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<document id="F6560FB4DEF3291706EE9D67ADFE5ED7" ID-DOI="10.37828/em.2023.69.4" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246886" ID-ZooBank="urn:lsid:zoobank.org:pub:A9914FEC-BD21-4088-9577-521C3A6B9776" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972195339" checkinUser="felipe" docAuthor="Pešić, Vladimir &amp; Goldschmidt, Tom" docDate="2023" docId="3B4417408E2CFF9AFF2DFAD6E7D7F9D9" docLanguage="en" docName="EcolMontenegrina.69.24-44.pdf" docOrigin="Ecologica Montenegrina 69" docSource="http://dx.doi.org/10.37828/em.2023.69.4" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Panisus torrenticolus Piersig 1898" docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="C77D6F388E24FF93FFBAFFE9E41FFFE7" masterDocTitle="New DNA-assisted records of water mites from Sardinia, with the description of a new species (Acari, Hydrachnidia)" masterLastPageNumber="44" masterPageNumber="24" pageNumber="32" updateTime="1722980808856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9F1E559A9F158F1B1EF3CF23477EE308">New DNA-assisted records of water mites from Sardinia, with the description of a new species (Acari, Hydrachnidia)</mods:title>
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<mods:namePart id="F25CC733BCE1F4013B2A3B18C0CF679E">Pešić, Vladimir</mods:namePart>
<mods:affiliation id="B828102AF394EA6A472DC1AA93EA5F2E">Department of Biology, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro.</mods:affiliation>
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<mods:namePart id="44290E6034E63714076DD4B86BB5C5D1">Goldschmidt, Tom</mods:namePart>
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<mods:affiliation id="9311488699850868DCF88FD20C129E28">SNSB - ZSM, Bavarian State Collection of Zoology, Section Arthropoda varia, Münchhausenstr. 21, 81247 Munich, Germany. E-mail: tomgoldschmidt 66 @ googlemail. com; https: // orcid. org / 0000 - 0002 - 5527 - 1719 * Corresponding author. E-mail: vladopesic @ gmail. com; https: // orcid. org / 0000 - 0002 - 9724 - 345 X</mods:affiliation>
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<mods:date id="AFAEDC654E02097FF4AB16BF0E9296DF">2023</mods:date>
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<treatment id="3B4417408E2CFF9AFF2DFAD6E7D7F9D9" ID-DOI="http://doi.org/10.5281/zenodo.13247490" ID-Zenodo-Dep="13247490" LSID="urn:lsid:plazi:treatment:3B4417408E2CFF9AFF2DFAD6E7D7F9D9" httpUri="http://treatment.plazi.org/id/3B4417408E2CFF9AFF2DFAD6E7D7F9D9" lastPageId="9" lastPageNumber="33" pageId="8" pageNumber="32">
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<paragraph id="B352A6568E2CFF9BFF2DFAD6E6B7FABE" blockId="8.[151,680,1343,1403]" box="[151,680,1343,1369]" pageId="8" pageNumber="32">
<heading id="E81A113A8E2CFF9BFF2DFAD6E6B7FABE" box="[151,680,1343,1369]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="32" reason="2">
<emphasis id="81997A448E2CFF9BFF2DFAD6E6B7FABE" bold="true" box="[151,680,1343,1369]" pageId="8" pageNumber="32">
<taxonomicName id="74EDDDD58E2CFF9BFF2DFAD6E65EFABE" authority="Piersig, 1898" authorityName="Piersig" authorityYear="1898" box="[151,577,1343,1369]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="torrenticolus">
<emphasis id="81997A448E2CFF9BFF2DFAD6E585FABE" bold="true" box="[151,410,1343,1369]" italics="true" pageId="8" pageNumber="32">Panisus torrenticolus</emphasis>
Piersig, 1898
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FBF7F5DD8E2CFF9BFF2DFA89E511FA9C" box="[151,270,1376,1403]" pageId="8" pageNumber="32" type="description">
<paragraph id="B352A6568E2CFF9BFF2DFA89E511FA9C" blockId="8.[151,680,1343,1403]" box="[151,270,1376,1403]" pageId="8" pageNumber="32">
<figureCitation id="2BD6BAD38E2CFF9BFF2DFA89E4C3FA9C" box="[151,220,1376,1403]" captionStart="Figure 5" captionStartId="9.[151,226,1004,1028]" captionTargetBox="[375,1196,189,985]" captionTargetId="figure-19@9.[375,1211,189,985]" captionTargetPageId="9" captionText="Figure 5A-C. Panisus torrenticolus group, ♀ [CCDB_39399_B10], ITALY, Sardinia, Comune di Fluminimaggiore, Sorgente Pubusinu, It 2022-1c: A photograph of dorsum; B dorsocentralia; C genital field. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13246902" httpUri="https://zenodo.org/record/13246902/files/figure.png" pageId="8" pageNumber="32">Fig. 5</figureCitation>
A-C
</paragraph>
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<subSubSection id="FBF7F5DD8E2CFF9BFF2DFA4DE064F9C3" pageId="8" pageNumber="32" type="materials_examined">
<paragraph id="B352A6568E2CFF9BFF2DFA4DE064F9C3" blockId="8.[151,1437,1443,2044]" pageId="8" pageNumber="32">
<emphasis id="81997A448E2CFF9BFF2DFA4DE599FA59" bold="true" box="[151,390,1444,1470]" pageId="8" pageNumber="32">Material examined</emphasis>
<materialsCitation id="0385AC0B8E2CFF9BFE0CFA4AE068F9C3" collectingDate="2021-08-07" collectionCode="RMNH" collectorName="Goldschmidt" country="Italy" county="Comune di Fluminimaggiore" elevation="206" latitude="39.4099" location="Sorgente Pubusinu" longLatPrecision="6" longitude="8.5275" municipality="It" pageId="8" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Sardegna">
<collectingCountry id="CBFAE6C68E2CFF9BFE0CFA4AE617FA59" box="[438,520,1443,1470]" name="Italy" pageId="8" pageNumber="32">ITALY</collectingCountry>
,
<collectingRegion id="712968B48E2CFF9BFDAFFA4AE665FA59" box="[533,634,1443,1470]" country="Italy" name="Sardegna" pageId="8" pageNumber="32">Sardinia</collectingRegion>
,
<collectingCounty id="5A33DEDA8E2CFF9BFD32FA4AE7FCFA59" box="[648,995,1443,1470]" pageId="8" pageNumber="32">Comune di Fluminimaggiore</collectingCounty>
,
<collectingMunicipality id="53363C2C8E2CFF9BFC4BFA4DE01CFA59" box="[1009,1027,1444,1470]" pageId="8" pageNumber="32">It</collectingMunicipality>
2022-1c,
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,
<locationDeviation id="A2E0BF748E2CFF9BFAD1FA4AE579FA07" location="Fluminimaggiore" pageId="8" pageNumber="32">S of Fluminimaggiore</locationDeviation>
, approx.
<quantity id="74150BB38E2CFF9BFE63FA2CE613FA07" box="[473,524,1477,1504]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="32" unit="km" value="5.0">5km</quantity>
gravel road from the junction of the SS 126 to the E, strong karst spring, moss carpet on broken dam,
<geoCoordinate id="D6D9C0918E2CFF9BFDB0FA0EE684F9E5" box="[522,667,1511,1538]" degrees="39.4099" direction="north" orientation="latitude" pageId="8" pageNumber="32" precision="5" value="39.4099">39.4099° N</geoCoordinate>
,
<geoCoordinate id="D6D9C0918E2CFF9BFD14FA0EE732F9E5" box="[686,813,1511,1538]" degrees="8.5275" direction="east" orientation="longitude" pageId="8" pageNumber="32" precision="5" value="8.5275">8.5275° E</geoCoordinate>
,
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<quantity id="74150BB38E2CFF9BFCFAFA0EE78EF9E5" box="[832,913,1511,1538]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="8" pageNumber="32" unit="m" value="206.0">206 m</quantity>
asl
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.,
<date id="C75380968E2CFF9BFC67FA01E09AF9E5" box="[989,1157,1511,1538]" pageId="8" pageNumber="32" value="2021-08-07">
<collectingDate id="D717797E8E2CFF9BFC67FA01E09AF9E5" box="[989,1157,1511,1538]" pageId="8" pageNumber="32" value="2021-08-07">7. Aug. 2021</collectingDate>
</date>
, leg.
