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<document ID-CLB-Dataset="154682" ID-DOI="http://dx.doi.org/10.3897/jor.31.82306" ID-GBIF-Dataset="c0b508a2-0e39-4d07-b633-48f6e32e987c" ID-Pensoft-Pub="1937-2426-2-143" ID-Pensoft-UUID="7C8205C212D35F86BC47E13AD1F0BA65" ID-ZooBank="42CAC6F064244883B0B5A81CE88A71BE" ModsDocID="1937-2426-31-2-143" checkinTime="1664560482712" checkinUser="pensoft" docAuthor="Braun, Holger &amp; Morris, Glenn K." docDate="2022" docId="6AE6E9B88E415EBAB9F5D5D3FD578F1C" docLanguage="en" docName="JourOrthoptRes 31(2): 143-156" docOrigin="Journal of Orthoptera Research 31 (2)" docPubDate="2022-09-30" docSource="http://dx.doi.org/10.3897/jor.31.82306" docTitle="Cestrophorus ditachus Braun &amp; Morris 2022, sp. nov." docType="treatment" docUuid="39B53563-0D85-42CA-8DF2-80F9C45018AB" docUuidSource="ZooBank" docVersion="5" id="7C8205C212D35F86BC47E13AD1F0BA65" lastPageNumber="143" masterDocId="7C8205C212D35F86BC47E13AD1F0BA65" masterDocTitle="New species of awl-head katydids, Cestrophorus and Acanthacara, from the Andes of Ecuador (Orthoptera, Conocephalinae, Cestrophorini)" masterLastPageNumber="156" masterPageNumber="143" pageNumber="143" updateTime="1732743010087" updateUser="ExternalLinkService">
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<mods:title id="D65862817A9C04B0FDAB9A579244C80C">New species of awl-head katydids, Cestrophorus and Acanthacara, from the Andes of Ecuador (Orthoptera, Conocephalinae, Cestrophorini)</mods:title>
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<mods:name id="8307FC548A2366B24A9F12B0A8439F29" type="personal">
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<mods:namePart id="817068142216FC8EF6F71C46AEEFC56E">Braun, Holger</mods:namePart>
<mods:nameIdentifier id="EA59514B84522B96D140DB965209D979" type="ORCID">https://orcid.org/0000-0002-1069-8794</mods:nameIdentifier>
<mods:affiliation id="28DFB4BA0486994A212395A0D3DD4DD7">Division Entomologia, Museo de La Plata, Paseo del Bosque s / n °, 1900 La Plata, Argentina.</mods:affiliation>
<mods:nameIdentifier id="6A1E353ADDF911236D00D06CA9B5670A" type="email">braun@fcnym.unlp.edu.ar</mods:nameIdentifier>
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<mods:namePart id="111C1E7958CF30DC887C8EC4EB5B71C7">Morris, Glenn K.</mods:namePart>
<mods:affiliation id="322549103EEDFA88122A08C18671FB12">Biology Department, University of Toronto at Mississauga, 3359 Mississauga Rd, ON, Canada.</mods:affiliation>
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<mods:title id="ED642A17F7E7BB98754E0B4044CC2A8D">Journal of Orthoptera Research</mods:title>
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<mods:part id="40F6B6565CDF87B63A326D414660A9EE">
<mods:date id="77451E4BE1E3C41C50606E2DEE82FA8F">2022</mods:date>
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<mods:number id="B2A025E5E9B8BFE2E2B3282060CF30C3">2022-09-30</mods:number>
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<mods:detail id="86E13B6F77B4FC0C4EA1D7B6F1239092" type="volume">
<mods:number id="60C85567D50EB711E42090D919535BED">31</mods:number>
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<mods:classification id="22D9A7B20A15167C9DD8F1324728D8C0">journal article</mods:classification>
<mods:identifier id="747183246C5E0B0605D060AE5E640DEF" type="DOI">http://dx.doi.org/10.3897/jor.31.82306</mods:identifier>
