254 lines
36 KiB
XML
254 lines
36 KiB
XML
<document id="A7467E76889F1B1D0DADE19F990D37BB" ID-CLB-Dataset="27512" ID-DOI="10.11646/zootaxa.4576.2.12" ID-GBIF-Dataset="53b96271-789f-4155-aa9f-995572842e98" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624852" ID-ZooBank="64A844F9-210B-4758-8A19-4ACE45B7C29B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554273205939" checkinUser="plazi" docAuthor="Jaschhof, Mathias & Ševčík, Jan" docDate="2019" docId="03DC87D0FF9EFFFC06833DD0306B5755" docLanguage="en" docName="zootaxa.4576.2.12.pdf" docOrigin="Zootaxa 4576 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cabamofa orientalis Jaschhof & Ševčík 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="394" masterDocId="FFE5FFA8FF9EFFFE0614381F33315522" masterDocTitle="Cabamofa orientalis sp. nov. from Thailand, congeneric with C. mira Jaschhof from Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis)" masterLastPageNumber="394" masterPageNumber="392" pageNumber="392" updateTime="1698716636453" updateUser="ExternalLinkService">
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<mods:title id="C1FFE15F7E34F867A72EA058E3CB3E98">Cabamofa orientalis sp. nov. from Thailand, congeneric with C. mira Jaschhof from Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis)</mods:title>
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<mods:namePart id="6A8503F7929135644E99EBE3DD54BFF7">Jaschhof, Mathias</mods:namePart>
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<mods:namePart id="56F656791AD7BA0AAD48501B8E5194BC">Ševčík, Jan</mods:namePart>
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<mods:typeOfResource id="A25B0A948FA0DBEF5F722CF5E5BF634A">text</mods:typeOfResource>
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<mods:title id="356DE691F7E68D7D2FAEFA532AEC5B33">Zootaxa</mods:title>
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<mods:part id="18D686E3F483ABEF75D59EF1702FAC78">
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<mods:date id="93B8C0FB1B2A216C163541094DFC2B24">2019</mods:date>
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<mods:number id="DE5938020CC073C1F465C817093CC6BB">2019-04-02</mods:number>
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<mods:number id="F37014640B6B721235CEC5AF8B263407">4576</mods:number>
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<treatment id="03DC87D0FF9EFFFC06833DD0306B5755" ID-DOI="http://doi.org/10.5281/zenodo.5932649" ID-GBIF-Taxon="154533432" ID-Zenodo-Dep="5932649" LSID="urn:lsid:plazi:treatment:03DC87D0FF9EFFFC06833DD0306B5755" httpUri="http://treatment.plazi.org/id/03DC87D0FF9EFFFC06833DD0306B5755" lastPageId="2" lastPageNumber="394" pageId="0" pageNumber="392">
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<subSubSection id="C36F654DFF9EFFFE06833DD0314550CB" box="[151,628,1487,1513]" pageId="0" pageNumber="392" type="nomenclature">
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<paragraph id="8BCA36C6FF9EFFFE06833DD0314550CB" blockId="0.[151,1436,1487,2035]" box="[151,628,1487,1513]" pageId="0" pageNumber="392">
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<heading id="D08281AAFF9EFFFE06833DD0314550CB" bold="true" box="[151,628,1487,1513]" fontSize="11" level="1" pageId="0" pageNumber="392" reason="1">
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<emphasis id="B901EAD4FF9EFFFE06833DD0314550CB" bold="true" box="[151,628,1487,1513]" pageId="0" pageNumber="392">
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<taxonomicName id="4C754D45FF9EFFFE06833DD032BC50CB" authority="Jaschhof & Ševčík, 2019" authorityName="Jaschhof & Ševčík" authorityYear="2019" box="[151,397,1487,1513]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
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<emphasis id="B901EAD4FF9EFFFE06833DD032BC50CB" bold="true" box="[151,397,1487,1513]" italics="true" pageId="0" pageNumber="392">Cabamofa orientalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257AFFF9EFFFE07803DD032C050CB" box="[404,497,1487,1513]" pageId="0" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="134E2A43FF9EFFFE04153DD0315A50CB" box="[513,619,1487,1513]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Figs 1–6</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F654DFF9EFFFC06833DED31B0558D" lastPageId="2" lastPageNumber="394" pageId="0" pageNumber="392" type="diagnosis">
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<paragraph id="8BCA36C6FF9EFFFE06833DED378C5223" blockId="0.[151,1436,1487,2035]" pageId="0" pageNumber="392">
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<emphasis id="B901EAD4FF9EFFFE06833DED323D532B" bold="true" box="[151,268,1522,1545]" pageId="0" pageNumber="392">Diagnosis.</emphasis>
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The new species fits largely the description of
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<taxonomicName id="4C754D45FF9EFFFE05313DEE30A5532B" authorityName="Jaschhof" authorityYear="2005" box="[805,916,1521,1545]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="genus">
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<emphasis id="B901EAD4FF9EFFFE05313DEE30A5532B" box="[805,916,1521,1545]" italics="true" pageId="0" pageNumber="392">Cabamofa</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EFE44B37FF9EFFFE05D03DED3744532B" author="Jaschhof, M." box="[964,1141,1522,1545]" pageId="0" pageNumber="392" pagination="323 - 332" refId="ref1698" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof (2005)</bibRefCitation>
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, with the exception of the following peculiarities in the wing (
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<figureCitation id="134E2A43FF9EFFFE04053E0B31405309" box="[529,625,1556,1579]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Figs 2, 4</figureCitation>
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): vein Rs is situated basal (not distal) of M
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<subScript id="17F13483FF9EFFFE02233E3B370F5312" attach="left" box="[1079,1086,1572,1584]" fontSize="5" pageId="0" pageNumber="392">4</subScript>
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; R
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<subScript id="17F13483FF9EFFFE024A3E3B37545312" attach="left" box="[1118,1125,1572,1584]" fontSize="5" pageId="0" pageNumber="392">1</subScript>
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joins the costa at (not clearly beyond) the midpoint of wing; and the stem of M
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<subScript id="17F13483FF9EFFFE04AE3E5631E15377" attach="left" box="[698,720,1609,1621]" fontSize="5" pageId="0" pageNumber="392">1+2</subScript>
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is completely absent (not even present as short vestige apically). These characters alone differentiate
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<taxonomicName id="4C754D45FF9EFFFE040A3E4031955354" authorityName="Jaschhof & Ševčík" authorityYear="2019" box="[542,676,1630,1654]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="B901EAD4FF9EFFFE040A3E4031955354" box="[542,676,1630,1654]" italics="true" pageId="0" pageNumber="392">C. orientalis</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C754D45FF9EFFFE04FC3E40300D5354" authorityName="Jaschhof" authorityYear="2005" box="[744,828,1631,1654]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="mira">
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<emphasis id="B901EAD4FF9EFFFE04FC3E40300D5354" box="[744,828,1631,1654]" italics="true" pageId="0" pageNumber="392">C. mira</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFE44B37FF9EFFFE055E3E40375D5354" author="Jaschhof, M." box="[842,1132,1631,1654]" pageId="0" pageNumber="392" pagination="323 - 332" refId="ref1698" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof 2005: figs 24–25</bibRefCitation>
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). Other distinctions concern male genitalic structures, as follows (see
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<bibRefCitation id="EFE44B37FF9EFFFE044C3E9D305153BB" author="Amorim, D. S. & Rindal, E." box="[600,864,1666,1689]" pageId="0" pageNumber="392" pagination="1 - 92" refId="ref1561" refString="Amorim, D. S. & Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92. https: // doi. org / 10.11646 / zootaxa. 1535.1.1" type="journal article" year="2007">Amorim & Rindal 2007</bibRefCitation>
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: figs 44–47 for genitalic characters of
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<taxonomicName id="4C754D45FF9EFFFE03133E9E366A53BB" authorityName="Jaschhof" authorityYear="2005" box="[1287,1371,1665,1689]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="mira">
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<emphasis id="B901EAD4FF9EFFFE03133E9E366A53BB" box="[1287,1371,1665,1689]" italics="true" pageId="0" pageNumber="392">C. mira</emphasis>
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</taxonomicName>
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). The ninth tergite has a convex, broadly rounded (rather than concave, V-shaped) posterior edge and no posterolateral lobes (
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<figureCitation id="134E2A43FF9EFFFE068B3ED833EE53FC" box="[159,223,1735,1758]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Fig. 6</figureCitation>
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); the gonostylus is not trilobate (as it is in
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<taxonomicName id="4C754D45FF9EFFFE04B73ED831C553FC" authorityName="Jaschhof" authorityYear="2005" box="[675,756,1735,1758]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="mira">
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<emphasis id="B901EAD4FF9EFFFE04B73ED831C553FC" box="[675,756,1735,1758]" italics="true" pageId="0" pageNumber="392">C. mira</emphasis>
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</taxonomicName>
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) and bears a small, blunt-ended tooth medially (which is absent in
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<taxonomicName id="4C754D45FF9EFFFE06A63EF632325223" authorityName="Jaschhof" authorityYear="2005" box="[178,259,1769,1793]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="mira">
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<emphasis id="B901EAD4FF9EFFFE06A63EF632325223" box="[178,259,1769,1793]" italics="true" pageId="0" pageNumber="392">C. mira</emphasis>
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</taxonomicName>
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) (
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<figureCitation id="134E2A43FF9EFFFE070F3EF5326D5223" box="[283,348,1770,1793]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Fig. 5</figureCitation>
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); and the ejaculatory apodeme is not furcate apically (as it is in
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<taxonomicName id="4C754D45FF9EFFFE02113EF637665223" authorityName="Jaschhof" authorityYear="2005" box="[1029,1111,1769,1793]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="mira">
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<emphasis id="B901EAD4FF9EFFFE02113EF637665223" box="[1029,1111,1769,1793]" italics="true" pageId="0" pageNumber="392">C. mira</emphasis>
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</taxonomicName>
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) (
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<figureCitation id="134E2A43FF9EFFFE027A3EF5379E5223" box="[1134,1199,1770,1793]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Fig. 5</figureCitation>
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).
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</paragraph>
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<paragraph id="8BCA36C6FF9EFFFF06D33F1333DC54C5" blockId="0.[151,1436,1487,2035]" lastBlockId="1.[151,1436,152,487]" lastPageId="1" lastPageNumber="393" pageId="0" pageNumber="392">
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<emphasis id="B901EAD4FF9EFFFE06D33F1332E05201" bold="true" box="[199,465,1804,1827]" pageId="0" pageNumber="392">Other male characters.</emphasis>
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Body length
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<quantity id="4C8D9B23FF9EFFFE04723F13318C5201" box="[614,701,1804,1827]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="0" pageNumber="392" unit="mm" value="2.8">2.8 mm</quantity>
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.
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<emphasis id="B901EAD4FF9EFFFE04DC3F13303D5201" bold="true" box="[712,780,1804,1827]" pageId="0" pageNumber="392">Head.</emphasis>
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Clypeus non-setose. Scape slightly larger than pedicel, both lighter than flagellum. Fourth flagellomere cylindrical, with short neck, node 2.5 longer than wide, with irregular cover of fine setae whose lengths equal the flagellomeral width, interspersed with a few larger setae arising from distinct sockets (
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<figureCitation id="134E2A43FF9EFFFE06E23F6B320752A9" box="[246,310,1908,1931]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Fig. 1</figureCitation>
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). Compound eyes touching at vertex, eye bridge about 7 ommatidia long dorsally. Palpus with 5 setaebearing segments, third segment conspicuously swollen, with sensory pit, fifth segment conspicuously long and thin (
|
||
<figureCitation id="134E2A43FF9EFFFE068B3FA533EE52F3" box="[159,223,1978,2001]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="0" pageNumber="392">Fig. 3</figureCitation>
|
||
).
