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<document id="7E60258C86062686AC4936CEC514F3CE" ID-CLB-Dataset="26862" ID-DOI="10.11646/zootaxa.4603.3.12" ID-GBIF-Dataset="7a36859e-2b95-4aed-b54b-86c5a1cbc2f3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2702255" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557468686060" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2019" docId="F22987B82E196872FF725BB4FAB6F853" docLanguage="en" docName="zootaxa.4603.3.12.pdf" docOrigin="Zootaxa 4603 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Grapholita diehli Heppner &amp; Bae 2019, new species" docType="treatment" docVersion="8" lastPageNumber="590" masterDocId="0E10FFC02E196873FFE55F48FFA0FFAF" masterDocTitle="A new Grapholita species from Sumatra, Indonesia (Lepidoptera: Tortricidae: Olethreutinae: Grapholitini)" masterLastPageNumber="591" masterPageNumber="589" pageNumber="589" updateTime="1698726907191" updateUser="ExternalLinkService">
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<mods:title id="CCBB22083FE084828772E301FCB26F45">A new Grapholita species from Sumatra, Indonesia (Lepidoptera: Tortricidae: Olethreutinae: Grapholitini)</mods:title>
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<mods:name id="2BE68F7F3862F7E0645CAFAADB1B5E94" type="personal">
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<mods:namePart id="D48E63271F3F114087D6C918F19E7D77">Heppner, John B.</mods:namePart>
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<mods:namePart id="B72B58595474099A664B2A069DD44CDA">Bae, Yang-Seop</mods:namePart>
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<mods:title id="7C269F49C09786E78A09B6E96A10A5C6">Zootaxa</mods:title>
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<mods:date id="E60B344167CAE7D7955004A31C068A22">2019</mods:date>
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<mods:number id="107A7804C4E69223B53D84C00F728B88">2019-05-10</mods:number>
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<treatment id="F22987B82E196872FF725BB4FAB6F853" ID-DOI="http://doi.org/10.5281/zenodo.5935029" ID-GBIF-Taxon="157119486" ID-Zenodo-Dep="5935029" LSID="urn:lsid:plazi:treatment:F22987B82E196872FF725BB4FAB6F853" httpUri="http://treatment.plazi.org/id/F22987B82E196872FF725BB4FAB6F853" lastPageId="1" lastPageNumber="590" pageId="0" pageNumber="589">
<subSubSection id="329A65252E196873FF725BB4FCB0FABA" box="[151,784,1276,1301]" pageId="0" pageNumber="589" type="nomenclature">
<paragraph id="7A3F36AE2E196873FF725BB4FCB0FABA" blockId="0.[151,1437,1240,2021]" box="[151,784,1276,1301]" pageId="0" pageNumber="589">
<emphasis id="48F4EABC2E196873FF725BB4FD3EFABB" bold="true" box="[151,670,1276,1301]" pageId="0" pageNumber="589">
<taxonomicName id="BD804D2D2E196873FF725BB4FDAFFABA" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2019" box="[151,527,1276,1301]" class="Insecta" family="Tortricidae" genus="Grapholita" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="589" phylum="Arthropoda" rank="species" species="diehli" status="sp. nov.">
<emphasis id="48F4EABC2E196873FF725BB4FEF4FABB" bold="true" box="[151,340,1276,1300]" italics="true" pageId="0" pageNumber="589">Grapholita diehli</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="53C757C72E196873FDFE5BB5FD3EFABB" box="[539,670,1276,1301]" pageId="0" pageNumber="589" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="E2BB2A2B2E196873FD485BB5FCA6FABA" box="[685,774,1277,1301]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,255,699,723]" captionStartId-1="1.[151,255,1443,1467]" captionStartId-2="2.[151,255,807,831]" captionTargetBox-0="[228,1365,191,631]" captionTargetBox-1="[174,1436,723,1399]" captionTargetBox-2="[340,1246,189,771]" captionTargetId-0="figure@1.