<collectorName id="1E18C3808E2CFF9BFB6BFA0EE176F9E5" box="[1233,1385,1511,1538]" pageId="8" pageNumber="32">Goldschmidt</collectorName>
<specimenCount id="A5EB6DDF8E2CFF9BFACDFA0EE184F9E5" box="[1399,1435,1511,1538]" count="1" pageId="8" pageNumber="32" type="female">1♀</specimenCount>
(sequenced; voucher ID: CCDB-44301-G06), dissected and slide mounted (
<collectionCode id="D5FC3E938E2CFF9BFBAAF9E0E070F9C3" box="[1040,1135,1545,1572]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="32" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="FBF7F5DD8E2CFF9AFF4DF9C2E7D7F9D9" lastPageId="9" lastPageNumber="33" pageId="8" pageNumber="32" type="discussion">
<paragraph id="B352A6568E2CFF9BFF4DF9C2E6A5F909" blockId="8.[151,1437,1443,2044]" pageId="8" pageNumber="32">
<emphasis id="81997A448E2CFF9BFF4DF9C2E576F9A2" bold="true" box="[247,361,1579,1605]" pageId="8" pageNumber="32">Remarks</emphasis>
— The single female from
<collectingRegion id="712968B48E2CFF9BFD4DF9C2E746F9A1" box="[759,857,1579,1606]" country="Italy" name="Sardegna" pageId="8" pageNumber="32">Sardinia</collectingRegion>
, used in this study for molecular analysis, morphologically matches the description of
<emphasis id="81997A448E2CFF9BFD04F9A4E7D7F980" box="[702,968,1612,1639]" italics="true" pageId="8" pageNumber="32">
<taxonomicName id="74EDDDD58E2CFF9BFD04F9A4E7DCF980" authorityName="Piersig" authorityYear="1898" box="[702,963,1612,1639]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="torrenticolus">Panisus torrenticolus</taxonomicName>
.
</emphasis>
The latter species is diagnosed by a characteristic porosity of the muscle attachment sclerites, having a centre of fine pores surrounded by a circle of large pores and an outer area of finer pores (
<figureCitation id="2BD6BAD38E2CFF9BFD7BF979E75EF94C" box="[705,833,1680,1707]" captionStart="Figure 5" captionStartId="9.[151,226,1004,1028]" captionTargetBox="[375,1196,189,985]" captionTargetId="figure-19@9.[375,1211,189,985]" captionTargetPageId="9" captionText="Figure 5A-C. Panisus torrenticolus group, ♀ [CCDB_39399_B10], ITALY, Sardinia, Comune di Fluminimaggiore, Sorgente Pubusinu, It 2022-1c: A photograph of dorsum; B dorsocentralia; C genital field. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13246902" httpUri="https://zenodo.org/record/13246902/files/figure.png" pageId="8" pageNumber="32">Fig. 5A, B</figureCitation>
). Further features include the frontal shield as long as wide, or longer than wide, and dorsolateral plates without or with only slightly projecting lateral extensions (
<figureCitation id="2BD6BAD38E2CFF9BFE99F93AE560F909" box="[291,383,1747,1774]" captionStart="Figure 5" captionStartId="9.[151,226,1004,1028]" captionTargetBox="[375,1196,189,985]" captionTargetId="figure-19@9.[375,1211,189,985]" captionTargetPageId="9" captionText="Figure 5A-C. Panisus torrenticolus group, ♀ [CCDB_39399_B10], ITALY, Sardinia, Comune di Fluminimaggiore, Sorgente Pubusinu, It 2022-1c: A photograph of dorsum; B dorsocentralia; C genital field. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13246902" httpUri="https://zenodo.org/record/13246902/files/figure.png" pageId="8" pageNumber="32">Fig. 5A</figureCitation>
) (
<bibRefCitation id="D77CDBA78E2CFF9BFE2DF93AE6B1F909" author="Di Sabatino, A. &amp; Gerecke, R. &amp; Gledhill, T. &amp; Smit, H." box="[407,686,1747,1774]" pageId="8" pageNumber="32" pagination="1 - 234" refId="ref8807" refString="Di Sabatino, A., Gerecke, R., Gledhill, T. &amp; Smit, H. (2010) Acari: Hydrachnidia II. In: Gerecke, R. (Ed.), Chelicerata: Acari II. Susswasserfauna von Mitteleuropa, Vol. 7, 2 - 2, Elsevier Spektrum Akademischer Verlag, Heidelberg, pp. 1 - 234." type="book chapter" year="2010">
Di Sabatino
<emphasis id="81997A448E2CFF9BFD96F93DE640F909" box="[556,607,1747,1774]" italics="true" pageId="8" pageNumber="32">et al</emphasis>
. 2010
</bibRefCitation>
).