|
||
<emphasis id="B901EAD4FF9EFFFE06E03FA5320152F3" bold="true" box="[244,304,1978,2001]" pageId="0" pageNumber="392">Legs.</emphasis>
|
||
Lengths of coxae relative to thoracal height: forecoxa, 0.8, midcoxa, 0.6, hindcoxa, 0.5. Edge of foretibial anteroapical depression with comb of about 15 setae. Mid- and hindlegs broken behind coxae. Claws small, strong, crescent-shaped, toothless. Empodia small, barely claw-long.
|
||
<emphasis id="B901EAD4FF9FFFFF055D38873088558D" bold="true" box="[841,953,152,175]" pageId="1" pageNumber="393">Genitalia.</emphasis>
|
||
Gonocoxae largely merged ventromedially except for a wide, shallow emargination medioposteriorly; lateral setae large, medial setae mostly smaller; a triangular portion below the emargination non-setose; a pair of small, setose protuberances inside (not illustrated); dorsal apodemes short (
|
||
<figureCitation id="134E2A43FF9FFFFF06C8391F322F5435" box="[220,286,256,279]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393">Fig. 5</figureCitation>
|
||
). Gonostylus with narrow, stem-like base; apical portion somewhat angular, ventrodorsally flattened, with long setae laterally; a bulbous, rounded lobe medially covered with numerous short, thick setae (
|
||
<figureCitation id="134E2A43FF9FFFFF02C4393D3625541B" box="[1232,1300,290,313]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393">Fig. 5</figureCitation>
|
||
). Parameres merged to form a subrectangular, weakly sclerotized plate (tegmen); parameral apodemes slender (
|
||
<figureCitation id="134E2A43FF9FFFFF0334395A365B547E" box="[1312,1386,325,348]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393">Fig. 5</figureCitation>
|
||
). A subtriangular area of fine aedeagal teeth near the tegminal edge (
|
||
<figureCitation id="134E2A43FF9FFFFF056C3977308C545D" box="[888,957,360,383]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393">Fig. 5</figureCitation>
|
||
). Ejaculatory apodeme as long as tegmen, sclerotized, slightly broadened apically, apex faintly contoured (
|
||
<figureCitation id="134E2A43FF9FFFFF054A399530925483" box="[862,931,394,417]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393">Fig. 5</figureCitation>
|
||
). Cerci small, setose, closely associated with hypoproct (the tenth tergite of
|
||
<bibRefCitation id="EFE44B37FF9FFFFF07F439B231D754E6" author="Amorim, D. S. & Rindal, E." box="[480,742,429,452]" pageId="1" pageNumber="393" pagination="1 - 92" refId="ref1561" refString="Amorim, D. S. & Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92. https: // doi. org / 10.11646 / zootaxa. 1535.1.1" type="journal article" year="2007">Amorim & Rindal 2007</bibRefCitation>
|
||
: fig. 47), the latter large, slightly bilobed, smooth, microtrichose (
|
||
<figureCitation id="134E2A43FF9FFFFF068B39CF33D054C5" box="[159,225,464,487]" captionStart="FIGURES 1–6" captionStartId="1.[151,258,1941,1963]" captionTargetBox="[225,1365,582,1882]" captionTargetId="figure@1.[187,1403,541,1924]" captionTargetPageId="1" captionText="FIGURES 1–6. Cabamofa orientalis sp. nov., male holotype. 1: Fourth flagellomere, lateral. 2: Wing, dorsal. 3: Palpus, ventrolateral. 4: Base of wing, dorsal. 5: Genitalia, ventral, dorsal parts omitted. 6: Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm." httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0A664EFF9FFFFF06833F8A306052CE" httpUri="https://zenodo.org/record/2624854/files/figure.png" pageId="1" pageNumber="393" startId="1.[151,258,1941,1963]" targetBox="[225,1365,582,1882]" targetPageId="1">
|
||
<paragraph id="8BCA36C6FF9FFFFF06833F8A306052CE" blockId="1.[151,1437,1940,2028]" pageId="1" pageNumber="393">
|
||
<emphasis id="B901EAD4FF9FFFFF06833F8A32005289" bold="true" box="[151,305,1941,1963]" pageId="1" pageNumber="393">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C754D45FF9FFFFF07223F8B32CD5288" authority="Jaschhof & Ševčík, 2019" authorityName="Jaschhof & Ševčík" authorityYear="2019" box="[310,508,1940,1962]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B901EAD4FF9FFFFF07223F8B32CD5288" box="[310,508,1940,1962]" italics="true" pageId="1" pageNumber="393">Cabamofa orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901EAD4FF9FFFFF04153F8A317C5289" bold="true" box="[513,589,1941,1963]" pageId="1" pageNumber="393">
|
||
<taxonomicNameLabel id="A23257AFFF9FFFFF04153F8A317C5289" box="[513,589,1941,1963]" pageId="1" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype.
|
||
<emphasis id="B901EAD4FF9FFFFF04F93F8A30305289" bold="true" box="[749,769,1941,1963]" pageId="1" pageNumber="393">1:</emphasis>
|
||
Fourth flagellomere, lateral.
|
||
<emphasis id="B901EAD4FF9FFFFF020C3F8A371D5289" bold="true" box="[1048,1068,1941,1963]" pageId="1" pageNumber="393">2:</emphasis>
|
||
Wing, dorsal.
|
||
<emphasis id="B901EAD4FF9FFFFF02AC3F8A37FA5289" bold="true" box="[1208,1227,1941,1963]" pageId="1" pageNumber="393">3:</emphasis>
|
||
Palpus, ventrolateral.
|
||
<emphasis id="B901EAD4FF9FFFFF06833FAA339B52E9" bold="true" box="[151,170,1973,1995]" pageId="1" pageNumber="393">4:</emphasis>
|
||
Base of wing, dorsal.
|
||
<emphasis id="B901EAD4FF9FFFFF076B3FAA32A552E9" bold="true" box="[383,404,1973,1995]" pageId="1" pageNumber="393">5:</emphasis>
|
||
Genitalia, ventral, dorsal parts omitted.
|
||
<emphasis id="B901EAD4FF9FFFFF05073FAA301652E9" bold="true" box="[787,807,1973,1995]" pageId="1" pageNumber="393">6:</emphasis>
|
||
Ninth tergite with hypoproct and cerci indicated, dorsal. Scale for 1 and 3, 0.05 mm, for 2, 0.50 mm, for 4, 0.20 mm, for 5 and 6, 0.10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA36C6FF9CFFFC06D3388731B0558D" blockId="2.[151,1436,152,631]" box="[199,641,152,175]" pageId="2" pageNumber="394">Female and preimaginal stages unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F654DFF9CFFFC06D338A530F355F3" box="[199,962,186,209]" pageId="2" pageNumber="394" type="etymology">
|
||
<paragraph id="8BCA36C6FF9CFFFC06D338A530F355F3" blockId="2.[151,1436,152,631]" box="[199,962,186,209]" pageId="2" pageNumber="394">
|
||
<emphasis id="B901EAD4FF9CFFFC06D338A5327655F3" bold="true" box="[199,327,186,209]" pageId="2" pageNumber="394">Etymology.</emphasis>
|
||
The name refers to the Oriental distribution of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F654DFF9CFFFC06D338C23629541B" pageId="2" pageNumber="394" type="materials_examined">
|
||
<paragraph id="8BCA36C6FF9CFFFC06D338C23629541B" blockId="2.[151,1436,152,631]" pageId="2" pageNumber="394">
|
||
<emphasis id="B901EAD4FF9CFFFC06D338C2324455D6" bold="true" box="[199,373,221,244]" pageId="2" pageNumber="394">Type material.</emphasis>
|
||
<materialsCitation id="3B1D3C9BFF9CFFFC079138C23629541B" ID-GBIF-Occurrence="2234219366" collectingDate="2008-07-11" collectingDateMax="2008-07-18" collectingDateMin="2008-07-11" collectionCode="TIGER" collectorName="Somboon & Chatchawan & Queen Sirikit Botanic Garden" country="Canada" elevation="210" latitude="14.6356" location="Khuean Srinagarindra National Park" longLatPrecision="1" longitude="98.997284" municipality="Male" pageId="2" pageNumber="394" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="holotype">
|
||
<typeStatus id="54CE8864FF9CFFFC079138C232D955D6" box="[389,488,221,244]" pageId="2" pageNumber="394" type="holotype">Holotype</typeStatus>
|
||
dissected and mounted on microscope slide in
|
||
<collectingCountry id="F3627656FF9CFFFC023338C2374655D6" box="[1063,1143,221,244]" name="Canada" pageId="2" pageNumber="394">Canada</collectingCountry>
|
||
balsam.