[218,1370,181,671]" captionTargetId-1="figure@1.[151,1436,744,1399]" captionTargetId-2="figure@2.[340,1247,189,771]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="2" captionText-0="FIGURE 1. Grapholita diehli n. sp., male holotype (18.5 mm wingspan)." captionText-1="FIGURE 2. Head views of Grapholita diehli n. sp., male: 2) lateral view, and 2a) dorsal view." captionText-2="FIGURE 3. Grapholita diehli n. sp., male genitalia, with details: a) central features enlarged, and b) posterior of abdomen, with sternal plate and lateral scent tufts (right tuft setae removed) (gen. slide JBH 3378)." httpUri-0="https://zenodo.org/record/2702257/files/figure.png" httpUri-1="https://zenodo.org/record/2702259/files/figure.png" httpUri-2="https://zenodo.org/record/2702261/files/figure.png" pageId="0" pageNumber="589">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="329A65252E196873FF725A69FD41FA97" box="[151,737,1312,1337]" pageId="0" pageNumber="589" type="materials_examined">
<paragraph id="7A3F36AE2E196873FF725A69FD41FA97" blockId="0.[151,1437,1240,2021]" box="[151,737,1312,1337]" pageId="0" pageNumber="589">
<materialsCitation id="CAE83CF32E196873FF725A69FD7EFA97" ID-GBIF-Occurrence="2239631041" box="[151,734,1312,1337]" country="Indonesia" location="Prapat" pageId="0" pageNumber="589" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="48F4EABC2E196873FF725A69FE92FA97" bold="true" box="[151,306,1312,1337]" pageId="0" pageNumber="589">
<typeStatus id="A53B880C2E196873FF725A69FF6FFA96" box="[151,207,1313,1337]" pageId="0" pageNumber="589">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="0297763E2E196873FEDD5A69FE02FA96" box="[312,418,1313,1337]" name="Indonesia" pageId="0" pageNumber="589">Indonesia</collectingCountry>
, near
<location id="7F5F60752E196873FE075A69FD89FA97" LSID="urn:lsid:plazi:treatment:F22987B82E196872FF725BB4FAB6F853:7F5F60752E196873FE075A69FD89FA97" box="[482,553,1313,1336]" country="Indonesia" name="Prapat" pageId="0" pageNumber="589" stateProvince="Sumatera Utara">Prapat</location>
,
<collectingRegion id="B844F84C2E196873FDD65A69FD7EFA97" box="[563,734,1313,1337]" country="Indonesia" name="Sumatera Utara" pageId="0" pageNumber="589">Sumatera Utara</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="329A65252E196873FF225A0CFB37FA2E" pageId="0" pageNumber="589" type="diagnosis">
<paragraph id="7A3F36AE2E196873FF225A0CFB37FA2E" blockId="0.[151,1437,1240,2021]" pageId="0" pageNumber="589">
<emphasis id="48F4EABC2E196873FF225A0CFE95FAF3" bold="true" box="[199,309,1348,1372]" pageId="0" pageNumber="589">Diagnosis</emphasis>
. This species is distinctive with its apical hindwing yellow patch, whereas both forewings and hindwings are otherwise mostly brown. This superficial character distinguishes it from all other congeners.
</paragraph>
</subSubSection>
<subSubSection id="329A65252E196872FF225AC4FB48F987" lastPageId="1" lastPageNumber="590" pageId="0" pageNumber="589" type="description">
<paragraph id="7A3F36AE2E196873FF225AC4FE10F84A" blockId="0.[151,1437,1240,2021]" pageId="0" pageNumber="589">
<emphasis id="48F4EABC2E196873FF225AC4FEEBFA0B" bold="true" box="[199,331,1420,1444]" pageId="0" pageNumber="589">Description</emphasis>
. Male (
<figureCitation id="E2BB2A2B2E196873FE465AC5FE44FA0A" box="[419,484,1421,1445]" captionStart="FIGURE 1" captionStartId="1.[151,255,699,723]" captionTargetBox="[228,1365,191,631]" captionTargetId="figure@1.[218,1370,181,671]" captionTargetPageId="1" captionText="FIGURE 1. Grapholita diehli n. sp., male holotype (18.5 mm wingspan)." httpUri="https://zenodo.org/record/2702257/files/figure.png" pageId="0" pageNumber="589">Fig. 1</figureCitation>
).
<emphasis id="48F4EABC2E196873FE185AC5FD97FA0A" box="[509,567,1421,1445]" italics="true" pageId="0" pageNumber="589">Head</emphasis>
(
<figureCitation id="E2BB2A2B2E196873FDAD5AC5FD2CFA0A" box="[584,652,1421,1445]" captionStart="FIGURE 2" captionStartId="1.[151,255,1443,1467]" captionTargetBox="[174,1436,723,1399]" captionTargetId="figure@1.[151,1436,744,1399]" captionTargetPageId="1" captionText="FIGURE 2. Head views of Grapholita diehli n. sp., male: 2) lateral view, and 2a) dorsal view." httpUri="https://zenodo.org/record/2702259/files/figure.png" pageId="0" pageNumber="589">Fig. 2</figureCitation>
): Vertex dark brown; frons pale brown; venter tan; labial palpus brown (mesally pale tan); antenna gray with pale white ventral bristles.
<emphasis id="48F4EABC2E196873FCA45AF9FC2EFA66" box="[833,910,1457,1481]" italics="true" pageId="0" pageNumber="589">Thorax</emphasis>
: Brown; patagia brown; tegulae brown, with posterior half silver; venter of thorax light tan; leg coxae dull light tan, with golden iridescence; femora same (ventrally pale tan); tibiae tan (with median tibia having tan spines), and mesally all dull white-tan; tarsi pale brown with tan segmental ends, and ventrally tan. Forewing (
<figureCitation id="E2BB2A2B2E196873FDF55955FDEFF99A" box="[528,591,1565,1589]" captionStart="FIGURE 1" captionStartId="1.[151,255,699,723]" captionTargetBox="[228,1365,191,631]" captionTargetId="figure@1.[218,1370,181,671]" captionTargetPageId="1" captionText="FIGURE 1. Grapholita diehli n. sp., male holotype (18.5 mm wingspan)." httpUri="https://zenodo.org/record/2702257/files/figure.png" pageId="0" pageNumber="589">Fig. 1</figureCitation>
) expanse 18.5 mm (n = 1); forewing dark brown, with brown-gray costa with short black strigulae (angled to tornus); scattered black spots on terminal 1/5; black-brown terminal line as fringe base; fringe gray; venter brown, with yellow patch at radial sector, terminal 1/4 with scattered tiny pale tan spots, and costal margin tan with dorsal strigulae repeated in dark brown, and pale white along dorsal margin. Hindwing (
<figureCitation id="E2BB2A2B2E196873FAE459C1FAE0F90E" box="[1281,1344,1673,1697]" captionStart="FIGURE 1" captionStartId="1.[151,255,699,723]" captionTargetBox="[228,1365,191,631]" captionTargetId="figure@1.[218,1370,181,671]" captionTargetPageId="1" captionText="FIGURE 1. Grapholita diehli n. sp., male holotype (18.5 mm wingspan)." httpUri="https://zenodo.org/record/2702257/files/figure.png" pageId="0" pageNumber="589">Fig. 1</figureCitation>
) brown, with apical 2/5 and dorsally to mid-wing yellow, with scattered brown spots on yellow patch near apex; fringe yellow on apex and termen to mid-wing, then pale brown; venter brown with yellow apical patch repeated.
