563 lines
71 KiB
XML
563 lines
71 KiB
XML
<document id="02368F1EB56B7788E44B537D6E32B28B" ID-ISSN="1536-9307" ID-ZooBank="9CFB25E3-7300-497E-884D-E73163E22267" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638243321932" checkinUser="felipe" docAuthor="Teruel, Rolando & Kovařík, František" docDate="2014" docId="3B2287D3FFF3FFFEFF1DF918230BFC2B" docLanguage="en" docName="Euscorpius.186.1-15.pdf" docOrigin="Euscorpius 186" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Androctonus turieli Teruel et Kovarik 2014, n. sp." docType="treatment" docUuid="5EFE22DA-DAB2-48C1-920A-4573754E774B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="13" masterDocId="C71BFFABFFFAFFF0FFC3FFB7277EFFCC" masterDocTitle="Redescription of Androctonus bicolor Ehrenberg, 1828, and description of Androctonus turieli sp. n. from Tunisia (Scorpiones: Buthidae)" masterLastPageNumber="15" masterPageNumber="1" pageNumber="8" updateTime="1699435022351" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="F57171C5487082EDB47EBA893C5FA02F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7B1B0EE6B37480CD615468884FFA1C87">Redescription of Androctonus bicolor Ehrenberg, 1828, and description of Androctonus turieli sp. n. from Tunisia (Scorpiones: Buthidae)</mods:title>
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<mods:name id="617BC8CEC0FF9490DE6A1185800C0994" type="personal">
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<mods:namePart id="E412AEE8865AE68DF60C17009CF13E19">Teruel, Rolando</mods:namePart>
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<mods:roleTerm id="C1694CE401EFD8032C08B401BAAA4A43">Author</mods:roleTerm>
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<mods:namePart id="CD05112D502B07639AD2C866594F2982">Kovařík, František</mods:namePart>
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<mods:typeOfResource id="2C0FC5DC9CE3FBC79624DD42458007B4">text</mods:typeOfResource>
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<mods:title id="6DF30F6E71CD6FB52EC04D6FB7D6DB0A">Euscorpius</mods:title>
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</mods:titleInfo>
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<mods:part id="04A08270F727BDFE00C9560CE3EE7D87">
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<mods:date id="A9DB78198C2160710D90681DC5D6B578">2014</mods:date>
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<mods:number id="33DE6B41A208E3282937275D8DD3ED10">186</mods:number>
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<mods:classification id="05CBFBF64F59AF0AECFC7982C0196715">journal article</mods:classification>
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<paragraph id="B33436C5FFF3FFF9FF1DF918247BF905" blockId="9.[195,799,1711,1827]" box="[222,773,1711,1737]" pageId="9" pageNumber="8">
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<emphasis id="81FFEAD7FFF3FFF9FF1DF918247BF905" bold="true" box="[222,773,1711,1737]" pageId="9" pageNumber="8">
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<taxonomicName id="748B4D46FFF3FFF9FF1DF91825CBF905" authority="Teruel et Kovarik" authorityName="Teruel et Kovarik" authorityYear="2014" box="[222,693,1711,1737]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
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<emphasis id="81FFEAD7FFF3FFF9FF1DF91826B5F905" bold="true" box="[222,459,1711,1737]" italics="true" pageId="9" pageNumber="8">Androctonus turieli</emphasis>
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Teruel et Kovařík
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</taxonomicName>
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,
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<taxonomicNameLabel id="9ACC57ACFFF3FFF9FD01F918247BF905" box="[706,773,1711,1737]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FB91654EFFF3FFF9FE9BF97825B7F8EF" pageId="9" pageNumber="8" type="description">
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<paragraph id="B33436C5FFF3FFF9FE9BF9782461F8C9" blockId="9.[195,799,1711,1827]" pageId="9" pageNumber="8">
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(
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<figureCitation id="2BB02A40FFF3FFF9FE9CF9782682F92B" box="[351,508,1743,1767]" captionStart-0="Figures 19–25" captionStart-1="Figures 26–31" captionStart-2="Figures 32–35" captionStartId-0="10.[192,278,1837,1861]" captionStartId-1="11.[192,278,1328,1352]" captionStartId-2="12.[192,278,1114,1138]" captionTargetBox-0="[229,1416,203,1790]" captionTargetBox-1="[203,1421,183,1283]" captionTargetBox-2="[202,1426,193,1053]" captionTargetId-0="figure-63@10.[212,1416,192,1802]" captionTargetId-1="figure-316@11.[192,1440,168,1293]" captionTargetId-2="figure-481@12.[192,1440,192,1080]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." captionText-1="Figures 26–31: Androctonus turieli sp. n., pedipalp chela dorsal (26) and external (27), pedipalp patela dorsal (28) and external (29), pedipalp femur dorsal (30), and right movable finger internal (31), ♀ holotype." captionText-2="Figures 32–35: Figures 32–34. Androctonus turieli sp. n., metasoma and telson dorsal (32), ventral (33) and lateral (34) views, ♀ holotype. Figure 35. Androctonus sergenti Vachon, 1948, metasoma and telson ventral, ♂, Morocco, Anti Atlas Mts., 62 km southeast of Agadir, 30°03.35'N 09°04.4'W, 798 m a. s. l. (FKCP)." pageId="9" pageNumber="8">Figures 19–34</figureCitation>
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,
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<figureCitation id="2BB02A40FFF3FFF9FDC4F978255BF92B" box="[519,549,1743,1767]" captionStart="Figure 36" captionStartId="13.[192,267,846,870]" captionTargetBox="[203,1426,167,811]" captionTargetId="figure-551@13.[204,1425,168,810]" captionTargetPageId="13" captionText="Figure 36: Type locality of Androctonus turieli sp.n." pageId="9" pageNumber="8">36</figureCitation>
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;
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<tableCitation id="FE09037EFFF3FFF9FDF3F97825FFF92B" box="[560,641,1743,1767]" captionStart="Table 2" captionStartId="9.[192,257,793,817]" captionText="Table 2: Measurements (mm) of two adult types of Androctonus turieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (H), left (L), right (R). Note: telson vesicle length is not given because it narrows evenly into aculeus, without any clear definition point." pageId="9" pageNumber="8">Table 2</tableCitation>
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)
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<emphasis id="81FFEAD7FFF3FFF9FF00F95A2461F8C9" bold="true" box="[195,799,1773,1797]" pageId="9" pageNumber="8">http://zoobank.org/urn:lsid:zoobank.org:act:5EFE22</emphasis>
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</paragraph>
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<paragraph id="B33436C5FFF3FFF9FEDAF8BC25B7F8EF" blockId="9.[195,799,1711,1827]" box="[281,713,1803,1827]" pageId="9" pageNumber="8">
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<emphasis id="81FFEAD7FFF3FFF9FEDAF8BC25B7F8EF" bold="true" box="[281,713,1803,1827]" pageId="9" pageNumber="8">DA-DAB2-48C1-920A-4573754E774B</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="FB91654EFFF3FFF9FF03F8F22379FC38" pageId="9" pageNumber="8" type="materials_examined">
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<paragraph id="B33436C5FFF3FFF9FF03F8F226BBF8B7" blockId="9.[192,802,1860,1915]" pageId="9" pageNumber="8">
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<materialsCitation id="03E33C98FFF3FFF9FF03F8F226D7F897" box="[192,425,1861,1884]" collectionCode="LOCALITY, AND" pageId="9" pageNumber="8" specimenCount="1" typeStatus="holotype">
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<collectionCode id="D59AAE00FFF3FFF9FEC6F8F026D7F897" box="[261,425,1863,1883]" pageId="9" pageNumber="8">LOCALITY AND</collectionCode>
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</materialsCitation>
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<materialsCitation id="03E33C98FFF3FFF9FE76F8FF278FF8B7" collectionCode="DEPOSITORY" country="Tunisia" location="Sfax" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Sfax" typeStatus="holotype">
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<typeStatus id="6C308867FFF3FFF9FE76F8FF2694F897" box="[437,490,1864,1883]" pageId="9" pageNumber="8">TYPE</typeStatus>
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.