|
||
<collectingMunicipality id="6BAEACBCFF9CFFFC02FD38C2361355D6" box="[1257,1314,221,244]" pageId="2" pageNumber="394">Male</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F3627656FF9CFFFC032238C236A655D6" box="[1334,1431,221,244]" name="Thailand" pageId="2" pageNumber="394">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="49B1F824FF9CFFFC0683391F321E5435" box="[151,303,256,279]" country="Thailand" name="Kanchanaburi" pageId="2" pageNumber="394">Kanchanaburi</collectingRegion>
|
||
,
|
||
<location id="8EAA601DFF9CFFFC072E391F31F45435" LSID="urn:lsid:plazi:treatment:03DC87D0FF9EFFFC06833DD0306B5755:8EAA601DFF9CFFFC072E391F31F45435" box="[314,709,256,279]" country="Canada" latitude="14.6356" longLatPrecision="1" longitude="98.997284" municipality="Male" name="Khuean Srinagarindra National Park" pageId="2" pageNumber="394" stateProvince="Kanchanaburi">Khuean Srinagarindra National Park</location>
|
||
, tourist center,
|
||
<geoCoordinate id="EE415001FF9CFFFC057E391F30C45435" box="[874,1013,256,279]" direction="north" orientation="latitude" pageId="2" pageNumber="394" precision="1" value="14.6356">14°38.136´N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE415001FF9CFFFC0218391F37A75435" box="[1036,1174,256,279]" direction="east" orientation="longitude" pageId="2" pageNumber="394" precision="1" value="98.997284">98°59.837´E</geoCoordinate>
|
||
,
|
||
<quantity id="4C8D9B23FF9CFFFC02B6391F37D95435" box="[1186,1256,256,279]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="394" unit="m" value="210.0">
|
||
<elevation id="0058D1F5FF9CFFFC02B6391F37D95435" box="[1186,1256,256,279]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="394" unit="m" value="210.0">210 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFCB1006FF9CFFFC02E0391F36A65435" box="[1268,1431,256,279]" pageId="2" pageNumber="394" value="2008-07-11" valueMax="2008-07-18" valueMin="2008-07-11">
|
||
<collectingDate id="EF8FE9EEFF9CFFFC02E0391F36A65435" box="[1268,1431,256,279]" pageId="2" pageNumber="394" value="2008-07-11" valueMax="2008-07-18" valueMin="2008-07-11">11–18.vii.2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26805310FF9CFFFC0683393D33CC541B" box="[151,253,290,313]" pageId="2" pageNumber="394">Somboon</collectorName>
|
||
&
|
||
<collectorName id="26805310FF9CFFFC070B393D3292541B" box="[287,419,290,313]" pageId="2" pageNumber="394">Chatchawan</collectorName>
|
||
(
|
||
<collectionCode id="ED64AE03FF9CFFFC07A5393D3130541B" box="[433,513,290,313]" pageId="2" pageNumber="394">TIGER</collectionCode>
|
||
sample T3432) (in
|
||
<collectorName id="26805310FF9CFFFC04C7393D3722541B" box="[723,1043,290,313]" pageId="2" pageNumber="394">Queen Sirikit Botanic Garden</collectorName>
|
||
,
|
||
<collectingRegion id="49B1F824FF9CFFFC020A393D37AE541B" box="[1054,1183,290,313]" country="Thailand" name="Chiang Mai" pageId="2" pageNumber="394">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3627656FF9CFFFC02BE393D363C541B" box="[1194,1293,290,313]" name="Thailand" pageId="2" pageNumber="394">Thailand</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F654DFF9CFFFC06D3395A306B5755" pageId="2" pageNumber="394" type="discussion">
|
||
<paragraph id="8BCA36C6FF9CFFFC06D3395A306B5755" blockId="2.[151,1436,152,631]" pageId="2" pageNumber="394">
|
||
<emphasis id="B901EAD4FF9CFFFC06D3395A3277547E" bold="true" box="[199,326,325,348]" pageId="2" pageNumber="394">Discussion.</emphasis>