<emphasis id="48F4EABC2E196873FB7B5999FAA3F946" box="[1182,1283,1745,1769]" italics="true" pageId="0" pageNumber="589">Abdomen</emphasis>
(
<figureCitation id="E2BB2A2B2E196873FAF45999FAF0F946" box="[1297,1360,1745,1769]" captionStart="FIGURE 1" captionStartId="1.[151,255,699,723]" captionTargetBox="[228,1365,191,631]" captionTargetId="figure@1.[218,1370,181,671]" captionTargetPageId="1" captionText="FIGURE 1. Grapholita diehli n. sp., male holotype (18.5 mm wingspan)." httpUri="https://zenodo.org/record/2702257/files/figure.png" pageId="0" pageNumber="589">Fig. 1</figureCitation>
): Dark brown; venter gray-brown; genital tufts dark brown and tan ventrally; pregenital segment with large coremata (
<figureCitation id="E2BB2A2B2E196873FAA559BDFA2FF8A2" box="[1344,1423,1781,1805]" captionStart="FIGURE 3" captionStartId="2.[151,255,807,831]" captionTargetBox="[340,1246,189,771]" captionTargetId="figure@2.[340,1247,189,771]" captionTargetPageId="2" captionText="FIGURE 3. Grapholita diehli n. sp., male genitalia, with details: a) central features enlarged, and b) posterior of abdomen, with sternal plate and lateral scent tufts (right tuft setae removed) (gen. slide JBH 3378)." httpUri="https://zenodo.org/record/2702261/files/figure.png" pageId="0" pageNumber="589">Fig. 3b</figureCitation>
). Male genitalia (
<figureCitation id="E2BB2A2B2E196873FEA45851FE21F89E" box="[321,385,1817,1841]" captionStart="FIGURE 3" captionStartId="2.[151,255,807,831]" captionTargetBox="[340,1246,189,771]" captionTargetId="figure@2.[340,1247,189,771]" captionTargetPageId="2" captionText="FIGURE 3. Grapholita diehli n. sp., male genitalia, with details: a) central features enlarged, and b) posterior of abdomen, with sternal plate and lateral scent tufts (right tuft setae removed) (gen. slide JBH 3378)." httpUri="https://zenodo.org/record/2702261/files/figure.png" pageId="0" pageNumber="589">Fig. 3</figureCitation>
) with tegumen simple, triangular, with rounded basal ends and pedunculus; juxta triangular, with thin cucullus (subequal in length to width of juxta); valva elongated with bulbous and rounded apical third, from quadrate base and large basal cavity, with a constricted narrow central portion due to an extreme invagination of the ventral margin, and with apical portion bent ventrally; apex of valva with numerous strong spines near to terminal margin; aedeagus (
<figureCitation id="E2BB2A2B2E196873FA9458CDFF11F86E" captionStart="FIGURE 3" captionStartId="2.[151,255,807,831]" captionTargetBox="[340,1246,189,771]" captionTargetId="figure@2.[340,1247,189,771]" captionTargetPageId="2" captionText="FIGURE 3. Grapholita diehli n. sp., male genitalia, with details: a) central features enlarged, and b) posterior of abdomen, with sternal plate and lateral scent tufts (right tuft setae removed) (gen. slide JBH 3378)." httpUri="https://zenodo.org/record/2702261/files/figure.png" pageId="0" pageNumber="589">Fig. 3a</figureCitation>
) relatively narrow, slightly curved near middle, with elongated phallobase; coremata as spinose line on distal vesica margin. Female unknown.
</paragraph>
<caption id="2EFF66262E186872FF725DF3FC0EFD7C" box="[151,942,699,723]" httpUri="https://zenodo.org/record/2702257/files/figure.png" pageId="1" pageNumber="590" startId="1.[151,255,699,723]" targetBox="[228,1365,191,631]" targetPageId="1">
<paragraph id="7A3F36AE2E186872FF725DF3FC0EFD7C" blockId="1.[151,942,699,723]" box="[151,942,699,723]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF725DF3FEB9FD7C" bold="true" box="[151,281,699,723]" pageId="1" pageNumber="590">FIGURE 1.</emphasis>
<taxonomicName id="BD804D2D2E186872FEC55DF3FE7AFD7C" authority="Heppner &amp; Bae, 2019" authorityName="Heppner &amp; Bae" authorityYear="2019" box="[288,474,699,723]" class="Insecta" family="Tortricidae" genus="Grapholita" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="diehli" status="sp. nov.">
<emphasis id="48F4EABC2E186872FEC55DF3FE7AFD7C" box="[288,474,699,723]" italics="true" pageId="1" pageNumber="590">Grapholita diehli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53C757C72E186872FE045DF3FDBCFD7D" box="[481,540,699,723]" pageId="1" pageNumber="590" rank="species">
<emphasis id="48F4EABC2E186872FE045DF3FDB6FD7C" bold="true" box="[481,534,699,723]" pageId="1" pageNumber="590">n. sp</emphasis>
.
</taxonomicNameLabel>
, male holotype (18.5 mm wingspan).
</paragraph>
</caption>
<caption id="2EFF66262E186872FF725AEBFB2AFA13" box="[151,1162,1443,1468]" httpUri="https://zenodo.org/record/2702259/files/figure.png" pageId="1" pageNumber="590" startId="1.[151,255,1443,1467]" targetBox="[174,1436,723,1399]" targetPageId="1">
<paragraph id="7A3F36AE2E186872FF725AEBFB2AFA13" blockId="1.[151,1162,1443,1468]" box="[151,1162,1443,1468]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF725AEBFEB9FA13" bold="true" box="[151,281,1443,1468]" pageId="1" pageNumber="590">FIGURE 2.</emphasis>
Head views of
<taxonomicName id="BD804D2D2E186872FE245AECFDDBFA13" authority="Heppner &amp; Bae, 2019" authorityName="Heppner &amp; Bae" authorityYear="2019" box="[449,635,1444,1468]" class="Insecta" family="Tortricidae" genus="Grapholita" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="diehli" status="sp. nov.">
<emphasis id="48F4EABC2E186872FE245AECFDDBFA13" box="[449,635,1444,1468]" italics="true" pageId="1" pageNumber="590">Grapholita diehli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53C757C72E186872FD675AECFD1DFA13" box="[642,701,1444,1468]" pageId="1" pageNumber="590" rank="species">
<emphasis id="48F4EABC2E186872FD675AECFD1DFA13" bold="true" box="[642,701,1444,1468]" pageId="1" pageNumber="590">n. sp.</emphasis>
</taxonomicNameLabel>
, male: 2) lateral view, and 2a) dorsal view.
</paragraph>
</caption>
<paragraph id="7A3F36AE2E186872FF225AA3FB48F987" blockId="1.[151,1437,1514,2044]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF225AA3FE3CF9AC" bold="true" box="[199,412,1515,1539]" pageId="1" pageNumber="590">Specimens studied</emphasis>
.