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<collectingCountry id="CB9C7655FFF3FFF9FD53F8F3259CF890" box="[656,738,1860,1884]" name="Tunisia" pageId="9" pageNumber="8">Tunisia</collectingCountry>
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, near
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<collectingRegion id="714FF827FFF3FFF9FF03F8D4278FF8B7" box="[192,241,1891,1915]" country="Tunisia" name="Sfax" pageId="9" pageNumber="8">Sfax</collectingRegion>
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</materialsCitation>
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<materialsCitation id="03E33C98FFF3FFF9FF34F8D426BFF8B7" box="[247,449,1891,1915]" collectionCode="FKCP" country="Tunisia" isTailEnumeration="true" location="Sfax" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Sfax" typeStatus="holotype">
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[=
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||
<collectingRegion id="714FF827FFF3FFF9FED5F8D42614F8B7" box="[278,362,1891,1915]" country="Tunisia" name="Sfax" pageId="9" pageNumber="8">Safaqis</collectingRegion>
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];
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<collectionCode id="D59AAE00FFF3FFF9FEB8F8D426BFF8B7" box="[379,449,1891,1915]" pageId="9" pageNumber="8">FKCP</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="B33436C5FFF3FFF9FCFCFC342379FC38" blockId="9.[831,1440,898,1013]" pageId="9" pageNumber="8">
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<collectionCode id="D59AAE00FFF3FFF9FCFCFC342490FC56" box="[831,1006,899,922]" pageId="9" pageNumber="8">TYPE MATERIAL</collectionCode>
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.
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<materialsCitation id="03E33C98FFF3FFF9FC3CFC34237DFC38" collectionCode="FKCP, RTOC" country="Tunisia" location="Sfax" pageId="9" pageNumber="8" specimenCount="6" specimenCount-female="3" specimenCount-juvenile="2" specimenCount-male="1" stateProvince="Sfax" typeStatus="holotype">
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<emphasis id="81FFEAD7FFF3FFF9FC3CFC34232BFC57" bold="true" box="[1023,1109,899,923]" pageId="9" pageNumber="8">
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<collectingCountry id="CB9C7655FFF3FFF9FC3CFC34232BFC57" box="[1023,1109,899,923]" name="Tunisia" pageId="9" pageNumber="8">Tunisia</collectingCountry>
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</emphasis>
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||
, near
|
||
<collectingRegion id="714FF827FFF3FFF9FB5CFC3523AEFC56" box="[1183,1232,898,922]" country="Tunisia" name="Sfax" pageId="9" pageNumber="8">Sfax</collectingRegion>
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||
[=
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||
<collectingRegion id="714FF827FFF3FFF9FB3FFC35222EFC56" box="[1276,1360,898,922]" country="Tunisia" name="Sfax" pageId="9" pageNumber="8">Safaqis</collectingRegion>
|
||
], 1980,
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||
<specimenCount id="A58DFD4CFFF3FFF9FCFCFC17241EFC74" box="[831,864,928,952]" pageId="9" pageNumber="8" type="female">1♀</specimenCount>
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||
<typeStatus id="6C308867FFF3FFF9FCA8FC1724B6FC74" box="[875,968,928,952]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="2BB02A40FFF3FFF9FC18FC17231CFC74" box="[987,1122,928,952]" captionStart-0="Figures 19–25" captionStart-1="Figures 26–31" captionStart-2="Figures 32–35" captionStartId-0="10.[192,278,1837,1861]" captionStartId-1="11.[192,278,1328,1352]" captionStartId-2="12.[192,278,1114,1138]" captionTargetBox-0="[229,1416,203,1790]" captionTargetBox-1="[203,1421,183,1283]" captionTargetBox-2="[202,1426,193,1053]" captionTargetId-0="figure-63@10.[212,1416,192,1802]" captionTargetId-1="figure-316@11.[192,1440,168,1293]" captionTargetId-2="figure-481@12.[192,1440,192,1080]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." captionText-1="Figures 26–31: Androctonus turieli sp. n., pedipalp chela dorsal (26) and external (27), pedipalp patela dorsal (28) and external (29), pedipalp femur dorsal (30), and right movable finger internal (31), ♀ holotype." captionText-2="Figures 32–35: Figures 32–34. Androctonus turieli sp. n., metasoma and telson dorsal (32), ventral (33) and lateral (34) views, ♀ holotype. Figure 35. Androctonus sergenti Vachon, 1948, metasoma and telson ventral, ♂, Morocco, Anti Atlas Mts., 62 km southeast of Agadir, 30°03.35'N 09°04.4'W, 798 m a. s. l. (FKCP)." pageId="9" pageNumber="8">Figs. 21–34</figureCitation>
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||
,
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||
<collectionCode id="D59AAE00FFF3FFF9FBB2FC1723C6FC74" box="[1137,1208,928,952]" pageId="9" pageNumber="8">FKCP</collectionCode>
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||
),
|
||
<specimenCount id="A58DFD4CFFF3FFF9FB0CFC17238EFC74" box="[1231,1264,928,952]" pageId="9" pageNumber="8" type="female">1♀</specimenCount>
|
||
<typeStatus id="6C308867FFF3FFF9FB38FC162229FC75" box="[1275,1367,929,953]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="2BB02A40FFF3FFF9FAA9FC1724FAFC1A" captionStart="Figures 19–25" captionStartId="10.[192,278,1837,1861]" captionTargetBox="[229,1416,203,1790]" captionTargetId="figure-63@10.[212,1416,192,1802]" captionTargetPageId="10" captionText="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." pageId="9" pageNumber="8">Figs. 19–20</figureCitation>
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||
,
|
||
<collectionCode id="D59AAE00FFF3FFF9FC5BFC0924A1FC1A" box="[920,991,958,982]" pageId="9" pageNumber="8">FKCP</collectionCode>
|
||
),
|
||
<specimenCount id="A58DFD4CFFF3FFF9FC38FC092362FC1A" box="[1019,1052,958,982]" pageId="9" pageNumber="8" type="female">1♀</specimenCount>
|
||
<typeStatus id="6C308867FFF3FFF9FBEFFC0823F9FC1B" box="[1068,1159,959,983]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="D59AAE00FFF3FFF9FB63FC092393FC1A" box="[1184,1261,958,982]" pageId="9" pageNumber="8">RTOC</collectionCode>
|
||
),
|
||
<specimenCount id="A58DFD4CFFF3FFF9FACBFC092205FC1A" box="[1288,1403,958,982]" pageId="9" pageNumber="8" type="juvenile">2 juvenile</specimenCount>
|
||
<specimenCount id="A58DFD4CFFF3FFF9FA48FC0822E1FC1A" box="[1419,1439,959,982]" pageId="9" pageNumber="8" type="male">♂</specimenCount>
|
||
<typeStatus id="6C308867FFF3FFF9FCFCFC6A24DBFC39" box="[831,933,989,1013]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="D59AAE00FFF3FFF9FC70FC6B2482FC38" box="[947,1020,988,1012]" pageId="9" pageNumber="8">FKCP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB91654EFFF3FFF9FCFCFBA424B0FB48" pageId="9" pageNumber="8" type="etymology">
|
||
<paragraph id="B33436C5FFF3FFF9FCFCFBA424B0FB48" blockId="9.[831,1440,1042,1156]" pageId="9" pageNumber="8">
|
||
ETYMOLOGY. It is a pleasure to name this species after our friend Carlos Turiel (Neuss,
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||
<collectingCountry id="CB9C7655FFF3FFF9FB79FB87225FFB84" box="[1210,1313,1072,1096]" name="Germany" pageId="9" pageNumber="8">Germany</collectingCountry>
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||
), who is
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||
<taxonomicName id="748B4D46FFF3FFF9FA57FB8624FFFBAB" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="7" phylum="Arthropoda" rank="species" species="young">a young</taxonomicName>
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||
scorpiologist especially devoted to the genus
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||
<taxonomicName id="748B4D46FFF3FFF9FCFCFBDB24B6FB48" authorityName="Ehrenberg" authorityYear="1828" box="[831,968,1132,1156]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81FFEAD7FFF3FFF9FCFCFBDB24B6FB48" box="[831,968,1132,1156]" italics="true" pageId="9" pageNumber="8">Androctonus</emphasis>
|
||
</taxonomicName>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB91654EFFF3FFF9FCFCFB1324FAF8B1" pageId="9" pageNumber="8" type="diagnosis">
|
||
<paragraph id="B33436C5FFF3FFF9FCFCFB1324FAF8B1" blockId="9.[831,1441,1188,1917]" pageId="9" pageNumber="8">
|
||
DIAGNOSIS. Adults large (females
|
||
<quantity id="74739B20FFF3FFF9FB15FB132228FB77" box="[1238,1366,1188,1212]" metricMagnitude="-2" metricUnit="m" metricValue="8.05" metricValueMax="8.8" metricValueMin="7.3" pageId="9" pageNumber="8" unit="mm" value="80.5" valueMax="88.0" valueMin="73.0">73–88 mm</quantity>
|
||
, adult males unknown) for the genus. Coloration basically dark reddish brown; pedipalps and legs apically yellowish. Pedipalps long and slender, with hand conspicuously narrower than patella and weakly carinate; fixed and movable fingers with 14–15 principal rows of denticles, basal lobe/notch combination absent. Carapace densely granulose, with anterior and posterior median carinae strong. Sternite V with a smooth patch in both sexes (larger and bulkier in males), III without carinae, IV–V with two carinae, VI–VII with four carinae. Metasoma robust and deep (females have segments I–III wider than long, and all segments wider than deep), with 10-10-8-8- 5 complete to essentially complete carinae, most of which are coarsely serrate to crenulate; segments II–IV dorsolateral carinae with 1–2 enlarged terminal denticles; segment V ventrolateral carinae with 4–5 acutely flared denticles and anal arch with 3–5 poorly defined crenate lobes; all intercarinal spaces densely granulose. Telson slender, with vesicle small but somewhat globose, and aculeus remarkably longer than vesicle. Leg bristlecombs well developed on I–III, poorly defined on IV. Pectines with 28 teeth in males and
|
||
<quantity id="74739B20FFF3FFF9FAD1F8F12208F892" box="[1298,1398,1862,1886]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="5.842" metricValueMin="5.334" pageId="9" pageNumber="8" unit="in" value="22.0" valueMax="23.0" valueMin="21.0">21–23 in</quantity>
|
||
females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="E7F4664DFFF0FFFAFF03F89A26E7F8B0" pageId="10" pageNumber="9" startId="10.[192,278,1837,1861]" targetBox="[229,1416,203,1790]" targetPageId="10">
|
||
<paragraph id="B33436C5FFF0FFFAFF03F89A26E7F8B0" blockId="10.[192,1441,1837,1916]" pageId="10" pageNumber="9">
|
||
<emphasis id="81FFEAD7FFF0FFFAFF03F89A247DF888" bold="true" box="[192,771,1837,1861]" pageId="10" pageNumber="9">
|
||
Figures 19–25:
|
||
<taxonomicName id="748B4D46FFF0FFFAFEACF898254FF888" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[367,561,1839,1860]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF0FFFAFEACF898254FF888" bold="true" box="[367,561,1839,1860]" italics="true" pageId="10" pageNumber="9">Androctonus turieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9ACC57ACFFF0FFFAFDF4F8982519F888" box="[567,615,1839,1860]" pageId="10" pageNumber="9" rank="species">sp.n.</taxonomicNameLabel>
|
||
Figures 19–20.
|
||
</emphasis>
|
||
Dorsal (19) and ventral (20) views, ♀ paratype (FKCP).
|
||
<emphasis id="81FFEAD7FFF0FFFAFAE9F89827A1F8AC" bold="true" pageId="10" pageNumber="9">Figures 21– 25.</emphasis>
|
||
Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E7F4664DFFF1FFFBFF03FA872340FAA8" pageId="11" pageNumber="10" startId="11.[192,278,1328,1352]" targetBox="[203,1421,183,1283]" targetPageId="11">
|
||
<paragraph id="B33436C5FFF1FFFBFF03FA872340FAA8" blockId="11.[192,1440,1328,1380]" pageId="11" pageNumber="10">
|
||
<emphasis id="81FFEAD7FFF1FFFBFF03FA8725FFFA8B" bold="true" box="[192,641,1328,1352]" pageId="11" pageNumber="10">
|
||
Figures 26–31:
|
||
<taxonomicName id="748B4D46FFF1FFFBFEB5FA852542FA8B" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[374,572,1330,1351]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF1FFFBFEB5FA852542FA8B" bold="true" box="[374,572,1330,1351]" italics="true" pageId="11" pageNumber="10">Androctonus turieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9ACC57ACFFF1FFFBFD84FA8525FFFA8B" box="[583,641,1330,1351]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, pedipalp chela dorsal (26) and external (27), pedipalp patela dorsal (28) and external (29), pedipalp femur dorsal (30), and right movable finger internal (31), ♀ holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="FB91654EFFF1FFFEFF03FA2F26DAF8EF" lastPageId="14" lastPageNumber="13" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="B33436C5FFF1FFFBFF03FA2F255BF908" blockId="11.[192,802,1432,1916]" pageId="11" pageNumber="10">
|
||
DESCRIPTION (adult female
|
||
<typeStatus id="6C308867FFF1FFFBFE31FA2F252AFA7C" box="[498,596,1432,1456]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
). Coloration (
|
||
<figureCitation id="2BB02A40FFF1FFFBFD28FA2F267AFA02" captionStart-0="Figures 19–25" captionStart-1="Figures 26–31" captionStart-2="Figures 32–35" captionStartId-0="10.[192,278,1837,1861]" captionStartId-1="11.[192,278,1328,1352]" captionStartId-2="12.[192,278,1114,1138]" captionTargetBox-0="[229,1416,203,1790]" captionTargetBox-1="[203,1421,183,1283]" captionTargetBox-2="[202,1426,193,1053]" captionTargetId-0="figure-63@10.[212,1416,192,1802]" captionTargetId-1="figure-316@11.[192,1440,168,1293]" captionTargetId-2="figure-481@12.[192,1440,192,1080]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." captionText-1="Figures 26–31: Androctonus turieli sp. n., pedipalp chela dorsal (26) and external (27), pedipalp patela dorsal (28) and external (29), pedipalp femur dorsal (30), and right movable finger internal (31), ♀ holotype." captionText-2="Figures 32–35: Figures 32–34. Androctonus turieli sp. n., metasoma and telson dorsal (32), ventral (33) and lateral (34) views, ♀ holotype. Figure 35. Androctonus sergenti Vachon, 1948, metasoma and telson ventral, ♂, Morocco, Anti Atlas Mts., 62 km southeast of Agadir, 30°03.35'N 09°04.4'W, 798 m a. s. l. (FKCP)." pageId="11" pageNumber="10">Figs. 19–34</figureCitation>
|
||
) uniformly dark reddish brown to unaided eye; ocular tubercles, eyes and distal part of aculeus blackish. Chelicerae very deeply infuscate. Pedipalps and legs progressively lighter apically, becoming yellowish, with carinae subtly infuscate and very faint traces of dark reticulations between carinae. Pectines yellow. Sternite V with the smooth patch yellow. Metasoma with carinae infuscate. Telson uniformly dark reddish brown, only with distal half of aculeus darker.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF1FFFBFF03F97D2571F8F2" blockId="11.[192,802,1432,1916]" pageId="11" pageNumber="10">
|
||
CHELICERAE (
|
||
<figureCitation id="2BB02A40FFF1FFFBFE9DF97D26CDF92E" box="[350,435,1738,1762]" captionStart="Figures 19–25" captionStartId="10.[192,278,1837,1861]" captionTargetBox="[229,1416,203,1790]" captionTargetId="figure-63@10.[212,1416,192,1802]" captionTargetPageId="10" captionText="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." pageId="11" pageNumber="10">Fig. 21</figureCitation>
|
||
). With dentition typical for the genus; tegument polished and smooth except on dorsodistal portion of manus, which possesses coarse granules arranged in longitudinal ridges.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF1FFFBFF03F8F22213F985" blockId="11.[192,802,1432,1916]" lastBlockId="11.[831,1441,1432,1916]" pageId="11" pageNumber="10">
|
||
CARAPACE (
|
||
<figureCitation id="2BB02A40FFF1FFFBFE89F8F226DEF891" box="[330,416,1861,1885]" captionStart="Figures 19–25" captionStartId="10.[192,278,1837,1861]" captionTargetBox="[229,1416,203,1790]" captionTargetId="figure-63@10.[212,1416,192,1802]" captionTargetPageId="10" captionText="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." pageId="11" pageNumber="10">Fig. 21</figureCitation>
|
||
). Trapezoidal, slightly wider than long and with all carinae strongly granulose, but not fused into any particular configuration; tegument very densely covered by granules of all sizes; anterior margin very shallowly V-shaped (almost straight), with one pair of stout macrosetae; median eyes separate by much more than one ocular diameter; five pairs of lateral eyes: three large and aligned, plus two small and slightly offset.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF1FFFCFCFCF9E6266EF948" blockId="11.[831,1441,1432,1916]" lastBlockId="12.[192,802,1245,1913]" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
|
||
MESOSOMA (
|
||
<figureCitation id="2BB02A40FFF1FFFBFC12F9E72327F9A4" box="[977,1113,1616,1640]" captionStart="Figures 19–25" captionStartId="10.[192,278,1837,1861]" captionTargetBox="[229,1416,203,1790]" captionTargetId="figure-63@10.[212,1416,192,1802]" captionTargetPageId="10" captionText="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." pageId="11" pageNumber="10">Figs. 22–23</figureCitation>
|
||
). Tergites very densely covered by granules of all sizes; I without carinae, II–VI with three carinae (median and submedians), which are moderate to strong and granulose, but not projected beyond posterior margin; VII with five carinae (median, submedians and laterals) which are strong and serrate. Sternum standard for the genus:
|
||
<typeStatus id="6C308867FFF1FFFBFB63F8BE23B0F8ED" box="[1184,1230,1801,1825]" pageId="11" pageNumber="10">type</typeStatus>
|
||
1, relatively small, and markedly triangular in shape. Pectines long (reaching leg IV coxa/trochanter joint), narrow and densely setose; tooth count 22/22; basal plate heavily sclerotized and wider than long, anterior margin with strong median indentation, posterior margin widely convex. Sternites very sparsely setose; III–VI smooth and glossy, with well-developed granulose lateral areas and spiracles very elongate and slit-like, VI finely and very densely granulose; III without carinae, IV–V with a pair of lateral carinae that surrounds spiracles anteriorly and internally reaching posterior margin of sternite, VI with two pairs of carinae (a very weakly granulose to subcostate ventrosubmedian pair and a serrate lateral pair), VII with the same two pairs of carinae, all strongly serrate; sternite V with smooth patch medium-sized, subtriangular to subcordate, moderately bulky, sulcate along midline, and yellow.
|
||
</paragraph>
|
||
<caption id="E7F4664DFFF6FFFCFF03FBED24FAFB65" pageId="12" pageNumber="11" startId="12.[192,278,1114,1138]" targetBox="[202,1426,193,1053]" targetPageId="12">
|
||
<paragraph id="B33436C5FFF6FFFCFF03FBED24FAFB65" blockId="12.[192,1440,1114,1193]" pageId="12" pageNumber="11">
|
||
<emphasis id="81FFEAD7FFF6FFFCFF03FBED245CFBBD" bold="true" box="[192,802,1114,1138]" pageId="12" pageNumber="11">
|
||
Figures 32–35: Figures 32–34.
|
||
<taxonomicName id="748B4D46FFF6FFFCFDDAFBEB25A1FBBD" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[537,735,1116,1137]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF6FFFCFDDAFBEB25A1FBBD" bold="true" box="[537,735,1116,1137]" italics="true" pageId="12" pageNumber="11">Androctonus turieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9ACC57ACFFF6FFFCFD2BFBEB245CFBBD" box="[744,802,1116,1137]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, metasoma and telson dorsal (32), ventral (33) and lateral (34) views, ♀ holotype.
|
||
<emphasis id="81FFEAD7FFF6FFFCFEBCFBCF2696FB41" bold="true" box="[383,488,1144,1165]" pageId="12" pageNumber="11">Figure 35.</emphasis>
|
||
<taxonomicName id="748B4D46FFF6FFFCFE2CFBCF242DFB42" authority="Vachon, 1948" authorityName="Vachon" authorityYear="1948" box="[495,851,1144,1166]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="sergenti">
|
||
<emphasis id="81FFEAD7FFF6FFFCFE2CFBCF25BBFB41" bold="true" box="[495,709,1144,1165]" italics="true" pageId="12" pageNumber="11">Androctonus sergenti</emphasis>
|
||
Vachon, 1948
|
||
</taxonomicName>
|
||
, metasoma and telson ventral, ♂, Morocco, Anti Atlas Mts., 62 km southeast of Agadir, 30°03.35'N 09°04.4'W, 798 m a. s. l. (FKCP).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B33436C5FFF6FFFDFF03F93C2471FC23" blockId="12.[192,802,1245,1913]" lastBlockId="13.[192,802,922,1805]" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
|
||
METASOMA AND TELSON (
|
||
<figureCitation id="2BB02A40FFF6FFFCFDC1F93C25EEF96F" box="[514,656,1675,1699]" captionStart="Figures 32–35" captionStartId="12.[192,278,1114,1138]" captionTargetBox="[202,1426,193,1053]" captionTargetId="figure-481@12.[192,1440,192,1080]" captionTargetPageId="12" captionText="Figures 32–35: Figures 32–34. Androctonus turieli sp. n., metasoma and telson dorsal (32), ventral (33) and lateral (34) views, ♀ holotype. Figure 35. Androctonus sergenti Vachon, 1948, metasoma and telson ventral, ♂, Morocco, Anti Atlas Mts., 62 km southeast of Agadir, 30°03.35'N 09°04.4'W, 798 m a. s. l. (FKCP)." pageId="12" pageNumber="11">Figs. 32–34</figureCitation>
|
||
). Metasoma essentially glabrous, with all segments robust and deep (I–III wider than long, IV–V longer than wide; all segments wider than deep); intercarinal tegument of dorsal surface of I–IV smooth but with a large arrowheadshaped granulose patches medially, flanked by a few finely granulose reticulations, of lateral and ventral surfaces densely granulose and forming a clearly reticulate pattern; dorsal furrow moderately shallow and narrow on all segments; I–II with ten complete to almost complete carinae (lateral inframedian carinae becomes less defined on the basal third of II), III–IV with eight (even though two coarse granules remain as vestiges of the lateral inframedian carinae on the distal portion of II), and V with five: dorsolateral carinae on I–IV strongly serrato-crenulate to subdentate and progressively raised with 1–2 enlarged terminal denticles, lateral supramedian carinae strongly serrato-crenulate to subdentate and progressively raised on I–IV, moderately granulose and even on V, lateral inframedian carinae strongly serratocrenulate on I, moderately granulose to denticulate and progressively raised on II, ventrolateral carinae strongly serrato-crenulate to subdentate and even on I–IV, strongly dentate and progressively raised distally on V with 4–5 acutely flared denticles, anal arch with 3– 5 very poorly defined round lobes, ventrosubmedian carina on V strongly serrate to denticulate. Telson slen- der and sparsely setose; vesicle small but somewhat globose (1.34 times longer than wide, 1.13 times wider than deep), tegument glossy, with abundant coarse but weak granules and a coarse ventromedian carinae; subaculear tubercle absent; aculeus very long and moderately thick, markedly longer than vesicle, and evenly curved.
|
||
</paragraph>
|
||
<caption id="E7F4664DFFF7FFFDFF03FCF925AAFCA9" box="[192,724,846,870]" pageId="13" pageNumber="12" startId="13.[192,267,846,870]" targetBox="[203,1426,167,811]" targetPageId="13">
|
||
<paragraph id="B33436C5FFF7FFFDFF03FCF925AAFCA9" blockId="13.[192,724,846,870]" box="[192,724,846,870]" pageId="13" pageNumber="12">
|
||
<emphasis id="81FFEAD7FFF7FFFDFF03FCF92648FCAA" bold="true" box="[192,310,846,870]" pageId="13" pageNumber="12">Figure 36:</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="748B4D46FFF7FFFDFE18FCE725E0FCA9" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[475,670,848,869]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF7FFFDFE18FCE725E0FCA9" bold="true" box="[475,670,848,869]" italics="true" pageId="13" pageNumber="12">Androctonus turieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9ACC57ACFFF7FFFDFD67FCE725AAFCA9" box="[676,724,848,869]" pageId="13" pageNumber="12" rank="species">sp.n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B33436C5FFF7FFFDFF03FC402475FACF" blockId="13.[192,802,922,1805]" pageId="13" pageNumber="12">
|
||
LEGS (
|
||
<figureCitation id="2BB02A40FFF7FFFDFECBFC4126F7FBC2" box="[264,393,1014,1038]" captionStart="Figures 19–25" captionStartId="10.[192,278,1837,1861]" captionTargetBox="[229,1416,203,1790]" captionTargetId="figure-63@10.[212,1416,192,1802]" captionTargetPageId="10" captionText="Figures 19–25: Androctonus turieli sp.n. Figures 19–20. Dorsal (19) and ventral (20) views, ♀ paratype (FKCP). Figures 21– 25. Carapace with chelicerae and tergite I (21), pectinal area (22), sternites III–VII (23), tarsomeres of third (24) and fourth (25) right legs, ♀ holotype." pageId="13" pageNumber="12">Figs. 24–25</figureCitation>
|
||
). Legs long, slender, and only sparsely setose; bristlecombs well-developed on I–III, poorly defined on IV; all carinae smooth to subserrate; intercarinal tegument coriaceous; tibial spurs absent from I– II, very large on III–IV; prolateral and retrolateral pedal spurs long and slender in all legs; ventral surface of all tarsomere II sharply edged and with two very irregular ventrosubmedian rows of thin spiniform setae, without median row of spinules; claws long and evenly curved.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF7FFFDFF03FABC2652F8C1" blockId="13.[192,802,922,1805]" pageId="13" pageNumber="12">
|
||
PEDIPALPS (
|
||
<figureCitation id="2BB02A40FFF7FFFDFE84FABD26B0FAEE" box="[327,462,1290,1314]" captionStart="Figures 26–31" captionStartId="11.[192,278,1328,1352]" captionTargetBox="[203,1421,183,1283]" captionTargetId="figure-316@11.[192,1440,168,1293]" captionTargetPageId="11" captionText="Figures 26–31: Androctonus turieli sp. n., pedipalp chela dorsal (26) and external (27), pedipalp patela dorsal (28) and external (29), pedipalp femur dorsal (30), and right movable finger internal (31), ♀ holotype." pageId="13" pageNumber="12">Figs. 26–31</figureCitation>
|
||
). Pedipalps long and slender, essentially glabrous and orthobothriotaxic A-β. Femur slender and straight; all carinae moderate and finely granulose to denticulate; intercarinal tegument very finely and densely granulose. Patella slender and straight; all carinae moderate and finely granulose to costate; intercarinal tegument irregularly but finely granulose. Chela with hand narrower than patella, oval (1.58 times longer than wide) and with all carinae obsolete to weakly costate, intercarinal tegument coriaceous, with abundant minute granules on all surfaces; fingers long and slender (movable finger 2.50 times longer than underhand), evenly curved and with basal lobe/notch combination absent, fixed fingers with 15 principal rows of granules; movable fingers with 15 principal rows of granules and three terminal granules (large terminal denticle not included).
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF7FFFDFF03F88423F5FC1C" blockId="13.[192,801,1842,1897]" lastBlockId="13.[831,1440,922,977]" pageId="13" pageNumber="12">
|
||
MALE (juvenile
|
||
<typeStatus id="6C308867FFF7FFFDFEB1F88526A3F886" box="[370,477,1842,1866]" pageId="13" pageNumber="12" type="paratype">paratypes</typeStatus>
|
||
). Primary characters (i.e., not maturity-dependent) that evidence sexual dimorphism are:
|
||
<emphasis id="81FFEAD7FFF7FFFDFCB1FC2D24F7FC7E" bold="true" box="[882,905,922,946]" pageId="13" pageNumber="12">1)</emphasis>
|
||
genital papillae present;
|
||
<emphasis id="81FFEAD7FFF7FFFDFB69FC2D23BEFC7E" bold="true" box="[1194,1216,922,946]" pageId="13" pageNumber="12">2)</emphasis>
|
||
pectines larger and with much higher tooth counts.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF7FFFDFCFCFC4023EEFA8D" blockId="13.[831,1440,1014,1498]" pageId="13" pageNumber="12">
|
||
VARIATION. The size differences among adults studied by us indicate the existence of two size-classes in females (
|
||
<tableCitation id="FE09037EFFF7FFFDFC51FB84249CFB87" box="[914,994,1075,1099]" captionStart="Table 2" captionStartId="9.[192,257,793,817]" captionText="Table 2: Measurements (mm) of two adult types of Androctonus turieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (H), left (L), right (R). Note: telson vesicle length is not given because it narrows evenly into aculeus, without any clear definition point." pageId="13" pageNumber="12">Tab. 2</tableCitation>
|
||
). Diagnostically relevant morphometric ratios revealed to be positively correlated to size: the
|
||
<typeStatus id="6C308867FFF7FFFDFCFCFBC624E2FB45" box="[831,924,1137,1161]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
and same-sized
|
||
<typeStatus id="6C308867FFF7FFFDFBA9FBC623B8FB45" box="[1130,1222,1137,1161]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
|
||
(RTOC) are very similar, but the larger
|
||
<typeStatus id="6C308867FFF7FFFDFBFAFB2723EBFB64" box="[1081,1173,1168,1192]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
|
||
(FKCP) has metasomal segments and pedipalps slightly more robust (
|
||
<tableCitation id="FE09037EFFF7FFFDFA85FB1922ECFB0A" box="[1350,1426,1198,1222]" captionStart="Table 3" captionStartId="14.[192,257,645,669]" captionText="Table 3: Comparison between Androctonus bicolor and its closest relatives Androctonus tenuissimus and Androctonus turieli sp. n., based upon selected morphometric ratios of adult females (adult males of the latter are still known), and including typespecimens. Abbreviations: length (L), width (W)." pageId="13" pageNumber="12">Tab. 3</tableCitation>
|
||
). This correlation is a very common occurrence among scorpions and has already been documented in other members of this genus studied recently (Kovařík & Ah- med, 2013; Teruel et al., 2013).
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF7FFFDFCACFAF023A2FA16" blockId="13.[831,1440,1014,1498]" pageId="13" pageNumber="12">Count of teeth per pecten varied in females as follows: 21 (1), 22 (3), and 23 (2); both juvenile male possess 28/28. Most fingers possess 15 principal rows of denticles, but in a few the two basalmost rows are poorly defined or fused to yield a count of 14.</paragraph>
|
||
<paragraph id="B33436C5FFF7FFFDFCFCFA48233AF8A5" blockId="13.[831,1441,1535,1897]" pageId="13" pageNumber="12">
|
||
AFFINITIES. Amongst the 22 species currently recognized as valid in this genus (Turiel, 2013; Teruel
|
||
<emphasis id="81FFEAD7FFF7FFFDFAA1F9A822DFF9FA" box="[1378,1441,1566,1590]" italics="true" pageId="13" pageNumber="12">et al.,</emphasis>
|
||
2013), only
|
||
<taxonomicName id="748B4D46FFF7FFFDFC0BF98A2348F999" authorityName="Ehrenberg" authorityYear="1828" box="[968,1078,1597,1621]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF7FFFDFC0BF98A2348F999" box="[968,1078,1597,1621]" italics="true" pageId="13" pageNumber="12">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="748B4D46FFF7FFFDFB8BF98A2398F999" box="[1096,1254,1597,1621]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="tenuissimus">
|
||
<emphasis id="81FFEAD7FFF7FFFDFB8BF98A2398F999" box="[1096,1254,1597,1621]" italics="true" pageId="13" pageNumber="12">A. tenuissimus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="748B4D46FFF7FFFDFAEAF98A22DEF999" authorityName="Vachon" authorityYear="1948" box="[1321,1440,1597,1621]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="sergenti">
|
||
<emphasis id="81FFEAD7FFF7FFFDFAEAF98A22DEF999" box="[1321,1440,1597,1621]" italics="true" pageId="13" pageNumber="12">A. sergenti</emphasis>
|
||
</taxonomicName>
|
||
do share with
|
||
<taxonomicName id="748B4D46FFF7FFFDFC19F9EC2345F9BF" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[986,1083,1627,1651]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF7FFFDFC19F9EC2345F9BF" box="[986,1083,1627,1651]" italics="true" pageId="13" pageNumber="12">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF7FFFDFB87F9EB2307F9B8" bold="true" box="[1092,1145,1628,1652]" pageId="13" pageNumber="12">
|
||
<taxonomicNameLabel id="9ACC57ACFFF7FFFDFB87F9EB2307F9B8" box="[1092,1145,1628,1652]" pageId="13" pageNumber="12" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the following combination of three characters in adults of both sexes:
|
||
<emphasis id="81FFEAD7FFF7FFFDFAD6F9CD2252F95E" bold="true" box="[1301,1324,1658,1682]" pageId="13" pageNumber="12">1)</emphasis>
|
||
coloration uniformly dark reddish brown to blackish;
|
||
<emphasis id="81FFEAD7FFF7FFFDFAD9F92E224EF97D" bold="true" box="[1306,1328,1689,1713]" pageId="13" pageNumber="12">2)</emphasis>
|
||
pedipalps very slender, with chelae much narrower than patella;
|
||
<emphasis id="81FFEAD7FFF7FFFDFA49F90F22DEF91C" bold="true" box="[1418,1440,1720,1744]" pageId="13" pageNumber="12">3)</emphasis>
|
||
pedipalp fingers without basal lobe/notch combination. All other species of
|
||
<taxonomicName id="748B4D46FFF7FFFDFBE0F94223D2F8C1" authorityName="Ehrenberg" authorityYear="1828" box="[1059,1196,1781,1805]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81FFEAD7FFF7FFFDFBE0F94223D2F8C1" box="[1059,1196,1781,1805]" italics="true" pageId="13" pageNumber="12">Androctonus</emphasis>
|
||
</taxonomicName>
|
||
which have the same coloration also possess adult pedipalp chelae wider than patella, and with basal lobe/notch combination of fingers present at least in males.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF4FFFEFE5EFF6D223FFDA8" pageId="14" pageNumber="13">
|
||
<table id="C18BC465FFF4000FFF28FF7C2221FDA8" box="[235,1375,203,612]" gridcols="4" gridrows="12" pageId="14" pageNumber="13">
|
||
<tr id="0DBB3487FFF4000FFF28FF7C2221FECB" box="[235,1375,203,263]" gridrow="0" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FF7C25F1FECB" box="[235,655,203,263]" gridcol="0" gridrow="0" pageId="14" pageNumber="13">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE5EFF6D2693FF38" bold="true" box="[413,493,218,244]" pageId="14" pageNumber="13">Ratios</emphasis>
|
||
</th>
|
||
<th id="4E6A5DFBFFF4000FFD26FF7C2423FECB" box="[741,861,203,263]" gridcol="1" gridrow="0" pageId="14" pageNumber="13">
|
||
<emphasis id="81FFEAD7FFF4FFFEFD26FF7C242CFECB" bold="true" pageId="14" pageNumber="13">
|
||
<taxonomicName id="748B4D46FFF4FFFEFD26FF7C2422FF29" authorityName="Ehrenberg" authorityYear="1828" box="[741,860,203,229]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFD26FF7C2422FF29" bold="true" box="[741,860,203,229]" italics="true" pageId="14" pageNumber="13">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
(n = 10)
|
||
</emphasis>
|
||
</th>
|
||
<th id="4E6A5DFBFFF4000FFC7AFF7C2310FECB" box="[953,1134,203,263]" gridcol="2" gridrow="0" pageId="14" pageNumber="13">
|
||
<emphasis id="81FFEAD7FFF4FFFEFC7AFF7C2342FECB" bold="true" pageId="14" pageNumber="13">
|
||
<taxonomicName id="748B4D46FFF4FFFEFC7AFF7C235AFF29" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[953,1060,203,229]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFC7AFF7C235AFF29" bold="true" box="[953,1060,203,229]" italics="true" pageId="14" pageNumber="13">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFBEFFF7C2310FF29" box="[1068,1134,203,229]" pageId="14" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
(n = 3)
|
||
</emphasis>
|
||
</th>
|
||
<th id="4E6A5DFBFFF4000FFB6EFF7C2221FECB" box="[1197,1375,203,263]" gridcol="3" gridrow="0" pageId="14" pageNumber="13">
|
||
<emphasis id="81FFEAD7FFF4FFFEFB6EFF7C2251FECB" bold="true" box="[1197,1375,203,263]" pageId="14" pageNumber="13">
|
||
<taxonomicName id="748B4D46FFF4FFFEFB6EFF7C2221FF29" box="[1197,1375,203,229]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="tenuissimus">
|
||
<emphasis id="81FFEAD7FFF4FFFEFB6EFF7C2221FF29" bold="true" box="[1197,1375,203,229]" italics="true" pageId="14" pageNumber="13">A. tenuissimus</emphasis>
|
||
</taxonomicName>
|
||
(n = 6)
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FEA62221FEE6" box="[235,1375,273,298]" gridrow="1" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FEA625F1FEE6" box="[235,655,273,298]" gridcol="0" gridrow="1" pageId="14" pageNumber="13">Metasomal segment I (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FEA62423FEE6" box="[741,861,273,298]" gridcol="1" gridrow="1" pageId="14" pageNumber="13">0.80</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFEA62310FEE6" box="[953,1134,273,298]" gridcol="2" gridrow="1" pageId="14" pageNumber="13">0.87–0.88</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFEA62221FEE6" box="[1197,1375,273,298]" gridcol="3" gridrow="1" pageId="14" pageNumber="13">1.02</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FE872221FE85" box="[235,1375,304,329]" gridrow="2" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FE8725F1FE85" box="[235,655,304,329]" gridcol="0" gridrow="2" pageId="14" pageNumber="13">Metasomal segment II (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FE872423FE85" box="[741,861,304,329]" gridcol="1" gridrow="2" pageId="14" pageNumber="13">0.76–0.87</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFE872310FE85" box="[953,1134,304,329]" gridcol="2" gridrow="2" pageId="14" pageNumber="13">0.90–0.91</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFE872221FE85" box="[1197,1375,304,329]" gridcol="3" gridrow="2" pageId="14" pageNumber="13">1.02–1.08</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FEF92221FEAB" box="[235,1375,334,359]" gridrow="3" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FEF925F1FEAB" box="[235,655,334,359]" gridcol="0" gridrow="3" pageId="14" pageNumber="13">Metasomal segment III (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FEF92423FEAB" box="[741,861,334,359]" gridcol="1" gridrow="3" pageId="14" pageNumber="13">0.75–0.79</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFEF92310FEAB" box="[953,1134,334,359]" gridcol="2" gridrow="3" pageId="14" pageNumber="13">0.91–0.93</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFEF92221FEAB" box="[1197,1375,334,359]" gridcol="3" gridrow="3" pageId="14" pageNumber="13">1.01–1.07</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FEDA2221FE4A" box="[235,1375,365,390]" gridrow="4" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FEDA25F1FE4A" box="[235,655,365,390]" gridcol="0" gridrow="4" pageId="14" pageNumber="13">Metasomal segment IV (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FEDA2423FE4A" box="[741,861,365,390]" gridcol="1" gridrow="4" pageId="14" pageNumber="13">0.96–0.99</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFEDA2310FE4A" box="[953,1134,365,390]" gridcol="2" gridrow="4" pageId="14" pageNumber="13">1.10–1.17</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFEDA2221FE4A" box="[1197,1375,365,390]" gridcol="3" gridrow="4" pageId="14" pageNumber="13">1.17–1.31</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FE3B2221FE68" box="[235,1375,396,420]" gridrow="5" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FE3B25F1FE68" box="[235,655,396,420]" gridcol="0" gridrow="5" pageId="14" pageNumber="13">Metasomal segment V (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FE3B2423FE68" box="[741,861,396,420]" gridcol="1" gridrow="5" pageId="14" pageNumber="13">1.07–1.23</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFE3B2310FE68" box="[953,1134,396,420]" gridcol="2" gridrow="5" pageId="14" pageNumber="13">1.24–1.35</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFE3B2221FE68" box="[1197,1375,396,420]" gridcol="3" gridrow="5" pageId="14" pageNumber="13">1.38–1.63</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FE1D2221FE0E" box="[235,1375,426,450]" gridrow="6" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FE1D25F1FE0E" box="[235,655,426,450]" gridcol="0" gridrow="6" pageId="14" pageNumber="13">Total (L) / Pedipalp (L)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FE1D2423FE0E" box="[741,861,426,450]" gridcol="1" gridrow="6" pageId="14" pageNumber="13">2.35–2.61</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFE1D2310FE0E" box="[953,1134,426,450]" gridcol="2" gridrow="6" pageId="14" pageNumber="13">2.48–2.88</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFE1D2221FE0E" box="[1197,1375,426,450]" gridcol="3" gridrow="6" pageId="14" pageNumber="13">2.26–2.49</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FE7E2221FE2D" box="[235,1375,457,481]" gridrow="7" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FE7E25F1FE2D" box="[235,655,457,481]" gridcol="0" gridrow="7" pageId="14" pageNumber="13">Pedipalp femur (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FE7E2423FE2D" box="[741,861,457,481]" gridcol="1" gridrow="7" pageId="14" pageNumber="13">3.20–3.43</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFE7E2310FE2D" box="[953,1134,457,481]" gridcol="2" gridrow="7" pageId="14" pageNumber="13">3.36–3.41</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFE7E2221FE2D" box="[1197,1375,457,481]" gridcol="3" gridrow="7" pageId="14" pageNumber="13">4.05–4.40</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FE5F2221FDCC" box="[235,1375,488,512]" gridrow="8" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FE5F25F1FDCC" box="[235,655,488,512]" gridcol="0" gridrow="8" pageId="14" pageNumber="13">Pedipalp patella (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FE5F2423FDCC" box="[741,861,488,512]" gridcol="1" gridrow="8" pageId="14" pageNumber="13">2.84–3.08</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFE5F2310FDCC" box="[953,1134,488,512]" gridcol="2" gridrow="8" pageId="14" pageNumber="13">3.07–3.18</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFE5F2221FDCC" box="[1197,1375,488,512]" gridcol="3" gridrow="8" pageId="14" pageNumber="13">3.71–4.04</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FDB12221FDD2" box="[235,1375,518,542]" gridrow="9" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FDB125F1FDD2" box="[235,655,518,542]" gridcol="0" gridrow="9" pageId="14" pageNumber="13">Pedipalp chela (L/W)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FDB12423FDD2" box="[741,861,518,542]" gridcol="1" gridrow="9" pageId="14" pageNumber="13">5.64–6.45</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFDB12310FDD2" box="[953,1134,518,542]" gridcol="2" gridrow="9" pageId="14" pageNumber="13">5.48–5.54</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFDB12221FDD2" box="[1197,1375,518,542]" gridcol="3" gridrow="9" pageId="14" pageNumber="13">7.74–8.32</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FD922221FDF1" box="[235,1375,549,573]" gridrow="10" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FD9225F1FDF1" box="[235,655,549,573]" gridcol="0" gridrow="10" pageId="14" pageNumber="13">Pedipalp chela (L) / movable finger (L)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FD922423FDF1" box="[741,861,549,573]" gridcol="1" gridrow="10" pageId="14" pageNumber="13">1.33–1.38</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFD922310FDF1" box="[953,1134,549,573]" gridcol="2" gridrow="10" pageId="14" pageNumber="13">1.36–1.40</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFD922221FDF1" box="[1197,1375,549,573]" gridcol="3" gridrow="10" pageId="14" pageNumber="13">1.31–1.32</td>
|
||
</tr>
|
||
<tr id="0DBB3487FFF4000FFF28FDFD2221FDA8" box="[235,1375,586,612]" gridrow="11" pageId="14" pageNumber="13">
|
||
<th id="4E6A5DFBFFF4000FFF28FDFD25F1FDA8" box="[235,655,586,612]" gridcol="0" gridrow="11" pageId="14" pageNumber="13">Total (L)</th>
|
||
<td id="4E6A5DFBFFF4000FFD26FDFD2423FDA8" box="[741,861,586,612]" gridcol="1" gridrow="11" pageId="14" pageNumber="13">64.2–80.9</td>
|
||
<td id="4E6A5DFBFFF4000FFC7AFDFD2310FDA8" box="[953,1134,586,612]" gridcol="2" gridrow="11" pageId="14" pageNumber="13">73.5–87.7</td>
|
||
<td id="4E6A5DFBFFF4000FFB6EFDFD2221FDA8" box="[1197,1375,586,612]" gridcol="3" gridrow="11" pageId="14" pageNumber="13">62.7–86.5</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="E7F4664DFFF4FFFEFF03FD3225E5FD02" pageId="14" pageNumber="13" startId="14.[192,257,645,669]">
|
||
<paragraph id="B33436C5FFF4FFFEFF03FD3225E5FD02" blockId="14.[192,1441,645,718]" pageId="14" pageNumber="13">
|
||
<emphasis id="81FFEAD7FFF4FFFEFF03FD32265EFD51" bold="true" box="[192,288,645,669]" pageId="14" pageNumber="13">Table 3:</emphasis>
|
||
Comparison between
|
||
<taxonomicName id="748B4D46FFF4FFFEFE3CFD3025B9FD51" authorityName="Ehrenberg" authorityYear="1828" box="[511,711,647,669]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE3CFD3025B9FD51" box="[511,711,647,669]" italics="true" pageId="14" pageNumber="13">Androctonus bicolor</emphasis>
|
||
</taxonomicName>
|
||
and its closest relatives
|
||
<taxonomicName id="748B4D46FFF4FFFEFC7FFD3023D1FD50" box="[956,1199,647,669]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="tenuissimus">
|
||
<emphasis id="81FFEAD7FFF4FFFEFC7FFD3023D1FD50" box="[956,1199,647,669]" italics="true" pageId="14" pageNumber="13">Androctonus tenuissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="748B4D46FFF4FFFEFB20FD3022DEFD51" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[1251,1440,647,669]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFB20FD3022DEFD51" box="[1251,1440,647,669]" italics="true" pageId="14" pageNumber="13">Androctonus turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF4FFFEFF03FD172786FD79" bold="true" box="[192,248,672,693]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFF03FD172786FD79" box="[192,248,672,693]" pageId="14" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, based upon selected morphometric ratios of adult females (adult males of the latter are still known), and including typespecimens. Abbreviations: length (L), width (W).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B33436C5FFF4FFFEFF33FD4E2583FC2B" blockId="14.[192,802,761,1827]" pageId="14" pageNumber="13">
|
||
Of them, the easiest to separate is
|
||
<taxonomicName id="748B4D46FFF4FFFEFD46FD4E2581FCDD" authorityName="Vachon" authorityYear="1948" box="[645,767,761,785]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="sergenti">
|
||
<emphasis id="81FFEAD7FFF4FFFEFD46FD4E2581FCDD" box="[645,767,761,785]" italics="true" pageId="14" pageNumber="13">A. sergenti</emphasis>
|
||
</taxonomicName>
|
||
: it differs remarkably from
|
||
<taxonomicName id="748B4D46FFF4FFFEFE16FCA0253AFCE3" authorityName="Ehrenberg" authorityYear="1828" box="[469,580,791,815]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE16FCA0253AFCE3" box="[469,580,791,815]" italics="true" pageId="14" pageNumber="13">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
by having tergite I with three carinae, metasoma smooth and coarsely punctate at least in males (
|
||
<figureCitation id="2BB02A40FFF4FFFEFE71FCE3257AFCA0" box="[434,516,852,876]" captionStart="Figures 32–35" captionStartId="12.[192,278,1114,1138]" captionTargetBox="[202,1426,193,1053]" captionTargetId="figure-481@12.[192,1440,192,1080]" captionTargetPageId="12" captionText="Figures 32–35: Figures 32–34. Androctonus turieli sp. n., metasoma and telson dorsal (32), ventral (33) and lateral (34) views, ♀ holotype. Figure 35. Androctonus sergenti Vachon, 1948, metasoma and telson ventral, ♂, Morocco, Anti Atlas Mts., 62 km southeast of Agadir, 30°03.35'N 09°04.4'W, 798 m a. s. l. (FKCP)." pageId="14" pageNumber="13">Fig. 35</figureCitation>
|
||
), pedipalps less attenuate, and much more reduced granulation and carination on the body and appendages. Furthermore,
|
||
<taxonomicName id="748B4D46FFF4FFFEFD49FC25247DFC66" authorityName="Vachon" authorityYear="1948" box="[650,771,914,938]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="sergenti">
|
||
<emphasis id="81FFEAD7FFF4FFFEFD49FC25247DFC66" box="[650,771,914,938]" italics="true" pageId="14" pageNumber="13">A. sergenti</emphasis>
|
||
</taxonomicName>
|
||
is locally endemic in a small area of the high Anti-Atlas Mountains of south-central
|
||
<collectingCountry id="CB9C7655FFF4FFFEFE24FC672536FC2B" box="[487,584,976,999]" name="Morocco" pageId="14" pageNumber="13">Morocco</collectingCountry>
|
||
(Vachon, 1952).
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF4FFFEFF33FC592662FB53" blockId="14.[192,802,761,1827]" pageId="14" pageNumber="13">
|
||
In fact,
|
||
<taxonomicName id="748B4D46FFF4FFFEFE85FC5926D8FBCA" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[326,422,1006,1030]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE85FC5926D8FBCA" box="[326,422,1006,1030]" italics="true" pageId="14" pageNumber="13">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF4FFFEFE73FC59269BFBCA" bold="true" box="[432,485,1006,1030]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFE73FC59269BFBCA" box="[432,485,1006,1030]" pageId="14" pageNumber="13" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="748B4D46FFF4FFFEFD76FC59245CFBCA" authorityName="Ehrenberg" authorityYear="1828" box="[693,802,1006,1030]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFD76FC59245CFBCA" box="[693,802,1006,1030]" italics="true" pageId="14" pageNumber="13">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="748B4D46FFF4FFFEFF33FBBB26F3FBE8" box="[240,397,1036,1060]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="tenuissimus">
|
||
<emphasis id="81FFEAD7FFF4FFFEFF33FBBB26F3FBE8" box="[240,397,1036,1060]" italics="true" pageId="14" pageNumber="13">A. tenuissimus</emphasis>
|
||
</taxonomicName>
|
||
: apart from exhibiting its own diagnostic attributes, it shares some diagnostic characters with each of these two species and it is also essentially intermediate between them in most morphometric ratios (
|
||
<tableCitation id="FE09037EFFF4FFFEFF0BFB302671FB53" box="[200,271,1159,1183]" captionStart="Table 3" captionStartId="14.[192,257,645,669]" captionText="Table 3: Comparison between Androctonus bicolor and its closest relatives Androctonus tenuissimus and Androctonus turieli sp. n., based upon selected morphometric ratios of adult females (adult males of the latter are still known), and including typespecimens. Abbreviations: length (L), width (W)." pageId="14" pageNumber="13">Tab. 3</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF4FFFEFF33FB112524F9E2" blockId="14.[192,802,761,1827]" pageId="14" pageNumber="13">
|
||
The best characters that unambiguously separate
|
||
<taxonomicName id="748B4D46FFF4FFFEFCC9FB112673FB10" authorityName="Ehrenberg" authorityYear="1828" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFCC9FB112673FB10" italics="true" pageId="14" pageNumber="13">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="748B4D46FFF4FFFEFE96FB7326C9FB10" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[341,439,1220,1244]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE96FB7326C9FB10" box="[341,439,1220,1244]" italics="true" pageId="14" pageNumber="13">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF4FFFEFE01FB722689FB11" bold="true" box="[450,503,1221,1245]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFE01FB722689FB11" box="[450,503,1221,1245]" pageId="14" pageNumber="13" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are:
|
||
<emphasis id="81FFEAD7FFF4FFFEFDF7FB722534FB11" bold="true" box="[564,586,1221,1245]" pageId="14" pageNumber="13">1)</emphasis>
|
||
pedipalp fixed and movable fingers with 12–13 and 13–14 principal rows of denticles, respectively;
|
||
<emphasis id="81FFEAD7FFF4FFFEFE7DFAB526AAFAD6" bold="true" box="[446,468,1282,1306]" pageId="14" pageNumber="13">2)</emphasis>
|
||
tergite I with three carinae;
|
||
<emphasis id="81FFEAD7FFF4FFFEFCC8FAB5245CFAD6" bold="true" box="[779,802,1282,1306]" pageId="14" pageNumber="13">3)</emphasis>
|
||
metasoma deeper and more robust, with carinae coarser and intercarinal tegument much less granulose and glossy;
|
||
<emphasis id="81FFEAD7FFF4FFFEFED9FAE9264EFABA" bold="true" box="[282,304,1374,1398]" pageId="14" pageNumber="13">4)</emphasis>
|
||
ventrolateral carinae on female metasomal segment V with the flared denticles much more irregular, larger, and lobate in shape;
|
||
<emphasis id="81FFEAD7FFF4FFFEFDE6FA2B2545FA78" bold="true" box="[549,571,1436,1460]" pageId="14" pageNumber="13">5)</emphasis>
|
||
metasomal segments II–III with lateral inframedian carinae complete and very strong, coarsely granulose. Further, there is a slight but consistent trend in
|
||
<taxonomicName id="748B4D46FFF4FFFEFE51FA402681F9C3" authorityName="Ehrenberg" authorityYear="1828" box="[402,511,1527,1551]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE51FA402681F9C3" box="[402,511,1527,1551]" italics="true" pageId="14" pageNumber="13">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
to be smaller, darker, and to possess lower pectinal tooth counts.
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF4FFFEFF33F98226DAF8EF" blockId="14.[192,802,761,1827]" pageId="14" pageNumber="13">
|
||
On the other hand,
|
||
<taxonomicName id="748B4D46FFF4FFFEFE17F982250AF981" box="[468,628,1589,1613]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="tenuissimus">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE17F982250AF981" box="[468,628,1589,1613]" italics="true" pageId="14" pageNumber="13">A. tenuissimus</emphasis>
|
||
</taxonomicName>
|
||
can be readily distinguished from
|
||
<taxonomicName id="748B4D46FFF4FFFEFE50F9E4268AF9A7" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[403,500,1619,1643]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFE50F9E4268AF9A7" box="[403,500,1619,1643]" italics="true" pageId="14" pageNumber="13">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF4FFFEFE3FF9E3254FF9A0" bold="true" box="[508,561,1620,1644]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFE3FF9E3254FF9A0" box="[508,561,1620,1644]" pageId="14" pageNumber="13" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by:
|
||
<emphasis id="81FFEAD7FFF4FFFEFDA6F9E32505F9A0" bold="true" box="[613,635,1620,1644]" pageId="14" pageNumber="13">1)</emphasis>
|
||
pedipalps conspicuously more attenuate, with femur, patella, and chela much longer:
|
||
<emphasis id="81FFEAD7FFF4FFFEFE9BF9262611F965" bold="true" box="[344,367,1681,1705]" pageId="14" pageNumber="13">2)</emphasis>
|
||
pedipalps with carinae stronger and intercarinal tegument of chela more densely granulose;
|
||
<emphasis id="81FFEAD7FFF4FFFEFCC8F907245CF904" bold="true" box="[779,802,1712,1736]" pageId="14" pageNumber="13">3)</emphasis>
|
||
tergite I with three carinae;
|
||
<emphasis id="81FFEAD7FFF4FFFEFE28F979257CF92A" bold="true" box="[491,514,1742,1766]" pageId="14" pageNumber="13">4)</emphasis>
|
||
metasoma less deeper and more slender (all segments longer than wide in females), with all carinae finer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB91654EFFF4FFFEFF03F8F123B9FC82" pageId="14" pageNumber="13" type="distribution">
|
||
<paragraph id="B33436C5FFF4FFFEFF03F8F1245FF8B0" blockId="14.[192,801,1862,1917]" pageId="14" pageNumber="13">
|
||
DISTRIBUTION (
|
||
<figureCitation id="2BB02A40FFF4FFFEFEB0F8F126B1F892" box="[371,463,1862,1886]" captionStart="Figure 36" captionStartId="13.[192,267,846,870]" captionTargetBox="[203,1426,167,811]" captionTargetId="figure-551@13.[204,1425,168,810]" captionTargetPageId="13" captionText="Figure 36: Type locality of Androctonus turieli sp.n." pageId="14" pageNumber="13">Fig. 36</figureCitation>
|
||
). This scorpion is currently known only from the
|
||
<typeStatus id="6C308867FFF4FFFEFE64F8D226ABF8B1" box="[423,469,1893,1917]" pageId="14" pageNumber="13">type</typeStatus>
|
||
locality, in the eastern coast of
|
||
</paragraph>
|
||
<paragraph id="B33436C5FFF4FFFEFCFCFD4E23B9FC82" blockId="14.[831,1441,761,846]" pageId="14" pageNumber="13">
|
||
<collectingCountry id="CB9C7655FFF4FFFEFCFCFD4E24ECFCDD" box="[831,914,761,785]" name="Tunisia" pageId="14" pageNumber="13">Tunisia</collectingCountry>
|
||
. Most of the records attributed by Vachon (1952) to
|
||
<taxonomicName id="748B4D46FFF4FFFEFC76FCA02304FCE3" authority="aeneas" box="[949,1146,791,815]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="aeneas" subSpecies="aeneas">
|
||
<emphasis id="81FFEAD7FFF4FFFEFC76FCA02304FCE3" box="[949,1146,791,815]" italics="true" pageId="14" pageNumber="13">A. aeneas aeneas</emphasis>
|
||
</taxonomicName>
|
||
most likely belong to
|
||
<taxonomicName id="748B4D46FFF4FFFEFA4AFCA024FEFC82" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFA4AFCA024FEFC82" italics="true" pageId="14" pageNumber="13">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF4FFFEFC45FC8124BCFC82" bold="true" box="[902,962,822,846]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFC45FC8124BCFC82" box="[902,962,822,846]" pageId="14" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but need to be verified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB91654EFFF4FFFEFCFCFCC4230BFC2B" pageId="14" pageNumber="13" type="discussion">
|
||
<paragraph id="B33436C5FFF4FFFEFCFCFCC4230BFC2B" blockId="14.[831,1440,883,999]" pageId="14" pageNumber="13">
|
||
COMMENTS. The taxonomic interpretation of
|
||
<taxonomicName id="748B4D46FFF4FFFEFAF6FCC424F7FC66" authority="aeneas" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="aeneas" subSpecies="aeneas">
|
||
<emphasis id="81FFEAD7FFF4FFFEFAF6FCC424F7FC66" italics="true" pageId="14" pageNumber="13">A. aeneas aeneas</emphasis>
|
||
</taxonomicName>
|
||
made by Vachon (1952: 126-128) partially matches
|
||
<taxonomicName id="748B4D46FFF4FFFEFCB5FC0624A9FC05" authority="Teruel & Kovařík, 2014" authorityName="Teruel & Kovařík" authorityYear="2014" box="[886,983,945,969]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
|
||
<emphasis id="81FFEAD7FFF4FFFEFCB5FC0624A9FC05" box="[886,983,945,969]" italics="true" pageId="14" pageNumber="13">A. turieli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81FFEAD7FFF4FFFEFC23FC062363FC05" bold="true" box="[992,1053,945,969]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="9ACC57ACFFF4FFFEFC23FC062363FC05" box="[992,1053,945,969]" pageId="14" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but the former is a
|
||
<emphasis id="81FFEAD7FFF4FFFEFAC2FC0522DEFC05" box="[1281,1440,945,969]" italics="true" pageId="14" pageNumber="13">nomen dubium</emphasis>
|
||
(see above under
|
||
<taxonomicName id="748B4D46FFF4FFFEFC39FC78231AFC2B" authorityName="Ehrenberg" authorityYear="1828" box="[1018,1124,975,999]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="81FFEAD7FFF4FFFEFC39FC78231AFC2B" box="[1018,1124,975,999]" italics="true" pageId="14" pageNumber="13">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |