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<document ID-DOI="10.25221/fee.398.2" ID-ISSN="2713-2196" ID-Zenodo-Dep="7165240" approvalRequired="53" approvalRequired_for_textStreams="53" checkinTime="1665248349315" checkinUser="felipe" docAuthor="Hongsuwong, T., Jaitrong, W., Tasen, W. &amp; Zorn, C." docDate="2020" docId="C01F94366E70FFB8A5CFD86DFE6FF50C" docLanguage="en" docName="FarEastEntomol.398.8-17.pdf" docOrigin="Far Eastern Entomologist 398" docSource="http://dx.doi.org/10.25221/fee.398.2" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Rutelarcha quadrimaculata Waterhouse 1874" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="3C26EC4E6E71FFBAA52CDC18FF87F04D" masterDocTitle="New records of the chining leaf chafer beetles (Coleoptera: Scarabaeidae, Rutelinae) from Thailand and Laos" masterLastPageNumber="17" masterPageNumber="8" pageNumber="9" updateTime="1665262470503" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New records of the chining leaf chafer beetles (Coleoptera: Scarabaeidae, Rutelinae) from Thailand and Laos</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hongsuwong, T.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaitrong, W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tasen, W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zorn, C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-01-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>398</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>8</mods:start>
<mods:end>17</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.398.2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.398.2</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7165240</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:C01F94366E70FFB8A5CFD86DFE6FF50C" httpUri="http://treatment.plazi.org/id/C01F94366E70FFB8A5CFD86DFE6FF50C" lastPageId="2" lastPageNumber="10" pageId="1" pageNumber="9">
<subSubSection box="[227,679,1141,1163]" pageId="1" pageNumber="9" type="nomenclature">
<paragraph blockId="1.[227,1128,1141,1328]" box="[227,679,1141,1163]" pageId="1" pageNumber="9">
<heading box="[227,679,1141,1163]" fontSize="9" level="2" pageId="1" pageNumber="9" reason="2">
<taxonomicName authority="Waterhouse, 1874" authorityName="Waterhouse" authorityYear="1874" box="[227,679,1141,1163]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis bold="true" box="[227,679,1141,1163]" pageId="1" pageNumber="9">
<emphasis bold="true" box="[227,501,1141,1163]" italics="true" pageId="1" pageNumber="9">Rutelarcha quadrimaculata</emphasis>
Waterhouse, 1874
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,310,1168,1190]" pageId="1" pageNumber="9" type="description">
<paragraph blockId="1.[227,1128,1141,1328]" box="[227,310,1168,1190]" pageId="1" pageNumber="9">
<figureCitation box="[227,310,1168,1190]" captionStart="Figs 16" captionStartId="2.[265,306,1103,1125]" captionTargetBox="[230,1099,388,1074]" captionTargetId="figure-33@2.[227,1114,368,1081]" captionTargetPageId="2" captionText="Figs 16. Rutelarcha quadrimaculata, Thailand: Trang province, female: 1 habitus," figureDoi="http://doi.org/10.5281/zenodo.7165242" httpUri="https://zenodo.org/record/7165242/files/figure.png" pageId="1" pageNumber="9">Figs 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="9" type="reference_group">
<paragraph blockId="1.[227,1128,1141,1328]" box="[265,1126,1196,1218]" pageId="1" pageNumber="9">
<taxonomicName authority="Waterhouse, 1874: 52" authorityName="Waterhouse" authorityPageNumber="52" authorityYear="1874" box="[265,775,1196,1218]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis box="[265,536,1196,1217]" italics="true" pageId="1" pageNumber="9">Rutelarcha quadrimaculata</emphasis>
<materialsCitation box="[549,775,1196,1218]" country="Malaysia" location="Penang" pageId="1" pageNumber="9" specimenCount="1" stateProvince="Pulau Pinang">Waterhouse, 1874: 52</materialsCitation>
</taxonomicName>
.
<typeStatus box="[790,840,1196,1217]" pageId="1" pageNumber="9">Type</typeStatus>
Locality:
<collectingRegion box="[950,1023,1196,1217]" country="Malaysia" name="Pulau Pinang" pageId="1" pageNumber="9">Penang</collectingRegion>
,
<collectingCountry box="[1038,1126,1196,1218]" name="Malaysia" pageId="1" pageNumber="9">Malaysia</collectingCountry>
</paragraph>
<paragraph blockId="1.[227,1128,1141,1328]" box="[227,322,1224,1245]" pageId="1" pageNumber="9">(BMNH).</paragraph>
<paragraph blockId="1.[227,1128,1141,1328]" box="[265,1128,1251,1273]" pageId="1" pageNumber="9">
<taxonomicName authority=": Westwood, 1875: 235" authorityName=": Westwood" authorityPageNumber="235" authorityYear="1875" box="[265,768,1251,1273]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis box="[265,533,1251,1272]" italics="true" pageId="1" pageNumber="9">Rutelarcha quadrimaculata</emphasis>
: Westwood, 1875: 235
</taxonomicName>
; Nonfried, 1891: 355; Ohaus, 1900:
</paragraph>
<paragraph blockId="1.[227,1128,1141,1328]" box="[227,1127,1279,1301]" pageId="1" pageNumber="9">260; Ohaus, 1918: 39; Ohaus, 1932a: 122; Kuijten, 1988: 79; Kuijten, 1992: 19; Wada, 1988: 5,</paragraph>
</subSubSection>
<subSubSection box="[227,391,1306,1328]" pageId="1" pageNumber="9" type="description">
<paragraph blockId="1.[227,1128,1141,1328]" box="[227,391,1306,1328]" pageId="1" pageNumber="9">2015: table III-2.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="9" type="materials_examined">
<paragraph blockId="1.[227,1127,1361,1659]" pageId="1" pageNumber="9">
<materialsCitation collectingDate="2019-03-06" collectionCode="THNHM" collectorName="S. Amuntaikul &amp; P. Wannaput" country="Thailand" location="Huai Yot Subdistrict" municipality="Huai Yot District" pageId="1" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Trang">
MATERIAL EXAMINED.
<emphasis bold="true" box="[536,629,1361,1382]" pageId="1" pageNumber="9">
<collectingCountry box="[536,629,1361,1382]" name="Thailand" pageId="1" pageNumber="9">Thailand</collectingCountry>
</emphasis>
:
<collectingRegion box="[643,794,1361,1383]" country="Thailand" name="Trang" pageId="1" pageNumber="9">Trang province</collectingRegion>
,
<collectingMunicipality box="[804,975,1361,1383]" pageId="1" pageNumber="9">Huai Yot District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C01F94366E70FFB8A5CFD86DFE6FF50C:4D6973FB6E70FFBBA6F5D949FEAFF5CE" country="Thailand" municipality="Huai Yot District" name="Huai Yot Subdistrict" pageId="1" pageNumber="9" stateProvince="Trang">Huai Yot Subdistrict</location>
,
<date box="[306,404,1389,1411]" pageId="1" pageNumber="9" value="2019-03-06">
<collectingDate box="[306,404,1389,1411]" pageId="1" pageNumber="9" value="2019-03-06">6.III 2019</collectingDate>
</date>
,
<specimenCount box="[414,496,1389,1411]" pageId="1" pageNumber="9" type="female">1 female</specimenCount>
(
<collectionCode box="[509,600,1389,1411]" pageId="1" pageNumber="9">THNHM</collectionCode>
I09976),
<collectorName box="[711,851,1389,1411]" pageId="1" pageNumber="9">S. Amuntaikul</collectorName>
&amp;
<collectorName box="[881,1006,1390,1411]" pageId="1" pageNumber="9">P. Wannaput</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[265,1062,1416,1438]" pageId="1" pageNumber="9" type="description">
<paragraph blockId="1.[227,1127,1361,1659]" box="[265,1062,1416,1438]" pageId="1" pageNumber="9">
MEASUREMENTS.
<emphasis box="[473,545,1417,1438]" italics="true" pageId="1" pageNumber="9">Female</emphasis>
: TL
<quantity box="[593,681,1416,1438]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="1" pageNumber="9" unit="mm" value="15.6">15.6 mm</quantity>
; EW
<quantity box="[734,811,1417,1438]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="1" pageNumber="9" unit="mm" value="9.4">9.4 mm</quantity>
; PW
<quantity box="[862,938,1416,1438]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="9" unit="mm" value="5.1">5.1 mm</quantity>
; PL
<quantity box="[982,1058,1417,1438]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="9" unit="mm" value="8.3">8.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="10" pageId="1" pageNumber="9" type="diagnosis">
<paragraph blockId="1.[227,1127,1361,1659]" box="[265,1126,1444,1466]" pageId="1" pageNumber="9">
DIAGNOSIS.
<taxonomicName authorityName=": Westwood" authorityYear="1875" box="[407,674,1444,1465]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis box="[407,674,1444,1465]" italics="true" pageId="1" pageNumber="9">Rutelarcha quadrimaculata</emphasis>
</taxonomicName>
is a conspicuous species, which can be distin-
</paragraph>
<paragraph blockId="1.[227,1127,1361,1659]" box="[227,1126,1472,1494]" pageId="1" pageNumber="9">
guished from the only other known species in this genus (
<taxonomicName authority="Ohaus, 1915" authorityName="Ohaus" authorityYear="1915" box="[809,1033,1472,1494]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis box="[809,900,1472,1493]" italics="true" pageId="1" pageNumber="9">R. bakeri</emphasis>
Ohaus, 1915
</taxonomicName>
from the
</paragraph>
<paragraph blockId="1.[227,1127,1361,1659]" box="[227,1126,1499,1521]" pageId="1" pageNumber="9">
<collectingCountry box="[227,337,1499,1521]" name="Philippines" pageId="1" pageNumber="9">Philippines</collectingCountry>
) and similar species in the closely related genera
<taxonomicName authority="Waterhouse, 1875" authorityName="Waterhouse" authorityYear="1875" box="[834,1126,1499,1521]" class="Insecta" family="Rutelidae" genus="Cyphelytra" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis box="[834,941,1499,1520]" italics="true" pageId="1" pageNumber="9">Cyphelytra</emphasis>
Waterhouse, 1875
</taxonomicName>
</paragraph>
<paragraph blockId="1.[227,1127,1361,1659]" box="[227,1127,1527,1549]" pageId="1" pageNumber="9">
and
<taxonomicName authority="Westwood, 1875" authorityName="Westwood" authorityYear="1875" box="[272,513,1527,1549]" class="Insecta" family="Rutelidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis box="[272,336,1528,1549]" italics="true" pageId="1" pageNumber="9">Lutera</emphasis>
Westwood, 1875
</taxonomicName>
by the combination of the following characters: head black,
</paragraph>
<paragraph blockId="1.[227,1127,1361,1659]" pageId="1" pageNumber="9">anterior clypeal margin medially with two weak denticles; lateral face of pronotum densely longitudinally striated. Pronotum largely black with its lateral face yellowish orange, with black spot located in middle of orange area; elytra with several rows of rounded punctures and smooth and shiny interspaces.</paragraph>
<paragraph blockId="2.[227,1127,263,340]" pageId="2" pageNumber="10">
Ground color of elytra yellowish orange, with small black spot behind humeral callus and large transverse spot in posterior half (
<figureCitation box="[628,720,291,313]" captionStart="Figs 16" captionStartId="2.[265,306,1103,1125]" captionTargetBox="[230,1099,388,1074]" captionTargetId="figure-33@2.[227,1114,368,1081]" captionTargetPageId="2" captionText="Figs 16. Rutelarcha quadrimaculata, Thailand: Trang province, female: 1 habitus," figureDoi="http://doi.org/10.5281/zenodo.7165242" httpUri="https://zenodo.org/record/7165242/files/figure.png" pageId="2" pageNumber="10">Figs 12</figureCitation>
); mesosternal process extending antero-
</paragraph>
<paragraph blockId="2.[227,1127,263,340]" box="[227,650,318,340]" pageId="2" pageNumber="10">
ventrally beyond base of mesocoxa (
<figureCitation box="[578,636,318,340]" captionStart="Figs 16" captionStartId="2.[265,306,1103,1125]" captionTargetBox="[230,1099,388,1074]" captionTargetId="figure-33@2.[227,1114,368,1081]" captionTargetPageId="2" captionText="Figs 16. Rutelarcha quadrimaculata, Thailand: Trang province, female: 1 habitus," figureDoi="http://doi.org/10.5281/zenodo.7165242" httpUri="https://zenodo.org/record/7165242/files/figure.png" pageId="2" pageNumber="10">Fig. 3</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7165242" ID-Zenodo-Dep="7165242" box="[265,1127,1102,1125]" httpUri="https://zenodo.org/record/7165242/files/figure.png" pageId="2" pageNumber="10" startId="2.[265,306,1103,1125]" targetBox="[230,1099,388,1074]" targetPageId="2">
<paragraph blockId="2.[227,1127,1102,1180]" box="[265,1127,1102,1125]" pageId="2" pageNumber="10">
Figs 16.
<taxonomicName authorityName=": Westwood" authorityYear="1875" box="[369,639,1103,1124]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="10" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis box="[369,639,1103,1124]" italics="true" pageId="2" pageNumber="10">Rutelarcha quadrimaculata</emphasis>
</taxonomicName>
, Thailand: Trang province, female: 1 habitus,
</paragraph>
</caption>
<paragraph blockId="2.[227,1127,1102,1180]" box="[227,1127,1130,1152]" pageId="2" pageNumber="10">dorsal view; 2 habitus, lateral view; 3 habitus, ventral view; 4 fore leg; 5 middle leg;</paragraph>
<paragraph blockId="2.[227,1127,1102,1180]" box="[227,348,1158,1180]" pageId="2" pageNumber="10">6 hind leg.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="10" type="distribution">
<paragraph blockId="2.[227,1127,1213,1345]" box="[265,1127,1213,1235]" pageId="2" pageNumber="10">
DISTRIBUTION.
<collectingCountry box="[448,533,1213,1235]" name="Thailand" pageId="2" pageNumber="10">Thailand</collectingCountry>
(
<emphasis bold="true" box="[550,669,1213,1234]" pageId="2" pageNumber="10">new record</emphasis>
), Borneo (Ohaus, 1900, 1932a; Wada, 2015),
</paragraph>
<paragraph blockId="2.[227,1127,1213,1345]" box="[227,1099,1240,1262]" pageId="2" pageNumber="10">
West
<collectingCountry box="[281,367,1240,1262]" name="Malaysia" pageId="2" pageNumber="10">Malaysia</collectingCountry>
(Kuijten, 1988; Ohaus, 1900; Waterhouse, 1874) and Sumatra (Ohaus, 1918).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="10" type="discussion">
<paragraph blockId="2.[227,1127,1213,1345]" pageId="2" pageNumber="10">
REMARKS.
<taxonomicName authorityName=": Westwood" authorityYear="1875" box="[393,660,1268,1289]" class="Insecta" family="Rutelidae" genus="Rutelarcha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="10" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis box="[393,660,1268,1289]" italics="true" pageId="2" pageNumber="10">Rutelarcha quadrimaculata</emphasis>
</taxonomicName>
has a typical sundaic distribution. The record in the present study is the northernmost in its distribution range. A larva of this species was collected from a rotten log.
</paragraph>
</subSubSection>
</treatment>
</document>