|
||
<bibRefCitation id="EFE44B37FF9CFFFC0759395A32CB547E" author="Jaschhof, M." box="[333,506,325,348]" pageId="2" pageNumber="394" pagination="323 - 332" refId="ref1698" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof (2005)</bibRefCitation>
|
||
postulated a sistergroup relationship between
|
||
<taxonomicName id="4C754D45FF9CFFFC05E4395B376E547E" authorityName="Jaschhof" authorityYear="2005" box="[1008,1119,324,348]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAD4FF9CFFFC05E4395B376E547E" box="[1008,1119,324,348]" italics="true" pageId="2" pageNumber="394">Cabamofa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C754D45FF9CFFFC0281395B3622547E" authorityName="Jaschhof" authorityYear="2005" box="[1173,1299,324,348]" class="Insecta" family="Rangomaramidae" genus="Rogambara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAD4FF9CFFFC0281395B3622547E" box="[1173,1299,324,348]" italics="true" pageId="2" pageNumber="394">Rogambara</emphasis>
|
||
</taxonomicName>
|
||
on the basis of two synapomorphies: the presence of holoptic eyes and the absence (loss) of ocelli. As recognized here, other synapomorphous characters supporting this clade are the foreshortening of the medial fork (i.e., the fork made of M
|
||
<subScript id="17F13483FF9CFFFC03733985365F5484" attach="left" box="[1383,1390,410,422]" fontSize="5" pageId="2" pageNumber="394">1</subScript>
|
||
and M
|
||
<subScript id="17F13483FF9CFFFC06BA39A0338454E9" attach="left" box="[174,181,447,459]" fontSize="5" pageId="2" pageNumber="394">2</subScript>
|
||
is shorter than the stem M
|
||
<subScript id="17F13483FF9CFFFC07C639A032D854E9" attach="left" box="[466,489,447,459]" fontSize="5" pageId="2" pageNumber="394">1+2</subScript>
|
||
) and the merging of the parameres into a plate (tegmen). In distinction from
|
||
<taxonomicName id="4C754D45FF9CFFFC030A39B036AD54E5" authorityName="Jaschhof" authorityYear="2005" box="[1310,1436,431,455]" class="Insecta" family="Rangomaramidae" genus="Rogambara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAD4FF9CFFFC030A39B036AD54E5" box="[1310,1436,431,455]" italics="true" pageId="2" pageNumber="394">Rogambara</emphasis>
|
||
</taxonomicName>
|
||
(character states in parentheses),
|
||
<taxonomicName id="4C754D45FF9CFFFC07E239CB315454CE" authorityName="Jaschhof" authorityYear="2005" box="[502,613,468,492]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAD4FF9CFFFC07E239CB315454CE" box="[502,613,468,492]" italics="true" pageId="2" pageNumber="394">Cabamofa</emphasis>
|
||
</taxonomicName>
|
||
have five setae-bearing palpal segments (three segments, the first and second reduced); the anepisternum is of normal size (enlarged); the pattern of tibial spurs is 1:2:2 (1:1:1); and the female cercus is two-segmented (one-segmented) (
|
||
<bibRefCitation id="EFE44B37FF9CFFFC040D3A0531885713" author="Jaschhof, M." box="[537,697,538,561]" pageId="2" pageNumber="394" pagination="323 - 332" refId="ref1698" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof 2005</bibRefCitation>
|
||
). Further distinctions are found in the male genitalia: the gonocoxites are largely merged ventromedially (largely separate); and hypoproct and cerci are closely associated with each other as well as with the ninth tergite (all these structures clearly separate).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |