refId="ref12677" refString="Wee, Y. C. &amp; Ng, P. K. L. (Eds.) (1994) A first look at biodiversity in Singapore. National council on the Environment, Singapore, 163 pp." type="book" year="1994">Wee &amp; Ng (1994)</bibRefCitation>
, are based on specimen records in the ZRC, are here referred to 
<taxonomicName id="8BE1D6E8FFE6FFC5FC59FF29FBB5E9FC" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[1022,1124,151,177]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="234" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE6FFC5FC59FF29FBC0E9FC" box="[1022,1041,151,177]" italics="true" pageId="17" pageNumber="234">C</emphasis>
. 
<emphasis id="7E957179FFE6FFC5FBB9FF29FBB5E9FC" box="[1054,1124,151,177]" italics="true" pageId="17" pageNumber="234">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE6FFC5FBCBFF29FB7AE9FC" bold="true" box="[1132,1195,151,177]" pageId="17" pageNumber="234">
<taxonomicNameLabel id="65A6CC02FFE6FFC5FBCBFF29FB7AE9FC" box="[1132,1195,151,177]" pageId="17" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
instead. A number of records of “ 
<taxonomicName id="8BE1D6E8FFE6FFC5FEB9FF05FE4EE998" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[286,415,187,213]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="234" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE6FFC5FEB9FF05FEE0E998" box="[286,305,187,213]" italics="true" pageId="17" pageNumber="234">C</emphasis>
. 
<emphasis id="7E957179FFE6FFC5FEE7FF05FE4EE998" box="[320,415,187,213]" italics="true" pageId="17" pageNumber="234">villosum</emphasis>
</taxonomicName>
” from the IWP, however, are in doubt as we have neither specimens from these areas nor good descriptions and/or figures on which to base our decisions (see later).
</paragraph>
<paragraph id="4C5EAD6BFFE6FFC5FF60FEBCFCE9E8B8" blockId="17.[151,1437,151,609]" pageId="17" pageNumber="234">
<emphasis id="7E957179FFE6FFC5FF60FEBCFE41E850" bold="true" box="[199,400,258,285]" pageId="17" pageNumber="234">Ecological notes.</emphasis>
<taxonomicName id="8BE1D6E8FFE6FFC5FE3FFEBDFD59E850" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[408,648,259,285]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="234" phylum="Arthropoda" rank="species" species="nobile" status="sp. nov.">
<emphasis id="7E957179FFE6FFC5FE3FFEBDFD59E850" box="[408,648,259,285]" italics="true" pageId="17" pageNumber="234">Clistocoeloma nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFE6FFC5FD37FEBDFD00E850" bold="true" box="[656,721,259,285]" pageId="17" pageNumber="234">
<taxonomicNameLabel id="65A6CC02FFE6FFC5FD37FEBDFD00E850" box="[656,721,259,285]" pageId="17" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is not a mangal species, although it has been found in habitats near mangroves. Most of the 
<collectingCountry id="34F6EDFBFFE6FFC5FE4DFE99FD8DE80C" box="[490,604,295,321]" name="Singapore" pageId="17" pageNumber="234">Singapore</collectingCountry>
specimens were found in muddy substrates amongst the rocky intertidal areas in the supralittoral zone, typically found under large rocks with muddy substrate. For the Japanese specimens (as 
<taxonomicName id="8BE1D6E8FFE6FFC5FF19FED1FEEDE8C4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[190,316,367,393]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="234" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFE6FFC5FF19FED1FF00E8C4" box="[190,209,367,393]" italics="true" pageId="17" pageNumber="234">C</emphasis>
. 
<emphasis id="7E957179FFE6FFC5FF7AFED1FEEDE8C4" box="[221,316,367,393]" italics="true" pageId="17" pageNumber="234">villosum</emphasis>
</taxonomicName>
), it was observed by 
<bibRefCitation id="2870D09AFFE6FFC5FD80FED1FCD0E8C4" author="Komai, T. &amp; Nagai, T. &amp; Yogi, A. &amp; Naruse, T. &amp; Fujita, Y. &amp; Shokita, S." box="[551,769,367,393]" pageId="17" pageNumber="234" pagination="33 - 63" refId="ref10699" refString="Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. &amp; Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63." type="journal article" year="2004">
Komai 
<emphasis id="7E957179FFE6FFC5FDDEFED1FD61E8C4" box="[633,688,367,393]" italics="true" pageId="17" pageNumber="234">et al.</emphasis>
(2004)
</bibRefCitation>
that it can be found underneath rocks or fallen leaves within the landward side of the mangrove, while 
<bibRefCitation id="2870D09AFFE6FFC5FDCCFE2DFC9AE8E0" author="Yuhara, T. &amp; Yokooka, H. &amp; Taru, M." box="[619,843,403,429]" pageId="17" pageNumber="234" pagination="19" refId="ref12778" refString="Yuhara, T., Yokooka, H. &amp; Taru, M. (2017) Range extension of the sesarmid crab Clistocoeloma villosum along the eastern Pacific coast of the Izu Peninsula, Japan. Marine Biodiversity Records, 10, 19. https: // doi. org / 10.1186 / s 41200 - 017 - 0122 - 1" type="journal article" year="2017">
Yuhara 
<emphasis id="7E957179FFE6FFC5FD65FE2DFD2BE8E0" box="[706,762,403,429]" italics="true" pageId="17" pageNumber="234">et al.</emphasis>
(2017)
</bibRefCitation>
noted that this species was found underneath gravels and cobble stones around the edges of mangrove forests. For the Korean material, 
<bibRefCitation id="2870D09AFFE6FFC5FB9AFE09FB2AE89C" author="Lee, S. - K. &amp; Jung, J. &amp; Kim, W." box="[1085,1275,439,465]" pageId="17" pageNumber="234" pagination="179 - 181" refId="ref11086" refString="Lee, S. - K., Jung, J. &amp; Kim, W. (2010) A new report on sesarmid crab Clistocoeloma villosum (Crustacea: Decapoda: Brachyura) from Korea. Korean Journal of Systematic Zoology, 26 (2), 179 - 181. https: // doi. org / 10.5635 / KJSZ. 2010.26.2.179" type="journal article" year="2010">
Lee 
<emphasis id="7E957179FFE6FFC5FBC8FE09FB79E89C" box="[1135,1192,439,465]" italics="true" pageId="17" pageNumber="234">et al.</emphasis>
(2010)
</bibRefCitation>
noted that the species occurred in “slightly moist soil under fallen leaves”.
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFE6FFC5FF60FE40FBF7EB2C" pageId="17" pageNumber="234" type="distribution">
<paragraph id="4C5EAD6BFFE6FFC5FF60FE40FBF7EB2C" blockId="17.[151,1437,151,609]" pageId="17" pageNumber="234">
<emphasis id="7E957179FFE6FFC5FF60FE40FEB0EB55" bold="true" box="[199,353,510,536]" pageId="17" pageNumber="234">Distribution.</emphasis>
<collectingCountry id="34F6EDFBFFE6FFC5FEC0FE41FE08EB54" box="[359,473,511,537]" name="Singapore" pageId="17" pageNumber="234">Singapore</collectingCountry>
(
<typeStatus id="935A13C9FFE6FFC5FE40FE41FDC8EB54" box="[487,537,511,537]" pageId="17" pageNumber="234">type</typeStatus>
locality), Sumatra (
<bibRefCitation id="2870D09AFFE6FFC5FD51FE41FC42EB54" author="De Man, J. G." box="[758,915,511,537]" pageId="17" pageNumber="234" pagination="75 - 218" refId="ref10460" refString="De Man, J. G. (1897) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 9 (1), 75 - 218. [in German]" type="journal article" year="1897">De Man 1897</bibRefCitation>
, 
<bibRefCitation id="2870D09AFFE6FFC5FC39FE41FC0BEB54" author="De Man, J. G." box="[926,986,511,537]" pageId="17" pageNumber="234" pagination="677 - 708" refId="ref10528" refString="De Man, J. G. (1898) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Sechster (Schluss-) Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (4), 677 - 708, pls. 28 - 38." type="journal article" year="1898">1898</bibRefCitation>
), 
<collectingCountry id="34F6EDFBFFE6FFC5FC4AFE41FB72EB54" box="[1005,1187,511,537]" name="Micronesia" pageId="17" pageNumber="234">Caroline Islands</collectingCountry>
(
<bibRefCitation id="2870D09AFFE6FFC5FB16FE41FA85EB54" author="Rathbun, M. J." box="[1201,1364,511,537]" pageId="17" pageNumber="234" pagination="25 - 74" refId="ref11841" refString="Rathbun, M. J. (1907) Reports on the scientific results of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. IX. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. X: The Brachyura. Memoirs of the Museum of Comparative Zoology at Harvard College, 35 (2), 25 - 74." type="journal article" year="1907">Rathbun 1907</bibRefCitation>
), New 
<collectingCountry id="34F6EDFBFFE6FFC5FF30FD9DFF39EB70" box="[151,232,547,573]" name="Guinea-Bissau" pageId="17" pageNumber="234">Guinea</collectingCountry>
(
<bibRefCitation id="2870D09AFFE6FFC5FF50FD9DFEACEB70" author="Tesch, J. J." box="[247,381,547,573]" pageId="17" pageNumber="234" pagination="127 - 260" refId="ref12609" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, Leiden, 3 (2 - 3), 127 - 260, pls. 15 - 17. https: // doi. org / 10.5962 / bhl. title. 10640" type="journal article" year="1917">Tesch 1917</bibRefCitation>
), the 
<collectingCountry id="34F6EDFBFFE6FFC5FE1CFD9DFDE8EB70" box="[443,569,547,573]" name="Philippines" pageId="17" pageNumber="234">Philippines</collectingCountry>
(
<bibRefCitation id="2870D09AFFE6FFC5FDEFFD9DFD3BEB70" author="McNeill, F. A." box="[584,746,547,573]" pageId="17" pageNumber="234" pagination="1 - 98" refId="ref11406" refString="McNeill, F. A. (1968) Crustacea, Decapoda and Stomatopoda. Great Barrier Expedition 1928 - 29, Scientific Reports, 7 (1), 1 - 98, pls. 1 + 2." type="journal article" year="1968">McNeill 1968</bibRefCitation>
), 
<collectingCountry id="34F6EDFBFFE6FFC5FD59FD9DFC81EB70" box="[766,848,547,573]" name="Taiwan" pageId="17" pageNumber="234">Taiwan</collectingCountry>
(
<bibRefCitation id="2870D09AFFE6FFC5FCF8FD9DFC6DEB70" author="Li, J. - J." box="[863,956,546,573]" pageId="17" pageNumber="234" pagination="84 - 91" refId="ref10890" refString="Li, J. - J. (2015) Land crabs of Christmas Island and Taiwan. Taiwan Natural Science, 126, 84 - 91. [in Chinese]" type="journal article" year="2015">Li 2015</bibRefCitation>
; 
<bibRefCitation id="2870D09AFFE6FFC5FC6FFD9DFBB5EB70" author="Li, J. - J. &amp; Hsu, J. - W. &amp; Ng, N. K. &amp; Shih, H. - T." box="[968,1124,547,573]" pageId="17" pageNumber="234" pagination="1207 - 1230" refId="ref10922" refString="Li, J. - J., Hsu, J. - W., Ng, N. K. &amp; Shih, H. - T. (2019) Eight new records of crabs (Decapoda, Brachyura: Sesarmidae, Varunidae) from the coasts of Taiwan. Crustaceana, 92 (10), 1207 - 1230. https: // doi. org / 10.1163 / 15685403 - 00003935" type="journal article" year="2019">