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<treatment id="6AE6E9B88E415EBAB9F5D5D3FD578F1C" ID-GBIF-Taxon="201149833" LSID="urn:lsid:zoobank.org:act:39B53563-0D85-42CA-8DF2-80F9C45018AB" httpUri="http://treatment.plazi.org/id/6AE6E9B88E415EBAB9F5D5D3FD578F1C" lastPageNumber="143" pageId="0" pageNumber="143" scope_order="Orthoptera" scope_subFamily="Conocephalinae" scope_tribe="Cestrophorini">
<subSubSection id="99A1616FB8DA7DE5A9E527A1850D68A0" pageId="0" pageNumber="143" type="nomenclature">
<paragraph id="A45D038719776142A5F62A0D23E3747C" pageId="0" pageNumber="143">
<taxonomicName id="CEA1C5B7FBC44D1F247853BCBE78F18F" LSID="https://zoobank.org/39B53563-0D85-42CA-8DF2-80F9C45018AB" authority="Braun &amp; Morris, 2022" authorityName="Braun &amp; Morris" authorityYear="2022" class="Insecta" family="Tettigoniidae" genus="Cestrophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cestrophorus ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus" status="sp. nov.">Cestrophorus ditachus</taxonomicName>
<taxonomicNameLabel id="E098195FCA94DD9B001CA8406C8A46BF" pageId="0" pageNumber="143">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="28050B4B62686C50ECBF506C88826005" pageId="0" pageNumber="143" type="description">
<paragraph id="D77E0DF5BAA564207513C36285E07CB4" pageId="0" pageNumber="143">
<figureCitation id="9DE89F1EE850A3F84F9F68EA99C589B4" captionStart="Fig. 8" captionStartId="F8" captionText="Fig. 8. Cestrophorus spp. male genitalia compared. A. White pronotal maculae of C. paradoxus; B. Parameres of C. paradoxus ' upside-down dancing anteaters'; C. Dorsal aspect C. ditachus parameres; D. Lateral view of C. ditachus paramere; E. Photo of terminalia of C. amplitenuis; F. Lateral aspect of C. amplitenuis paramere." figureDoi="10.3897/jor.31.82306.fig8" httpUri="https://binary.pensoft.net/fig/750937" pageId="0" pageNumber="143">Figs 8C, D</figureCitation>
<figureCitation id="72F96C821929C55E945ECCC81D46FEFF" captionStart="Fig. 13" captionStartId="F13" captionText="Fig. 13. Field-recorded song (Sony Walkman, audio range only) of C. ditachus: trains of pulses characterized by two different pulse rates, slower (SPR) and faster (FPR). A. Two complete calls taken from an ongoing sequence. Single SPR train prefaces a 9 FPR-train sequence; B, D. FPR trains at higher time resolutions showing waveform of rapid-decay pulses; C, E. SPR train at higher time resolution showing waveform of rapid-decay pulses; F, G. Audio-limited spectra of FPR and SPR respectively; spectra unaffected by scraper speed, peaking at 6 - 10 kHz." figureDoi="10.3897/jor.31.82306.fig13" httpUri="https://binary.pensoft.net/fig/750942" pageId="0" pageNumber="143">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="480ABDDF7961C8B8F8BC0B7BFD31F471" pageId="0" pageNumber="143" type="reference_group">
<paragraph id="21EE5B11349AF65820C78C90D6173F67" pageId="0" pageNumber="143">
<taxonomicName id="CD02E35A233634B83FFFB929571F51B4" authority="Braun &amp; Morris, 2022" authorityName="Braun &amp; Morris" authorityYear="2022" class="Insecta" family="Tettigoniidae" genus="Gymnacoustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnacoustes ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus" status="sp. nov.">Gymnacoustes ditachus</taxonomicName>
('
<taxonomicName id="CA607F916D1095346AC8903D96898BFA" authority="Braun &amp; Morris, 2022" authorityName="Braun &amp; Morris" authorityYear="2022" class="Insecta" family="Tettigoniidae" genus="Gymnacoustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnacoustes ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus" status="sp. nov.">Gymnacoustes ditachus</taxonomicName>
' Morris, 1987)
</paragraph>
</subSubSection>
<subSubSection id="8926FEFCE1AF8D028BC92AA41DF22326" pageId="0" pageNumber="143" type="materials_examined">
<paragraph id="67667401EB521B15FEB3E9052103EAEF" pageId="0" pageNumber="143">
<emphasis id="75B4754D935CB54FDA442087C7E78ECF" italics="true" pageId="0" pageNumber="143">Material examined.</emphasis>
-
</paragraph>
<paragraph id="7CA48EDF6D4FE6E2EF1BB32A888061CE" pageId="0" pageNumber="143">
<materialsCitation id="D893A22EA7255AAE47F6824496D04ED9" collectingDate="1985-07-11" collectorName="G. K. Morris" country="ECUADOR" location="ECUADOR" specimenCount="♂" typeStatus="Holotype">
<emphasis id="1AE7A3B0F0A8B5D508A8A9FCF1936EB4" bold="true" pageId="0" pageNumber="143">
<typeStatus id="040623B0338224A37DDB879BF19047D3">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="D733AAF8EA0470CEF5D04A00421A2F08" name="Ecuador">ECUADOR</collectingCountry>
<specimenCount id="2A1C107EFF233299F0C60766D179A91B"></specimenCount>
; old Baeza;
<collectingDate id="6DAF882A5314A478555085541E5F1D29" value="1985-07-11">11 Jul. 1985</collectingDate>
;
<collectorName id="8A5B1C003A474A11B92E2E5CEB663D9F">G.K. Morris</collectorName>
leg.; SN-2, MLP
</materialsCitation>
.
<materialsCitation id="911786B8A3052E8049BCCF0B0DE89342" collectingDate="1985-07-10" collectorName="G. K. Morris, Glassy Tegmina" country="ECUADOR" location="Baeza" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
<emphasis id="B164B1C5401360B2EF228D982C7CDBDC" bold="true" pageId="0" pageNumber="143">
<typeStatus id="1D77318B9845F04A169DCB9B75B251E9">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="33EA73C805CA8F5D438C987D0218252D" name="Ecuador">ECUADOR</collectingCountry>
<specimenCount id="7A30A8306C9CCB92809ECD4932FECD43" type="male">1 ♂</specimenCount>
; old
<location id="87EC3987068041B62985BD3D458C38BB" LSID="urn:lsid:plazi:treatment:6AE6E9B88E415EBAB9F5D5D3FD578F1C:87EC3987068041B62985BD3D458C38BB" country="ECUADOR" name="Baeza">Baeza</location>
;
<collectingDate id="16EBE03577DE839A9195D164AB6A6BE9" value="1985-07-10">10 Jul. 1985</collectingDate>
;
<collectorName id="C810DAF42A7433E0A227D68D29576216">G.K. Morris</collectorName>
leg.; 85-1,
<collectorName id="1809C7FE0FEE750F462B2219E3D5043B">Glassy Tegmina</collectorName>
, ANSP
</materialsCitation>
<materialsCitation id="21915CE0976798923E27F776ECDB79F3" collectingDate="1985-02-25" collectorName="G. K. Morris" country="Ecuador" location="Tungurahua" specimenCount="1" specimenCount-male="1" stateProvince="Tungurahua" typeStatus="paratype">
<specimenCount id="8D73CDF78AC952FF7039D3A12047CA97" type="male">1 ♂</specimenCount>
;
<collectingRegion id="15A613787C1569FA9057652766E8DA35" country="Ecuador" name="Tungurahua">Tungurahua</collectingRegion>
;
<collectingDate id="B06182381A8AA0F409A5DDB01BF64867" value="1985-02-25">25 Feb. 1985</collectingDate>
;
<collectorName id="63E0F513186FEDDCCD3E9607FF876060">G.K. Morris</collectorName>
leg.; GLASSY TEG, SN-3, ANSP
</materialsCitation>
<materialsCitation id="D83147A981923509DABE4A4AB6BE57F1" collectingDate="1988-02-19" collectorName="G. K. Morris" country="Ecuador" location="Cosanga" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="68F9767239B0B50848E6E3E51B20B849" type="female">1 ♀</specimenCount>
;
<location id="3E1E6E1DF07038C8A58FA21444918A02" LSID="urn:lsid:plazi:treatment:6AE6E9B88E415EBAB9F5D5D3FD578F1C:3E1E6E1DF07038C8A58FA21444918A02" country="Ecuador" name="Cosanga">Cosanga</location>
;
<collectingDate id="55A36D6C1AB120CBFB28D7CE5CF6389C" value="1988-02-19">19 Feb. 1988</collectingDate>
;
<collectorName id="946C194CBD700E458AAD84A2E603412F">G.K. Morris</collectorName>
leg.; MLP
</materialsCitation>
<materialsCitation id="0E6C25C63FB26ACCEC0DAB366181566D" collectingDate="1988-02-19" collectorName="G. K. Morris" country="Ecuador" location="Cosanga" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="EBC0060DC14C4A3904223DA207BB520D" type="female">1 ♀</specimenCount>
;
<location id="9825CA35C5D229192782253CC5CBBB36" LSID="urn:lsid:plazi:treatment:6AE6E9B88E415EBAB9F5D5D3FD578F1C:9825CA35C5D229192782253CC5CBBB36" country="Ecuador" name="Cosanga">Cosanga</location>
;
<collectingDate id="C5D8005A77680A858352499F1E8150B9" value="1988-02-19">19 Feb. 1988</collectingDate>
;
<collectorName id="F45769E2B3A5E7C4E15C15DC95E68AE7">G.K. Morris</collectorName>
; ANSP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7EF671F9436A8EBBEFC87863B2597EF" pageId="0" pageNumber="143" type="etymology">
<paragraph id="1520402840BC0585BEF68F6478729252" pageId="0" pageNumber="143">
<emphasis id="DB1D7EBAC37239DE193DA0F2AEB2A62B" italics="true" pageId="0" pageNumber="143">Etymology.</emphasis>
-
</paragraph>
<paragraph id="FA5866EEAE8DB49B3F42BA694CBC8B2C" pageId="0" pageNumber="143">
Named for its two-part, two-pulse-rate, song structure, readily apparent to the human ear in real time. Greek
<normalizedToken id="945390478B27CCD79F6289A9D00086E2" originalValue="δι">δι</normalizedToken>
di [two],
<normalizedToken id="B2741B92FCDEE470A9FE313E00827F5F" originalValue="ταξηοσ">ταξηοσ</normalizedToken>
tachos [speed]. This insect is also named (inadvertently but deservedly) for Dita Klimas, katydid field photographer extraordinaire.
</paragraph>
</subSubSection>
<subSubSection id="B298D546887CCB174B2B2CC4DFB593A6" pageId="0" pageNumber="143" type="diagnosis">
<paragraph id="09C36FE0ECDB5DDC651599C8FCC5BB87" pageId="0" pageNumber="143">
<emphasis id="F7496AD0535FB3050D85D60F33259037" italics="true" pageId="0" pageNumber="143">Diagnosis.</emphasis>
-
</paragraph>
<paragraph id="C22D15C1E2D75B8831BAC751744AC58E" pageId="0" pageNumber="143">
More slender and uniformly colored than
<taxonomicName id="FA174A041624D0249385D8CB70B13ADA" class="Insecta" kingdom="Animalia" lsidName="C. paradoxus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="D6F62F7A9FA2C8D019C141AE9D25E04F" italics="true" pageId="0" pageNumber="143">C. paradoxus</emphasis>
</taxonomicName>
, with shorter tegmina. General coloration pale greenish; in males, metazona of pronotum with brown trapezoidal spot fringed by elongate whitish lateral spots. Male tegmina leaving abdomen tip exposed. Females uniformly greenish and apterous. Male cerci with obtusely triangular dorsal lobe (internally with inconspicuous toothlet at the tip) and robust inward-curved acuminate spine, baso-ventral spine with broad base and tapering in perpendicularly upcurved tip; above it on same lobe, another short spine (difficult to see). Styli completely reduced.
