210 lines
16 KiB
XML
210 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.46.32823" ID-GBIF-Dataset="e546c2a6-4668-4253-b2e2-b462f23dab8c" ID-PMC="PMC6379322" ID-Pensoft-Pub="1314-4049-46-1" ID-PubMed="30787668" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-46-1" ModsDocOrigin="MycoKeys 46" ModsDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" checkinTime="1555333436509" checkinUser="pensoft" docAuthor="Cripps, Cathy L., Eberhardt, Ursula, Schuetz, Nicole, Beker, Henry J., Vera S. Evenson, & Horak, Egon" docDate="2019" docId="3B3AD62F743DD88E27810322CCA913AA" docLanguage="en" docName="MycoKeys 46: 1-54" docOrigin="MycoKeys 46" docSource="http://dx.doi.org/10.3897/mycokeys.46.32823" docTitle="Hebeloma oreophilum Beker & U. Eberh., Mycologia 107: 1295 (2016 2015" docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="AC18AA047D78FFBFD06BFFFDFF948577" masterDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" masterLastPageNumber="54" masterPageNumber="1" pageNumber="35" updateTime="1668136151891" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Hebeloma in the Rocky Mountain Alpine Zone</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cripps, Cathy L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Eberhardt, Ursula</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schuetz, Nicole</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beker, Henry J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vera S. Evenson,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Horak, Egon</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>46</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.46.32823</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.46.32823</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-46-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156202110" LSID="urn:lsid:plazi:treatment:3B3AD62F743DD88E27810322CCA913AA" httpUri="http://treatment.plazi.org/id/3B3AD62F743DD88E27810322CCA913AA" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<subSubSection pageId="34" pageNumber="35" type="nomenclature">
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<paragraph pageId="34" pageNumber="35">
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13.
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<taxonomicName authority="Beker & U. Eberh., Mycologia 107: 1295 (2016) [2015]" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma oreophilum" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="oreophilum">Hebeloma oreophilum Beker & U. Eberh., Mycologia 107: 1295 (2016) [2015]</taxonomicName>
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Figures 6A, 19, 23 (13)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="etymology">
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<paragraph pageId="34" pageNumber="35">Etymology.</paragraph>
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<paragraph pageId="34" pageNumber="35">From oreophilus, mountain loving to emphasize its presence in alpine habitats.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="description">
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<paragraph pageId="34" pageNumber="35">Description.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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Cortina present. Pileus 15-30 mm in diameter, convex, hemispherical, not umbonate, smooth, dry or greasy, medium brown, bay brown, reddish brown, dark black brown, with white to cream rim of fibrillose veil remnants at margin, with hoary coating; margin even or weakly scalloped. Thick waxy pellicle mentioned in one collection. Lamellae emarginate, subdistant, L = 40-50 plus lamellulae, cream at first then milk coffee color, pinkish cinnamon; margin floccose, white. Stipe 15-60
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<normalizedToken originalValue="×">x</normalizedToken>
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3-8 mm, equal or slightly enlarged at base, a bit curved or undulating, whitich, tan, brown, in top part and darkening to blackish brown at base, pruinose in top half and fibrous below, with patches of fibrils. Context watery buff with yellow tint, and blackish brown in base, stipe hollow. Odor raphanoid. Exsiccate pale brown all over, not dark.
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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Basidiospores amygdaliform, with a small snout, apiculate, not guttulate, finely verrucose (O1, O2), distinctly dextrinoid (D2, D3), no perispore loosening observed (P0), 10-14
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<normalizedToken originalValue="×">x</normalizedToken>
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6-8
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, on average 11.7
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<normalizedToken originalValue="×">x</normalizedToken>
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6.9
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, Q = 1.68. Basidia clavate, 25-35
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<normalizedToken originalValue="×">x</normalizedToken>
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8-10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, mostly four-spored. Cheilocystidia lageniform, with subcapitate apex, long neck (sometimes wiggly), with gradually swollen base, sometimes septate, length 30-70
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<normalizedToken originalValue="×">x</normalizedToken>
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4-7
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<normalizedToken originalValue="µm">µm</normalizedToken>
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at apex, 3-6.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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in middle, and up to 13
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<normalizedToken originalValue="µm">µm</normalizedToken>
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at base, no thickening noticed. Pleurocystidia absent. Epicutis thickness 40-75
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, with no encrusted hyphae recorded.
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</paragraph>
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<caption pageId="34" pageNumber="35">
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<paragraph pageId="34" pageNumber="35">
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Figure 19.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma oreophilum" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="oreophilum">Hebeloma oreophilum</taxonomicName>
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, DBG-F-027674 and ZT12733.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="rocky mountain ecology">
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<paragraph pageId="34" pageNumber="35">Rocky Mountain ecology.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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In low alpine with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">Salix</taxonomicName>
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species in Montana and Colorado.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="materials_examined">
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<paragraph pageId="34" pageNumber="35">Rocky Mountain specimens examined.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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U.S.A. COLORADO: Clear Creek County, Denver Mountain Park, Summit Lake, 3911 m, in
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix arctica" order="Malpighiales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="arctica">Salix arctica</taxonomicName>
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and
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<taxonomicName lsidName="S. glauca" pageId="34" pageNumber="35" rank="species" species="glauca">S. glauca</taxonomicName>
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, 20 Aug 2013, DBG-F-027674, L, Gillman; Summit Lake Park, 3912 m, with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., 22 Aug 2012, DBG-F-022788, L. Gillman; Arapaho National Forest, Nature Trail, Mount Goliath, 3658 m, in
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp, 1 Sept 1999, DBG-F-020558, V.S. Evenson; Pitkin County, White River National Forest, junction of Montezuma Basin and Pearl Pass, in
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., 3658 m, 6 Aug 1999, DBG-F-020053, V.S. Evenson. MONTANA: Carbon County, Beartooth Plateau, Frozen Lakes, with dwarf
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">Salix</taxonomicName>
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, 26 July 1997, 3200 m, CLC1102 (MONT), C. Cripps; site 2, 3100 m, 8 Aug 2002, CLC1937 (MONT), with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix planifolia" order="Malpighiales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="planifolia">Salix planifolia</taxonomicName>
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, C. Cripps; Billings Fen, in moss near
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<taxonomicName lsidName="S. planifolia" pageId="34" pageNumber="35" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, 3048 m, 23 Aug 2017, CLC 3607 (MONT). WYOMING: Beartooth Plateau, Wyoming Creek, with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix planifolia" order="Malpighiales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="planifolia">Salix planifolia</taxonomicName>
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, 3176 m, 6 Aug 2008, HJB12449, C. Cripps; Beartooth Plateau, Hell-roaring Plateau, near
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., 14 Aug 2007, ZT12733 (ETH), E. Horak.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
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<paragraph pageId="34" pageNumber="35">Discussion.</paragraph>
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<paragraph lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma oreophilum" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="oreophilum">Hebeloma oreophilum</taxonomicName>
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is a member of the
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<taxonomicName lsidName="H. nigellum" pageId="34" pageNumber="35" rank="species" species="nigellum">H. nigellum</taxonomicName>
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complex that cannot always be distinguished from
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<taxonomicName lsidName="H. nigellum" pageId="34" pageNumber="35" rank="species" species="nigellum">H. nigellum</taxonomicName>
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based on ITS data (Fig. 6A). In terms
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<pageBreakToken pageId="35" pageNumber="36" start="start">of</pageBreakToken>
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differences, the
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<taxonomicName lsidName="H. oreophilum" pageId="35" pageNumber="36" rank="species" species="oreophilum">H. oreophilum</taxonomicName>
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sequences from the sample (9 RM, 10 FE) differ by 0-9 [0-3] bp; 0-8 [0-1] bp within the RM sample. Most similar to
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<taxonomicName lsidName="H. oreophilum" pageId="35" pageNumber="36" rank="species" species="oreophilum">H. oreophilum</taxonomicName>
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is
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<taxonomicName lsidName="H. clavulipes" pageId="35" pageNumber="36" rank="species" species="clavulipes">H. clavulipes</taxonomicName>
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, which in this sample differs by 1-11 [0-3] bp. The two species do not share the same habitats. The differences between species sharing the same habitats (
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<taxonomicName lsidName="H. nigellum" pageId="35" pageNumber="36" rank="species" species="nigellum">H. nigellum</taxonomicName>
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and
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<taxonomicName lsidName="H. spetsbergense" pageId="35" pageNumber="36" rank="species" species="spetsbergense">H. spetsbergense</taxonomicName>
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) are 3-10 [0-5] bp. Morphologically, the easiest way to separate
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<taxonomicName lsidName="H. oreophilum" pageId="35" pageNumber="36" rank="species" species="oreophilum">H. oreophilum</taxonomicName>
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from
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<taxonomicName lsidName="H. hygrophilum" pageId="35" pageNumber="36" rank="species" species="hygrophilum">H. hygrophilum</taxonomicName>
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and
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<taxonomicName lsidName="H. nigellum" pageId="35" pageNumber="36" rank="species" species="nigellum">H. nigellum</taxonomicName>
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is by the number of full length lamellae, always at least 40 for
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<taxonomicName lsidName="H. oreophilum" pageId="35" pageNumber="36" rank="species" species="oreophilum">H. oreophilum</taxonomicName>
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and less than 36 for the others.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma clavulipes" order="Agaricales" pageId="35" pageNumber="36" phylum="Basidiomycota" rank="species" species="clavulipes">Hebeloma clavulipes</taxonomicName>
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is not known from arctic-alpine habitats and has spores with an average width at most 6.6
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<normalizedToken originalValue="µm">µm</normalizedToken>
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while the average spore width for
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<taxonomicName lsidName="H. oreophilum" pageId="35" pageNumber="36" rank="species" species="oreophilum">H. oreophilum</taxonomicName>
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is on average at least 6.8
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma oreophilum" order="Agaricales" pageId="35" pageNumber="36" phylum="Basidiomycota" rank="species" species="oreophilum">Hebeloma oreophilum</taxonomicName>
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has a persisting cortina and the lageniform/ventricose cheilocystidia of
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<taxonomicName infraspecific-rank="sect." lsidName="H." pageId="35" pageNumber="36" rank="section" section="Hebeloma">H. sect. Hebeloma</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="35" pageNumber="36">
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This species was first described from the western Carpathians (Slovakia) with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix reticulata" order="Malpighiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="reticulata">Salix reticulata</taxonomicName>
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,
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<taxonomicName lsidName="S. retusa" pageId="35" pageNumber="36" rank="species" species="retusa">S. retusa</taxonomicName>
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, or
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<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Dryas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dryas octopetala" order="Rosales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="octopetala">Dryas octopetala</taxonomicName>
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on calcareous soil (
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<bibRefCitation author="Eberhardt, U" journalOrPublisher="Mycologia" pageId="43" pageNumber="44" pagination="1285 - 1303" title="The genus Hebeloma in the alpine belt of the Carpathians including two new species." url="https://doi.org/10.3852/15-097" volume="107" year="2015 b">Eberhardt et al. 2015b</bibRefCitation>
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). It has since been reported from Canada, Greenland, Scandinavia, Svalbard, and the Rocky Mountains (
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<bibRefCitation pageId="35" pageNumber="36">Beker et al. 2016</bibRefCitation>
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;
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<bibRefCitation pageId="35" pageNumber="36">Beker et al. 2018</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |