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<document id="61613C987EA5EFBE7F1073BDE460A95D" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626287244211" checkinUser="felipe" docAuthor="Phillips-Rodríguez, Eugenie &amp; Powell, Jerry A." docDate="2007" docId="7464879A1F4CFFCDFF48FB7C3907DF6D" docLanguage="en" docName="zt01670p109.pdf" docOrigin="Zootaxa 1670 (1670)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Amorbia exustana" docType="treatment" docVersion="2" lastPageNumber="34" masterDocId="885DFFE21F6CFFECFFDFFFB33C5EDA5B" masterDocTitle="Phylogenetic relationships, systematics, and biology of the species of Amorbia Clemens (Lepidoptera: Tortricidae: Sparganothini)." masterLastPageNumber="109" masterPageNumber="1" pageNumber="33" updateTime="1699009087976" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C3A4F2878AA5E9346C916FC64C8E02ED">Phylogenetic relationships, systematics, and biology of the species of Amorbia Clemens (Lepidoptera: Tortricidae: Sparganothini).</mods:title>
</mods:titleInfo>
<mods:name id="47CC30E40A26BB7D61D442AEC5054570" type="personal">
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<mods:namePart id="7FBBE69DDD856E2F71F131D5069AF3CF">Phillips-Rodríguez, Eugenie</mods:namePart>
</mods:name>
<mods:name id="1E6C48E576C73B2761D2D66FCA5434D9" type="personal">
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<mods:namePart id="7B9EB1D1686505CC67F64ED0E52FE005">Powell, Jerry A.</mods:namePart>
</mods:name>
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<mods:relatedItem id="4CA1063417BABE3FCD4492EE7DFB23BA" type="host">
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<mods:title id="DFAA4364CDC79808B3D2DA9207112255">Zootaxa</mods:title>
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<mods:part id="8355192C96E4A0619E92A398ED437886">
<mods:date id="F77F84E21B9605EDD5C1C96970DBEEDD">2007</mods:date>
<mods:detail id="307146D25A2B37785DCBE1EE2C536615" type="pubDate">
<mods:number id="B879EEE90EC394221F8EFFE2367D8EEC">2007-12-21</mods:number>
</mods:detail>
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<mods:number id="5C8AC0FB47332607BC328CA8662A9340">1670</mods:number>
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<mods:number id="35796C94281A4E9C9EAC008EA60E334C">1670</mods:number>
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<mods:classification id="5CF57D754E49F44AF8D79BCE94812880">journal article</mods:classification>
<mods:identifier id="855BEB1590BDC386D1EC041E066AB30A" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="0516D9F1E68DCC9D15145807477BC487" type="ZooBank">28818E29-DB9A-4423-8488-648C45E607EF</mods:identifier>
</mods:mods>
<treatment id="7464879A1F4CFFCDFF48FB7C3907DF6D" LSID="urn:lsid:plazi:treatment:7464879A1F4CFFCDFF48FB7C3907DF6D" httpUri="http://treatment.plazi.org/id/7464879A1F4CFFCDFF48FB7C3907DF6D" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="B4D765071F4CFFCCFF48FB7C3E12DEB2" box="[151,588,1231,1257]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="FC72368C1F4CFFCCFF48FB7C3E12DEB2" blockId="32.[151,588,1231,1292]" box="[151,588,1231,1257]" pageId="32" pageNumber="33">
<heading id="A73A81E01F4CFFCCFF48FB7C3E12DEB2" bold="true" box="[151,588,1231,1257]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="CEB9EA9E1F4CFFCCFF48FB7C3E12DEB2" bold="true" box="[151,588,1231,1257]" pageId="32" pageNumber="33">
18.
<taxonomicName id="3BCD4D0F1F4CFFCCFF1CFB7C3E12DEB2" ID-CoL="CWWP" authority="(Zeller, 1866)" baseAuthorityName="Zeller" baseAuthorityYear="1866" box="[195,588,1231,1257]" class="Insecta" family="Tortricidae" genus="Amorbia" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="exustana">
<emphasis id="CEB9EA9E1F4CFFCCFF1CFB7C3DFFDEB2" bold="true" box="[195,417,1231,1257]" italics="true" pageId="32" pageNumber="33">Amorbia exustana</emphasis>
(
<bibRefCitation id="985C4B7D1F4CFFCCFE6FFB7C3E1ADEB2" author="Zeller, P. C." box="[432,580,1231,1257]" pageId="32" pageNumber="33" pagination="137 - 157" refId="ref62738" refString="Zeller, P. C. (1866) Beschreibung einger amerikanischen Wickler und Crambien. Stettiner Entomologische Zeitung, 27, 137 - 157." type="journal article" year="1866">Zeller, 1866</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4D765071F4CFFCCFF48FB413D9CDF57" box="[151,450,1266,1292]" pageId="32" pageNumber="33" type="description">
<paragraph id="FC72368C1F4CFFCCFF48FB413D9CDF57" blockId="32.[151,588,1231,1292]" box="[151,450,1266,1292]" pageId="32" pageNumber="33">
<figureCitation id="64F62A091F4CFFCCFF48FB413D7ADF57" box="[151,292,1266,1292]" captionStart="FIGURES 41–45" captionStartId="67.[151,269,1545,1569]" captionTargetBox="[264,1326,188,1504]" captionTargetId="figure-88@67.[254,1341,146,1521]" captionTargetPageId="67" captionText="FIGURES 41–45. Adults. 41, Amorbia exustana male (Costa Rica: Cartago: Paraíso, P.N. Tapantí-Macizo de la Muerte, Costado Casa Administrativa, 1200 m, xi.1999, R. Delgado); 42, A. exustana female (Costa Rica: Alajuela: NE slope Volcán Poás, 8 km N Vara Blanca, 1400 m, 6.vi.1988, J. Brown &amp; J. Powell); 43, A. cocori male (holoype); 44, A. cocori female (paratype); 45, A. exsectana male (Brazil: Rio De Janeiro, Teresópolis 1000 m. 15.i.1985, V. Becker)." pageId="32" pageNumber="33">Figs. 41, 42</figureCitation>
,
<figureCitation id="64F62A091F4CFFCCFEEFFB413D0EDF57" box="[304,336,1266,1292]" captionStart="FIGURES 80–82" captionStartId="77.[151,269,1588,1612]" captionTargetBox="[238,1356,210,1518]" captionTargetId="figure-39@77.[201,1414,152,1564]" captionTargetPageId="77" captionText="FIGURES 80–82. Male genitalia of Amorbia, valvae spread, aedeagus removed. 80, A. exustana; 81, A. cocori; 82, A. exsectana." pageId="32" pageNumber="33">80</figureCitation>
,
<figureCitation id="64F62A091F4CFFCCFE83FB413DD4DF57" box="[348,394,1266,1292]" captionStart="FIGURES 103–105" captionStartId="85.[151,269,1851,1875]" captionTargetBox="[266,1320,192,1815]" captionTargetId="figure-35@85.[235,1351,170,1828]" captionTargetPageId="85" captionText="FIGURES 103–105. Female genitalia of Amorbia. 103, A. exustana; 104, A. cocori; 105, A. exsectana." pageId="32" pageNumber="33">103</figureCitation>
,
<figureCitation id="64F62A091F4CFFCCFE49FB413D9CDF57" box="[406,450,1266,1292]" captionStart="FIGURE 135" captionStartId="95.[151,254,1860,1884]" captionTargetBox="[317,1277,151,1846]" captionTargetId="figure-83@95.[317,1315,105,1864]" captionTargetPageId="95" captionText="FIGURE 135. Geographical distribution of Amorbia. A. laterculana: open circles; A. knudsoni: filled oval; A. osmotris: triangle; A. colubrana: open diamonds; A. cacao: filled diamond; A. nuptana: solid squares; A. exustana: solid circles; A. cocori: open square." pageId="32" pageNumber="33">135</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B4D765071F4CFFCCFF48FAF13FB7DF22" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="FC72368C1F4CFFCCFF48FAF13F4FDF02" blockId="32.[151,1001,1346,1401]" box="[151,785,1346,1369]" pageId="32" pageNumber="33">
<taxonomicName id="3BCD4D0F1F4CFFCCFF48FAF13D8EDF02" ID-CoL="98LLQ" authority="Zeller, 1866" authorityName="Zeller" authorityYear="1866" box="[151,464,1346,1369]" class="Insecta" family="Tortricidae" genus="Tortrix" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="exustana">
<emphasis id="CEB9EA9E1F4CFFCCFF48FAF13D1BDF02" box="[151,325,1346,1369]" italics="true" pageId="32" pageNumber="33">Tortrix exustana</emphasis>
<bibRefCitation id="985C4B7D1F4CFFCCFE93FAF13D8EDF02" author="Zeller, P. C." box="[332,464,1346,1369]" pageId="32" pageNumber="33" pagination="137 - 157" refId="ref62738" refString="Zeller, P. C. (1866) Beschreibung einger amerikanischen Wickler und Crambien. Stettiner Entomologische Zeitung, 27, 137 - 157." type="journal article" year="1866">Zeller, 1866</bibRefCitation>
</taxonomicName>
, Stett. Entomol. Zeit. 27: 146.
</paragraph>
<paragraph id="FC72368C1F4CFFCCFF48FAD13FB7DF22" blockId="32.[151,1001,1346,1401]" box="[151,1001,1378,1401]" pageId="32" pageNumber="33">
<taxonomicName id="3BCD4D0F1F4CFFCCFF48FAD13D07DF22" ID-CoL="CWWP" baseAuthorityName="Zeller" baseAuthorityYear="1866" box="[151,345,1378,1401]" class="Insecta" family="Tortricidae" genus="Amorbia" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="exustana">
<emphasis id="CEB9EA9E1F4CFFCCFF48FAD13D07DF22" box="[151,345,1378,1401]" italics="true" pageId="32" pageNumber="33">Amorbia exustana</emphasis>
</taxonomicName>
;
<bibRefCitation id="985C4B7D1F4CFFCCFEB8FAD13E6FDF22" author="Powell, J. A. &amp; Razowski, J. &amp; Brown, J. W." box="[359,561,1378,1401]" pageId="32" pageNumber="33" pagination="138 - 150" refId="ref62377" refString="Powell, J. A., Razowski, J. and Brown, J. W. (1995) Tortricidae: Tortricinae, pp. 138 - 150. In Heppner, J. B. (ed.), Atlas of Neotropical Lepidoptera, Checklist: part 2, Hyblaeoidea-Pyraloidea-Tortricoidea. Association for Tropical Lepidoptera, Gainesville, Florida." type="book chapter" year="1995">Powell et al., 1995</bibRefCitation>
, Atlas of Neotropical Lepid., part 2: 148.
</paragraph>
</subSubSection>
<subSubSection id="B4D765071F4CFFCCFF48FA193D53DC4F" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="FC72368C1F4CFFCCFF48FA193D53DC4F" blockId="32.[151,1436,1450,2036]" pageId="32" pageNumber="33">
DIAGNOSIS. This species is most similar to
<emphasis id="CEB9EA9E1F4CFFCCFD6BFA183F1CDF9F" box="[692,834,1451,1476]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="3BCD4D0F1F4CFFCCFD6BFA183F60DF9F" authorityName="Meyrick" authorityYear="1932" box="[692,830,1451,1476]" class="Insecta" family="Tortricidae" genus="Amorbia" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="osmotris">A. osmotris</taxonomicName>
,
</emphasis>
but
<taxonomicName id="3BCD4D0F1F4CFFCCFCA7FA18385CDF9F" baseAuthorityName="Zeller" baseAuthorityYear="1866" box="[888,1026,1451,1476]" class="Insecta" family="Tortricidae" genus="Amorbia" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="exustana">
<emphasis id="CEB9EA9E1F4CFFCCFCA7FA18385CDF9F" box="[888,1026,1451,1476]" italics="true" pageId="32" pageNumber="33">A. exustana</emphasis>
</taxonomicName>
can be distinguished by the modified scaling on the vertex, male forewing costal fold, and conspicuous cubital pecten at the base of Cu of the hindwing.
</paragraph>
</subSubSection>
<subSubSection id="B4D765071F4CFFCDFF19F9913F7BDB59" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="FC72368C1F4CFFCCFF19F991380ADDFF" blockId="32.[151,1436,1450,2036]" pageId="32" pageNumber="33">
<materialsCitation id="4CA53CD11F4CFFCCFF19F991380EDDFF" collectionCode="INBio" country="Costa Rica" county="Estacion Cacao" location="Valva" municipality="Transtilla" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Guanacaste">
REDESCRIPTION. MALE. FW length 10.5–11.5 mm (mean = 11.0; n = 7). HEAD: Frons concave, white scaled, vertex scaling in two groups, rust colored. Antenna with orange-brown scaling. Labial palpus at least 3.0X length of inter-antennal space, porrect, dark brown laterally, inner surface paler. THORAX: Dorsum smooth scaled, rust colored; tegula concolorous. FW ground reddish brown highly variable in intensity of color and shapes of fasciae, but basal, median and terminal fasciae dark brown. Costal fold well developed, reaching to middle of costa. HW white with gray mottling toward apex, conspicuous cubital pecten at the base of Cu. ABDOMEN: Pale brown. Genitalia as in
<figureCitation id="64F62A091F4CFFCCFD16F8A13F42DD77" box="[713,796,1810,1836]" captionStart="FIGURES 80–82" captionStartId="77.[151,269,1588,1612]" captionTargetBox="[238,1356,210,1518]" captionTargetId="figure-39@77.[201,1414,152,1564]" captionTargetPageId="77" captionText="FIGURES 80–82. Male genitalia of Amorbia, valvae spread, aedeagus removed. 80, A. exustana; 81, A. cocori; 82, A. exsectana." pageId="32" pageNumber="33">Fig. 80</figureCitation>
(slide # 1945218,
<collectionCode id="9ADCAE491F4CFFCCFC2BF8A1381EDD77" box="[1012,1088,1810,1836]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="32" pageNumber="33">INBio</collectionCode>
,
<collectingCountry id="84DA761C1F4CFFCCFB94F8A13893DD77" box="[1099,1229,1810,1836]" name="Costa Rica" pageId="32" pageNumber="33">Costa Rica</collectingCountry>
,
<collectingRegion id="3E09F86E1F4CFFCCFB07F8A1393CDD77" box="[1240,1378,1810,1836]" country="Costa Rica" name="Guanacaste" pageId="32" pageNumber="33">Guanacaste</collectingRegion>
,
<collectingCounty id="15134E001F4CFFCCFAB2F8A13D74DD0F" pageId="32" pageNumber="33">Estación Cacao</collectingCounty>
; n = 3).
<collectingMunicipality id="1C16ACF61F4CFFCCFE55F8893DA7DD0F" box="[394,505,1850,1876]" pageId="32" pageNumber="33">Transtilla</collectingMunicipality>
bluntly pointed in middle anterior, numerous fine spines over posterior margin.
<location id="F91260571F4CFFCCFF48F8D13C84DD27" LSID="urn:lsid:plazi:treatment:7464879A1F4CFFCDFF48FB7C3907DF6D:F91260571F4CFFCCFF48F8D13C84DD27" box="[151,218,1890,1916]" country="Costa Rica" county="Estacion Cacao" municipality="Transtilla" name="Valva" pageId="32" pageNumber="33" stateProvince="Guanacaste">Valva</location>
subrectangular.
<location id="F91260571F4CFFCCFE45F8D13E5CDD27" LSID="urn:lsid:plazi:treatment:7464879A1F4CFFCDFF48FB7C3907DF6D:F91260571F4CFFCCFE45F8D13E5CDD27" box="[410,514,1890,1916]" country="Costa Rica" county="Estacion Cacao" municipality="Transtilla" name="Sacculus" pageId="32" pageNumber="33" stateProvince="Guanacaste">Sacculus</location>
sclerotized, 0.65X length of valva, emarginate in basal half, without free distal lobe.
<location id="F91260571F4CFFCCFF07F8393D15DDFF" LSID="urn:lsid:plazi:treatment:7464879A1F4CFFCDFF48FB7C3907DF6D:F91260571F4CFFCCFF07F8393D15DDFF" box="[216,331,1930,1956]" country="Costa Rica" county="Estacion Cacao" municipality="Transtilla" name="Aedeagus" pageId="32" pageNumber="33" stateProvince="Guanacaste">Aedeagus</location>
0.66X length of valva, slender, curved at 0.66 distance from base
</materialsCitation>
.
</paragraph>
<paragraph id="FC72368C1F4CFFCDFF19F8013F7BDB59" blockId="32.[151,1436,1450,2036]" lastBlockId="33.[151,1436,152,1334]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<materialsCitation id="4CA53CD11F4CFFCDFF19F8013F7FDB59" collectionCode="INBio" country="Costa Rica" county="Orosi" lastPageId="33" lastPageNumber="34" location="Ductus" municipality="Sterigma" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Cartago">
FEMALE. FW length 12.0–13.0 mm (mean = 12.5; n = 10). HEAD and THORAX: As described for male, except patch of cream colored scales in center of FW. ABDOMEN: Genitalia as in
<figureCitation id="64F62A091F4CFFCCFB0BF8693966DDAF" box="[1236,1336,2010,2036]" captionStart="FIGURES 103–105" captionStartId="85.[151,269,1851,1875]" captionTargetBox="[266,1320,192,1815]" captionTargetId="figure-35@85.[235,1351,170,1828]" captionTargetPageId="85" captionText="FIGURES 103–105. Female genitalia of Amorbia. 103, A. exustana; 104, A. cocori; 105, A. exsectana." pageId="32" pageNumber="33">Fig. 103</figureCitation>
(slide # 2427538,
<collectionCode id="9ADCAE491F4DFFCDFED1FF2B3D04DAE9" box="[270,346,152,178]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
,
<collectingCountry id="84DA761C1F4DFFCDFEB7FF2B3DB3DAE9" box="[360,493,152,178]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
,
<collectingRegion id="3E09F86E1F4DFFCDFE25FF2B3E07DAE9" box="[506,601,152,178]" country="Costa Rica" name="Cartago" pageId="33" pageNumber="34">Cartago</collectingRegion>
,
<collectingCounty id="15134E001F4DFFCDFDB8FF2B3EF2DAE9" box="[615,684,152,178]" pageId="33" pageNumber="34">Orosi</collectingCounty>
; n = 2).
<collectingMunicipality id="1C16ACF61F4DFFCDFCCBFF2B3F23DAE9" box="[788,893,152,178]" pageId="33" pageNumber="34">Sterigma</collectingMunicipality>
developed into small lateral pockets, pointed anteriorly; median sinus rounded.
<location id="F91260571F4DFFCDFDE3FF733ED1DA81" LSID="urn:lsid:plazi:treatment:7464879A1F4CFFCDFF48FB7C3907DF6D:F91260571F4DFFCDFDE3FF733ED1DA81" box="[572,655,192,218]" country="Costa Rica" county="Orosi" municipality="Sterigma" name="Ductus" pageId="33" pageNumber="34" stateProvince="Cartago">Ductus</location>
bursae 2.0X length of papillae anales, conspicuously widening towards corpus bursae, sclerotized in the posterior 0.25
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4D765071F4DFFCDFF19FEA33DC4DE1C" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="FC72368C1F4DFFCDFF19FEA33FC8DB71" blockId="33.[151,1436,152,1334]" box="[198,918,272,298]" pageId="33" pageNumber="34">
<materialsCitation id="4CA53CD11F4DFFCDFF19FEA33FC8DB71" box="[198,918,272,298]" collectionCode="BMNH" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
TYPE MATERIAL.
<typeStatus id="2376882E1F4DFFCDFE65FEA33E75DB71" box="[442,555,272,298]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
: “Columbien”, 1866 (
<collectionCode id="9ADCAE491F4DFFCDFCF4FEA33FD4DB71" box="[811,906,272,298]" pageId="33" pageNumber="34">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FC72368C1F4DFFCDFF19FE8B3DC4DE1C" blockId="33.[151,1436,152,1334]" pageId="33" pageNumber="34">
ADDITIONAL MATERIAL.
<collectingCountry id="84DA761C1F4DFFCDFDF6FE8B3E87DB09" box="[553,729,312,338]" name="Costa Rica" pageId="33" pageNumber="34">COSTA RICA</collectingCountry>
:
<collectingRegion id="3E09F86E1F4DFFCDFD39FE8B3FDEDB09" box="[742,896,312,338]" country="Costa Rica" name="Alajuela" pageId="33" pageNumber="34">ALAJUELA</collectingRegion>
: NE slope Volcán Poás,
<quantity id="3B359B691F4DFFCDFB7CFE8B3881DB09" box="[1187,1247,312,338]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="34" unit="km" value="8.0">8 km</quantity>
N Vara Blanca,
<quantity id="3B359B691F4DFFCDFF48FEEC3CADDB22" box="[151,243,351,377]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="33" pageNumber="34" unit="m" value="1400.0">1400 m</quantity>
, 1F,
<date id="8873104C1F4DFFCDFEF3FEEC3DC5DB22" box="[300,411,351,377]" pageId="33" pageNumber="34" value="1988-06-06">6.vi.1988</date>
(J. Brown &amp; J. Powell,
<collectionCode id="9ADCAE491F4DFFCDFD64FEEC3EA0DB22" box="[699,766,351,377]" pageId="33" pageNumber="34">EME</collectionCode>
).
<collectingRegion id="3E09F86E1F4DFFCDFCCDFEEC3FFEDB22" box="[786,928,351,377]" country="Costa Rica" name="Cartago" pageId="33" pageNumber="34">CARTAGO</collectingRegion>
: Tapantí, Río Grande de Orosi,
<quantity id="3B359B691F4DFFCDFAC7FEEC3CEEDBFA" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="33" pageNumber="34" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400 m</quantity>
, 1M,
<date id="8873104C1F4DFFCDFF27FE343D38DBFA" box="[248,358,391,417]" pageId="33" pageNumber="34" value="1985-01-23">23.i.1985</date>
(D. Janzen &amp; W. Hallwachs,
<collectionCode id="9ADCAE491F4DFFCDFD10FE343F43DBFA" box="[719,797,391,417]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
); R. Grande de Orosi, desde Puente Río Dos Amigos hasta La Represa,
<quantity id="3B359B691F4DFFCDFEB3FE1C3E4EDB92" box="[364,528,431,457]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="33" pageNumber="34" unit="m" value="1600.0" valueMax="1800.0" valueMin="1400.0">1400–1800 m</quantity>
, LN 186600, 562000, 1F,
<date id="8873104C1F4DFFCDFC9EFE1C3FCBDB92" box="[833,917,431,457]" pageId="33" pageNumber="34" value="1995-02">ii.1995</date>
, 1F, 22.viii./
<date id="8873104C1F4DFFCDFBF5FE1C38F9DB92" box="[1066,1191,431,457]" pageId="33" pageNumber="34" value="1995-09-15">15.ix.1995</date>
, 1F,
<date id="8873104C1F4DFFCDFB02FE1C3968DB92" box="[1245,1334,431,457]" pageId="33" pageNumber="34" value="1997-11">xi.1997</date>
(R. Delgado,
<collectionCode id="9ADCAE491F4DFFCDFF3EFE643D71DBAA" box="[225,303,471,497]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
); Río Aquiares, nr. Sta. Cruz, 9 air km NW Turrialba,
<quantity id="3B359B691F4DFFCDFC1EFE64384BDBAA" box="[961,1045,471,497]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="m" value="1500.0">1500m</quantity>
, 1F,
<date id="8873104C1F4DFFCDFB8FFE6438E1DBAA" box="[1104,1215,471,497]" pageId="33" pageNumber="34" value="1988-06-08">8.vi.1988</date>
(
<collectionCode id="9ADCAE491F4DFFCDFB0FFE64395DDBAA" box="[1232,1283,471,497]" pageId="33" pageNumber="34">JAP</collectionCode>
88F26, confined, no oviposition) (J. Powell,
<collectionCode id="9ADCAE491F4DFFCDFDFCFE4C3E38D842" box="[547,614,511,537]" pageId="33" pageNumber="34">EME</collectionCode>
); Paraíso, P.N. Tapantí, Sect. La Represa,
<quantity id="3B359B691F4DFFCDFB87FE4C38FDD842" box="[1112,1187,511,537]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="34" unit="m" value="300.0">300 m</quantity>
S del puente del Río Porras,
<quantity id="3B359B691F4DFFCDFF2EFD943D1BD81A" box="[241,325,551,577]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="33" pageNumber="34" unit="m" value="1660.0">1660m</quantity>
, LN 186550, 560600, 1F,
<date id="8873104C1F4DFFCDFDAEFD943EBED81A" box="[625,736,551,577]" pageId="33" pageNumber="34" value="1998-08">viii.1998</date>
, 3F,
<date id="8873104C1F4DFFCDFCC6FD943F30D81A" box="[793,878,551,577]" pageId="33" pageNumber="34" value="1999-02">ii.1999</date>
, 1F,
<date id="8873104C1F4DFFCDFC78FD94385CD81A" box="[935,1026,551,577]" pageId="33" pageNumber="34" value="1999-06">vi.1999</date>
, 3M, 3F,
<date id="8873104C1F4DFFCDFBABFD94388ED81A" box="[1140,1232,551,577]" pageId="33" pageNumber="34" value="1999-09">ix.1999</date>
, 1M, 2F
<date id="8873104C1F4DFFCDFAE3FD9439C9D81A" box="[1340,1431,551,577]" pageId="33" pageNumber="34" value="2001-11">xi.2001</date>
, 1M,
<date id="8873104C1F4DFFCDFF0FFDFC3D73D832" box="[208,301,591,617]" pageId="33" pageNumber="34" value="2002-03">iii.2002</date>
, 2F,
<date id="8873104C1F4DFFCDFEB7FDFC3DE6D832" box="[360,440,591,617]" pageId="33" pageNumber="34" value="2002-10">x.2002</date>
(R. Delgado,
<collectionCode id="9ADCAE491F4DFFCDFDBCFDFC3EEFD832" box="[611,689,591,617]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
); Paraíso, Parque Nacional Tapantí-Macizo de la Muerte, Costado Casa Administrativa 1F,
<quantity id="3B359B691F4DFFCDFE27FDC53E08D8CB" box="[504,598,630,656]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="1200.0">1200 m</quantity>
,
<date id="8873104C1F4DFFCDFDBCFDC53EE5D8CB" box="[611,699,630,656]" pageId="33" pageNumber="34" value="1999-11">xi.1999</date>
(R. Delgado,
<collectionCode id="9ADCAE491F4DFFCDFCBAFDC53FEDD8CB" box="[869,947,630,656]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
). Tapantí,
<quantity id="3B359B691F4DFFCDFBF0FDC5388BD8CB" box="[1071,1237,630,656]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="33" pageNumber="34" unit="m" value="1450.0" valueMax="1700.0" valueMin="1200.0">1200–1700 m</quantity>
, 1M, 1F, 20.viii/
<date id="8873104C1F4DFFCDFF48FD2D3D4AD8E3" box="[151,276,670,696]" pageId="33" pageNumber="34" value="1999-09-15">15.ix.1999</date>
(V. Becker,
<collectionCode id="9ADCAE491F4DFFCDFE74FD2D3DB2D8E3" box="[427,492,670,696]" pageId="33" pageNumber="34">VBC</collectionCode>
). P.N. Tapantí,
<quantity id="3B359B691F4DFFCDFD7CFD2D3EA1D8E3" box="[675,767,670,696]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="34" unit="m" value="1150.0">1150 m</quantity>
, 1F,
<date id="8873104C1F4DFFCDFCE5FD2D3FD5D8E3" box="[826,907,670,696]" pageId="33" pageNumber="34" value="1994-10">x.1994</date>
(R. Delgado,
<collectionCode id="9ADCAE491F4DFFCDFBEAFD2D38DDD8E3" box="[1077,1155,670,696]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
). Tapantí, 1F,
<date id="8873104C1F4DFFCDFAF2FD2D39C5D8E3" box="[1325,1435,670,696]" pageId="33" pageNumber="34" value="1999-09-09">9.ix.1999</date>
(V. Becker,
<collectionCode id="9ADCAE491F4DFFCDFEFAFD753D2ED8BB" box="[293,368,710,736]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
) (1M, r.
<taxonomicName id="3BCD4D0F1F4DFFCDFE0BFD753E7BD8BB" box="[468,549,710,736]" form="eggs" pageId="33" pageNumber="34" rank="form">f. eggs</taxonomicName>
on synthetic diet, em: i.
<date id="8873104C1F4DFFCDFC99FD753FF2D8BB" box="[838,940,710,736]" pageId="33" pageNumber="34" value="1999-12">xii.1999</date>
, EPR-103-99,
<collectionCode id="9ADCAE491F4DFFCDFB81FD7538F2D8BB" box="[1118,1196,710,736]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
). Tapantí, Quebrada Segunda,
<quantity id="3B359B691F4DFFCDFED0FD5D3D30D953" box="[271,366,750,776]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="34" unit="m" value="1300.0">1300 m</quantity>
,
<date id="8873104C1F4DFFCDFEA0FD5D3D89D953" box="[383,471,750,776]" pageId="33" pageNumber="34" value="2000-11">xi.2000</date>
(R. Delgado,
<collectionCode id="9ADCAE491F4DFFCDFD55FD5D3E86D953" box="[650,728,750,776]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
).
<collectingRegion id="3E09F86E1F4DFFCDFD2FFD5D3F2BD953" box="[752,885,750,776]" country="Costa Rica" name="Heredia" pageId="33" pageNumber="34">HEREDIA</collectingRegion>
:
<quantity id="3B359B691F4DFFCDFC5BFD5D3F9DD953" box="[900,963,750,776]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="34" unit="km" value="6.0">6 km</quantity>
ENE Vara Blanca
<quantity id="3B359B691F4DFFCDFB6CFD5D3902D953" box="[1203,1372,750,776]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="33" pageNumber="34" unit="m" value="2000.0" valueMax="2050.0" valueMin="1950.0">1950–2050 m</quantity>
, 1M,
<date id="8873104C1F4DFFCDFF48FCA53D50D96B" box="[151,270,790,816]" pageId="33" pageNumber="34" value="2002-02-17">17.ii.2002</date>
(J. Powell,
<collectionCode id="9ADCAE491F4DFFCDFE7DFCA53DBAD96B" box="[418,484,790,816]" pageId="33" pageNumber="34">EME</collectionCode>
), 1M,
<date id="8873104C1F4DFFCDFDE9FCA53EF3D96B" box="[566,685,790,816]" pageId="33" pageNumber="34" value="2002-12-07">7.xii.2002</date>
(K. Nishida,
<collectionCode id="9ADCAE491F4DFFCDFC8AFCA53FF1D96B" box="[853,943,790,816]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34" type="Museum">USNM</collectionCode>
).
<collectingRegion id="3E09F86E1F4DFFCDFC19FCA538CFD96B" box="[966,1169,790,816]" country="Costa Rica" name="Guanacaste" pageId="33" pageNumber="34">GUANACASTE</collectingRegion>
: Estación Cacao, lado sureste del Volcán Cacao,
<quantity id="3B359B691F4DFFCDFE14FC8D3E78D903" box="[459,550,830,856]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="34" unit="m" value="1100.0">1100 m</quantity>
, LN 323300, 375700, 1M, 8/
<date id="8873104C1F4DFFCDFCA1FC8D3FA5D903" box="[894,1019,830,856]" pageId="33" pageNumber="34" value="1991-06-30">30.vi.1991</date>
(C. Chaves,
<collectionCode id="9ADCAE491F4DFFCDFB4BFC8D38BCD903" box="[1172,1250,830,856]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
); Fila Orosilito, Est. Pitilla,
<quantity id="3B359B691F4DFFCDFEFEFCD53D02D9DB" box="[289,348,870,896]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="km" value="9.0">9 km</quantity>
S Santa Cecilia,
<quantity id="3B359B691F4DFFCDFDF8FCD53EE3D9DB" box="[551,701,870,896]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="33" pageNumber="34" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</quantity>
, LN 328650, 378600, 1M,
<date id="8873104C1F4DFFCDFBDFFCD53808D9DB" box="[1024,1110,870,896]" pageId="33" pageNumber="34" value="1995-04">iv.1995</date>
(C. Moraga,
<collectionCode id="9ADCAE491F4DFFCDFB29FCD5391AD9DB" box="[1270,1348,870,896]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
). PUN-
<collectingRegion id="3E09F86E1F4DFFCDFF48FC3E3D7FD9FC" box="[151,289,909,935]" country="Costa Rica" name="TARENAS" pageId="33" pageNumber="34">TARENAS</collectingRegion>
: Monteverde
<quantity id="3B359B691F4DFFCDFE61FC3E3E46D9FC" box="[446,536,909,935]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="m" value="1500.0">1500 m</quantity>
, 1M, 1/
<date id="8873104C1F4DFFCDFDB0FC3E3EBED9FC" box="[623,736,909,935]" pageId="33" pageNumber="34" value="1999-09-04">4.ix.1999</date>
(V. Becker,
<collectionCode id="9ADCAE491F4DFFCDFCAFFC3E3FEFD9FC" box="[880,945,909,935]" pageId="33" pageNumber="34">VBC</collectionCode>
).
<collectingRegion id="3E09F86E1F4DFFCDFC1BFC3E3812D9FC" box="[964,1100,909,935]" country="Costa Rica" name="San Jose" pageId="33" pageNumber="34">SAN JOSE</collectingRegion>
: San Gerardo de Dota, Cerro de la Muerte, 3M,
<date id="8873104C1F4DFFCDFEA7FC063DB1D994" box="[376,495,949,975]" pageId="33" pageNumber="34" value="1996-02-20">20.ii.1996</date>
(D. &amp; J. Powell,
<collectionCode id="9ADCAE491F4DFFCDFD1AFC063F56D994" box="[709,776,949,975]" pageId="33" pageNumber="34">EME</collectionCode>
). ACLAP [Area de Conservación la Amistad Pacífico], Parque Internacional La Amistad, Sector Altamira, Cerro Frantzius, Sendero al Valle del Silencio, LS 334200N 574 500E,
<quantity id="3B359B691F4DFFCDFE55FBB63DBBDE44" box="[394,485,1029,1055]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="33" pageNumber="34" unit="m" value="2100.0">2100 m</quantity>
, 1M, 8/
<date id="8873104C1F4DFFCDFDE1FBB63EEADE44" box="[574,692,1029,1055]" pageId="33" pageNumber="34" value="2004-10-20">20.x.2004</date>
(voucher number RGT-00766.1, pupa on
<taxonomicName id="3BCD4D0F1F4DFFCDFB44FBB63946DE44" box="[1179,1304,1029,1055]" class="Bivalvia" family="Pharidae" genus="Pharus" kingdom="Animalia" order="Poales" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="CEB9EA9E1F4DFFCDFB44FBB638AEDE45" box="[1179,1264,1029,1054]" italics="true" pageId="33" pageNumber="34">Pharus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="3BCD4D0F1F4DFFCDFAF8FBB639D4DE44" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1319,1418,1029,1055]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
)) (R. Gonzalez,
<collectionCode id="9ADCAE491F4DFFCDFE9FFB9E3DD0DE1C" box="[320,398,1069,1095]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="34">INBio</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B4D765071F4DFFCDFF19FBE63907DF6D" pageId="33" pageNumber="34" type="distribution">
<paragraph id="FC72368C1F4DFFCDFF19FBE63E73DEE4" blockId="33.[151,1436,152,1334]" pageId="33" pageNumber="34">
DISTRIBUTION AND BIOLOGY. This species is known from
<collectingCountry id="84DA761C1F4DFFCDFC67FBE63872DE34" box="[952,1068,1109,1135]" name="Colombia" pageId="33" pageNumber="34">Colombia</collectingCountry>
and
<collectingCountry id="84DA761C1F4DFFCDFBBBFBE638BDDE34" box="[1124,1251,1109,1135]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
where it occurs from
<quantity id="3B359B691F4DFFCDFF06FBCE3DD9DECC" box="[217,391,1149,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="33" pageNumber="34" unit="m" value="1400.0" valueMax="2000.0" valueMin="800.0">800 to 2000 m</quantity>
elevation (
<figureCitation id="64F62A091F4DFFCDFDD2FBCE3E2BDECC" box="[525,629,1149,1175]" captionStart="FIGURE 135" captionStartId="95.[151,254,1860,1884]" captionTargetBox="[317,1277,151,1846]" captionTargetId="figure-83@95.[317,1315,105,1864]" captionTargetPageId="95" captionText="FIGURE 135. Geographical distribution of Amorbia. A. laterculana: open circles; A. knudsoni: filled oval; A. osmotris: triangle; A. colubrana: open diamonds; A. cacao: filled diamond; A. nuptana: solid squares; A. exustana: solid circles; A. cocori: open square." pageId="33" pageNumber="34">Fig. 135</figureCitation>
). Records show flight activity in nine months of the year, including both the dry and the rainy seasons.
</paragraph>
<paragraph id="FC72368C1F4DFFCDFF19FB7F3907DF6D" blockId="33.[151,1436,152,1334]" pageId="33" pageNumber="34">IMMATURE STAGES. Eggs are green. Larvae hatched after an average of 12 days (n = 6) and fed on synthetic diet. The head is tan with dark lateral bands; the prothoracic shield is tan with narrow lateral and dorsal dark brown bands. Development required ca. 10 weeks (E. Phillips voucher record EPR-103-99).</paragraph>
</subSubSection>
</treatment>
</document>
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