Li 
<emphasis id="7E957179FFE6FFC5FC4FFD9DFBCBEB70" box="[1000,1050,547,573]" italics="true" pageId="17" pageNumber="234">et al</emphasis>
. 2019
</bibRefCitation>
), 
<collectingCountry id="34F6EDFBFFE6FFC5FBDFFD9DFB69EB70" box="[1144,1208,547,573]" name="Japan" pageId="17" pageNumber="234">Japan</collectingCountry>
(
<bibRefCitation id="2870D09AFFE6FFC5FB60FD9DFA49EB70" author="Komai, T. &amp; Nagai, T. &amp; Yogi, A. &amp; Naruse, T. &amp; Fujita, Y. &amp; Shokita, S." box="[1223,1432,547,573]" pageId="17" pageNumber="234" pagination="33 - 63" refId="ref10699" refString="Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. &amp; Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63." type="journal article" year="2004">
Komai 
<emphasis id="7E957179FFE6FFC5FABBFD9DFA9FEB70" box="[1308,1358,547,573]" italics="true" pageId="17" pageNumber="234">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="2870D09AFFE6FFC5FF30FDF9FEB6EB2C" author="Yuhara, T. &amp; Yokooka, H. &amp; Taru, M." box="[151,359,583,609]" pageId="17" pageNumber="234" pagination="19" refId="ref12778" refString="Yuhara, T., Yokooka, H. &amp; Taru, M. (2017) Range extension of the sesarmid crab Clistocoeloma villosum along the eastern Pacific coast of the Izu Peninsula, Japan. Marine Biodiversity Records, 10, 19. https: // doi. org / 10.1186 / s 41200 - 017 - 0122 - 1" type="journal article" year="2017">
Yuhara 
<emphasis id="7E957179FFE6FFC5FF4AFDF9FECEEB2C" box="[237,287,583,609]" italics="true" pageId="17" pageNumber="234">et al</emphasis>
. 2017
</bibRefCitation>
; 
<bibRefCitation id="2870D09AFFE6FFC5FED4FDF9FD5FEB2C" author="Takakura, H. &amp; Komai, T." box="[371,654,583,609]" pageId="17" pageNumber="234" pagination="25 - 31" refId="ref12498" refString="Takakura, H. &amp; Komai, T. (2019) New records of a semiterrestrial sesarmid crab Clistocoeloma villosum (A. Milne-Edwards, 1869) (Crustacea: Decapoda: Brachyura: Grapsoidea) from Izu and Boso Peninsulas, Japan. Cancer, 28, 25 - 31. [in Japanese with English abstract]" type="journal article" year="2019">Takakura &amp; Komai 2019</bibRefCitation>
), and 
<collectingCountry id="34F6EDFBFFE6FFC5FD77FDF9FC8CEB2C" box="[720,861,583,609]" name="South Korea" pageId="17" pageNumber="234">South Korea</collectingCountry>
(
<bibRefCitation id="2870D09AFFE6FFC5FCCBFDF9FBCBEB2C" author="Lee, S. - K. &amp; Jung, J. &amp; Kim, W." box="[876,1050,583,609]" pageId="17" pageNumber="234" pagination="179 - 181" refId="ref11086" refString="Lee, S. - K., Jung, J. &amp; Kim, W. (2010) A new report on sesarmid crab Clistocoeloma villosum (Crustacea: Decapoda: Brachyura) from Korea. Korean Journal of Systematic Zoology, 26 (2), 179 - 181. https: // doi. org / 10.5635 / KJSZ. 2010.26.2.179" type="journal article" year="2010">
Lee 
<emphasis id="7E957179FFE6FFC5FC39FDF9FC1EEB2C" box="[926,975,583,609]" italics="true" pageId="17" pageNumber="234">et al</emphasis>
. 2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>