</paragraph>
</subSubSection>
<subSubSection id="05D937B30B75E08EAC99D74BA24DEE3E" pageId="0" pageNumber="143" type="description">
<paragraph id="F65BBAE646DAEB1F7C8F9998928C110B" pageId="0" pageNumber="143">
<emphasis id="CCDF00DCEE1A9733F14A665CCEAF578C" italics="true" pageId="0" pageNumber="143">Measurements.</emphasis>
-
</paragraph>
<paragraph id="285FAA7192BBFD15CA373A78AC2ECB54" pageId="0" pageNumber="143">Male: midline pronotum 4.3 mm, fastigium verticis 1.7 mm, tegmen 7.5 mm, hind tibiae 8.6 mm; female: midline pronotum 4.1 mm, hind tibiae 9.7 mm, ovipositor 8.7 mm (based on holotype and one paratype).</paragraph>
</subSubSection>
<subSubSection id="4D29E1E591B43CD2289DB816367F717A" pageId="0" pageNumber="143" type="field notes.">
<paragraph id="9452C52C59DC46E62A1FE4358242D81C" pageId="0" pageNumber="143">
<emphasis id="86BBEAEBA40D6822A6E9313E57FF878C" italics="true" pageId="0" pageNumber="143">Field notes.</emphasis>
-
</paragraph>
<paragraph id="D8B29360C20DC0DB4F30FA0A5339AE79" pageId="0" pageNumber="143">
We hunted in light rain at the [historic pre-earthquake] townsite of Baeza, along a muddy rock-strewn trail fenced by barbed wire, uphill beyond the hospital and cemetery on the night of July 10, 1985. We recorded and captured one male
<normalizedToken id="14A458DEDE021FF6689121F222CDF895" originalValue="agraeciine">'agraeciine'</normalizedToken>
of &quot;pearly pale cast&quot;, [like
<taxonomicName id="310700D85A88B0BBD231A8A084CE9A44" class="Insecta" kingdom="Animalia" lsidName="C. paradoxus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="52703CC3F2A9FFBB6EF5DAE8AFCA5467" italics="true" pageId="0" pageNumber="143">C. paradoxus</emphasis>
</taxonomicName>
] with white maculae prominent laterad on its pronotal metazona, its very pale greenish tegmina shorter than the abdomen, and its eardrums not recessed behind slits [field name 'glassy
<normalizedToken id="CAE5C972F3457DFD35F9647F80CB3C12" originalValue="tegmina">tegmina'</normalizedToken>
]. This specimen is now the holotype male. On July 11, GKM recorded another singer up the eastern branch of the trail above Baeza but failed to capture him.
</paragraph>
<paragraph id="9A62E382A4C57F951330EED50E2FEB7F" pageId="0" pageNumber="143">
On July 25, 1985, we drove from Quito and parked part way up
<normalizedToken id="CC7DD11B76D1BD01FFA9099476437EE2" originalValue="Volcán">Volcan</normalizedToken>
Tungurahua at signed &quot;ecological reserve&quot; hut under construction, below Pondua. Many
<taxonomicName id="6273813A8B3C639FC626EB6B4D009BF4" class="Insecta" kingdom="Animalia" lsidName="C. ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus">
<emphasis id="DF9FC344598CA7DBA83824F40CB509E2" italics="true" pageId="0" pageNumber="143">C. ditachus</emphasis>
</taxonomicName>
sp. nov. were heard singing from shrubby vegetation scattered about a pasture (never a bovid seen). GKM &quot;recorded a male singing from well above my head in a clump of bamboo and one small hardwood tree. After recording, we bent the bamboo down and searched the foliage with our lights,&quot; finding and capturing the presumed singer. &quot;As we left the immediate vicinity of the road and climbed several hundred feet, the incidence of [
<emphasis id="851A7027B6C9AB50AB8B048E87809A70" italics="true" pageId="0" pageNumber="143">C. ditacus sp. nov.</emphasis>
] singers declined to zero. They had ceased calling by 10:30 pm. Chilly and soon one could only hear single quiet lisps recurring at intervals of several seconds, presumably
<taxonomicName id="1BAD1A9BC0BB446CC9ACD1F5A62B67A8" class="Insecta" family="Tettigoniidae" genus="Acanthacara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthacara incisa." order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="incisa.">
<emphasis id="4383515F364053ED240A00928C5C2244" italics="true" pageId="0" pageNumber="143">Acanthacara incisa.</emphasis>
</taxonomicName>
&quot;
</paragraph>
</subSubSection>
</treatment>
</document>