1425 lines
206 KiB
XML
1425 lines
206 KiB
XML
<document id="8D0CD3C1022E4B7C56FC52695DC0DBFF" ID-DOI="10.5852/ejt.2023.914.2377" ID-ISSN="2118-9773" ID-Zenodo-Dep="10407071" ID-ZooBank="2ECE4DA6-97CA-4917-97F1-FE506998A2D0" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1703002881060" checkinUser="plazi" docAuthor="Basílio, Daniel Silva, Vaz-De-Mello, Fernando Zagury, Cherman, Mariana Alejandra & Almeida, Lúcia Massutti de" docDate="2023" docId="EA056F037438FFECB80FFB1F4578FABE" docLanguage="en" docName="EJT.2023.914.1-53.pdf" docOrigin="European Journal of Taxonomy 914 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2377/10437" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Coilodes Westwood 1846" docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="163C177B743AFFEBBA58FFE4400FFFA7" masterDocTitle="A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species" masterLastPageNumber="53" masterPageNumber="1" pageNumber="3" updateTime="1704204554843" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="49BC52F7FC1ED843C8FBBB209A626007" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5B7779E38C28AD903730DFCB1618845E">
|
||
<mods:title id="0469A8A802082A9EB5E92CFF39DF1181">A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="2B959FCBEF7C6271C16AD25188194FFE" type="personal">
|
||
<mods:role id="3F498DEEE98B34DF0BF2976F33A6A636">
|
||
<mods:roleTerm id="764474E54BAEB4EFB1A8A9B343C2D8DC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3AD671399A48C65A5AC9BF57B96D74F1">Basílio, Daniel Silva</mods:namePart>
|
||
<mods:nameIdentifier id="22B4999366B6600E186DFC6179E6DDBD" type="LSID">A036BC96-736C-46C7-9959-A5980DECFAC4</mods:nameIdentifier>
|
||
<mods:affiliation id="D6055905253C1C981C517089A8BE2400">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="0927F2EB33B41C7CB7EF3A712D7648BA" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="E0130BD0266616DD93C865AB51CDE905" type="personal">
|
||
<mods:role id="615298B9669159DF22F732359BF27FCF">
|
||
<mods:roleTerm id="94CEB1B23184DA6E1959527C82E81D84">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DE8A7A614CAB3036D8EA31CF87FF6171">Vaz-De-Mello, Fernando Zagury</mods:namePart>
|
||
<mods:nameIdentifier id="7207CF2F9233BA5EF20FA9F76676BD5A" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
|
||
<mods:affiliation id="7A8579A7E74F92999B6BD2715F9C3909">Laboratório de Scarabaeoidologia, Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="B0EBAA432B179B30DE329CFA11542A9F" type="email">vazdemello@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="095E8290370FAD30A6035140C9824629" type="personal">
|
||
<mods:role id="B116BC258CFB8167FD1CDDC265F26A19">
|
||
<mods:roleTerm id="F1A6507FBD6B781F091A4AB28AED3A44">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="26D24D0290D281C2D0D2A3195727B576">Cherman, Mariana Alejandra</mods:namePart>
|
||
<mods:nameIdentifier id="D2F1B4B581949DF7D7DA456B32E901E4" type="LSID">EAF28053-7ABC-4F2D-9C46-CA7B1AEC2C5A</mods:nameIdentifier>
|
||
<mods:affiliation id="8B1A1B2FF1AE307F6E20D109A524AB7F">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="EC69B44CDC25D3AA51E3CB185F6EEF31" type="email">marianabioar@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="D5DCB9961B26022E20FC21A042364BA6" type="personal">
|
||
<mods:role id="DB0917218BBEADEC37AE997461A0506D">
|
||
<mods:roleTerm id="95979B6B184AD038FBF996A1B1C63D1E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8D29B9007FCFDBE69E963201C4F25D27">Almeida, Lúcia Massutti de</mods:namePart>
|
||
<mods:nameIdentifier id="94EF7D367D4D24526554FEFE077E3894" type="LSID">B3B2EF56-E27B-4822-B854-CB01A1D8E64B</mods:nameIdentifier>
|
||
<mods:affiliation id="DF2C883468E89AB37E67AAE06BF4183C">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="342710EDBF032D59B285AEB1FD38C188" type="email">lalmeida51@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D898385C0EE629C624384CFBBAB7FB2A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="927A3CFF4E296ABAC2D48795E8A29A38" type="host">
|
||
<mods:titleInfo id="CB2F5D52EBCC9E3F74951322CFBACF3F">
|
||
<mods:title id="4A3021DEC70351AAAE3DD1C00DE80E45">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="37673F6B27094B22B7F39CCB42DE7936">
|
||
<mods:date id="851942AF97ACF348E6E94C93F1B8303D">2023</mods:date>
|
||
<mods:detail id="8E801C8C1F2F2C4E77947DBFE5F7B1DD" type="pubDate">
|
||
<mods:number id="EA6F48B2F7302DBF3F97927B9AC384DE">2023-12-19</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="824906B02E2787E67C47EE6A988DF762" type="volume">
|
||
<mods:number id="91E77676332425C75887C07BC9738557">914</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="753C15A871CE31CADA33770956379F11" type="issue">
|
||
<mods:number id="A21E3C8818126BF076389E698B82872E">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="787688CCE49FF8021AE76CF6EB8F4BE1" unit="page">
|
||
<mods:start id="2D6CA39572CDE4D8F9758477AE0F2C90">1</mods:start>
|
||
<mods:end id="1485252B08C96CD101CAF0D2ACA5F788">53</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="FC4A5EA77A350F26C836DB9784013B20">
|
||
<mods:url id="3EAD49A640242C1AA417A38DEECF6197">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2377/10437</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="8059618B7C33C6FB50DDA990EE154717">journal article</mods:classification>
|
||
<mods:identifier id="10F9B0B1F2D690395BAAC19EC379A25E" type="DOI">10.5852/ejt.2023.914.2377</mods:identifier>
|
||
<mods:identifier id="184FF7DD01F33F30B70AEB43BB44630D" type="ISSN">2118-9773</mods:identifier>
|
||
<mods:identifier id="848099401B3E7DDE6B08D7842517DC19" type="Zenodo-Dep">10407071</mods:identifier>
|
||
<mods:identifier id="0C0A5CA3CEC83492E07DD65C7C969B10" type="ZooBank">2ECE4DA6-97CA-4917-97F1-FE506998A2D0</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="EA056F037438FFECB80FFB1F4578FABE" LSID="urn:lsid:plazi:treatment:EA056F037438FFECB80FFB1F4578FABE" httpUri="http://treatment.plazi.org/id/EA056F037438FFECB80FFB1F4578FABE" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
|
||
<subSubSection id="2AB68D9E7438FFE9B80FFB1F43D2FAB2" box="[599,989,1274,1301]" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="6213DE157438FFE9B80FFB1F43D2FAB2" blockId="2.[599,989,1274,1337]" box="[599,989,1274,1301]" pageId="2" pageNumber="3">
|
||
<heading id="395B69797438FFE9B80FFB1F43D2FAB2" box="[599,989,1274,1301]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
|
||
<emphasis id="50D802077438FFE9B80FFB1F42ADFAB2" box="[599,674,1275,1301]" italics="true" pageId="2" pageNumber="3">Genus</emphasis>
|
||
<taxonomicName id="A5ACA5967438FFE9B8F1FB1E43D2FAB2" ID-CoL="3S4K" authority="Westwood, 1846" authorityName="Westwood" authorityYear="1846" box="[681,989,1274,1301]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077438FFE9B8F1FB1E431EFAB2" bold="true" box="[681,785,1274,1301]" italics="true" pageId="2" pageNumber="3">Coilodes</emphasis>
|
||
<bibRefCitation id="063DA3E47438FFE9B940FB1F43D2FAB2" author="Westwood J. O." box="[792,989,1275,1301]" pageId="2" pageNumber="3" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7438FFE9B887FAFB435BFA9E" box="[735,852,1311,1337]" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="6213DE157438FFE9B887FAFB435BFA9E" blockId="2.[599,989,1274,1337]" box="[735,852,1311,1337]" pageId="2" pageNumber="3">
|
||
<figureCitation id="FA97C2907438FFE9B887FAFB435BFA9E" box="[735,852,1311,1337]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig" captionStart-11="Fig" captionStart-12="Fig" captionStart-13="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="8.[189,232,1857,1883]" captionStartId-1="10.[189,232,1857,1883]" captionStartId-10="40.[189,232,1822,1848]" captionStartId-11="42.[189,233,1892,1918]" captionStartId-12="45.[189,232,1822,1848]" captionStartId-13="47.[189,232,1787,1813]" captionStartId-2="15.[189,233,1892,1918]" captionStartId-3="18.[189,232,1787,1813]" captionStartId-4="24.[189,232,1822,1848]" captionStartId-5="27.[189,232,1857,1883]" captionStartId-6="29.[189,231,1892,1918]" captionStartId-7="32.[189,232,1822,1848]" captionStartId-8="35.[189,232,1857,1883]" captionStartId-9="37.[191,235,1857,1883]" captionTargetBox-0="[189,1398,730,1795]" captionTargetBox-1="[194,1397,736,1817]" captionTargetBox-10="[189,1398,703,1775]" captionTargetBox-11="[189,1398,766,1834]" captionTargetBox-12="[189,1398,698,1763]" captionTargetBox-13="[272,1315,265,1739]" captionTargetBox-2="[190,1398,774,1820]" captionTargetBox-3="[189,1398,661,1744]" captionTargetBox-4="[189,1397,698,1750]" captionTargetBox-5="[189,1398,731,1793]" captionTargetBox-6="[198,1398,774,1811]" captionTargetBox-7="[189,1398,698,1768]" captionTargetBox-8="[197,1398,733,1807]" captionTargetBox-9="[189,1398,731,1779]" captionTargetId-0="figure-14@8.[189,1398,729,1816]" captionTargetId-1="figure-92@10.[189,1398,731,1819]" captionTargetId-10="figure-14@40.[189,1398,695,1784]" captionTargetId-11="figure-14@42.[189,1398,765,1853]" captionTargetId-12="figure-11@45.[189,1398,697,1784]" captionTargetId-13="figure-12@47.[272,1315,265,1739]" captionTargetId-2="figure-11@15.[189,1398,768,1855]" captionTargetId-3="figure-15@18.[189,1398,660,1748]" captionTargetId-4="figure-14@24.[189,1398,697,1784]" captionTargetId-5="figure-196@27.[189,1398,731,1819]" captionTargetId-6="figure-12@29.[189,1398,766,1854]" captionTargetId-7="figure-14@32.[189,1398,697,1784]" captionTargetId-8="figure-231@35.[189,1398,732,1819]" captionTargetId-9="figure-80@37.[189,1398,730,1817]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-10="40" captionTargetPageId-11="42" captionTargetPageId-12="45" captionTargetPageId-13="47" captionTargetPageId-2="15" captionTargetPageId-3="18" captionTargetPageId-4="24" captionTargetPageId-5="27" captionTargetPageId-6="29" captionTargetPageId-7="32" captionTargetPageId-8="35" captionTargetPageId-9="37" captionText-0="Fig. 1. Coilodes bezerrai Basílio & Vaz-de-Mello sp. nov., paratype ♂ at DZUP (A, B, D), holotype ♂ (C, E–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-1="Fig. 2. Coilodes castaneus Westwood, 1846, ♂ at CEMT from Cerro Campana, Panama (A, C–E), ♂ at DZUP from Puntarenas, Costa Rica (B), ♂ at DZUP from Canal Zone, Panamá (F–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-10="Fig. 11. Coilodes punctipennis Arrow, 1909, ♂ at UCCC from Tropica Region Chapare, Bolivia (A, C–E), ♂ at CEMT from Cotococha, Napo, Ecuador (B), ♂ at DZUP from Tugurahua, Ecuador (F–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-11="Fig. 12. Coilodes ravii Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-12="Fig. 13. Coilodes skelleyi Basílio & Vaz-de-Mello sp. nov., paratype ♂ at CEMT from Pico da Neblina, Amazonas, Brazil (A, F–G), paratype ♀ at DZUP from Loreto, Peru (B), paratype ♂ at DZUP from Benjamin Constant, Amazonas, Brazil (C, E), holotype ♂ (D, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-13="Fig. 14. Distribution map of species of Coilodes Westwood, 1846. A. Coilodes bezerrai Basílio & Vazde-Mello sp. nov.; Coilodes castaneus Westwood, 1846; and Coilodes edeiltae Basílio & Vaz-de-Mello sp. nov. B. Coilodes fumipennis Arrow, 1909; Coilodes humeralis (Mannerheim, 1829); and Coilodes lunae Basílio & Vaz-de-Mello sp. nov. C. Coilodes mayae Basílio & Vaz-de-Mello sp. nov.;Coilodes niger (Mannerheim, 1829) stat. rev.; Coilodes ovalis Robinson, 1948; and Coilodes parvulus Westwood 1846. D. Coilodes punctipennis Arrow, 1909; Coilodes ravii Basílio & Vaz-de-Mello sp. nov.; and Coilodes skelleyi Basílio & Vaz-de-Mello sp. nov." captionText-2="Fig. 3. Coilodes edeiltae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-3="Fig. 4. Coilodes fumipennis Arrow, 1909, ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (A, C, E), ♂ at CERPE from Rondônia, Brazil (B), ♂ at CERPE from Alta Floresta, Mato Grosso, Brazil (D); ♂ at CEMT from Rondônia, Brazil (F–G), graphical representation (H–I), and another ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-4="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-5="Fig. 6. Coilodes lunae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–C, E–G, J–K), paratype ♀ at DZUP from Itatiaia, Rio de Janeiro (D), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-6="Fig. 7. Coilodes mayae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-7="Fig. 8. Coilodes niger (Mannerheim, 1829) stat. rev., two ♂♂ from Macaé de Cima, Nova Friburgo, Rio de Janeiro, Brazil, at CERPE (A–E) and at CEMT (F–G); another ♂ at CEMT from Nova Friburgo, Rio de Janeiro, Brazil (J–K); and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-8="Fig. 9. Coilodes ovalis Robinson, 1948, three ♂♂ from Jaji, Sucre, Venezuela, two at CEMT (first A, C–D; and second B, E), and one at DZUP (F–G, J–K); and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view.C. Head, dorsal view. D. Head, ventral view.E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." captionText-9="Fig. 10. Coilodes parvulus Westwood, 1846, Three ♂♂ at CEMT, two from Ubajara, Ceará, Brazil (A–B, E) and (C), and one from Igrapiuna, Bahia, Brazil (D, F–G, J–K); graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10407073" figureDoi-1="http://doi.org/10.5281/zenodo.10407075" figureDoi-10="http://doi.org/10.5281/zenodo.10407103" figureDoi-11="http://doi.org/10.5281/zenodo.10407105" figureDoi-12="http://doi.org/10.5281/zenodo.10407109" figureDoi-13="http://doi.org/10.5281/zenodo.10407111" figureDoi-2="http://doi.org/10.5281/zenodo.10407081" figureDoi-3="http://doi.org/10.5281/zenodo.10407085" figureDoi-4="http://doi.org/10.5281/zenodo.10407089" figureDoi-5="http://doi.org/10.5281/zenodo.10407093" figureDoi-6="http://doi.org/10.5281/zenodo.10407095" figureDoi-7="http://doi.org/10.5281/zenodo.10407097" figureDoi-8="http://doi.org/10.5281/zenodo.10407099" figureDoi-9="http://doi.org/10.5281/zenodo.10407101" httpUri-0="https://zenodo.org/record/10407073/files/figure.png" httpUri-1="https://zenodo.org/record/10407075/files/figure.png" httpUri-10="https://zenodo.org/record/10407103/files/figure.png" httpUri-11="https://zenodo.org/record/10407105/files/figure.png" httpUri-12="https://zenodo.org/record/10407109/files/figure.png" httpUri-13="https://zenodo.org/record/10407111/files/figure.png" httpUri-2="https://zenodo.org/record/10407081/files/figure.png" httpUri-3="https://zenodo.org/record/10407085/files/figure.png" httpUri-4="https://zenodo.org/record/10407089/files/figure.png" httpUri-5="https://zenodo.org/record/10407093/files/figure.png" httpUri-6="https://zenodo.org/record/10407095/files/figure.png" httpUri-7="https://zenodo.org/record/10407097/files/figure.png" httpUri-8="https://zenodo.org/record/10407099/files/figure.png" httpUri-9="https://zenodo.org/record/10407101/files/figure.png" pageId="2" pageNumber="3">Figs 1–14</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7438FFE9BAE5FA8644D2F8D1" pageId="2" pageNumber="3" type="reference_group">
|
||
<paragraph id="6213DE157438FFE9BAE5FA86433BFADB" blockId="2.[189,820,1378,1404]" box="[189,820,1378,1404]" pageId="2" pageNumber="3">
|
||
<treatmentCitationGroup id="42BCF93B7438FFE9BAE5FA86433BFADB" box="[189,820,1378,1404]" pageId="2" pageNumber="3">
|
||
<treatmentCitation id="E30DF8047438FFE9BAE5FA864223FADB" author="Westwood J. O." box="[189,556,1378,1404]" page="163" pageId="2" pageNumber="3" year="1846">
|
||
<taxonomicName id="A5ACA5967438FFE9BAE5FA864223FADB" ID-CoL="3S4K" authority="Westwood, 1846: 163" authorityName="Westwood" authorityPageNumber="163" authorityYear="1846" box="[189,556,1378,1404]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077438FFE9BAE5FA86412AFADB" box="[189,293,1378,1404]" italics="true" pageId="2" pageNumber="3">Coilodes</emphasis>
|
||
<bibRefCitation id="063DA3E47438FFE9BB74FA864223FADB" author="Westwood J. O." box="[300,556,1378,1404]" pageId="2" pageNumber="3" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846: 163</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(original description).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="6213DE157438FFE9BAE5FA414198F918" blockId="2.[189,1401,1445,1910]" pageId="2" pageNumber="3">
|
||
<treatmentCitationGroup id="42BCF93B7438FFE9BAE5FA414198F918" pageId="2" pageNumber="3">
|
||
<taxonomicName id="A5ACA5967438FFE9BAE5FA41412AFA18" ID-CoL="3S4K" authorityName="Westwood" authorityYear="1846" box="[189,293,1445,1471]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077438FFE9BAE5FA41412AFA18" box="[189,293,1445,1471]" italics="true" pageId="2" pageNumber="3">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="E30DF8047438FFE9BB1AFA414228FA18" author="Erichson W. F." box="[322,551,1445,1471]" page="717" pageId="2" pageNumber="3" year="1847">
|
||
<bibRefCitation id="063DA3E47438FFE9BB1AFA414228FA18" author="Erichson W. F." box="[322,551,1445,1471]" pageId="2" pageNumber="3" pagination="67 - 185" refId="ref29306" refString="Erichson W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185." type="journal article" year="1847">Erichson 1847: 717</bibRefCitation>
|
||
</treatmentCitation>
|
||
(taxonomy). —
|
||
<treatmentCitation id="E30DF8047438FFE9B8B2FA4143E9FA67" author="Lacordaire J. T." box="[746,998,1445,1472]" page="135" pageId="2" pageNumber="3" year="1856">
|
||
<bibRefCitation id="063DA3E47438FFE9B8B2FA4143E9FA67" author="Lacordaire J. T." box="[746,998,1445,1472]" pageId="2" pageNumber="3" refId="ref29690" refString="Lacordaire J. T. 1856. Histoire naturelle des Insectes: Genera des Coleopteres ou Expose methodique et critique de tous les Genres proposes jusqu'ici dans cet Ordre d'Insectes. Tome 3. Contenant les Familles des Pectinicornes et Lamellicornes. Librairie encyclopedique de Roret, Paris." type="book" year="1856">Lacordaire 1856: 135</bibRefCitation>
|
||
</treatmentCitation>
|
||
(redescription). —
|
||
<treatmentCitation id="E30DF8047438FFE9BE95FA414108FA43" author="Scudder S. H." page="73" pageId="2" pageNumber="3" year="1882">
|
||
<bibRefCitation id="063DA3E47438FFE9BE95FA414108FA43" author="Scudder S. H." pageId="2" pageNumber="3" refId="ref30822" refString="Scudder S. H. 1882. Nomenclator Zoologicus: an Alphabetical List of All Generic Names that have been Employed by Naturalists for Recent and Fossil Animals from the Earliest Times to the Close of the Year 1879. n. 19. Government Printing Office, Washington DC. https: // doi. org / 10.5962 / bhl. title. 1143" type="book" year="1882">Scudder 1882: 73</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="E30DF8047438FFE9BB90FA2E4281FA43" author="Lucas R." box="[456,654,1481,1508]" page="195" pageId="2" pageNumber="3" year="1920">
|
||
<bibRefCitation id="063DA3E47438FFE9BB90FA2E4281FA43" author="Lucas R." box="[456,654,1481,1508]" pageId="2" pageNumber="3" refId="ref29739" refString="Lucas R. 1920. Catalogus Alphabeticus Generum et Subgenerum Coleopterorum Orbis Terrarum Totius (Famil., Trib., Subtr., Sect. Incl.). Pars I. R. Stricker, Berlin." type="book" year="1920">Lucas 1920: 195</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="E30DF8047438FFE9B917FA2D446DFA43" author="Blackwelder R. E." box="[847,1122,1481,1508]" page="217" pageId="2" pageNumber="3" year="1944">
|
||
<bibRefCitation id="063DA3E47438FFE9B917FA2D446DFA43" author="Blackwelder R. E." box="[847,1122,1481,1508]" pageId="2" pageNumber="3" pagination="2" refId="ref29063" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Parts 1 and 2. Bulletin 185, United States National Museum. United States Government Printing Office, Washington DC." type="journal article" year="1944">Blackwelder 1944: 217</bibRefCitation>
|
||
</treatmentCitation>
|
||
(checklist). —
|
||
<treatmentCitation id="E30DF8047438FFE9BF41FA2D4151F9AF" author="Allsopp P. G." page="106" pageId="2" pageNumber="3" year="1984">
|
||
<bibRefCitation id="063DA3E47438FFE9BF41FA2D4151F9AF" author="Allsopp P. G." pageId="2" pageNumber="3" pagination="105 - 117" refId="ref28631" refString="Allsopp P. G. 1984. Checklist of the Hybosoridae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 38 (2): 105 - 117. Available from https: // www. jstor. org / stable / 4008153 [accessed 23 Nov. 2023]." type="journal article" year="1984">Allsopp 1984: 106</bibRefCitation>
|
||
</treatmentCitation>
|
||
(checklist). —
|
||
<treatmentCitation id="E30DF8047438FFE9B854FA0A42EEF9AF" author="Escobar F." box="[524,737,1518,1544]" page="422" pageId="2" pageNumber="3" year="1997">
|
||
<bibRefCitation id="063DA3E47438FFE9B854FA0A42EEF9AF" author="Escobar F." box="[524,737,1518,1544]" pageId="2" pageNumber="3" pagination="419 - 430" refId="ref29332" refString="Escobar F. 1997. Estudios de la comunidad de coleopteros coprofagos (Scarabaeidae) en un remaente de bosque seco al norte del Tolima, Colombia. Caldasia 19 (3): 419 - 430." type="journal article" year="1997">Escobar 1997: 422</bibRefCitation>
|
||
</treatmentCitation>
|
||
(ecology). —
|
||
<treatmentCitation id="E30DF8047438FFE9B9DDFA0A44EBF9AE" author="Ratcliffe B. C. & Ocampo F. C." box="[901,1252,1518,1545]" page="352" pageId="2" pageNumber="3" year="2001">
|
||
<bibRefCitation id="063DA3E47438FFE9B9DDFA0A44EBF9AE" author="Ratcliffe B. C. & Ocampo F. C." box="[901,1252,1518,1545]" pageId="2" pageNumber="3" pagination="351 - 355" refId="ref30552" refString="Ratcliffe B. C. & Ocampo F. C. 2001. Tyrannosorus rex Ratcliffe and Ocampo, a new genus and species of Miocene hybosorid in amber from the Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae). The Coleopterists Bulletin 55 (3): 351 - 355. https: // doi. org / 10.1649 / 0010 - 065 X (2001) 055 [0351: TRRAOA] 2.0. CO; 2" type="journal article" year="2001">Ratcliffe & Ocampo 2001: 352</bibRefCitation>
|
||
</treatmentCitation>
|
||
(taxonomy). —
|
||
<treatmentCitation id="E30DF8047438FFE9BB48F9F741FBF98A" author="Ocampo F. C." box="[272,500,1554,1581]" page="125" pageId="2" pageNumber="3" year="2002">
|
||