<materialsCitation id="CAE83CF32E186872FE495AA4FB48F987" ID-GBIF-Occurrence="2239631068" collectingDate="1992-08-30" collectionCode="MGCL" collectorName="E. Prapat &amp; J. B. Heppner &amp; E. W. Diehl" country="Indonesia" elevation="1050" location="Sumatera Utara" pageId="1" pageNumber="590" specimenCode="JBH 3378" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<typeStatus id="A53B880C2E186872FE495AA4FDAFF9AB" box="[428,527,1516,1540]" pageId="1" pageNumber="590" type="holotype">Holotype</typeStatus>
<quantity id="BD789B4B2E186872FDFC5AA2FD24F9AB" box="[537,644,1514,1540]" metricMagnitude="3" metricUnit="m" metricValue="3.18" pageId="1" pageNumber="590" unit="km" value="3.18">³, 18 km</quantity>
<collectorName id="D77553782E186872FD6B5AA4FD52F9AC" box="[654,754,1516,1539]" pageId="1" pageNumber="590">E. Prapat</collectorName>
(H[olz]W[eg]2),
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<quantity id="BD789B4B2E186872FC505AA3FBABF9AC" box="[949,1035,1515,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="590" unit="m" value="1050.0">1050 m</quantity>
</elevation>
,
<collectingRegion id="B844F84C2E186872FBFC5AA4FB66F9AC" box="[1049,1222,1516,1540]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="590">Sumatera Utara</collectingRegion>
,
<collectingCountry id="0297763E2E186872FB365AA4FA9DF9AB" box="[1235,1341,1516,1540]" name="Indonesia" pageId="1" pageNumber="590">Indonesia</collectingCountry>
,
<collectingDate id="1E7AE9862E186872FAAE5AA4FF6FF987" pageId="1" pageNumber="590" value="1992-08-30">
<date id="0E3E106E2E186872FAAE5AA4FF6FF987" pageId="1" pageNumber="590" value="1992-08-30">30 Aug 1992</date>
</collectingDate>
,
<collectorName id="D77553782E186872FF3F5958FECDF988" box="[218,365,1552,1575]" pageId="1" pageNumber="590">J. B. Heppner</collectorName>
&amp;
<collectorName id="D77553782E186872FE6A5958FDABF987" box="[399,523,1552,1576]" pageId="1" pageNumber="590">E. W. Diehl</collectorName>
(gen. slide
<specimenCode id="2A269ED52E186872FD625958FD57F987" box="[647,759,1552,1576]" collectionCode="JBH" pageId="1" pageNumber="590">JBH 3378</specimenCode>
; photo 11817) (
<collectionCode id="1C91AE6B2E186872FC785958FC4DF987" box="[925,1005,1552,1576]" pageId="1" pageNumber="590">MGCL</collectionCode>
) (deposited at
<collectionCode id="1C91AE6B2E186872FB6E5958FB7DF987" box="[1163,1245,1552,1576]" pageId="1" pageNumber="590">MGCL</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="329A65252E186872FF22597CFF40F9DF" pageId="1" pageNumber="590" type="etymology">
<paragraph id="7A3F36AE2E186872FF22597CFF40F9DF" blockId="1.[151,1437,1514,2044]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF22597CFEE2F9E3" bold="true" box="[199,322,1588,1612]" pageId="1" pageNumber="590">Etymology</emphasis>
. The species is named in honor of the late Dr. E. W. Diehl, long-time collector and specialist of Sumatran moths.
</paragraph>
</subSubSection>
<subSubSection id="329A65252E186872FF225933FE3BF93B" box="[199,411,1659,1684]" pageId="1" pageNumber="590" type="biology_ecology">
<paragraph id="7A3F36AE2E186872FF225933FE3BF93B" blockId="1.[151,1437,1514,2044]" box="[199,411,1659,1684]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF225933FEBDF93C" bold="true" box="[199,285,1659,1683]" pageId="1" pageNumber="590">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="329A65252E186872FF2259D7FCB4F8AF" pageId="1" pageNumber="590" type="distribution">
<paragraph id="7A3F36AE2E186872FF2259D7FCB4F8AF" blockId="1.[151,1437,1514,2044]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF2259D7FEF2F918" bold="true" box="[199,338,1695,1719]" pageId="1" pageNumber="590">Distribution</emphasis>
. The new species is known only from northern
<collectingRegion id="B844F84C2E186872FCBD59E8FC11F917" box="[856,945,1696,1720]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="590">Sumatra</collectingRegion>
in the Medan region. The collecting site data used by E. W. Dielh, “HW2” [Holzweg 2, or logging road 2], refers to one of his usual collecting sites east of the town of Prapat, in mountainous areas about
<quantity id="BD789B4B2E186872FDF159A0FDFEF8AF" box="[532,606,1768,1792]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="590" unit="km" value="100.0">100km</quantity>
south of Medan.
</paragraph>
</subSubSection>
<subSubSection id="329A65252E186872FF225843FAB6F853" pageId="1" pageNumber="590" type="discussion">
<paragraph id="7A3F36AE2E186872FF225843FAB6F853" blockId="1.[151,1437,1514,2044]" pageId="1" pageNumber="590">
<emphasis id="48F4EABC2E186872FF225843FE9FF88C" bold="true" box="[199,319,1803,1827]" pageId="1" pageNumber="590">Discussion</emphasis>
. The new species is unique among
<taxonomicName id="BD804D2D2E186872FD585844FC94F88B" authorityName="Treitschke" authorityYear="1829" box="[701,820,1804,1828]" class="Insecta" family="Tortricidae" genus="Grapholita" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="48F4EABC2E186872FD585844FC94F88B" box="[701,820,1804,1828]" italics="true" pageId="1" pageNumber="590">Grapholita</emphasis>
</taxonomicName>
species. Few species of
<taxonomicName id="BD804D2D2E186872FBD95844FB11F88B" box="[1084,1201,1804,1828]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
depart from the usual monotone hindwings, or with at most the basal region pale, and these alternate maculation forms either have a light-colored hindwing overall with margins dark, or with the apical portion dark, thus, the reverse of
<taxonomicName id="BD804D2D2E186872FB64581CFB7FF8C3" authority="Heppner &amp; Bae, 2019" authorityName="Heppner &amp; Bae" authorityYear="2019" box="[1153,1247,1876,1900]" class="Insecta" family="Tortricidae" genus="Grapholita" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="diehli" status="sp. nov.">
<emphasis id="48F4EABC2E186872FB64581CFB7FF8C3" box="[1153,1247,1876,1900]" italics="true" pageId="1" pageNumber="590">G. diehli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53C757C72E186872FB03581CFA83F8C3" box="[1254,1315,1876,1900]" pageId="1" pageNumber="590" rank="species">
<emphasis id="48F4EABC2E186872FB03581CFA83F8C3" bold="true" box="[1254,1315,1876,1900]" pageId="1" pageNumber="590">n. sp.</emphasis>
</taxonomicNameLabel>
, which has a light apical portion (
<bibRefCitation id="1E114B5F2E186872FE605830FDA9F83F" author="Horak, M." box="[389,521,1912,1936]" pageId="1" pageNumber="590" refId="ref1197" refString="Horak, M. (2006) Olethreutine Moths of Australia (Lepidoptera: Tortricidae). In: Monographs on Australian Lepidoptera. Vol. 10. CSIRO, Collingwood, 528 pp." type="book" year="2006">Horak 2006</bibRefCitation>
,
<bibRefCitation id="1E114B5F2E186872FDF15830FD3DF83F" author="Komai, F." box="[532,669,1912,1936]" pageId="1" pageNumber="590" pagination="1 - 226" refId="ref1232" refString="Komai, F. (1999) A taxonomic review of the genus Grapholita and allied genera (Lepidoptera: Tortricidae) in the Palaearctic region. Entomologica Scandinavica, 55 (Supplement), 1 - 226. [Copenhagen]" type="journal article" year="1999">Komai 1999</bibRefCitation>
). It is unfortunate that only a single specimen has been discovered thus far in
<collectingRegion id="B844F84C2E186872FF3258D4FE93F81B" box="[215,307,1948,1972]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="590">Sumatra</collectingRegion>
, even though one of us (JBH) has surveyed the region extensively for micro-moths on several expeditions with Dr. Diehl. After 27 years, and before that, 180 years of collected, no additional specimens have bee discovered, so the species is now named in the hope that it will foster the search for more specimens, including the female.
</paragraph>
</subSubSection>
</treatment>
</document>