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<document id="B2A256AD3B6EA4442D2993100264D103" ID-CLB-Dataset="26022" ID-DOI="10.11646/zootaxa.4656.2.2" ID-GBIF-Dataset="62912763-9710-4c81-88ea-5d8c9f9a3bdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3368143" ID-ZooBank="1CD78AA3-EC4F-4E7F-9005-5ED81CB1F72E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565768352213" checkinUser="plazi" docAuthor="Pérez-Gutiérrez, León Andrés" docDate="2019" docId="BC6E87EBFFBE071E24A6F905FF16E93D" docLanguage="en" docName="zootaxa.4656.2.2.pdf" docOrigin="Zootaxa 4656 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dactylobasis demarmelsi Pérez-Gutiérrez 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="240" masterDocId="4057FF93FFBD07162431FFEDFFCEEB02" masterDocTitle="Dactylobasis gen. nov. from Colombia, a new genus of Zygoptera with Teinobasini affinities (Odonata: Coenagrionidae)" masterLastPageNumber="242" masterPageNumber="232" pageNumber="235" updateTime="1732786376359" updateUser="ExternalLinkService">
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<mods:title id="F2DD5A0B338DDA9FEA5357249F833820">Dactylobasis gen. nov. from Colombia, a new genus of Zygoptera with Teinobasini affinities (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="7C3830C8E279A092ED174CE0D11338AB">Pérez-Gutiérrez, León Andrés</mods:namePart>
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<mods:title id="BD8F787B497A4CCD6EE5A903AF2F88CD">Zootaxa</mods:title>
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<mods:date id="8DADDBC1BAE7A5C1160A18ABB6270B10">2019</mods:date>
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<mods:number id="3B04C1F38D4E67D6770348BB3BA69DC2">2019-08-14</mods:number>
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<mods:identifier id="250B471797B217FEF84842A0A3D711E5" type="DOI">10.11646/zootaxa.4656.2.2</mods:identifier>
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<treatment id="BC6E87EBFFBE071E24A6F905FF16E93D" ID-GBIF-Taxon="159134097" LSID="urn:lsid:plazi:treatment:BC6E87EBFFBE071E24A6F905FF16E93D" httpUri="http://treatment.plazi.org/id/BC6E87EBFFBE071E24A6F905FF16E93D" lastPageId="8" lastPageNumber="240" pageId="3" pageNumber="235" scope_family="Coenagrionidae" scope_order="Odonata">
<subSubSection id="7CDD6576FFBE071524A6F905FDE7EC00" box="[151,553,1768,1795]" pageId="3" pageNumber="235" type="nomenclature">
<paragraph id="347836FDFFBE071524A6F905FDE7EC00" blockId="3.[151,553,1768,1795]" box="[151,553,1768,1795]" pageId="3" pageNumber="235">
<heading id="6F308191FFBE071524A6F905FDE7EC00" bold="true" box="[151,553,1768,1795]" fontSize="11" level="1" pageId="3" pageNumber="235" reason="1">
<emphasis id="06B3EAEFFFBE071524A6F905FDE7EC00" bold="true" box="[151,553,1768,1795]" pageId="3" pageNumber="235">
<taxonomicName id="F3C74D7EFFBE071524A6F905FE0AEC01" ID-CoL="9X2FS" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[151,452,1768,1795]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="235" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="06B3EAEFFFBE071524A6F905FE0AEC01" bold="true" box="[151,452,1768,1795]" italics="true" pageId="3" pageNumber="235">Dactylobasis demarmelsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D805794FFBE071525FAF905FDE7EC00" box="[459,553,1768,1794]" pageId="3" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7CDD6576FFBE071E24A6F8DDFF16E93D" lastPageId="8" lastPageNumber="240" pageId="3" pageNumber="235" type="materials_examined">
<paragraph id="347836FDFFBE071524A6F8DDFB3EEC6C" blockId="3.[151,1437,1840,2010]" pageId="3" pageNumber="235">
<emphasis id="06B3EAEFFFBE071524A6F8DDFED0EC48" bold="true" box="[151,286,1840,1866]" pageId="3" pageNumber="235">Etymology.</emphasis>
The specific epithet is named in honor of Professor Doctor Jürg Carl De Marmels for his scientific work inspiration and extensive contribution to Odonatology, especially concerning the neotropical fauna.
</paragraph>
<paragraph id="347836FDFFBE071224F6F89AFF0FEBF9" blockId="3.[151,1437,1840,2010]" lastBlockId="4.[151,1436,152,539]" lastPageId="4" lastPageNumber="236" pageId="3" pageNumber="235">
<emphasis id="06B3EAEFFFBE071524F6F89AFE05EC90" bold="true" box="[199,459,1911,1938]" pageId="3" pageNumber="235">Specimens examined.</emphasis>
<materialsCitation id="84AF3CA0FFBE071525E6F895FCF2ECB4" ID-GBIF-Occurrence="2332527395" collectingDate="2005-08-10" collectionCode="SAIA" collectorName="L. A. Perez" country="Colombia" county="Dept." elevation="129" lastPageId="3" lastPageNumber="236" latitude="5.3169446" location="Salero" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" pageId="3" pageNumber="235" specimenCount="1" specimenCount-female="18" specimenCount-male="25" stateProvince="Choco Department" typeStatus="holotype">
<typeStatus id="EB7C885FFFBE071525E6F895FDF1EC90" box="[471,575,1912,1938]" pageId="3" pageNumber="235" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="4CD0766DFFBE071526A4F895FCE3EC90" box="[661,813,1912,1938]" name="Colombia" pageId="3" pageNumber="235">COLOMBIA</collectingCountry>
,
<collectingRegion id="F603F81FFFBE0715270CF895FBD7EC90" box="[829,1049,1912,1938]" country="Colombia" name="Choco" pageId="3" pageNumber="235">Chocó Department</collectingRegion>
,
<location id="31186026FFBE07152018F895FBBFEC90" LSID="urn:lsid:plazi:treatment:BC6E87EBFFBE071E24A6F905FF16E93D:31186026FFBE07152018F895FBBFEC90" box="[1065,1137,1912,1938]" country="Colombia" county="Dept." latitude="5.3169446" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" name="Salero" pageId="3" pageNumber="235" stateProvince="Choco Department">Salero</location>
(
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<geoCoordinate id="51F3503AFFBE07152116F89AFF7BECB4" degrees="76" direction="west" minutes="36" orientation="longitude" pageId="3" pageNumber="235" precision="15" seconds="52" value="-76.61444">76°3652” W</geoCoordinate>
), alt.
<quantity id="F33F9B18FFBE071524C4F871FEF9ECB4" box="[245,311,1948,1974]" metricMagnitude="2" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="235" unit="m" value="129.0">
<elevation id="BFEAD1CEFFBE071524C4F871FEF9ECB4" box="[245,311,1948,1974]" metricMagnitude="2" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="235" unit="m" value="129.0">129m</elevation>
</quantity>
,
<date id="4079103DFFBE07152572F871FE14ECB4" box="[323,474,1947,1974]" pageId="3" pageNumber="235" value="2005-08-10">
<collectingDate id="503DE9D5FFBE07152572F871FE14ECB4" box="[323,474,1947,1974]" pageId="3" pageNumber="235" value="2005-08-10">10 VIII 2005</collectingDate>
</date>
,
<collectorName id="9932532BFFBE071525D7F871FDABECB4" box="[486,613,1948,1974]" pageId="3" pageNumber="235">L. A. Pérez</collectorName>
leg. (
<collectionCode id="52D6AE38FFBE07152697F871FD24ECB4" box="[678,746,1948,1974]" pageId="3" pageNumber="235">SAIA</collectionCode>
_0287)
</materialsCitation>
.
<materialsCitation id="84AF3CA0FFBE07152777F871FD8FECD8" ID-GBIF-Occurrence="2413161225" collectingDate="2005-08-10" collectionCode="SAIA, UARC" collectorName="L. A. Perez" country="Colombia" county="Dept." elevation="129" lastPageId="3" lastPageNumber="236" latitude="5.3169446" location="Salero" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" pageId="3" pageNumber="235" specimenCount="43" specimenCount-female="18" specimenCount-male="25" stateProvince="Choco" typeStatus="allotype">
<typeStatus id="EB7C885FFFBE07152777F871FC66ECB4" box="[838,936,1948,1974]" pageId="3" pageNumber="235" type="allotype">Allotype</typeStatus>
same holotype but (
<collectionCode id="52D6AE38FFBE071520BFF871FB1CECB4" box="[1166,1234,1948,1974]" pageId="3" pageNumber="235">SAIA</collectionCode>
_0288).
<typeStatus id="EB7C885FFFBE0715211EF871FA52ECB4" box="[1327,1436,1948,1974]" pageId="3" pageNumber="235" type="paratype">Paratypes</typeStatus>
<specimenCount id="22C1FD74FFBE071524A6F852FECFECD8" box="[151,257,1983,2010]" pageId="3" pageNumber="235" type="male">25 males</specimenCount>
,
<specimenCount id="22C1FD74FFBE0715253EF82DFE42ECD8" box="[271,396,1984,2010]" pageId="3" pageNumber="235" type="female">18 females</specimenCount>
(all in
<collectionCode id="52D6AE38FFBE071525D4F82DFDF7ECD8" box="[485,569,1984,2010]" pageId="3" pageNumber="235">UARC</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="84AF3CA0FFBE0712267FF82DFC60EBB1" ID-GBIF-Occurrence="2332527396" collectingDate="2008-08-10" collectionCode="SAIA" collectorName="L. A. Perez" country="Colombia" county="Dept." elevation="150" lastPageId="4" lastPageNumber="236" latitude="5.3169446" location="Tado" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" pageId="3" pageNumber="235" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Choco" typeStatus="holotype">
<specimenCount id="22C1FD74FFBE0715267FF82DFD47ECD8" box="[590,649,1984,2010]" pageId="3" pageNumber="235" type="male">6♂♂</specimenCount>
,
<specimenCount id="22C1FD74FFBE071526A6F82DFD1EECD8" box="[663,720,1984,2010]" pageId="3" pageNumber="235" type="female">4♀♀</specimenCount>
,
<collectingRegion id="F603F81FFFBE071526D1F82DFCE7ECD8" box="[736,809,1984,2010]" country="Colombia" name="Choco" pageId="3" pageNumber="235">Chocó</collectingRegion>
<collectingCounty id="DD194E71FFBE07152702F82DFCBAECD8" box="[819,884,1984,2010]" pageId="3" pageNumber="235">Dept.</collectingCounty>
,
<collectingMunicipality id="D41CAC87FFBE071527B3F82DFC17ECD8" box="[898,985,1984,2010]" pageId="3" pageNumber="235">Quibdó</collectingMunicipality>
, km 10 via
<location id="31186026FFBE07152057F82DFB6EECD8" LSID="urn:lsid:plazi:treatment:BC6E87EBFFBE071E24A6F905FF16E93D:31186026FFBE07152057F82DFB6EECD8" box="[1126,1184,1984,2010]" country="Colombia" county="Dept." latitude="5.3169446" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" name="Tado" pageId="3" pageNumber="235" stateProvince="Choco">Tadó</location>
, sector “Los Chalets” (
<geoCoordinate id="51F3503AFFB907122491FF75FEFFEBB1" box="[160,305,152,179]" degrees="05" direction="north" minutes="19" orientation="latitude" pageId="4" pageNumber="236" precision="15" seconds="01" value="5.3169446">05°1901” N</geoCoordinate>
<geoCoordinate id="51F3503AFFB907122509FF75FE1FEBB1" box="[312,465,152,179]" degrees="76" direction="west" minutes="36" orientation="longitude" pageId="4" pageNumber="236" precision="15" seconds="52" value="-76.61444">76°3652” W</geoCoordinate>
), alt.
<quantity id="F33F9B18FFB90712263EFF75FD9CEBB1" box="[527,594,152,179]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="236" unit="m" value="150.0">
<elevation id="BFEAD1CEFFB90712263EFF75FD9CEBB1" box="[527,594,152,179]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="236" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="4079103DFFB90712266CFF74FD3CEBB1" box="[605,754,153,179]" pageId="4" pageNumber="236" value="2008-08-10">
<collectingDate id="503DE9D5FFB90712266CFF74FD3CEBB1" box="[605,754,153,179]" pageId="4" pageNumber="236" value="2008-08-10">10 VIII 2008</collectingDate>
</date>
,
<collectorName id="9932532BFFB9071226CCFF74FCB5EBB1" box="[765,891,153,179]" pageId="4" pageNumber="236">L. A. Pérez</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="84AF3CA0FFB90712278BFF75FA59EBB1" ID-GBIF-Occurrence="2413161228" box="[954,1431,152,179]" collectingDate="2010-08-10" collectionCode="SAIA" collectorName="L. A. Perez" country="Colombia" county="Dept." elevation="129" lastPageId="4" lastPageNumber="236" latitude="5.3169446" location="Salero" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" pageId="4" pageNumber="235" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Choco" typeStatus="holotype">
<specimenCount id="22C1FD74FFB90712278BFF75FC3AEBB1" box="[954,1012,152,179]" pageId="4" pageNumber="236" type="male">5♂♂</specimenCount>
,
<specimenCount id="22C1FD74FFB907122031FF74FBF7EBB1" box="[1024,1081,153,179]" pageId="4" pageNumber="236" type="female">4♀♀</specimenCount>
,
<date id="4079103DFFB907122077FF74FB15EBB1" box="[1094,1243,153,179]" pageId="4" pageNumber="236" value="2010-08-10">
<collectingDate id="503DE9D5FFB907122077FF74FB15EBB1" box="[1094,1243,153,179]" pageId="4" pageNumber="236" value="2010-08-10">10 VIII 2010</collectingDate>
</date>
, L. A. Pérez leg.
</materialsCitation>
;
<materialsCitation id="84AF3CA0FFB9071224A6FF50FCE3EBD5" ID-GBIF-Occurrence="2413161227" box="[151,813,189,215]" collectingDate="2012-08-10" collectionCode="SAIA" collectorName="L. A. Perez" country="Colombia" county="Dept." elevation="129" lastPageId="4" lastPageNumber="236" latitude="5.3169446" location="Salero" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" pageId="4" pageNumber="235" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Choco" typeStatus="holotype">
<specimenCount id="22C1FD74FFB9071224A6FF50FF1CEBD5" box="[151,210,189,215]" pageId="4" pageNumber="236" type="male">6♂♂</specimenCount>
,
<specimenCount id="22C1FD74FFB9071224ECFF50FED9EBD5" box="[221,279,189,215]" pageId="4" pageNumber="236" type="female">6♀♀</specimenCount>
,
<date id="4079103DFFB907122512FF50FE76EBD5" box="[291,440,189,215]" pageId="4" pageNumber="236" value="2012-08-10">
<collectingDate id="503DE9D5FFB907122512FF50FE76EBD5" box="[291,440,189,215]" pageId="4" pageNumber="236" value="2012-08-10">10 VIII 2012</collectingDate>
</date>
, L. A. Pérez &amp; J. M. Montes leg.
</materialsCitation>
;
<materialsCitation id="84AF3CA0FFB907122708FF50FF0FEBF9" ID-GBIF-Occurrence="2413161226" collectingDate="2013-08-10" collectionCode="SAIA" collectorName="L. A. Perez" country="Colombia" county="Dept." elevation="129" lastPageId="4" lastPageNumber="236" latitude="5.3169446" location="Salero" longLatPrecision="21" longitude="-76.61444" municipality="Quibdo" pageId="4" pageNumber="235" specimenCount="12" specimenCount-female="4" specimenCount-male="8" stateProvince="Choco" typeStatus="holotype">
<specimenCount id="22C1FD74FFB907122708FF50FCBAEBD5" box="[825,884,189,215]" pageId="4" pageNumber="236" type="male">8♂♂</specimenCount>
,
<specimenCount id="22C1FD74FFB90712274EFF50FC76EBD5" box="[895,952,189,215]" pageId="4" pageNumber="236" type="female">4♀♀</specimenCount>
,
<date id="4079103DFFB9071227F7FF50FB92EBD5" box="[966,1116,189,215]" pageId="4" pageNumber="236" value="2013-08-10">
<collectingDate id="503DE9D5FFB9071227F7FF50FB92EBD5" box="[966,1116,189,215]" pageId="4" pageNumber="236" value="2013-08-10">10 VIII 2013</collectingDate>
</date>
. L. A. Pérez &amp; J. M. Montes leg.
</materialsCitation>
</paragraph>
<paragraph id="347836FDFFB9071224F6FEE9FDB4E919" blockId="4.[151,1436,152,539]" pageId="4" pageNumber="236">
<emphasis id="06B3EAEFFFB9071224F6FEE9FE27EA1D" bold="true" box="[199,489,260,287]" pageId="4" pageNumber="236">
Description of
<typeStatus id="EB7C885FFFB90712254DFEE8FE2BEA1D" box="[380,485,261,287]" pageId="4" pageNumber="236" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="06B3EAEFFFB9071225C2FEE8FDFEEA1D" box="[499,560,261,287]" italics="true" pageId="4" pageNumber="236">Head</emphasis>
(
<figureCitation id="ACFC2A78FFB907122670FEE8FD5FEA1D" box="[577,657,261,287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1589,1614]" captionTargetBox="[151,1436,227,1569]" captionTargetId="figure@3.[151,1436,227,1569]" captionTargetPageId="3" captionText="FIGURE 2. Dactylobasis demarmelsi sp. nov. (ab) male head and prothorax, lateral view; (b) female head, dorsal view; (c) thorax showing left projection of propleuron, lateral view; (d) thoracic color pattern; (e) male prothorax, dorsal view showing projections of propleuron (f) male thorax, dorsal view. Pp: Propleuron projections." httpUri="https://zenodo.org/record/3368147/files/figure.png" pageId="4" pageNumber="236">Figs. 2</figureCitation>
ab; 5): Labium pale blue, base of mandible black, tip pale; labrum shiny black; anteclypeus pale blue, postclypeus entirely dark metallic green, gena green, frons deeply sunk between antennal bases, angulated with indication of a frontal rim; occipital rim prolonged laterally into a curved carina running in direction to antennal base and stopping at level of anterior end of lateral ocellus; frons, vertex and top of head dark metallic green, black along inner border of compound eyes, antenna with scape pedicel and flagellum brown. Postocular spots and mark between lateral ocellus and antennal base absent. Rear of head pale green. Com- pound eyes in life dorsally aquamarine blue, below pale blue to lime green ventrally. Postocular lobe not surpassing compound eye posteriorly, occiput convex.
</paragraph>
<caption id="60B86675FFB9071224A6F865FE2BECEA" httpUri="https://zenodo.org/record/3368149/files/figure.png" pageId="4" pageNumber="236" startId="4.[151,250,1928,1953]" targetBox="[151,1437,561,1909]" targetPageId="4">
<paragraph id="347836FDFFB9071224A6F865FE2BECEA" blockId="4.[151,1436,1928,2025]" pageId="4" pageNumber="236">
<emphasis id="06B3EAEFFFB9071224A6F865FEDDECA3" bold="true" box="[151,275,1928,1953]" pageId="4" pageNumber="236">FIGURE 3.</emphasis>
<taxonomicName id="F3C74D7EFFB907122528F864FDD4ECA3" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[281,538,1929,1953]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="06B3EAEFFFB907122528F864FDD4ECA3" box="[281,538,1929,1953]" italics="true" pageId="4" pageNumber="236">Dactylobasis demarmelsi</emphasis>
</taxonomicName>
<emphasis id="06B3EAEFFFB907122611F864FDBEECA3" bold="true" box="[544,624,1929,1953]" pageId="4" pageNumber="236">
<taxonomicNameLabel id="1D805794FFB907122611F864FDBEECA3" box="[544,624,1929,1953]" pageId="4" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) male S10, lateral view; (b) cercus, dorsal view; (c) male S10 posterior view; (d) male cercus and paraproct, left lateral view; (e) male, cercus inner view; (f) male S10, ventral view; (g) female S810, lateral view.
<emphasis id="06B3EAEFFFB9071224E2F83FFF26ECEB" box="[211,232,2002,2025]" italics="true" pageId="4" pageNumber="236">vs</emphasis>
: ventral articulated spur.
</paragraph>
</caption>
<caption id="60B86675FFB8071324A6F918FCD9EC30" httpUri="https://zenodo.org/record/3368151/files/figure.png" pageId="5" pageNumber="237" startId="5.[151,250,1781,1806]" targetBox="[166,1425,196,1751]" targetPageId="5">
<paragraph id="347836FDFFB8071324A6F918FCD9EC30" blockId="5.[151,1436,1781,1842]" pageId="5" pageNumber="237">
<emphasis id="06B3EAEFFFB8071324A6F918FEDDEC0C" bold="true" box="[151,275,1781,1806]" pageId="5" pageNumber="237">FIGURE 4.</emphasis>
<taxonomicName id="F3C74D7EFFB80713252BF918FDD5EC0F" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[282,539,1781,1805]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="237" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="06B3EAEFFFB80713252BF918FDD5EC0F" box="[282,539,1781,1805]" italics="true" pageId="5" pageNumber="237">Dactylobasis demarmelsi</emphasis>
</taxonomicName>
<emphasis id="06B3EAEFFFB807132613F91BFDBDEC0C" bold="true" box="[546,627,1782,1806]" pageId="5" pageNumber="237">
<taxonomicNameLabel id="1D805794FFB807132613F91BFDBDEC0C" box="[546,627,1782,1806]" pageId="5" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) fore and hind left wings of male; (b) fore and hind left wings of female; (c) genital ligula, ectal view; (d) latero ental view; (e) lateral view.
</paragraph>
</caption>
<paragraph id="347836FDFFB8071024F6F88DFA91EA41" blockId="5.[151,1437,1888,2022]" lastBlockId="6.[151,1437,153,431]" lastPageId="6" lastPageNumber="238" pageId="5" pageNumber="237">
<emphasis id="06B3EAEFFFB8071324F6F88DFED6EC78" box="[199,280,1888,1914]" italics="true" pageId="5" pageNumber="237">Thorax</emphasis>
. Anterior lobe of pronotum inflated and pale yellow with dark brown anterior edge, rest of dorsum of pro- and pterothorax entirely dark metallic green (
<figureCitation id="ACFC2A78FFB8071326F7F869FCC9EC9D" box="[710,775,1924,1951]" captionStart="FIGURE 2" captionStartId="3.[151,250,1589,1614]" captionTargetBox="[151,1436,227,1569]" captionTargetId="figure@3.[151,1436,227,1569]" captionTargetPageId="3" captionText="FIGURE 2. Dactylobasis demarmelsi sp. nov. (ab) male head and prothorax, lateral view; (b) female head, dorsal view; (c) thorax showing left projection of propleuron, lateral view; (d) thoracic color pattern; (e) male prothorax, dorsal view showing projections of propleuron (f) male thorax, dorsal view. Pp: Propleuron projections." httpUri="https://zenodo.org/record/3368147/files/figure.png" pageId="5" pageNumber="237">Fig. 2</figureCitation>
cd). Propleuron sky-blue globose laterally, with coniform, prominent, lateral expansion with posteriorly directed spinulose texture (
<figureCitation id="ACFC2A78FFB8071327F7F845FBD6ECC0" box="[966,1048,1960,1986]" captionStart="FIGURE 2" captionStartId="3.[151,250,1589,1614]" captionTargetBox="[151,1436,227,1569]" captionTargetId="figure@3.[151,1436,227,1569]" captionTargetPageId="3" captionText="FIGURE 2. Dactylobasis demarmelsi sp. nov. (ab) male head and prothorax, lateral view; (b) female head, dorsal view; (c) thorax showing left projection of propleuron, lateral view; (d) thoracic color pattern; (e) male prothorax, dorsal view showing projections of propleuron (f) male thorax, dorsal view. Pp: Propleuron projections." httpUri="https://zenodo.org/record/3368147/files/figure.png" pageId="5" pageNumber="237">Fig. 2e</figureCitation>
), pronotal hind lobe rounded, dark brown in middle and pale blue laterally. Procoxa pale blue. Mesanepisternum entirely dark metallic green with a pair of horns directed cephalad, convergent in dorsal view, and with tips curved towards dorsal carina (
<figureCitation id="ACFC2A78FFBB071020E8FF74FAE9EBB6" box="[1241,1319,153,180]" captionStart="FIGURE 2" captionStartId="3.[151,250,1589,1614]" captionTargetBox="[151,1436,227,1569]" captionTargetId="figure@3.[151,1436,227,1569]" captionTargetPageId="3" captionText="FIGURE 2. Dactylobasis demarmelsi sp. nov. (ab) male head and prothorax, lateral view; (b) female head, dorsal view; (c) thorax showing left projection of propleuron, lateral view; (d) thoracic color pattern; (e) male prothorax, dorsal view showing projections of propleuron (f) male thorax, dorsal view. Pp: Propleuron projections." httpUri="https://zenodo.org/record/3368147/files/figure.png" pageId="6" pageNumber="238">Fig. 2f</figureCitation>
), in lateral view depressed; middorsal carina black. Dorsal third of mesokatepisternum dark metallic green, remainder pale green to skyblue. Mesanepimeron with dark metallic green not reaching interpleural suture except at dorsal fifth; metanepisternum sky blue, with ventrally incomplete, shiny brown black stripe along metapleural suture connecting above with black antealar carina above. Remainder of thorax sky-blue. Axillar and humeral sclerites skyblue.
</paragraph>
<paragraph id="347836FDFFBB071024F6FEA0FE05EAAD" blockId="6.[151,1437,153,431]" pageId="6" pageNumber="238">Legs short, pale, with tips of femora, tibiae and tarsomeres dark brown, hind femur with 45 spurs, hind tibia with 6 spurs externally, each of which are slightly shorter than intervals separating them; tarsal claws without tooth, but with very low swelling.</paragraph>
<caption id="60B86675FFBB071024A6F9C3FCBFED68" httpUri="https://zenodo.org/record/3368153/files/figure.png" pageId="6" pageNumber="238" startId="6.[151,250,1582,1607]" targetBox="[151,1436,453,1563]" targetPageId="6">
<paragraph id="347836FDFFBB071024A6F9C3FCBFED68" blockId="6.[151,1436,1582,1643]" pageId="6" pageNumber="238">
<emphasis id="06B3EAEFFFBB071024A6F9C3FEDDED45" bold="true" box="[151,275,1582,1607]" pageId="6" pageNumber="238">FIGURE 5.</emphasis>
<taxonomicName id="F3C74D7EFFBB07102528F9C2FDD4ED45" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[281,538,1583,1607]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="06B3EAEFFFBB07102528F9C2FDD4ED45" box="[281,538,1583,1607]" italics="true" pageId="6" pageNumber="238">Dactylobasis demarmelsi</emphasis>
</taxonomicName>
<emphasis id="06B3EAEFFFBB07102610F9C2FDBFED45" bold="true" box="[545,625,1583,1607]" pageId="6" pageNumber="238">
<taxonomicNameLabel id="1D805794FFBB07102610F9C2FDBFED45" box="[545,625,1583,1607]" pageId="6" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male at Quibdó, Salero, Chocó, Colombia. Unión Panamericana (5° 2200.17 N 76° 3603.11W) (Chocó, Colombia), 28 January 2013. Photo by LP.
</paragraph>
</caption>
<paragraph id="347836FDFFBB071024F6F977FAC3EC41" blockId="6.[151,1437,1689,2040]" pageId="6" pageNumber="238">
<emphasis id="06B3EAEFFFBB071024F6F977FEC5EDB1" box="[199,267,1690,1715]" italics="true" pageId="6" pageNumber="238">Wings</emphasis>
(
<figureCitation id="ACFC2A78FFBB0710252AF974FE93EDB6" box="[283,349,1689,1716]" captionStart="FIGURE 4" captionStartId="5.[151,250,1781,1806]" captionTargetBox="[166,1425,196,1751]" captionTargetId="figure@5.[151,1437,181,1760]" captionTargetPageId="5" captionText="FIGURE 4. Dactylobasis demarmelsi sp. nov. (a) fore and hind left wings of male; (b) fore and hind left wings of female; (c) genital ligula, ectal view; (d) latero ental view; (e) lateral view." httpUri="https://zenodo.org/record/3368151/files/figure.png" pageId="6" pageNumber="238">Fig. 4</figureCitation>
ab) narrow, hyaline, veins brown; 11 complete postnodals plus one incomplete (same in Fw and Hw), costal side of quadrangle less than half as long as distal side; vein that descends from quadrangle straight to posterior margin of wing; three cells between quadrangle and vein descending from subnodus, CuA extending 7 cells distal to vein descending from subnodus, IR1 branching at 10th Pxl, RP2 at 6
<superScript id="C3B29BB5FFBB07102009F8E9FB8BEC10" attach="left" box="[1080,1093,1796,1810]" fontSize="6" pageId="6" pageNumber="238">th</superScript>
Px, pterostigma pale brown, a parallelogram, from slightly shorter to slightly longer than underlying cell; petiolation ceasing at CuP.
</paragraph>
<paragraph id="347836FDFFBB071124F6F8A0FE4BEBD5" blockId="6.[151,1437,1689,2040]" lastBlockId="7.[151,1437,153,360]" lastPageId="7" lastPageNumber="239" pageId="6" pageNumber="238">
<emphasis id="06B3EAEFFFBB071024F6F8A0FEFFEC65" box="[199,305,1869,1895]" italics="true" pageId="6" pageNumber="238">Abdomen</emphasis>
. Dorsally shiny black, S12 laterally blue, S38 with bluish white basal ring interrupted by black at dorsal carina; S37 lateroventrally pale brown to bluish (S7), black on apical sixth, S8 blue laterally, S9 and 10 mostly black, postero-dorsal margins of S79 with robust denticles. Pruinescence lacking. Cercus and paraproct brown with dark tip (
<figureCitation id="ACFC2A78FFBB071025B5F854FE18ECD1" box="[388,470,1977,2003]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1953]" captionTargetBox="[151,1437,561,1909]" captionTargetId="figure@4.[151,1437,561,1909]" captionTargetPageId="4" captionText="FIGURE 3. Dactylobasis demarmelsi sp. nov. (a) male S10, lateral view; (b) cercus, dorsal view; (c) male S10 posterior view; (d) male cercus and paraproct, left lateral view; (e) male, cercus inner view; (f) male S10, ventral view; (g) female S810, lateral view. vs: ventral articulated spur." httpUri="https://zenodo.org/record/3368149/files/figure.png" pageId="6" pageNumber="238">Fig. 3a</figureCitation>
). Cercus short and ventrally concave (
<figureCitation id="ACFC2A78FFBB071027B2F854FBD9ECD6" box="[899,1047,1977,2004]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1953]" captionTargetBox="[151,1437,561,1909]" captionTargetId="figure@4.[151,1437,561,1909]" captionTargetPageId="4" captionText="FIGURE 3. Dactylobasis demarmelsi sp. nov. (a) male S10, lateral view; (b) cercus, dorsal view; (c) male S10 posterior view; (d) male cercus and paraproct, left lateral view; (e) male, cercus inner view; (f) male S10, ventral view; (g) female S810, lateral view. vs: ventral articulated spur." httpUri="https://zenodo.org/record/3368149/files/figure.png" pageId="6" pageNumber="238">Fig. 3b and e</figureCitation>
), a fleshy to moderately chitinized digitiform ventrobasal spur, articulated to a ventral membranous area at base of cercus (
<figureCitation id="ACFC2A78FFBB07102053F830FB28ECFA" box="[1122,1254,2013,2040]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1953]" captionTargetBox="[151,1437,561,1909]" captionTargetId="figure@4.[151,1437,561,1909]" captionTargetPageId="4" captionText="FIGURE 3. Dactylobasis demarmelsi sp. nov. (a) male S10, lateral view; (b) cercus, dorsal view; (c) male S10 posterior view; (d) male cercus and paraproct, left lateral view; (e) male, cercus inner view; (f) male S10, ventral view; (g) female S810, lateral view. vs: ventral articulated spur." httpUri="https://zenodo.org/record/3368149/files/figure.png" pageId="6" pageNumber="238">Fig. 3a, c, e</figureCitation>
); paraproct long with short, thick ventral process at half its length, basally wide and cylindrical, straight, directed caudad and dorsad towards incurved tip.
</paragraph>
<paragraph id="347836FDFFBA071124F6FF0CFB22EA1D" blockId="7.[151,1437,153,360]" pageId="7" pageNumber="239">
<emphasis id="06B3EAEFFFBA071124F6FF0CFEADEBF9" box="[199,355,225,251]" italics="true" pageId="7" pageNumber="239">Genital ligula</emphasis>
. Shaft of genital ligula with 57 soft setae on each side, medial segment broad, inner fold absent, external fold well developed, apex entire with distal lateral lobes triangular and pointed (
<figureCitation id="ACFC2A78FFBA0711204BFEE8FB72EA1D" box="[1146,1212,261,287]" captionStart="FIGURE 4" captionStartId="5.[151,250,1781,1806]" captionTargetBox="[166,1425,196,1751]" captionTargetId="figure@5.[151,1437,181,1760]" captionTargetPageId="5" captionText="FIGURE 4. Dactylobasis demarmelsi sp. nov. (a) fore and hind left wings of male; (b) fore and hind left wings of female; (c) genital ligula, ectal view; (d) latero ental view; (e) lateral view." httpUri="https://zenodo.org/record/3368151/files/figure.png" pageId="7" pageNumber="239">Fig. 4</figureCitation>
ce)
</paragraph>
<paragraph id="347836FDFFBA071124F6FEC4FD69EA65" blockId="7.[151,1437,153,360]" pageId="7" pageNumber="239">Measurements (in mm): Total length 39.4; Fw 19.6, Hw 18, abdomen (excluding appendages) 31.4, cercus 0.23 (61% longer than S10, dorsally), paraproct 1.2.</paragraph>
<caption id="60B86675FFBA071124A6F98EFCFCED9D" httpUri="https://zenodo.org/record/3368155/files/figure.png" pageId="7" pageNumber="239" startId="7.[151,250,1635,1660]" targetBox="[340,1247,381,1610]" targetPageId="7">
<paragraph id="347836FDFFBA071124A6F98EFCFCED9D" blockId="7.[151,1436,1635,1696]" pageId="7" pageNumber="239">
<emphasis id="06B3EAEFFFBA071124A6F98EFEDDED7E" bold="true" box="[151,275,1635,1660]" pageId="7" pageNumber="239">FIGURE 6.</emphasis>
<taxonomicName id="F3C74D7EFFBA0711252BF98EFDD4ED79" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[282,538,1635,1659]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="06B3EAEFFFBA0711252BF98EFDD4ED79" box="[282,538,1635,1659]" italics="true" pageId="7" pageNumber="239">Dactylobasis demarmelsi</emphasis>
</taxonomicName>
<emphasis id="06B3EAEFFFBA07112610F98EFDBCED79" bold="true" box="[545,626,1635,1659]" pageId="7" pageNumber="239">
<taxonomicNameLabel id="1D805794FFBA07112610F98EFDBCED79" box="[545,626,1635,1659]" pageId="7" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female at Salero, Unión Panamericana, Chocó, Colombia. (5° 2200.17N 76° 3603.11 W) (Chocó, Colombia), 28 January 2013. Photo by LP.
</paragraph>
</caption>
<paragraph id="347836FDFFBA071124F6F923FE6AECC2" blockId="7.[151,1436,1741,2020]" pageId="7" pageNumber="239">
<emphasis id="06B3EAEFFFBA071124F6F923FE50EDEA" bold="true" box="[199,414,1741,1768]" pageId="7" pageNumber="239">
Female (
<typeStatus id="EB7C885FFFBA0711251CF920FE5BEDEA" box="[301,405,1741,1768]" pageId="7" pageNumber="239" type="allotype">Allotype</typeStatus>
)
</emphasis>
(
<figureCitation id="ACFC2A78FFBA0711259CF923FE3CEDEB" box="[429,498,1742,1769]" captionStart="FIGURE 6" captionStartId="7.[151,250,1635,1660]" captionTargetBox="[340,1247,381,1610]" captionTargetId="figure@7.[340,1247,381,1610]" captionTargetPageId="7" captionText="FIGURE 6. Dactylobasis demarmelsi sp. nov., female at Salero, Unión Panamericana, Chocó, Colombia. (5° 2200.17N 76° 3603.11 W) (Chocó, Colombia), 28 January 2013. Photo by LP." httpUri="https://zenodo.org/record/3368155/files/figure.png" pageId="7" pageNumber="239">Fig. 6</figureCitation>
). General appearance as in male, but larger; no sexual dimorphism by color detectable, but abdomen with pale zones more expanded; S67 remarkably bright blue in lateral view, as well as sternum of S8; external valve of ovipositor blue in base towards middle of segment and black in tip. Pronotal hind lobe convex and low, propleuron as in male, the mesepisternal horns slightly more divergent in comparison with those of male. Posterior margin of S10 complete. Ovipositor surpassing tip of cercus, external valve finely serrate and setose (710 setae) along ventral border (
<figureCitation id="ACFC2A78FFBA0711262BF86FFDA3EC9E" box="[538,621,1922,1948]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1953]" captionTargetBox="[151,1437,561,1909]" captionTargetId="figure@4.[151,1437,561,1909]" captionTargetPageId="4" captionText="FIGURE 3. Dactylobasis demarmelsi sp. nov. (a) male S10, lateral view; (b) cercus, dorsal view; (c) male S10 posterior view; (d) male cercus and paraproct, left lateral view; (e) male, cercus inner view; (f) male S10, ventral view; (g) female S810, lateral view. vs: ventral articulated spur." httpUri="https://zenodo.org/record/3368149/files/figure.png" pageId="7" pageNumber="239">
Fig.
<quantity id="F33F9B18FFBA0711267FF86FFDA3EC9E" box="[590,621,1922,1948]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="7" pageNumber="239" unit="g" value="3.0">3g</quantity>
</figureCitation>
), styli as long as ventral margin of S10. Wing venation: postnodals 12+
<quantity id="F33F9B18FFBA071121A3F86FFF63ECC2" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="239" unit="in" value="1.0">1 in</quantity>
HW and 11+
<quantity id="F33F9B18FFBA07112576F84BFEA0ECC2" box="[327,366,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="239" unit="in" value="1.0">1 in</quantity>
FW.
</paragraph>
<paragraph id="347836FDFFBA071124F6F827FC10ECE6" blockId="7.[151,1436,1741,2020]" box="[199,990,1994,2020]" pageId="7" pageNumber="239">Measurements (in mm). Total length 42; abdomen 34; FW 23; HW 22.</paragraph>
<paragraph id="347836FDFFB5071E24F6FF75FDF7EA40" blockId="8.[151,1437,152,575]" pageId="8" pageNumber="240">
<emphasis id="06B3EAEFFFB5071E24F6FF75FE2EEBB1" bold="true" box="[199,480,152,179]" pageId="8" pageNumber="240">
Variation in
<typeStatus id="EB7C885FFFB5071E2552FF74FE15EBB1" box="[355,475,153,179]" pageId="8" pageNumber="240" type="paratype">paratypes</typeStatus>
.
</emphasis>
Some specimens show variation in the degree of inclination of the horns over the prothorax, but no major variation was detectable in the
<specimenCount id="22C1FD74FFB5071E2722FF51FC62EBD4" box="[787,940,188,214]" pageId="8" pageNumber="240" type="generic">46 specimens</specimenCount>
than I examined. In summary as follow; in males: Total length 38.239.3; Abdomen 3031; HW 18.5; 19.2. Females: Total length 4042; abdomen 32.334; FW21.522.7; HW 2021.8. CuA extending 8 cells distal to vein descending from subnodus (
<figureCitation id="ACFC2A78FFB5071E2095FEE9FB29EA1D" box="[1188,1255,260,287]" captionStart="FIGURE 4" captionStartId="5.[151,250,1781,1806]" captionTargetBox="[166,1425,196,1751]" captionTargetId="figure@5.[151,1437,181,1760]" captionTargetPageId="5" captionText="FIGURE 4. Dactylobasis demarmelsi sp. nov. (a) fore and hind left wings of male; (b) fore and hind left wings of female; (c) genital ligula, ectal view; (d) latero ental view; (e) lateral view." httpUri="https://zenodo.org/record/3368151/files/figure.png" pageId="8" pageNumber="240">Fig. 4</figureCitation>
ab). Coloration patterns without variation noticeable.
</paragraph>
<paragraph id="347836FDFFB5071E24F6FEA0FE43EA64" blockId="8.[151,1437,152,575]" box="[199,397,332,359]" pageId="8" pageNumber="240">Larva. Unknown.</paragraph>
<paragraph id="347836FDFFB5071E24F6FE9DFCA5EAF5" blockId="8.[151,1437,152,575]" pageId="8" pageNumber="240">
<emphasis id="06B3EAEFFFB5071E24F6FE9DFE77EA89" bold="true" box="[199,441,368,395]" pageId="8" pageNumber="240">Habitats and habits.</emphasis>
Both localities are located within pluvial tropical forest. Adults were relatively abundant mostly perching below leaves of shrubs near marshes within riparian forest (
<figureCitation id="ACFC2A78FFB5071E2039FE79FB94EAAD" box="[1032,1114,404,431]" captionStart="FIGURE 7" captionStartId="8.[151,250,1903,1928]" captionTargetBox="[160,1427,609,1877]" captionTargetId="figure@8.[151,1433,596,1882]" captionTargetPageId="8" captionText="FIGURE 7. Dactylobasis demarmelsi sp. nov., (a) Habitus female, resting in dorsal view; (b) Panoramic view of small streams inside the secondary forest in Parcela Permanente para estudios de Biodiversidad. Salero-Unión Panamericana (Choco- Colombia), semi pristine habitat of Dactylobasis demarmelsi. Photo By LP" httpUri="https://zenodo.org/record/3368157/files/figure.png" pageId="8" pageNumber="240">Fig. 7a</figureCitation>
) or flying slowly from near ground level up to
<quantity id="F33F9B18FFB5071E2558FE54FE5DEAD1" box="[361,403,441,467]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="240" unit="m" value="3.0">3 m</quantity>
near the trees, males and females fly slowly in densely arboreal areas close to small streams, slow current and with abundant retained plant material (
<figureCitation id="ACFC2A78FFB5071E273BFE31FC90EAF5" box="[778,862,476,503]" captionStart="FIGURE 7" captionStartId="8.[151,250,1903,1928]" captionTargetBox="[160,1427,609,1877]" captionTargetId="figure@8.[151,1433,596,1882]" captionTargetPageId="8" captionText="FIGURE 7. Dactylobasis demarmelsi sp. nov., (a) Habitus female, resting in dorsal view; (b) Panoramic view of small streams inside the secondary forest in Parcela Permanente para estudios de Biodiversidad. Salero-Unión Panamericana (Choco- Colombia), semi pristine habitat of Dactylobasis demarmelsi. Photo By LP" httpUri="https://zenodo.org/record/3368157/files/figure.png" pageId="8" pageNumber="240">Fig. 7b</figureCitation>
).
</paragraph>
<paragraph id="347836FDFFB5071E24F6FDEDFF16E93D" blockId="8.[151,1437,152,575]" pageId="8" pageNumber="240">
<emphasis id="06B3EAEFFFB5071E24F6FDEDFEAFE919" bold="true" box="[199,353,512,539]" pageId="8" pageNumber="240">Distribution.</emphasis>
This species is probably endemic to the Biogeographic Province
<collectingRegion id="F603F81FFFB5071E205AFDEDFB79E918" box="[1131,1207,512,538]" country="Colombia" name="Choco" pageId="8" pageNumber="240">Chocó</collectingRegion>
, Colombian Pacific coast.
</paragraph>
</subSubSection>
</treatment>
</document>