</paragraph>
<paragraph id="B352A6568E2CFF9BFF4DF91CE671F85F" blockId="8.[151,1437,1443,2044]" pageId="8" pageNumber="32">
The sequenced specimen from Sardinia forms a unique BIN (BOLD:AFC5560) with the nearest neighbouring BIN BOLD:AET0970, which includes a specimen of
<taxonomicName id="74EDDDD58E2CFF9BFC01F8F1E0C7F8D6" authority="Valdecasas" authorityName="Valdecasas" box="[955,1240,1814,1841]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="chicamoi">
<emphasis id="81997A448E2CFF9BFC01F8F1E057F8D6" box="[955,1096,1814,1841]" italics="true" pageId="8" pageNumber="32">P. chicamoi</emphasis>
Valdecasas
</taxonomicName>
, García- Jiménez &amp; Fernández- Campoamor, 2021(GBMNF37305-22) from
<collectingCountry id="CBFAE6C68E2CFF9BFCF6F8D1E78DF8B4" box="[844,914,1848,1875]" name="Spain" pageId="8" pageNumber="32">Spain</collectingCountry>
, a species recently described by
<bibRefCitation id="D77CDBA78E2CFF9BFAA2F8D1E537F892" author="Valdecasas, A. G. &amp; Jimenez, R. &amp; Campoamor, M. M." pageId="8" pageNumber="32" pagination="73 - 79" refId="ref11373" refString="Valdecasas, A. G., Garcia- Jimenez, R. &amp; Fernandez- Campoamor, M. M. (2021) Sobre una nueva especie de Panisus (Acari, Hydrachnidia) de la Peninsula Iberica. Revista iberica de aracnologia, 39, 73 - 79." type="journal article" year="2021">
Valdecasas
<emphasis id="81997A448E2CFF9BFF2DF8B2E4D4F892" box="[151,203,1882,1909]" italics="true" pageId="8" pageNumber="32">et al</emphasis>
. (2021)
</bibRefCitation>
from
<collectingRegion id="712968B48E2CFF9BFECBF8B3E5D6F892" box="[369,457,1882,1909]" country="Spain" name="Murcia" pageId="8" pageNumber="32">Murcia</collectingRegion>
,
<collectingCountry id="CBFAE6C68E2CFF9BFE6FF8B3E607F892" box="[469,536,1882,1909]" name="Spain" pageId="8" pageNumber="32">Spain</collectingCountry>
. In our COI tree, the specimen from Sardinia appeared as sister clade of the latter species from
<collectingCountry id="CBFAE6C68E2CFF9BFEC2F895E5A4F870" box="[376,443,1916,1943]" name="Spain" pageId="8" pageNumber="32">Spain</collectingCountry>
. The
<emphasis id="81997A448E2CFF9BFDBBF894E60FF871" box="[513,528,1917,1942]" italics="true" pageId="8" pageNumber="32">p</emphasis>
-distance between these two BINs was estimated at 15.54% indicating a long independent history of these two clades.
</paragraph>
<paragraph id="B352A6568E2CFF9BFF4DF829E021F81B" blockId="8.[151,1437,1443,2044]" pageId="8" pageNumber="32">
<materialsCitation id="0385AC0B8E2CFF9BFF4DF829E025F81B" location="Western Mediterranean" pageId="8" pageNumber="32" specimenCount="1" stateProvince="Sardegna">
In order to finally define the taxonomic status of the new lineage from
<collectingRegion id="712968B48E2CFF9BFBC3F856E0C1F83D" box="[1145,1246,1983,2010]" country="Italy" name="Sardegna" pageId="8" pageNumber="32">Sardinia</collectingRegion>
, the taxonomic situation of two other
<location id="B632F08D8E2CFF9BFE20F80BE6B1F81B" LSID="urn:lsid:plazi:treatment:3B4417408E2CFF9AFF2DFAD6E7D7F9D9:B632F08D8E2CFF9BFE20F80BE6B1F81B" box="[410,686,2017,2044]" name="Western Mediterranean" pageId="8" pageNumber="32" stateProvince="Sardegna">Western Mediterranean</location>
species needs to be clarified first
</materialsCitation>
:
</paragraph>
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<emphasis id="81997A448E2DFF9AFF2DFC05E535FBE3" bold="true" box="[151,298,1004,1028]" pageId="9" pageNumber="33">Figure 5A-C</emphasis>
.
<taxonomicName id="74EDDDD58E2DFF9AFE81FC05E63CFBE4" authorityName="Piersig" authorityYear="1898" box="[315,547,1003,1027]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="torrenticolus">
<emphasis id="81997A448E2DFF9AFE81FC05E63CFBE4" box="[315,547,1003,1027]" italics="true" pageId="9" pageNumber="33">Panisus torrenticolus</emphasis>
</taxonomicName>
group, ♀ [CCDB_39399_B10], ITALY, Sardinia, Comune di Fluminimaggiore, Sorgente Pubusinu, It 2022-1c: A photograph of dorsum; B dorsocentralia; C genital field. Scale bar = 100 μm.
</paragraph>
</caption>
<paragraph id="B352A6568E2DFF9AFF4DFB87E553FAD7" blockId="9.[151,1437,1133,1598]" pageId="9" pageNumber="33">
<taxonomicName id="74EDDDD58E2DFF9AFF4DFB87E684FB6F" authority="Maglio, 1909" authorityName="Maglio" authorityYear="1909" box="[247,667,1133,1160]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="clypeolatus">
<emphasis id="81997A448E2DFF9AFF4DFB87E5F8FB6F" box="[247,487,1133,1160]" italics="true" pageId="9" pageNumber="33">Panisus clypeolatus</emphasis>
<bibRefCitation id="D77CDBA78E2DFF9AFE4EFB84E684FB6F" author="Maglio, C." box="[500,667,1133,1160]" pageId="9" pageNumber="33" pagination="251 - 296" refId="ref9925" refString="Maglio, C. (1909) Idracarini del Trentino. (Contributo alla conoscenza dell'idracnofauna alpina.). Atti della Societa Italiana di Scienze Naturali, Pavia, 48, 251 - 296." type="journal article" year="1909">Maglio, 1909</bibRefCitation>
</taxonomicName>
, originally described from
<collectingRegion id="712968B48E2DFF9AFC55FB84E0F5FB6F" box="[1007,1258,1133,1160]" country="Italy" name="Trentino-Alto Adige" pageId="9" pageNumber="33">Trentino-Alto Adige</collectingRegion>
,
<collectingCountry id="CBFAE6C68E2DFF9AFB41FB84E12FFB6F" box="[1275,1328,1133,1160]" name="Italy" pageId="9" pageNumber="33">Italy</collectingCountry>
(
<bibRefCitation id="D77CDBA78E2DFF9AFAFCFB84E4CCFB4E" author="Maglio, C." pageId="9" pageNumber="33" pagination="251 - 296" refId="ref9925" refString="Maglio, C. (1909) Idracarini del Trentino. (Contributo alla conoscenza dell'idracnofauna alpina.). Atti della Societa Italiana di Scienze Naturali, Pavia, 48, 251 - 296." type="journal article" year="1909">Maglio 1909</bibRefCitation>
) and later on reported from the Western Mediterranean (e.g.,
<collectingCountry id="CBFAE6C68E2DFF9AFC62FB67E02AFB4E" box="[984,1077,1166,1193]" name="Algeria" pageId="9" pageNumber="33">Algeria</collectingCountry>
: K.
<bibRefCitation id="D77CDBA78E2DFF9AFBD7FB67E0EAFB4E" author="Viets, K." box="[1133,1269,1166,1193]" pageId="9" pageNumber="33" pagination="285 - 303" refId="ref11423" refString="Viets, K. (1951) Hydrachnellae (Acari) aus den franzosischen Alpen und aus Afrika. Zoologscher Anzeiger, 147 (11 / 12), 285 - 303." type="journal article" year="1951">Viets 1951</bibRefCitation>
, Bader 1989;
<collectingCountry id="CBFAE6C68E2DFF9AFF2DFB58E4F3FB2C" box="[151,236,1201,1227]" name="France" pageId="9" pageNumber="33">France</collectingCountry>
:
<bibRefCitation id="D77CDBA78E2DFF9AFF43FB58E589FB2C" author="Motas, C." box="[249,406,1200,1227]" pageId="9" pageNumber="33" pagination="150 - 186" refId="ref9961" refString="Motas, C. (1928 a) La faune hydracarienne des eaux courantes et des lacs eleves des Alpes du Dauphine. Compte rendu des seances de la Societe de biogeographie, 2, 150 - 186." type="journal article" year="1928">Motas 1928a</bibRefCitation>
, b,
<bibRefCitation id="D77CDBA78E2DFF9AFE7EFB59E663FB2C" author="Lundblad, O." box="[452,636,1200,1227]" pageId="9" pageNumber="33" pagination="1 - 306" refId="ref9898" refString="Lundblad, O. (1956) Zur Kenntnis sud- und mitteleuropaischer Hydrachnellen. Arkiv for Zoologi, 10 (1), 1 - 306." type="journal article" year="1956">Lundblad 1956</bibRefCitation>
;
<collectingCountry id="CBFAE6C68E2DFF9AFD30FB59E6DBFB2C" box="[650,708,1200,1227]" name="Italy" pageId="9" pageNumber="33">Italy</collectingCountry>
: Cicolani &amp; Di Sabatino 1985). It was synonymized with
<taxonomicName id="74EDDDD58E2DFF9AFA39FB5BE530FB0A" authorityName="Piersig" authorityYear="1898" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="torrenticolus">
<emphasis id="81997A448E2DFF9AFA39FB5BE530FB0A" italics="true" pageId="9" pageNumber="33">P. torrenticolus</emphasis>
</taxonomicName>
by
<bibRefCitation id="D77CDBA78E2DFF9AFEDDFB3BE6B9FB0A" author="Di Sabatino, A. &amp; Gerecke, R. &amp; Gledhill, T. &amp; Smit, H." box="[359,678,1234,1261]" pageId="9" pageNumber="33" pagination="1 - 34" refId="ref8741" refString="Di Sabatino, A., Gerecke, R., Gledhill, T. &amp; Smit, H. (2009) On the taxonomy of water mites (Acari: Hydrachnidia) described from the Palaearctic, part 2: Hydryphantoidea and Lebertioidea. Zootaxa, 2266, 1 - 34. https: // doi. org / 10.11646 / zootaxa. 2266.1.1" type="journal article" year="2009">
Di Sabatino
<emphasis id="81997A448E2DFF9AFDBCFB3AE65FFB0A" box="[518,576,1234,1261]" italics="true" pageId="9" pageNumber="33">et al</emphasis>
. (2009)
</bibRefCitation>
. The
<typeStatus id="6C5618F48E2DFF9AFD48FB3AE73BFB0A" box="[754,804,1235,1261]" pageId="9" pageNumber="33">type</typeStatus>
material of
<taxonomicName id="74EDDDD58E2DFF9AFC7CFB3AE06EFB0A" authorityName="Maglio" authorityYear="1909" box="[966,1137,1234,1261]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="clypeolatus">
<emphasis id="81997A448E2DFF9AFC7CFB3AE06EFB0A" box="[966,1137,1234,1261]" italics="true" pageId="9" pageNumber="33">P. clypeolatus</emphasis>
</taxonomicName>
is lost and the original description does not include original details that allow a definition of this taxon (see
<bibRefCitation id="D77CDBA78E2DFF9AFB32FB1DE182FAE8" author="Di Sabatino, A. &amp; Gerecke, R. &amp; Gledhill, T. &amp; Smit, H." box="[1160,1437,1268,1295]" pageId="9" pageNumber="33" pagination="1 - 34" refId="ref8741" refString="Di Sabatino, A., Gerecke, R., Gledhill, T. &amp; Smit, H. (2009) On the taxonomy of water mites (Acari: Hydrachnidia) described from the Palaearctic, part 2: Hydryphantoidea and Lebertioidea. Zootaxa, 2266, 1 - 34. https: // doi. org / 10.11646 / zootaxa. 2266.1.1" type="journal article" year="2009">
Di Sabatino
<emphasis id="81997A448E2DFF9AFAA4FB1CE14DFAE8" box="[1310,1362,1268,1295]" italics="true" pageId="9" pageNumber="33">et al</emphasis>
. 2009
</bibRefCitation>
for discussion).
</paragraph>
<paragraph id="B352A6568E2DFF9AFF4CFAD1E7D7F9D9" blockId="9.[151,1437,1133,1598]" pageId="9" pageNumber="33">
<taxonomicName id="74EDDDD58E2DFF9AFF4CFAD1E66BFAB5" authority="Bader, 1981" authorityName="Bader" authorityYear="1981" box="[246,628,1335,1362]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="81997A448E2DFF9AFF4CFAD1E5CEFAB5" box="[246,465,1335,1362]" italics="true" pageId="9" pageNumber="33">Panisus lundbladi</emphasis>
<bibRefCitation id="D77CDBA78E2DFF9AFE64FADEE66BFAB5" author="Bader, C." box="[478,628,1335,1362]" pageId="9" pageNumber="33" pagination="51 - 77" refId="ref8490" refString="Bader, C. (1981) Panisus-Studien: 1. Zur Revision der Gattung Panisus Koenike, 1896 (Acari, Actinedida, Hydrachnellae). Entomologica Basiliensia, 6, 51 - 77." type="journal article" year="1981">Bader, 1981</bibRefCitation>
</taxonomicName>
, a species originally described from
<collectingCountry id="CBFAE6C68E2DFF9AFBFDFADEE095FAB5" box="[1095,1162,1335,1362]" name="Spain" pageId="9" pageNumber="33">Spain</collectingCountry>
based on a specimen collected by Lundblad in an aqueduct in Girona,
<collectingRegion id="712968B48E2DFF9AFD64FAB0E751FA93" box="[734,846,1369,1396]" country="Spain" name="Catalunya" pageId="9" pageNumber="33">Catalonia</collectingRegion>
(
<bibRefCitation id="D77CDBA78E2DFF9AFCE4FAB0E7F1FA93" author="Bader, C." box="[862,1006,1369,1396]" pageId="9" pageNumber="33" pagination="51 - 77" refId="ref8490" refString="Bader, C. (1981) Panisus-Studien: 1. Zur Revision der Gattung Panisus Koenike, 1896 (Acari, Actinedida, Hydrachnellae). Entomologica Basiliensia, 6, 51 - 77." type="journal article" year="1981">Bader 1981</bibRefCitation>
). Morphologically, the latter species is somehow intermediate between
<taxonomicName id="74EDDDD58E2DFF9AFDFFFA95E792FA71" authority="Koenike, 1896" authorityName="Koenike" authorityYear="1896" box="[581,909,1402,1430]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="81997A448E2DFF9AFDFFFA95E6D2FA72" box="[581,717,1402,1429]" italics="true" pageId="9" pageNumber="33">P. michaeli</emphasis>
Koenike, 1896
</taxonomicName>
and
<taxonomicName id="74EDDDD58E2DFF9AFC6BFA95E094FA72" authorityName="Piersig" authorityYear="1898" box="[977,1163,1402,1429]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="torrenticolus">
<emphasis id="81997A448E2DFF9AFC6BFA95E094FA72" box="[977,1163,1402,1429]" italics="true" pageId="9" pageNumber="33">P. torrenticolus</emphasis>
</taxonomicName>
, resembling the latter species in the presence of a ring of larger pores on most muscle attachment sclerites, and to
<taxonomicName id="74EDDDD58E2DFF9AFB6BFA77E14CFA50" authorityName="Koenike" authorityYear="1896" box="[1233,1363,1436,1463]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="81997A448E2DFF9AFB6BFA77E14CFA50" box="[1233,1363,1436,1463]" italics="true" pageId="9" pageNumber="33">P. michaeli</emphasis>
</taxonomicName>
in the presence of an enlarged frontal shield and the extensive lateral ramifications of the dorsolateralia. For now, it seems most appropriate to leave the clade from Sardinia as “
<taxonomicName id="74EDDDD58E2DFF9AFC2CFA08E088FA1C" authorityName="Piersig" authorityYear="1898" box="[918,1175,1504,1531]" class="Arachnida" family="Hydryphantidae" genus="Panisus" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="torrenticolus">
<emphasis id="81997A448E2DFF9AFC2CFA08E088FA1C" box="[918,1175,1504,1531]" italics="true" pageId="9" pageNumber="33">Panisus torrenticolus</emphasis>
</taxonomicName>
group” until more specimens from Western Mediterranean, including specimens from the
<emphasis id="81997A448E2DFF9AFB91F9EBE0B6F9FB" box="[1067,1193,1538,1565]" italics="true" pageId="9" pageNumber="33">loci typici</emphasis>
of the two above mentioned species with uncertain taxonomic status become available.
</paragraph>
</subSubSection>
</treatment>
</document>