<bibRefCitation id="063DA3E47438FFE9BB48F9F741FBF98A" author="Ocampo F. C." box="[272,500,1554,1581]" pageId="2" pageNumber="3" pagination="316 - 322" refId="ref29848" refString="Ocampo F. C. 2002. Hybosorids of the United States and expanding distribution of the introduced species Hybosorus illigeri (Coleoptera: Scarabaeoidea: Hybosoridae). Annals of the Entomological Society of America 95 (3): 316 - 322. https: // doi. org / 10.1603 / 0013 - 8746 (2002) 095 [0316: HOTUSA] 2.0. CO; 2" type="journal article" year="2002">Ocampo 2002: 125</bibRefCitation>
|
||
</treatmentCitation>
|
||
(taxonomy). —
|
||
<treatmentCitation id="E30DF8047438FFE9B8E4F9F6438AF98A" author="Ratcliffe B. C." box="[700,901,1554,1581]" page="7" pageId="2" pageNumber="3" year="2002">
|
||
<bibRefCitation id="063DA3E47438FFE9B8E4F9F6438AF98A" author="Ratcliffe B. C." box="[700,901,1554,1581]" pageId="2" pageNumber="3" pagination="1 - 48" refId="ref30512" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32 (1): 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 7</bibRefCitation>
|
||
</treatmentCitation>
|
||
(checklist). —
|
||
<treatmentCitation id="E30DF8047438FFE9BE18F9F74119F9F6" author="Ocampo F. C. & Ballerio A." page="191" pageId="2" pageNumber="3" year="2006">
|
||
<bibRefCitation id="063DA3E47438FFE9BE18F9F74119F9F6" author="Ocampo F. C. & Ballerio A." pageId="2" pageNumber="3" pagination="178 - 209" refId="ref30003" refString="Ocampo F. C. & Ballerio A. 2006. Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Hybosoridae). Bulletin of the University of Nebraska State Museum 19: 178 - 209." type="journal article" year="2006">Ocampo & Ballerio 2006: 191</bibRefCitation>
|
||
</treatmentCitation>
|
||
(checklist). —
|
||
<treatmentCitation id="E30DF8047438FFE9BB94F9D34309F9F6" author="Ocampo F. C. & Hawks D. C." box="[460,774,1591,1617]" page="8" pageId="2" pageNumber="3" year="2006">
|
||
<bibRefCitation id="063DA3E47438FFE9BB94F9D34309F9F6" author="Ocampo F. C. & Hawks D. C." box="[460,774,1591,1617]" pageId="2" pageNumber="3" pagination="7 - 12" refId="ref30070" refString="Ocampo F. C. & Hawks D. C. 2006. Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae 2. Molecular phylogenetics and systematic placement of the Hybosoridae. Bulletin of the University of Nebraska State Museum 19: 7 - 12." type="journal article" year="2006">Ocampo & Hawks 2006: 8</bibRefCitation>
|
||
</treatmentCitation>
|
||
(molecular phylogeny). —
|
||
<treatmentCitation id="E30DF8047438FFE9BE14F9D34579F9F6" author="Orozco M. & Perez A." box="[1100,1398,1591,1617]" page="38" pageId="2" pageNumber="3" year="2008">
|
||
<bibRefCitation id="063DA3E47438FFE9BE14F9D34579F9F6" author="Orozco M. & Perez A." box="[1100,1398,1591,1617]" pageId="2" pageNumber="3" pagination="36 - 40" refId="ref30124" refString="Orozco M. & Perez A. 2008. Escarabajos coprofagos (Coleoptera, Scarabaeoidea) del Parque Nacional Los Estoraques (Norte de Santander, Colombia). Revista brasileira de Entomologia 52 (1): 36 - 40. https: // doi. org / 10.1590 / S 0085 - 56262008000100007" type="journal article" year="2008">Orozco & Pérez 2008: 38</bibRefCitation>
|
||
</treatmentCitation>
|
||
(ecology). —
|
||
<treatmentCitation id="E30DF8047438FFE9BBC9F9B84279F9D1" author="Ocampo F. C." box="[401,630,1628,1654]" page="201" pageId="2" pageNumber="3" year="2010">
|
||
<bibRefCitation id="063DA3E47438FFE9BBC9F9B84279F9D1" author="Ocampo F. C." box="[401,630,1628,1654]" pageId="2" pageNumber="3" pagination="201 - 207" refId="ref29965" refString="Ocampo F. C. 2010. A new species of the South American genus Metachaetodus (Coleoptera: Scarabaeoidea: Hybosoridae: Hybosorinae). Revista de la Sociedad entomologica Argentina 69: 201 - 207." type="journal article" year="2010">Ocampo 2010: 201</bibRefCitation>
|
||
</treatmentCitation>
|
||
(taxonomy). —
|
||
<treatmentCitation id="E30DF8047438FFE9B967F9B84442F9D1" author="Otavo S. F. & Parrado-Rosselli A. & Noriega J. A." box="[831,1101,1628,1654]" page="741" pageId="2" pageNumber="3" year="2013">
|
||
<bibRefCitation id="063DA3E47438FFE9B967F9B84442F9D1" author="Otavo S. F. & Parrado-Rosselli A. & Noriega J. A." box="[831,1101,1628,1654]" pageId="2" pageNumber="3" pagination="735 - 752" refId="ref30179" refString="Otavo S. F., Parrado-Rosselli A. & Noriega J. A. 2013. Superfamilia Scarabaeoidea (Insecta: Coleoptera) como elemento bioindicador de perturbacion antropogenica en un parque nacional amazonico. Revista de Biologia tropical 61 (2): 735 - 752." type="journal article" year="2013">
|
||
Otavo
|
||
<emphasis id="50D802077438FFE9B9C9F9B843C1F9D1" box="[913,974,1628,1654]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
|
||
2013: 741
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(ecology). —
|
||
<treatmentCitation id="E30DF8047438FFE9BEA5F9B841AFF93D" author="Rodrigues M. M. & Uchoa M. A. & Ide S." page="213" pageId="2" pageNumber="3" year="2013">
|
||
<bibRefCitation id="063DA3E47438FFE9BEA5F9B841AFF93D" author="Rodrigues M. M. & Uchoa M. A. & Ide S." pageId="2" pageNumber="3" pagination="211 - 220" refId="ref30683" refString="Rodrigues M. M., Uchoa M. A. & Ide S. 2013. Dung beetles (Coleoptera: Scarabaeoidea) in three landscapes in Mato Grosso do Sul, Brazil. Brazilian Journal of Biology 73 (1): 211 - 220. https: // doi. org / 10.1590 / S 1519 - 69842013000100023" type="journal article" year="2013">
|
||
Rodrigues
|
||
<emphasis id="50D802077438FFE9BAB4F965412BF93D" box="[236,292,1664,1690]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
|
||
2013: 213
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(ecology). —
|
||
<treatmentCitation id="E30DF8047438FFE9B811F96443DEF93D" author="Ballerio A. & Grebennikov V. V." box="[585,977,1664,1690]" page="49" pageId="2" pageNumber="3" year="2016">
|
||
<bibRefCitation id="063DA3E47438FFE9B811F96443DEF93D" author="Ballerio A. & Grebennikov V. V." box="[585,977,1664,1690]" pageId="2" pageNumber="3" pagination="23 - 52" refId="ref28890" refString="Ballerio A. & Grebennikov V. V. 2016. Rolling into a ball: phylogeny of the Ceratocanthinae (Coleoptera: Hybosoridae) inferred from adult morphology and origin of a unique body enrollment coaptation in terrestrial arthropods. Arthropod Systematics & Phylogeny 74 (1): 23 - 52." type="journal article" year="2016">Ballerio & Grebennikov 2016: 49</bibRefCitation>
|
||
</treatmentCitation>
|
||
(phylogeny). —
|
||
<treatmentCitation id="E30DF8047438FFE9BEC2F96440F6F967" author="Basilio D. S. & Cherman M. A. & Vaz-de-Mello F. Z. & Almeida L. M." httpUri="http://treatment.plazi.org/id/EA056F037438FFECB80FFB1F4578FABE" page="5" pageId="2" pageNumber="3" year="2023">
|
||
<bibRefCitation id="063DA3E47438FFE9BEC2F96440F6F967" author="Basilio D. S. & Cherman M. A. & Vaz-de-Mello F. Z. & Almeida L. M." pageId="2" pageNumber="3" pagination="1156 - 1170" refId="ref28944" refString="Basilio D. S., Cherman M. A., Vaz-de-Mello F. Z. & Almeida L. M. 2023. Phylogenetic relationships in Hybosoridae (Coleoptera: Scarabaeoidea). Zoological Journal of the Linnean Society 198 (4): 1156 - 1170. https: // doi. org / 10.1093 / zoolinnean / zlac 095" type="journal article" year="2023">
|
||
Basílio
|
||
<emphasis id="50D802077438FFE9BEAAF9654523F93D" box="[1266,1324,1664,1690]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
|
||
2023: 5
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(phylogeny).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="6213DE157438FFE9BAE5F92D42E8F8F6" blockId="2.[189,1401,1445,1910]" pageId="2" pageNumber="3">
|
||
<treatmentCitationGroup id="42BCF93B7438FFE9BAE5F92D42E8F8F6" pageId="2" pageNumber="3">
|
||
<taxonomicName id="A5ACA5967438FFE9BAE5F92D4123F944" ID-CoL="3S4K" authority="Westwood, 1846: 163" authorityName="Westwood" authorityPageNumber="163" authorityYear="1846" box="[189,300,1737,1763]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077438FFE9BAE5F92D4123F944" box="[189,300,1737,1763]" italics="true" pageId="2" pageNumber="3">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="E30DF8047438FFE9BB15F92D4236F943" author="Erichson W. F." box="[333,569,1737,1764]" page="717" pageId="2" pageNumber="3" year="1847">
|
||
<bibRefCitation id="063DA3E47438FFE9BB15F92D4236F943" author="Erichson W. F." box="[333,569,1737,1764]" pageId="2" pageNumber="3" pagination="67 - 185" refId="ref29306" refString="Erichson W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185." type="journal article" year="1847">Erichson 1847: 717</bibRefCitation>
|
||
</treatmentCitation>
|
||
(incorrect subsequent spelling: taxonomy). —
|
||
<treatmentCitation id="E30DF8047438FFE9BE2BF92D4579F943" author="Lacordaire J. T." box="[1139,1398,1737,1764]" page="135" pageId="2" pageNumber="3" year="1856">
|
||
<bibRefCitation id="063DA3E47438FFE9BE2BF92D4579F943" author="Lacordaire J. T." box="[1139,1398,1737,1764]" pageId="2" pageNumber="3" refId="ref29690" refString="Lacordaire J. T. 1856. Histoire naturelle des Insectes: Genera des Coleopteres ou Expose methodique et critique de tous les Genres proposes jusqu'ici dans cet Ordre d'Insectes. Tome 3. Contenant les Familles des Pectinicornes et Lamellicornes. Librairie encyclopedique de Roret, Paris." type="book" year="1856">Lacordaire 1856: 135</bibRefCitation>
|
||
</treatmentCitation>
|
||
(redescription). —
|
||
<treatmentCitation id="E30DF8047438FFE9BB97F90A436BF8AE" author="Gemminger M. & Harold E." box="[463,868,1774,1801]" page="1075" pageId="2" pageNumber="3" year="1869">
|
||
<bibRefCitation id="063DA3E47438FFE9BB97F90A436BF8AE" author="Gemminger M. & Harold E." box="[463,868,1774,1801]" pageId="2" pageNumber="3" refId="ref29416" refString="Gemminger M. & Harold E. 1869. Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus, Tom. IV: Scarabaeidae. E. H. Gumm, Munich." type="book" year="1869">Gemminger & Harold 1869: 1075</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="E30DF8047438FFE9BE7DF90A44ECF8AF" author="Arrow G. J." box="[1061,1251,1774,1800]" page="37" pageId="2" pageNumber="3" year="1912">
|
||
<bibRefCitation id="063DA3E47438FFE9BE7DF90A44ECF8AF" author="Arrow G. J." box="[1061,1251,1774,1800]" pageId="2" pageNumber="3" pagination="1 - 65" refId="ref28783" refString="Arrow G. J. 1912. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Coleopterorum Catalogus 43: 1 - 65." type="journal article" year="1912">Arrow 1912: 37</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="E30DF8047438FFE9BB48F8F641ECF88A" author="Paulian R." box="[272,483,1810,1837]" page="230" pageId="2" pageNumber="3" year="1938">
|
||
<bibRefCitation id="063DA3E47438FFE9BB48F8F641ECF88A" author="Paulian R." box="[272,483,1810,1837]" pageId="2" pageNumber="3" pagination="213 - 296" refId="ref30228" refString="Paulian R. 1938. Contribution a l'etude des canthonides americains. Annales de la Societe entomologique de France 107: 213 - 296." type="journal article" year="1938">Paulian 1938: 230</bibRefCitation>
|
||
</treatmentCitation>
|
||
(key). —
|
||
<treatmentCitation id="E30DF8047438FFE9B801F8F64397F88A" author="Howden H. F. & Gill B. D." box="[601,920,1810,1837]" page="2074" pageId="2" pageNumber="3" year="1987">
|
||
<bibRefCitation id="063DA3E47438FFE9B801F8F64397F88A" author="Howden H. F. & Gill B. D." box="[601,920,1810,1837]" pageId="2" pageNumber="3" pagination="201 - 224" refId="ref29567" refString="Howden H. F. & Gill B. D. 1987. New species and new records of Panamanian and Costa Rican Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 41 (3): 201 - 224. Available from https: // www. jstor. org / stable / 4008409 [accessed 23 Nov. 2023]." type="journal article" year="1987">Howden & Gill 1987: 2074</bibRefCitation>
|
||
</treatmentCitation>
|
||
(taxonomy). —
|
||
<treatmentCitation id="E30DF8047438FFE9BE01F8F744F6F88A" author="Feer F." box="[1113,1273,1811,1837]" page="33" pageId="2" pageNumber="3" year="2000">
|
||
<bibRefCitation id="063DA3E47438FFE9BE01F8F744F6F88A" author="Feer F." box="[1113,1273,1811,1837]" pageId="2" pageNumber="3" pagination="29 - 43" refId="ref29370" refString="Feer F. 2000. Les coleopteres coprophages et necrophages (Scarabaeidae s. str. et Aphodiidae) de la foret de Guyane francaise: composition specifique et structure des peuplements. Annales de la Societe entomologique de France 36: 29 - 43." type="journal article" year="2000">Feer 2000: 33</bibRefCitation>
|
||
</treatmentCitation>
|
||
(ecology). —
|
||
<treatmentCitation id="E30DF8047438FFE9BB56F8D34268F8F6" author="Uchoa M. A. & Rodrigues M. M." box="[270,615,1847,1873]" page="21" pageId="2" pageNumber="3" year="2019">
|
||
<bibRefCitation id="063DA3E47438FFE9BB56F8D34268F8F6" author="Uchoa M. A. & Rodrigues M. M." box="[270,615,1847,1873]" pageId="2" pageNumber="3" pagination="19 - 26" refId="ref30924" refString="Uchoa M. A. & Rodrigues M. M. 2019. Dung beetles (Coleoptera: Scarabaeoidea) population patterns in three environments in the Midwest of Brazil. EntomoBrasilis 12 (1): 19 - 26. https: // doi. org / 10.12741 / ebrasilis. v 12 i 1.825" type="journal article" year="2019">Uchoa & Rodrigues 2019: 21</bibRefCitation>
|
||
</treatmentCitation>
|
||
(ecology).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="6213DE157438FFE9BAE5F8B844D2F8D1" blockId="2.[189,1401,1445,1910]" box="[189,1245,1884,1910]" pageId="2" pageNumber="3">
|
||
<treatmentCitationGroup id="42BCF93B7438FFE9BAE5F8B844D2F8D1" box="[189,1245,1884,1910]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="A5ACA5967438FFE9BAE5F8B84226F8D1" ID-CoL="4PMS" authority="Prokofiev 2013" authorityName="Prokofiev" authorityYear="2013" box="[189,553,1884,1910]" class="Insecta" family="Hybosoridae" genus="Gnombolbus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077438FFE9BAE5F8B8415CF8D1" box="[189,339,1884,1910]" italics="true" pageId="2" pageNumber="3">Gnombolbus</emphasis>
|
||
–
|
||
<treatmentCitation id="E30DF8047438FFE9BB28F8B8425FF8D1" author="Prokofiev A. M." box="[368,592,1884,1910]" page="1" pageId="2" pageNumber="3" year="2013">
|
||
<bibRefCitation id="063DA3E47438FFE9BB28F8B8425FF8D1" author="Prokofiev A. M." box="[368,592,1884,1910]" pageId="2" pageNumber="3" pagination="1 - 2" refId="ref30456" refString="Prokofiev A. M. 2013 a. A new bolboceratine beetle from Mesoamerica (Coleoptera: Geotrupidae: Bolboceratinae). Calodema 244: 1 - 2." type="journal article" year="2013">Prokofiev 2013a: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(original description);
|
||
<treatmentCitation id="E30DF8047438FFE9B907F8B8444FF8D1" author="Prokofiev A. M." box="[863,1088,1884,1910]" page="1" pageId="2" pageNumber="3" year="2013">
|
||
<bibRefCitation id="063DA3E47438FFE9B907F8B8444FF8D1" author="Prokofiev A. M." box="[863,1088,1884,1910]" pageId="2" pageNumber="3" pagination="1" refId="ref30485" refString="Prokofiev A. M. 2013 b. Gnombolbus, a synonym of Coilodes (Coleoptera: Scarabaeoidea: Hybosoridae). Calodema 252: 1." type="journal article" year="2013">Prokofiev 2013b: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
(synonymy).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7438FFE9BAE5F87B4157F81F" box="[189,344,1950,1977]" pageId="2" pageNumber="3" type="type_taxon">
|
||
<paragraph id="6213DE157438FFE9BAE5F87B4157F81F" blockId="2.[189,344,1950,1977]" box="[189,344,1950,1977]" pageId="2" pageNumber="3">
|
||
<heading id="395B69797438FFE9BAE5F87B4157F81F" bold="true" box="[189,344,1950,1977]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="50D802077438FFE9BAE5F87B4157F81F" bold="true" box="[189,344,1950,1977]" pageId="2" pageNumber="3">
|
||
<typeStatus id="BD1760B77438FFE9BAE5F87B40F5F81E" box="[189,250,1951,1977]" pageId="2" pageNumber="3">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7438FFE9BAE5F82A4513F84F" box="[189,1308,1998,2025]" pageId="2" pageNumber="3" type="reference_group">
|
||
<paragraph id="6213DE157438FFE9BAE5F82A4513F84F" blockId="2.[189,1308,1998,2025]" box="[189,1308,1998,2025]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="A5ACA5967438FFE9BAE5F82A4210F84F" ID-CoL="3MYSL" authority="Perty, 1830" authorityName="Perty" authorityYear="1830" box="[189,543,1998,2025]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gibbus">
|
||
<emphasis id="50D802077438FFE9BAE5F82A419EF84F" box="[189,401,1998,2024]" italics="true" pageId="2" pageNumber="3">Hybosorus gibbus</emphasis>
|
||
<bibRefCitation id="063DA3E47438FFE9BBC1F82B4210F84F" author="Perty J. A. M." box="[409,543,1998,2025]" pageId="2" pageNumber="3" refId="ref30331" refString="Perty J. A. M. 1830. Delectus Animalium Articulatorum quae in Itinere per Brasiliam Annis MDCCCXVII- MDCCCXX Jussu et Auspiciis Maximiliani Josephi I Bavariae Regis Augustissimi Peracto. Munich. https: // doi. org / 10.5962 / bhl. title. 158694" type="book" year="1830">Perty, 1830</bibRefCitation>
|
||
</taxonomicName>
|
||
(junior synonym of
|
||
<taxonomicName id="A5ACA5967438FFE9B948F82A43FBF84F" ID-CoL="WWJ2" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[784,1012,1998,2024]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D802077438FFE9B948F82A43FBF84F" box="[784,1012,1998,2024]" italics="true" pageId="2" pageNumber="3">Coilodes humeralis</emphasis>
|
||
</taxonomicName>
|
||
) by original designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7439FFE8BAE5FEEC42BCFE47" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="6213DE157439FFE8BAE5FEEC4139FE85" blockId="3.[189,310,264,290]" box="[189,310,264,290]" pageId="3" pageNumber="4">
|
||
<heading id="395B69797439FFE8BAE5FEEC4139FE85" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="50D802077439FFE8BAE5FEEC4139FE85" bold="true" box="[189,310,264,290]" pageId="3" pageNumber="4">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5FED342BCFE47" blockId="3.[188,1399,311,480]" pageId="3" pageNumber="4">
|
||
<taxonomicName id="A5ACA5967439FFE8BAE5FED3412AFEF6" authorityName="Westwood" authorityYear="1846" box="[189,293,311,337]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077439FFE8BAE5FED3412AFEF6" box="[189,293,311,337]" italics="true" pageId="3" pageNumber="4">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from other Neotropical
|
||
<taxonomicName id="A5ACA5967439FFE8B8A5FED3439CFEF6" authorityName="Erichson" authorityYear="1847" box="[765,915,311,337]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Hybosorinae">Hybosorinae</taxonomicName>
|
||
in the body shape being strongly convex, the elytra having sparse twin lines composed of punctures and the mandible being dorsally excavated. In addition, this genus is distinguished from the other
|
||
<taxonomicName id="A5ACA5967439FFE8B941FE9A43A0FE3F" authorityName="Erichson" authorityYear="1847" box="[793,943,382,408]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
by the presence of sexual dimorphism both in the pronotum (convex in females and excavated in males) and in the tarsal claws (simple in females and with a middle tooth in males).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7439FFE8BAE5FDEC43DEFD3F" pageId="3" pageNumber="4" type="etymology">
|
||
<paragraph id="6213DE157439FFE8BAE5FDEC414BFD85" blockId="3.[189,324,520,546]" box="[189,324,520,546]" pageId="3" pageNumber="4">
|
||
<heading id="395B69797439FFE8BAE5FDEC414BFD85" bold="true" box="[189,324,520,546]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="50D802077439FFE8BAE5FDEC414BFD85" bold="true" box="[189,324,520,546]" pageId="3" pageNumber="4">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5FDD243DEFD3F" blockId="3.[189,1398,566,664]" pageId="3" pageNumber="4">
|
||
The name “
|
||
<taxonomicName id="A5ACA5967439FFE8BB12FDD241BCFDF7" authorityName="Westwood" authorityYear="1846" box="[330,435,566,592]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D802077439FFE8BB12FDD241BCFDF7" box="[330,435,566,592]" italics="true" pageId="3" pageNumber="4">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
” is masculine in gender, as it was treated in the original description (
|
||
<bibRefCitation id="063DA3E47439FFE8BEA0FDD240F1FDD3" author="Westwood J. O." pageId="3" pageNumber="4" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood 1846</bibRefCitation>
|
||
). This term comes from the Greek κοιλότης (koîlotes) derived from κοίλος (koîlos). Although it was stated by
|
||
<bibRefCitation id="063DA3E47439FFE8BB3AFD99423DFD30" author="Westwood J. O." box="[354,562,637,663]" pageId="3" pageNumber="4" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood (1846)</bibRefCitation>
|
||
as “convexitas”, it means concave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E7439FFEEBAE5FD5B43A6F8A8" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="6213DE157439FFE8BAE5FD5B4164FD7E" blockId="3.[189,363,703,729]" box="[189,363,703,729]" pageId="3" pageNumber="4">
|
||
<heading id="395B69797439FFE8BAE5FD5B4164FD7E" bold="true" box="[189,363,703,729]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="50D802077439FFE8BAE5FD5B4164FD7E" bold="true" box="[189,363,703,729]" pageId="3" pageNumber="4">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAB2FD0A4125FCAF" blockId="3.[189,1215,750,812]" box="[234,298,750,776]" pageId="3" pageNumber="4">
|
||
<emphasis id="50D802077439FFE8BAB2FD0A4125FCAF" bold="true" box="[234,298,750,776]" pageId="3" pageNumber="4">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5FCF644B0FC8B" blockId="3.[189,1215,750,812]" box="[189,1215,785,812]" pageId="3" pageNumber="4">
|
||
MEASUREMENTS. Length
|
||
<quantity id="A55473F07439FFE8BB86FCF64261FC8B" box="[478,622,786,812]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="8.7" metricValueMin="4.4" pageId="3" pageNumber="4" unit="mm" value="6.55" valueMax="8.7" valueMin="4.4">4.4–8.7 mm</quantity>
|
||
. Width
|
||
<quantity id="A55473F07439FFE8B890FCF54356FC8B" box="[712,857,785,812]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="5.1" metricValueMin="2.6" pageId="3" pageNumber="4" unit="mm" value="3.8499999999999996" valueMax="5.1" valueMin="2.6">2.6–5.1 mm</quantity>
|
||
. Body convex, oval and shiny.
|
||
</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5FCB04424FAF8" blockId="3.[187,1400,851,1376]" pageId="3" pageNumber="4">HEAD. Surface smooth or strigulate. Frons, in dorsal view, subrectangular; two small tubercles in the middle or one long tubercle formed by the fusion of the small ones; proximal border rounded; lateral margin variable. Canthus strong; distinct area expanded downwards with erect bristles; groove inconspicuous, medially extending up to the lateral margin of the frons. Clypeus subtrapezoidal or subrectangular; lateral margins weakly rounded; apex usually straight; thin and sparse setae throughout the anterior margin dorsally; frontoclypeal suture absent. Labrum semicircular; sparse setae spread along the entire dorsal margin; surface without punctures. Mandibles dorsally excavated, protruding beyond the apex of labrum; lateral margin smooth; setae on the basal half; one tooth at the inferior area of the mandibular apex. Labium with mentum subquadrate or subrectangular; labial palps with four palpomeres; basal palpomere twice as wide as long; second and third palpomeres subglobose; distal palpomere barrel-shaped, length equal to the second and third palpomeres combined. Maxilla subtriangular; long setae throughout surface; maxillary palp with four palpomeres; basal palpomere curved; second and third palpomeres longer than wide; distal palpomere barrel-shaped, length equivalent to the sum of the previous three. Antenna with 10 antennomeres; scape with long and erect setae; antennal club with three antennomeres; club with basal antennomere cupuliform, pubescent distally, sparse, short setae on its base.</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5FA6342B1F9AB" blockId="3.[189,1400,1415,1548]" pageId="3" pageNumber="4">PRONOTUM. Convex; excavated medially; usually subtrapezoidal, posterior margin wider than anterior margin or equal in size; medial length usually longer than the anterior and posterior margins; anterior margin with subacute angles; posterior margin rounded and weakly produced medially; lateral margin convex, setae and punctures usually absent.</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5F9D0454BF9E9" blockId="3.[189,1348,1587,1614]" box="[189,1348,1587,1614]" pageId="3" pageNumber="4">SCUTELLAR SHIELD. Subtriangular, twice as long as wide; mostly without punctures; apex punctate.</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5F99242C4F971" blockId="3.[189,1399,1653,1751]" pageId="3" pageNumber="4">ELYTRA. Double row of longitudinal punctures, sutural stria usually complete; elytral disc glabrous; external margins with short setae; elytral epipleuron complete, wider at the apex; posterior margin of elytra entirely covering tergite 8 (pygidium).</paragraph>
|
||
<paragraph id="6213DE157439FFE8BAE5F91B447DF89B" blockId="3.[187,1398,1790,1852]" pageId="3" pageNumber="4">VENTER. Hypomeral surface strigulate, setose; prosternum with midlength elevated, transversally grooved; anterior margin variable in shape, mesoventrite and metaventrite usually smooth.</paragraph>
|
||
<paragraph id="6213DE157439FFEFBAE5F8804485FE14" blockId="3.[187,1399,1892,2024]" lastBlockId="4.[188,1400,265,436]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">LEGS. Procoxa conical; surface strigulate. Protrochanter joint with the procoxa rounded, distally angulate. Profemur with posterior margin carinate; surface smooth, with sparse setae. Protibia with inner margin convex; surface with long setae; carinate; single spur; external margin with three larger teeth and series of smaller denticles along the entire margin. Protarsi with tarsal insertion beneath the protibial second tooth. Mesothoracic and metathoracic legs smooth. Mesotrochanter and metatrochanter subtriangular with bifurcated apex. Mesofemur and metafemur carinate on posterior margin. Mesotibia and metatibia with erect setae; apex weakly expanded; pair of spurs with sharp apex, inner spur as long as the protibial spur; outer spur ⅓ longer than the inner one. Tarsi with five tarsomeres, each tarsomere twice as long as wide, distal tarsomere twice as wide as the previous; tarsal claws toothed medially.</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FE3842D2FE51" blockId="4.[189,733,476,502]" box="[189,733,476,502]" pageId="4" pageNumber="5">ABDOMEN. Six ventrites with recumbent setae.</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FDFA4422FD9F" blockId="4.[189,1069,542,569]" box="[189,1069,542,569]" pageId="4" pageNumber="5">GENITALIA. Parameres asymmetrical; tegmen with or without a dorsal lobe.</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAB2FD84414AFDDD" blockId="4.[189,1399,608,706]" box="[234,325,608,634]" pageId="4" pageNumber="5">
|
||
<emphasis id="50D80207743EFFEFBAB2FD84414AFDDD" bold="true" box="[234,325,608,634]" pageId="4" pageNumber="5">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FD604150FD65" blockId="4.[189,1399,608,706]" pageId="4" pageNumber="5">
|
||
Length
|
||
<quantity id="A55473F0743EFFEFBB41FD6041A2FD38" box="[281,429,644,671]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" metricValueMax="8.3" metricValueMin="5.1" pageId="4" pageNumber="5" unit="mm" value="6.7" valueMax="8.3" valueMin="5.1">5.1–8.3 mm</quantity>
|
||
. Width 3.0–
|
||
<quantity id="A55473F0743EFFEFB865FD614292FD38" box="[573,669,645,671]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="5" unit="mm" value="4.7">4.7 mm</quantity>
|
||
. As the male except by pronotal excavation absent and tarsal claws simple.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FD0F4309FCA2" blockId="4.[189,774,746,773]" box="[189,774,746,773]" pageId="4" pageNumber="5">
|
||
<heading id="395B6979743EFFEFBAE5FD0F4309FCA2" bold="true" box="[189,774,746,773]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="50D80207743EFFEFBAE5FD0F4309FCA2" bold="true" box="[189,774,746,773]" pageId="4" pageNumber="5">
|
||
Taxonomic history of
|
||
<taxonomicName id="A5ACA596743EFFEFBB95FD0E4309FCA2" authority="Westwood, 1846" authorityName="Westwood" authorityYear="1846" box="[461,774,746,773]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBB95FD0E423AFCA2" bold="true" box="[461,565,746,773]" italics="true" pageId="4" pageNumber="5">Coilodes</emphasis>
|
||
<bibRefCitation id="063DA3E4743EFFEFB864FD0F4309FCA2" author="Westwood J. O." box="[572,774,747,773]" pageId="4" pageNumber="5" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FCFE4313FC06" blockId="4.[189,1400,794,929]" pageId="4" pageNumber="5">
|
||
<bibRefCitation id="063DA3E4743EFFEFBAE5FCFE41A0FC93" author="Mannerheim K. G." box="[189,431,794,820]" pageId="4" pageNumber="5" pagination="29 - 80" refId="ref29813" refString="Mannerheim K. G. 1829. Description de quarante nouvelles especes de scarabeides du Bresil: avec figures. Memoires de la Societe imperiale des Naturalistes de Moscou 7: 29 - 80." type="journal article" year="1829">Mannerheim (1829)</bibRefCitation>
|
||
described two
|
||
<taxonomicName id="A5ACA596743EFFEFB832FCFE42E4FC93" authorityName="MacLeay" authorityYear="1819" box="[618,747,794,820]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB832FCFE42E4FC93" box="[618,747,794,820]" italics="true" pageId="4" pageNumber="5">Hybosorus</emphasis>
|
||
</taxonomicName>
|
||
species:
|
||
<taxonomicName id="A5ACA596743EFFEFB903FCFF43F7FC93" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[859,1016,794,820]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743EFFEFB903FCFF43F7FC93" box="[859,1016,794,820]" italics="true" pageId="4" pageNumber="5">H. humeralis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743EFFEFBE6FFCFF4493FC93" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1079,1180,795,820]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="50D80207743EFFEFBE6FFCFF4493FC93" box="[1079,1180,795,820]" italics="true" pageId="4" pageNumber="5">H. niger</emphasis>
|
||
</taxonomicName>
|
||
in “Decription de quarante nouvelles espèces de Scarabéides du
|
||
<collectingCountry id="1ABB9E85743EFFEFB8B2FCDA433EFCFF" box="[746,817,830,856]" name="Brazil" pageId="4" pageNumber="5">Brésil</collectingCountry>
|
||
”, published in
|
||
<emphasis id="50D80207743EFFEFB9B0FCDB420DFCDB" italics="true" pageId="4" pageNumber="5">Mémoires de la Société impériale des Naturalistes de Moscou</emphasis>
|
||
. According to the author, even though the generic attribution was doubtful, he decided to include them in the genus
|
||
<taxonomicName id="A5ACA596743EFFEFB8CDFC63431BFC06" authorityName="MacLeay" authorityYear="1819" box="[661,788,903,929]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB8CDFC63431BFC06" box="[661,788,903,929]" italics="true" pageId="4" pageNumber="5">Hybosorus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FC2E42BEFB8C" blockId="4.[187,1398,969,1068]" pageId="4" pageNumber="5">
|
||
<bibRefCitation id="063DA3E4743EFFEFBAE5FC2E4142FC44" author="Perty J. A. M." box="[189,333,969,996]" pageId="4" pageNumber="5" refId="ref30331" refString="Perty J. A. M. 1830. Delectus Animalium Articulatorum quae in Itinere per Brasiliam Annis MDCCCXVII- MDCCCXX Jussu et Auspiciis Maximiliani Josephi I Bavariae Regis Augustissimi Peracto. Munich. https: // doi. org / 10.5962 / bhl. title. 158694" type="book" year="1830">Perty (1830)</bibRefCitation>
|
||
described
|
||
<emphasis id="50D80207743EFFEFBB9FFC2D42AFFC44" box="[455,672,969,995]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="A5ACA596743EFFEFBB9FFC2D4294FC44" authorityName="Perty" authorityYear="1830" box="[455,667,969,995]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gibbus">Hybosorus gibbus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
a species from
|
||
<collectingCountry id="1ABB9E85743EFFEFB90AFC2D4393FC44" box="[850,924,969,995]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
|
||
, but he also questioned its position in this genus. In 1840, Laporte (alias Castelnau) succinctly described four more species of
|
||
<taxonomicName id="A5ACA596743EFFEFBEDDFC09450BFBA0" authorityName="MacLeay" authorityYear="1819" box="[1157,1284,1005,1031]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBEDDFC09450BFBA0" box="[1157,1284,1005,1031]" italics="true" pageId="4" pageNumber="5">Hybosorus</emphasis>
|
||
</taxonomicName>
|
||
, amongst them
|
||
<taxonomicName id="A5ACA596743EFFEFBAA7FBF641A5FB8C" authorityName="Castelnau" authorityYear="1840" box="[255,426,1041,1067]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="50D80207743EFFEFBAA7FBF641A5FB8C" box="[255,426,1041,1067]" italics="true" pageId="4" pageNumber="5">H. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
, this also from
|
||
<collectingCountry id="1ABB9E85743EFFEFB839FBF542A3FB8C" box="[609,684,1041,1067]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5FBB743BCF9E4" blockId="4.[189,1400,1107,1603]" pageId="4" pageNumber="5">
|
||
The name
|
||
<taxonomicName id="A5ACA596743EFFEFBB64FBB741ABFBCA" authorityName="Westwood" authorityYear="1846" box="[316,420,1107,1133]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBB64FBB741ABFBCA" box="[316,420,1107,1133]" italics="true" pageId="4" pageNumber="5">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
appeared for the first time when
|
||
<bibRefCitation id="063DA3E4743EFFEFB976FBB743F0FBCA" author="Westwood J. O." box="[814,1023,1107,1133]" pageId="4" pageNumber="5" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood (1846)</bibRefCitation>
|
||
merged a small group of South American species previously included in
|
||
<taxonomicName id="A5ACA596743EFFEFB8C8FB9C4302FB35" authorityName="MacLeay" authorityYear="1819" box="[656,781,1144,1170]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB8C8FB9C4302FB35" box="[656,781,1144,1170]" italics="true" pageId="4" pageNumber="5">Hybosorus</emphasis>
|
||
</taxonomicName>
|
||
. According to this author, these two genera differ from each other, besides in their distribution, by the mandible shape and the presence of sexual dimorphism. The dimorphism involves the excavated pronotum and the tarsal claws with one medial tooth in males, and convex pronotum and simple tarsal claws in females.
|
||
<bibRefCitation id="063DA3E4743EFFEFB907FB004421FB59" author="Westwood J. O." box="[863,1070,1252,1278]" pageId="4" pageNumber="5" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood (1846)</bibRefCitation>
|
||
designated
|
||
<taxonomicName id="A5ACA596743EFFEFBEE1FB004579FB59" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[1209,1398,1252,1278]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gibbus">
|
||
<emphasis id="50D80207743EFFEFBEE1FB004579FB59" box="[1209,1398,1252,1278]" italics="true" pageId="4" pageNumber="5">Coilodes gibbus</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="BD1760B7743EFFEFBB50FAED4134FA84" box="[264,315,1289,1315]" pageId="4" pageNumber="5">type</typeStatus>
|
||
species, accompanied by drawings of mouthparts, protibia and the male tarsal claw. He also described three new species:
|
||
<taxonomicName id="A5ACA596743EFFEFB847FAC942A4FAE1" authorityName="Westwood" authorityYear="1846" box="[543,683,1324,1351]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="50D80207743EFFEFB847FAC942A4FAE1" box="[543,683,1324,1351]" italics="true" pageId="4" pageNumber="5">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="1ABB9E85743EFFEFB8E2FAC84307FAE1" box="[698,776,1324,1350]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
|
||
),
|
||
<taxonomicName id="A5ACA596743EFFEFB978FAC943B6FAE1" authorityName="Westwood" authorityYear="1846" box="[800,953,1325,1351]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="50D80207743EFFEFB978FAC943B6FAE1" box="[800,953,1325,1351]" italics="true" pageId="4" pageNumber="5">C. castaneus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="1ABB9E85743EFFEFB992FAC8444CFAE1" box="[970,1091,1324,1350]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
|
||
) and
|
||
<taxonomicName id="A5ACA596743EFFEFBED0FAC9451BFAE1" authorityName="Westwood" authorityYear="1846" box="[1160,1300,1324,1351]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="50D80207743EFFEFBED0FAC9451BFAE1" box="[1160,1300,1324,1351]" italics="true" pageId="4" pageNumber="5">C. chilensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="1ABB9E85743EFFEFBF7CFAC84565FAE1" box="[1316,1386,1324,1350]" name="Chile" pageId="4" pageNumber="5">Chile</collectingCountry>
|
||
). Westwood also synonymized
|
||
<taxonomicName id="A5ACA596743EFFEFB847FAB44321FACD" authorityName="Castelnau" authorityYear="1840" box="[543,814,1360,1386]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="50D80207743EFFEFB847FAB44321FACD" box="[543,814,1360,1386]" italics="true" pageId="4" pageNumber="5">Hybosorus brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743EFFEFB932FAB54407FACD" box="[874,1032,1361,1386]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="geminatus">
|
||
<emphasis id="50D80207743EFFEFB932FAB54407FACD" box="[874,1032,1361,1386]" italics="true" pageId="4" pageNumber="5">H. geminatus</emphasis>
|
||
</taxonomicName>
|
||
(the latter is a nomem nudum; this name was cited in Dejean’s catalogues, in 1833 and 1837, but without a description) with
|
||
<taxonomicName id="A5ACA596743EFFEFBEA7FA914561FA29" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[1279,1390,1396,1423]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gibbus">
|
||
<emphasis id="50D80207743EFFEFBEA7FA914561FA29" box="[1279,1390,1396,1423]" italics="true" pageId="4" pageNumber="5">C. gibbus</emphasis>
|
||
</taxonomicName>
|
||
. Based on the species distribution, he suggested that many other
|
||
<taxonomicName id="A5ACA596743EFFEFB996FA7C445FFA15" authorityName="MacLeay" authorityYear="1819" box="[974,1104,1432,1458]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB996FA7C445FFA15" box="[974,1104,1432,1458]" italics="true" pageId="4" pageNumber="5">Hybosorus</emphasis>
|
||
</taxonomicName>
|
||
from Central and South America should be transferred to
|
||
<emphasis id="50D80207743EFFEFB80FFA5842C7FA71" box="[599,712,1468,1494]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="A5ACA596743EFFEFB80FFA5842CAFA71" authorityName="Westwood" authorityYear="1846" box="[599,709,1468,1494]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Coilodes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The seven species transferred to
|
||
<taxonomicName id="A5ACA596743EFFEFBE38FA5844C5FA71" authorityName="Westwood" authorityYear="1846" box="[1120,1226,1468,1494]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBE38FA5844C5FA71" box="[1120,1226,1468,1494]" italics="true" pageId="4" pageNumber="5">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
were: the two described by
|
||
<bibRefCitation id="063DA3E4743EFFEFBB01FA054248FA5C" author="Mannerheim K. G." box="[345,583,1505,1531]" pageId="4" pageNumber="5" pagination="29 - 80" refId="ref29813" refString="Mannerheim K. G. 1829. Description de quarante nouvelles especes de scarabeides du Bresil: avec figures. Memoires de la Societe imperiale des Naturalistes de Moscou 7: 29 - 80." type="journal article" year="1829">Mannerheim (1829)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="A5ACA596743EFFEFB808FA0542E7FA5C" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[592,744,1505,1531]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743EFFEFB808FA0542E7FA5C" box="[592,744,1505,1531]" italics="true" pageId="4" pageNumber="5">H. humeralis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743EFFEFB978FA05438EFA5D" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[800,897,1505,1530]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="50D80207743EFFEFB978FA05438EFA5D" box="[800,897,1505,1530]" italics="true" pageId="4" pageNumber="5">H. niger</emphasis>
|
||
</taxonomicName>
|
||
(misspelled by Westwood as
|
||
<taxonomicName id="A5ACA596743EFFEFBE81FA05454FFA5D" authorityName="Westwood" authorityYear="1845" box="[1241,1344,1505,1530]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="auger">
|
||
<emphasis id="50D80207743EFFEFBE81FA05454FFA5D" box="[1241,1344,1505,1530]" italics="true" pageId="4" pageNumber="5">H. auger</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<taxonomicName id="A5ACA596743EFFEFBF02FA0541E2F9B8" authority="(Castelnau, 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rufulus">
|
||
<emphasis id="50D80207743EFFEFBF02FA05411FF9B8" italics="true" pageId="4" pageNumber="5">H. rufulus</emphasis>
|
||
(
|
||
<bibRefCitation id="063DA3E4743EFFEFBB78F9E141EBF9B8" author="Castelnau & Laporte F. D." box="[288,484,1541,1567]" pageId="4" pageNumber="5" refId="ref29111" refString="Castelnau [Laporte F. D.] 1840. Histoire naturelle des Animaux articules. Volume 2. Dumenil, Paris." type="book" year="1840">Castelnau, 1840</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; and the four cited in Dejean’s catalogues (
|
||
<bibRefCitation id="063DA3E4743EFFEFB9B3F9E1448AF9B8" author="Dejean P. F." box="[1003,1157,1541,1567]" pageId="4" pageNumber="5" refId="ref29235" refString="Dejean P. F. 1833. Catalogue des Coleopteres de la Collection de M. le Comte Dejean. Mequignon-Marvis pere et fils, Paris. https: // doi. org / 10.5962 / bhl. title. 8771" type="book" year="1833">Dejean 1833</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="063DA3E4743EFFEFBECAF9E144DFF9B8" author="Dejean P. F." box="[1170,1232,1541,1567]" pageId="4" pageNumber="5" refId="ref29276" refString="Dejean P. F. 1837. Catalogue des Coleopteres de la Collection de M. le Comte Dejean. Third edition. Mequignon-Marvis pere et fils, Paris." type="book" year="1837">1837</bibRefCitation>
|
||
),
|
||
<taxonomicName id="A5ACA596743EFFEFBEBFF9E24577F9B8" box="[1255,1400,1542,1567]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="testaceus">
|
||
<emphasis id="50D80207743EFFEFBEBFF9E24577F9B8" box="[1255,1400,1542,1567]" italics="true" pageId="4" pageNumber="5">H. testaceus</emphasis>
|
||
</taxonomicName>
|
||
and three nomina nuda,
|
||
<taxonomicName id="A5ACA596743EFFEFBB80F9CE424BF9E4" box="[472,580,1577,1603]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="discus">
|
||
<emphasis id="50D80207743EFFEFBB80F9CE424BF9E4" box="[472,580,1577,1603]" italics="true" pageId="4" pageNumber="5">H. discus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A5ACA596743EFFEFB80BF9CE42E4F9E4" box="[595,747,1578,1603]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="granarius">
|
||
<emphasis id="50D80207743EFFEFB80BF9CE42E4F9E4" box="[595,747,1578,1603]" italics="true" pageId="4" pageNumber="5">H. granarius</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A5ACA596743EFFEFB973F9CE43A3F9E4" box="[811,940,1578,1603]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutus">
|
||
<emphasis id="50D80207743EFFEFB973F9CE43A3F9E4" box="[811,940,1578,1603]" italics="true" pageId="4" pageNumber="5">H. minutus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5F98F4482F89D" blockId="4.[187,1399,1643,1850]" pageId="4" pageNumber="5">
|
||
<bibRefCitation id="063DA3E4743EFFEFBAE5F98F417AF922" author="Erichson W. F." box="[189,373,1643,1669]" pageId="4" pageNumber="5" pagination="67 - 185" refId="ref29306" refString="Erichson W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185." type="journal article" year="1847">Erichson (1847)</bibRefCitation>
|
||
proposed the name
|
||
<taxonomicName id="A5ACA596743EFFEFB80FF98F42E5F922" authorityName="Erichson" authorityYear="1847" box="[599,746,1643,1669]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
for the family, based on the number of antennal segments (10) and the presence of strongly fused abdominal segments. This author included six genera:
|
||
<taxonomicName id="A5ACA596743EFFEFBEAEF96B4177F96A" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBEAEF96B457AF90E" box="[1270,1397,1679,1705]" italics="true" pageId="4" pageNumber="5">Hybosorus</emphasis>
|
||
MacLeay, 1819
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743EFFEFBBDEF95742DFF96A" authority="Laporte, 1840" authorityName="Laporte" authorityYear="1840" box="[390,720,1715,1742]" class="Insecta" family="Hybosoridae" genus="Phaeochrous" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBBDEF957422FF96A" box="[390,544,1715,1741]" italics="true" pageId="4" pageNumber="5">Phaeochrous</emphasis>
|
||
Laporte, 1840
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743EFFEFB886F9574419F969" authority="Westwood, 1845" authorityName="Westwood" authorityYear="1845" box="[734,1046,1715,1742]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB886F9574349F96A" box="[734,838,1715,1741]" italics="true" pageId="4" pageNumber="5">Coilodes</emphasis>
|
||
Westwood, 1845
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743EFFEFBE7BF957457DF969" authority="Westwood, 1845" authorityName="Westwood" authorityYear="1845" box="[1059,1394,1715,1742]" class="Insecta" family="Hybosoridae" genus="Chaetodus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBE7BF95744AFF96A" box="[1059,1184,1715,1741]" italics="true" pageId="4" pageNumber="5">Chaetodus</emphasis>
|
||
Westwood, 1845
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743EFFEFBAE6F933420AF955" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" box="[190,517,1751,1778]" class="Insecta" family="Hybosoridae" genus="Dicraeodon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBAE6F9334145F956" box="[190,330,1751,1777]" italics="true" pageId="4" pageNumber="5">Dicraeodon</emphasis>
|
||
<bibRefCitation id="063DA3E4743EFFEFBB08F933420AF955" author="Erichson W. F." box="[336,517,1751,1778]" pageId="4" pageNumber="5" pagination="67 - 185" refId="ref29306" refString="Erichson W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185." type="journal article" year="1847">Erichson, 1847</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="A5ACA596743EFFEFB819F93343A9F955" authority="Westwood, 1845" authorityName="Westwood" authorityYear="1845" box="[577,934,1751,1778]" class="Insecta" family="Hybosoridae" genus="Apalonychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB819F93342D6F956" box="[577,729,1751,1777]" italics="true" pageId="4" pageNumber="5">Apalonychus</emphasis>
|
||
Westwood, 1845
|
||
</taxonomicName>
|
||
, and suggested changes in the names of two genera,
|
||
<taxonomicName id="A5ACA596743EFFEFBB0AF91F41B4F8B2" authorityName="Westwood" authorityYear="1845" box="[338,443,1787,1813]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBB0AF91F41B4F8B2" box="[338,443,1787,1813]" italics="true" pageId="4" pageNumber="5">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
to ‘
|
||
<taxonomicName id="A5ACA596743EFFEFBBB3F91F4257F8B2" box="[491,600,1787,1813]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFBBB3F91F4257F8B2" box="[491,600,1787,1813]" italics="true" pageId="4" pageNumber="5">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
’, and
|
||
<taxonomicName id="A5ACA596743EFFEFB8FEF91F4331F8B2" authorityName="Westwood" authorityYear="1845" box="[678,830,1787,1813]" class="Insecta" family="Hybosoridae" genus="Apalonychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB8FEF91F4331F8B2" box="[678,830,1787,1813]" italics="true" pageId="4" pageNumber="5">Apalonychus</emphasis>
|
||
</taxonomicName>
|
||
to ‘
|
||
<taxonomicName id="A5ACA596743EFFEFB928F91F4415F8B2" box="[880,1050,1787,1813]" class="Insecta" family="Hybosoridae" genus="Hapalonychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB928F91F4415F8B2" box="[880,1050,1787,1813]" italics="true" pageId="4" pageNumber="5">Hapalonychus</emphasis>
|
||
</taxonomicName>
|
||
’.
|
||
<bibRefCitation id="063DA3E4743EFFEFBE6CF91F4504F8B1" author="Lacordaire J. T." box="[1076,1291,1787,1814]" pageId="4" pageNumber="5" refId="ref29690" refString="Lacordaire J. T. 1856. Histoire naturelle des Insectes: Genera des Coleopteres ou Expose methodique et critique de tous les Genres proposes jusqu'ici dans cet Ordre d'Insectes. Tome 3. Contenant les Familles des Pectinicornes et Lamellicornes. Librairie encyclopedique de Roret, Paris." type="book" year="1856">Lacordaire (1856)</bibRefCitation>
|
||
used the same names proposed by Erichson and redescribed
|
||
<taxonomicName id="A5ACA596743EFFEFB944F8C443BDF89D" authorityName="Erichson" authorityYear="1847" box="[796,946,1824,1850]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
and its six genera.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743EFFEFBAE5F88741D2F84E" blockId="4.[187,1399,1890,2025]" pageId="4" pageNumber="5">
|
||
<bibRefCitation id="063DA3E4743EFFEFBAE5F8874172F8DB" author="MacLeay W. J." box="[189,381,1890,1917]" pageId="4" pageNumber="5" pagination="106 - 130" refId="ref29778" refString="MacLeay W. J. 1864. Descriptions of new genera and species of Coleoptera from Port Denison. The Transactions of the Entomological Society of New South Wales 1: 106 - 130." type="journal article" year="1864">MacLeay (1864)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="A5ACA596743EFFEFBBAFF88642AAF8DB" authorityName="MacLeay" authorityYear="1864" box="[503,677,1890,1916]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bimaculatus">
|
||
<emphasis id="50D80207743EFFEFBBAFF88642AAF8DB" box="[503,677,1890,1916]" italics="true" pageId="4" pageNumber="5">C. bimaculatus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="A5ACA596743EFFEFB89FF886433CF8DB" box="[711,819,1890,1916]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743EFFEFB89FF886433CF8DB" box="[711,819,1890,1916]" italics="true" pageId="4" pageNumber="5">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
[sic], for an Australian species from Port Denison. Despite the discrepant distribution, the author stated doubts about the generic position of this species due to the lack of sexual dimorphism in the pronotum and the tarsal claws, used by
|
||
<bibRefCitation id="063DA3E4743EFFEFBED1F84E4557F863" author="Westwood J. O." box="[1161,1368,1962,1988]" pageId="4" pageNumber="5" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood (1846)</bibRefCitation>
|
||
as diagnostic for the genus.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5FEEE422FFE50" blockId="5.[187,1400,265,503]" pageId="5" pageNumber="6">
|
||
In the catalogue of
|
||
<taxonomicName id="A5ACA596743FFFEEBBCFFEED4219FE84" authorityName="Linnaeus" authorityYear="1758" box="[407,534,265,291]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
|
||
by
|
||
<bibRefCitation id="063DA3E4743FFFEEB867FEED439CFE84" author="Gemminger M. & Harold E." box="[575,915,265,291]" pageId="5" pageNumber="6" refId="ref29416" refString="Gemminger M. & Harold E. 1869. Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus, Tom. IV: Scarabaeidae. E. H. Gumm, Munich." type="book" year="1869">Gemminger & Harold (1869)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="A5ACA596743FFFEEB9C3FEED440DFE84" authorityName="Westwood" authorityYear="1845" box="[923,1026,265,291]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB9C3FEED440DFE84" box="[923,1026,265,291]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
was composed by eight species:
|
||
<taxonomicName id="A5ACA596743FFFEEBAE3FEC94220FEE1" authority="MacLeay, 1864" authorityName="MacLeay" authorityYear="1864" box="[187,559,300,327]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bimaculatus">
|
||
<emphasis id="50D80207743FFFEEBAE3FEC94163FEE1" box="[187,364,300,327]" italics="true" pageId="5" pageNumber="6">C. bimaculatus</emphasis>
|
||
<bibRefCitation id="063DA3E4743FFFEEBB2DFEC94220FEE1" author="MacLeay W. J." box="[373,559,300,327]" pageId="5" pageNumber="6" pagination="106 - 130" refId="ref29778" refString="MacLeay W. J. 1864. Descriptions of new genera and species of Coleoptera from Port Denison. The Transactions of the Entomological Society of New South Wales 1: 106 - 130." type="journal article" year="1864">MacLeay, 1864</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEB862FEC94391FEE1" authority="Westwood, 1846" authorityName="Westwood" authorityYear="1846" box="[570,926,300,327]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="50D80207743FFFEEB862FEC942DFFEE1" box="[570,720,301,327]" italics="true" pageId="5" pageNumber="6">C. castaneus</emphasis>
|
||
<bibRefCitation id="063DA3E4743FFFEEB88FFEC84391FEE1" author="Westwood J. O." box="[727,926,300,326]" pageId="5" pageNumber="6" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEB9F2FEC944F5FEE1" authority="Westwood 1846" authorityName="Westwood" authorityYear="1846" box="[938,1274,300,327]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="50D80207743FFFEEB9F2FEC9443CFEE1" box="[938,1075,300,327]" italics="true" pageId="5" pageNumber="6">C. chilensis</emphasis>
|
||
<bibRefCitation id="063DA3E4743FFFEEBE62FEC844F5FEE1" author="Westwood J. O." box="[1082,1274,300,326]" pageId="5" pageNumber="6" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBF5EFEC94157FECD" authority="(Perty, 1830)" baseAuthorityName="Perty" baseAuthorityYear="1830" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gibbus">
|
||
<emphasis id="50D80207743FFFEEBF5EFEC94579FEE1" box="[1286,1398,300,327]" italics="true" pageId="5" pageNumber="6">C. gibbus</emphasis>
|
||
(
|
||
<bibRefCitation id="063DA3E4743FFFEEBA9DFEB44140FECD" author="Perty J. A. M." box="[197,335,336,362]" pageId="5" pageNumber="6" refId="ref30331" refString="Perty J. A. M. 1830. Delectus Animalium Articulatorum quae in Itinere per Brasiliam Annis MDCCCXVII- MDCCCXX Jussu et Auspiciis Maximiliani Josephi I Bavariae Regis Augustissimi Peracto. Munich. https: // doi. org / 10.5962 / bhl. title. 158694" type="book" year="1830">Perty, 1830</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBB3AFEB442FBFECD" authority="(Mannerheim, 1829)" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[354,756,335,362]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743FFFEEBB3AFEB441F7FECE" box="[354,504,335,362]" italics="true" pageId="5" pageNumber="6">C. humeralis</emphasis>
|
||
(
|
||
<bibRefCitation id="063DA3E4743FFFEEB85FFEAB42E4FECD" author="Mannerheim K. G." box="[519,747,335,362]" pageId="5" pageNumber="6" pagination="29 - 80" refId="ref29813" refString="Mannerheim K. G. 1829. Description de quarante nouvelles especes de scarabeides du Bresil: avec figures. Memoires de la Societe imperiale des Naturalistes de Moscou 7: 29 - 80." type="journal article" year="1829">Mannerheim, 1829</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEB8A7FEB44454FECD" authority="(Mannerheim, 1829)" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[767,1115,335,362]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="50D80207743FFFEEB8A7FEB44351FECE" box="[767,862,336,362]" italics="true" pageId="5" pageNumber="6">C. niger</emphasis>
|
||
(
|
||
<bibRefCitation id="063DA3E4743FFFEEB936FEAB445DFECD" author="Mannerheim K. G." box="[878,1106,335,362]" pageId="5" pageNumber="6" pagination="29 - 80" refId="ref29813" refString="Mannerheim K. G. 1829. Description de quarante nouvelles especes de scarabeides du Bresil: avec figures. Memoires de la Societe imperiale des Naturalistes de Moscou 7: 29 - 80." type="journal article" year="1829">Mannerheim, 1829</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBE3DFEB440F7FE2A" authority="Westwood, 1846" authorityName="Westwood" authorityYear="1846" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="50D80207743FFFEEBE3DFEB444E2FECE" box="[1125,1261,335,362]" italics="true" pageId="5" pageNumber="6">C. parvulus</emphasis>
|
||
<bibRefCitation id="063DA3E4743FFFEEBEABFEAB40F7FE2A" author="Westwood J. O." pageId="5" pageNumber="6" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743FFFEEBB69FE974270FE2A" authority="(Castelnau, 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[305,639,371,397]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rufulus">
|
||
<emphasis id="50D80207743FFFEEBB69FE9741A8FE2A" box="[305,423,371,397]" italics="true" pageId="5" pageNumber="6">C. rufulus</emphasis>
|
||
(
|
||
<bibRefCitation id="063DA3E4743FFFEEBBEEFE974278FE2A" author="Castelnau & Laporte F. D." box="[438,631,371,397]" pageId="5" pageNumber="6" refId="ref29111" refString="Castelnau [Laporte F. D.] 1840. Histoire naturelle des Animaux articules. Volume 2. Dumenil, Paris." type="book" year="1840">Castelnau, 1840</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(included in
|
||
<taxonomicName id="A5ACA596743FFFEEB943FE97438CFE2A" authorityName="Westwood" authorityYear="1845" box="[795,899,371,397]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB943FE97438CFE2A" box="[795,899,371,397]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
after suggestion of
|
||
<bibRefCitation id="063DA3E4743FFFEEBE34FE974521FE2A" author="Westwood J. O." box="[1132,1326,371,397]" pageId="5" pageNumber="6" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood 1846</bibRefCitation>
|
||
). Five years later,
|
||
<bibRefCitation id="063DA3E4743FFFEEBB18FE7241EAFE17" author="Harold E. F." box="[320,485,406,432]" pageId="5" pageNumber="6" pagination="26 - 51" refId="ref29521" refString="Harold E. F. 1874. Zur Kenntnis der kugelformigen Trogiden. Coleopterologischen Hefte 12: 26 - 51." type="journal article" year="1874">Harold (1874)</bibRefCitation>
|
||
transferred
|
||
<taxonomicName id="A5ACA596743FFFEEB835FE724312FE17" authorityName="MacLeay" authorityYear="1864" box="[621,797,406,432]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bimaculatus">
|
||
<emphasis id="50D80207743FFFEEB835FE724312FE17" box="[621,797,406,432]" italics="true" pageId="5" pageNumber="6">C. bimaculatus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="A5ACA596743FFFEEB919FE724486FE17" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[833,1161,406,432]" class="Insecta" family="Hybosoridae" genus="Liparochrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB919FE7243DFFE17" box="[833,976,406,432]" italics="true" pageId="5" pageNumber="6">Liparochrus</emphasis>
|
||
Erichson, 1848
|
||
</taxonomicName>
|
||
. Then,
|
||
<bibRefCitation id="063DA3E4743FFFEEBE83FE724170FE74" author="Preudhomme de Borre A." pageId="5" pageNumber="6" pagination="103 - 120" refId="ref30400" refString="Preudhomme de Borre A. 1886. Liste des lamellicornes laparostictiques recueillis par Feu Camille van Volxem pendant son voyage au Bresil et a la Plata en 1872, suivie de la description de dix huit especes nouvelles et un genre nouveau. Annales de la Societe entomologique de Belgique 30: 103 - 120." type="journal article" year="1886">Preudhomme de Borre (1886)</bibRefCitation>
|
||
transferred
|
||
<taxonomicName id="A5ACA596743FFFEEB84EFE5E429FFE74" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[534,656,441,468]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rufulus">
|
||
<emphasis id="50D80207743FFFEEB84EFE5E429FFE74" box="[534,656,441,468]" italics="true" pageId="5" pageNumber="6">C. rufulus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="A5ACA596743FFFEEB8E2FE5D435CFE74" authorityName="Westwood" authorityYear="1845" box="[698,851,441,467]" class="Insecta" family="Hybosoridae" genus="Apalonychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB8E2FE5D435CFE74" box="[698,851,441,467]" italics="true" pageId="5" pageNumber="6">Apalonychus</emphasis>
|
||
</taxonomicName>
|
||
, which was synonymized years later with
|
||
<taxonomicName id="A5ACA596743FFFEEBF07FE5E4143FE51" authorityName="Westwood" authorityYear="1845" class="Insecta" family="Hybosoridae" genus="Apalonychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="waterhousei">
|
||
<emphasis id="50D80207743FFFEEBF07FE5E4143FE51" italics="true" pageId="5" pageNumber="6">A. waterhousei</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="063DA3E4743FFFEEBB2EFE394213FE50" author="Arrow G. J." box="[374,540,477,503]" pageId="5" pageNumber="6" pagination="479 - 507" refId="ref28738" refString="Arrow G. J. 1909. On the characters and relationships of the less know groups of lamellicorn Coleoptera, with description of a new species of Hybosorinae, etc. Transactions of the Entomological Society of London 2: 479 - 507." type="journal article" year="1909">Arrow (1909)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5FDFA4490FDFC" blockId="5.[189,1398,542,604]" pageId="5" pageNumber="6">
|
||
<bibRefCitation id="063DA3E4743FFFEEBAE5FDFA4158FD9F" author="Bates H. W." box="[189,343,542,568]" pageId="5" pageNumber="6" refId="ref29007" refString="Bates H. W. 1887. Biologia Centrali Americana: Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. R. H. Porter, London. Available from https: // www. biodiversitylibrary. org / page / 42603494 [accessed 12 Nov. 2023]." type="url" year="1887">Bates (1887)</bibRefCitation>
|
||
cited new distribution records for
|
||
<taxonomicName id="A5ACA596743FFFEEB8A1FDFA439EFD9F" authorityName="Westwood" authorityYear="1846" box="[761,913,542,568]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="50D80207743FFFEEB8A1FDFA439EFD9F" box="[761,913,542,568]" italics="true" pageId="5" pageNumber="6">C. castaneus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="1ABB9E85743FFFEEB9F8FDFA4414FD9F" box="[928,1051,542,568]" name="Nicaragua" pageId="5" pageNumber="6">Nicaragua</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743FFFEEBE00FDFA44D3FD9F" box="[1112,1244,542,568]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
|
||
) and a small unidentified specimen from
|
||
<collectingCountry id="1ABB9E85743FFFEEB850FDA54285FDFC" box="[520,650,577,603]" name="Guatemala" pageId="5" pageNumber="6">Guatemala</collectingCountry>
|
||
, but without any further information about it.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5FD674300FC8D" blockId="5.[187,1398,643,810]" pageId="5" pageNumber="6">
|
||
<bibRefCitation id="063DA3E4743FFFEEBAE5FD674154FD3A" author="Arrow G. J." box="[189,347,643,669]" pageId="5" pageNumber="6" pagination="509 - 520" refId="ref28682" refString="Arrow G. J. 1903. On the laparostict lamellicorne Coleoptera of Granada and St. Vincent (W. Indies). Transactions of the Entomological Society of London 2: 509 - 520. https: // doi. org / 10.1111 / j. 1365 - 2311.1903. tb 01144. x" type="journal article" year="1903">Arrow (1903)</bibRefCitation>
|
||
described three other species of
|
||
<taxonomicName id="A5ACA596743FFFEEB892FD67433EFD3A" authorityName="Westwood" authorityYear="1845" box="[714,817,643,669]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB892FD67433EFD3A" box="[714,817,643,669]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
(still using the ‘
|
||
<taxonomicName id="A5ACA596743FFFEEB9B0FD67445CFD3A" box="[1000,1107,643,669]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB9B0FD67445CFD3A" box="[1000,1107,643,669]" italics="true" pageId="5" pageNumber="6">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
’ spelling):
|
||
<taxonomicName id="A5ACA596743FFFEEBE89FD67457AFD3B" authorityName="Arrow" authorityYear="1903" box="[1233,1397,643,669]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nigripennis">
|
||
<emphasis id="50D80207743FFFEEBE89FD67457AFD3B" box="[1233,1397,643,669]" italics="true" pageId="5" pageNumber="6">C. nigripennis</emphasis>
|
||
</taxonomicName>
|
||
(from
|
||
<collectingCountry id="1ABB9E85743FFFEEBB5EFD424184FD67" box="[262,395,678,704]" name="Saint Vincent and the Grenadines" pageId="5" pageNumber="6">St. Vincent</collectingCountry>
|
||
, Leeward side),
|
||
<taxonomicName id="A5ACA596743FFFEEB808FD424307FD67" authorityName="Arrow" authorityYear="1909" box="[592,776,678,704]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="punctipennis">
|
||
<emphasis id="50D80207743FFFEEB808FD424307FD67" box="[592,776,678,704]" italics="true" pageId="5" pageNumber="6">C. punctipennis</emphasis>
|
||
</taxonomicName>
|
||
(from
|
||
<collectingCountry id="1ABB9E85743FFFEEB902FD4243B1FD67" box="[858,958,678,704]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
|
||
) and
|
||
<taxonomicName id="A5ACA596743FFFEEBE58FD4244ACFD67" authorityName="Arrow" authorityYear="1909" box="[1024,1187,678,704]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fumipennis">
|
||
<emphasis id="50D80207743FFFEEBE58FD4244ACFD67" box="[1024,1187,678,704]" italics="true" pageId="5" pageNumber="6">C. fumipennis</emphasis>
|
||
</taxonomicName>
|
||
(from Amazonas,
|
||
<collectingCountry id="1ABB9E85743FFFEEBAE4FD2D4104FD44" box="[188,267,713,739]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
|
||
). In his catalogue of
|
||
<taxonomicName id="A5ACA596743FFFEEB84BFD2D4345FD44" authority="(Arrow 1912)" baseAuthorityName="Arrow" baseAuthorityYear="1912" box="[531,842,713,739]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="order">
|
||
Coleoptera (
|
||
<bibRefCitation id="063DA3E4743FFFEEB8F2FD2D434CFD44" author="Arrow G. J." box="[682,835,713,739]" pageId="5" pageNumber="6" pagination="1 - 65" refId="ref28783" refString="Arrow G. J. 1912. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Coleopterorum Catalogus 43: 1 - 65." type="journal article" year="1912">Arrow 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
the same author updated the composition of ‘
|
||
<taxonomicName id="A5ACA596743FFFEEBA9FFD08413BFCA1" box="[199,308,748,774]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBA9FFD08413BFCA1" box="[199,308,748,774]" italics="true" pageId="5" pageNumber="6">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
’ to nine species:
|
||
<taxonomicName id="A5ACA596743FFFEEB85EFD094292FCA1" authorityName="Westwood" authorityYear="1846" box="[518,669,749,775]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="50D80207743FFFEEB85EFD094292FCA1" box="[518,669,749,775]" italics="true" pageId="5" pageNumber="6">C. castaneus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEB8F5FD094338FCA1" authorityName="Westwood" authorityYear="1846" box="[685,823,748,775]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="50D80207743FFFEEB8F5FD094338FCA1" box="[685,823,748,775]" italics="true" pageId="5" pageNumber="6">C. chilensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEB91FFD0943B6FCA1" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[839,953,748,775]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gibbus">
|
||
<emphasis id="50D80207743FFFEEB91FFD0943B6FCA1" box="[839,953,748,775]" italics="true" pageId="5" pageNumber="6">C. gibbus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEB991FD09446DFCA1" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[969,1122,748,775]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743FFFEEB991FD09446DFCA1" box="[969,1122,748,775]" italics="true" pageId="5" pageNumber="6">C. humeralis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBE2BFD0944DBFCA1" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1139,1236,749,775]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="50D80207743FFFEEBE2BFD0944DBFCA1" box="[1139,1236,749,775]" italics="true" pageId="5" pageNumber="6">C. niger</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBEBCFD094560FCA1" authorityName="Westwood" authorityYear="1846" box="[1252,1391,748,775]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="50D80207743FFFEEBEBCFD094560FCA1" box="[1252,1391,748,775]" italics="true" pageId="5" pageNumber="6">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBAE5FCF4416FFC8D" authorityName="Arrow" authorityYear="1909" box="[189,352,784,810]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fumipennis">
|
||
<emphasis id="50D80207743FFFEEBAE5FCF4416FFC8D" box="[189,352,784,810]" italics="true" pageId="5" pageNumber="6">C. fumipennis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBB37FCF44227FC8E" authorityName="Arrow" authorityYear="1909" box="[367,552,784,810]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="punctipennis">
|
||
<emphasis id="50D80207743FFFEEBB37FCF44227FC8E" box="[367,552,784,810]" italics="true" pageId="5" pageNumber="6">C. punctipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743FFFEEB839FCF44306FC8E" authorityName="Arrow" authorityYear="1903" box="[609,777,784,810]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nigripennis">
|
||
<emphasis id="50D80207743FFFEEB839FCF44306FC8E" box="[609,777,784,810]" italics="true" pageId="5" pageNumber="6">C. nigripennis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5FCB644F1FBC5" blockId="5.[188,1399,849,1122]" pageId="5" pageNumber="6">
|
||
<bibRefCitation id="063DA3E4743FFFEEBAE5FCB64157FCCC" author="Lucas R." box="[189,344,849,876]" pageId="5" pageNumber="6" refId="ref29739" refString="Lucas R. 1920. Catalogus Alphabeticus Generum et Subgenerum Coleopterorum Orbis Terrarum Totius (Famil., Trib., Subtr., Sect. Incl.). Pars I. R. Stricker, Berlin." type="book" year="1920">Lucas (1920)</bibRefCitation>
|
||
redressed the spelling to “
|
||
<taxonomicName id="A5ACA596743FFFEEB8D6FCB542F9FCCC" authorityName="Westwood" authorityYear="1845" box="[654,758,849,875]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB8D6FCB542F9FCCC" box="[654,758,849,875]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
”. However,
|
||
<bibRefCitation id="063DA3E4743FFFEEB9D3FCB54409FCCC" author="Pic M." box="[907,1030,849,875]" pageId="5" pageNumber="6" refId="ref30378" refString="Pic M. 1928. Melanges exotico-entomologiques. Notes et Descriptions. Vol. 51. Les Imprimeries Reunies, Moulins." type="book" year="1928">Pic (1928)</bibRefCitation>
|
||
resumed the use of the spelling “
|
||
<taxonomicName id="A5ACA596743FFFEEBA92FC904138FC29" box="[202,311,884,910]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBA92FC904138FC29" box="[202,311,884,910]" italics="true" pageId="5" pageNumber="6">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
”, which was also reproduced by
|
||
<bibRefCitation id="063DA3E4743FFFEEB8E4FC904363FC29" author="Paulian R." box="[700,876,884,910]" pageId="5" pageNumber="6" pagination="213 - 296" refId="ref30228" refString="Paulian R. 1938. Contribution a l'etude des canthonides americains. Annales de la Societe entomologique de France 107: 213 - 296." type="journal article" year="1938">Paulian (1938)</bibRefCitation>
|
||
. The former also indicated the existence of a variety of
|
||
<taxonomicName id="A5ACA596743FFFEEBB69FC7C41AFFC16" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[305,416,919,946]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gibbus">
|
||
<emphasis id="50D80207743FFFEEBB69FC7C41AFFC16" box="[305,416,919,946]" italics="true" pageId="5" pageNumber="6">C. gibbus</emphasis>
|
||
</taxonomicName>
|
||
(“
|
||
<taxonomicName id="A5ACA596743FFFEEBBE6FC734270FC16" authority="Basílio & Vaz-De-Mello & Cherman & Almeida, 2023" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" box="[446,639,919,945]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gibbus" status="var. nov.">
|
||
<emphasis id="50D80207743FFFEEBBE6FC734270FC16" box="[446,639,919,945]" italics="true" pageId="5" pageNumber="6">Coelodes gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4BEBBF7C743FFFEEB8DDFC7C42DEFC15" box="[645,721,920,946]" pageId="5" pageNumber="6" rank="variety">v. nov.</taxonomicNameLabel>
|
||
<taxonomicName id="A5ACA596743FFFEEB88DFC7C434FFC16" box="[725,832,920,945]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="testaceus">
|
||
<emphasis id="50D80207743FFFEEB88DFC7C434FFC16" box="[725,832,920,945]" italics="true" pageId="5" pageNumber="6">testaceus</emphasis>
|
||
</taxonomicName>
|
||
”).
|
||
<bibRefCitation id="063DA3E4743FFFEEB93DFC734442FC15" author="Blackwelder R. E." box="[869,1101,919,946]" pageId="5" pageNumber="6" pagination="2" refId="ref29063" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Parts 1 and 2. Bulletin 185, United States National Museum. United States Government Printing Office, Washington DC." type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
|
||
used the original spelling for the genus in his checklist, and included within
|
||
<taxonomicName id="A5ACA596743FFFEEB8A7FC5F4368FC72" authorityName="Westwood" authorityYear="1845" box="[767,871,955,981]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB8A7FC5F4368FC72" box="[767,871,955,981]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
the same nine species cited by
|
||
<bibRefCitation id="063DA3E4743FFFEEBE96FC5F457CFC72" author="Arrow G. J." box="[1230,1395,955,981]" pageId="5" pageNumber="6" pagination="1 - 65" refId="ref28783" refString="Arrow G. J. 1912. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Coleopterorum Catalogus 43: 1 - 65." type="journal article" year="1912">Arrow (1912)</bibRefCitation>
|
||
. The author also attributed a feminine grammatical gender to its names (
|
||
<taxonomicName id="A5ACA596743FFFEEB9A6FC3A446DFC5F" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[1022,1122,990,1016]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gibba">
|
||
<emphasis id="50D80207743FFFEEB9A6FC3A446DFC5F" box="[1022,1122,990,1016]" italics="true" pageId="5" pageNumber="6">C. gibba</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A5ACA596743FFFEEBE28FC3A44E2FC5F" authorityName="Westwood" authorityYear="1845" box="[1136,1261,990,1016]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="parvula">
|
||
<emphasis id="50D80207743FFFEEBE28FC3A44E2FC5F" box="[1136,1261,990,1016]" italics="true" pageId="5" pageNumber="6">C. parvula</emphasis>
|
||
</taxonomicName>
|
||
). However, the name “
|
||
<taxonomicName id="A5ACA596743FFFEEBB61FBE541AEFBBC" authorityName="Westwood" authorityYear="1845" box="[313,417,1025,1051]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBB61FBE541AEFBBC" box="[313,417,1025,1051]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
” is masculine in gender in its original description (see also Art. 30.1.4.4 of the ICZN Code;
|
||
<bibRefCitation id="063DA3E4743FFFEEBB50FBC1419EFB98" author="ICZN" box="[264,401,1061,1087]" pageId="5" pageNumber="6" refId="ref29631" refString="ICZN 1999. International Code of Zoological Nomenclature. Fourth edition adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
). Four years later,
|
||
<bibRefCitation id="063DA3E4743FFFEEB83DFBC04325FB98" author="Robinson M." box="[613,810,1060,1087]" pageId="5" pageNumber="6" pagination="29 - 36" refId="ref30632" refString="Robinson M. 1948. Studies in the Scarabaeidae IV (Coleoptera). Transactions of the American Entomological Society 74 (1): 29 - 36. Available from https: // www. jstor. org / stable / 25077572 [accessed 23 Nov. 2023]." type="journal article" year="1948">Robinson (1948)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="A5ACA596743FFFEEB9FDFBC14404FB99" authorityName="Robinson" authorityYear="1948" box="[933,1035,1060,1087]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="50D80207743FFFEEB9FDFBC14404FB99" box="[933,1035,1060,1087]" italics="true" pageId="5" pageNumber="6">C. ovalis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="1ABB9E85743FFFEEBE08FBC044C3FB99" box="[1104,1228,1060,1086]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
|
||
, based only on female specimens, using once more the generic spelling suggested by Erichson (‘
|
||
<taxonomicName id="A5ACA596743FFFEEBE2EFBAC44ECFBC5" box="[1142,1251,1096,1122]" class="Insecta" family="Hybosoridae" genus="Coelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBE2EFBAC44ECFBC5" box="[1142,1251,1096,1122]" italics="true" pageId="5" pageNumber="6">Coelodes</emphasis>
|
||
</taxonomicName>
|
||
’).
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5FB6D41CEFA96" blockId="5.[188,1398,1161,1329]" pageId="5" pageNumber="6">
|
||
The first checklist of
|
||
<taxonomicName id="A5ACA596743FFFEEBBEEFB6D4243FB04" authorityName="Erichson" authorityYear="1847" box="[438,588,1161,1187]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
was published by
|
||
<bibRefCitation id="063DA3E4743FFFEEB97BFB6D43D6FB04" author="Allsopp P. G." box="[803,985,1161,1187]" pageId="5" pageNumber="6" pagination="105 - 117" refId="ref28631" refString="Allsopp P. G. 1984. Checklist of the Hybosoridae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 38 (2): 105 - 117. Available from https: // www. jstor. org / stable / 4008153 [accessed 23 Nov. 2023]." type="journal article" year="1984">Allsopp (1984)</bibRefCitation>
|
||
, where the author ranked the group as a subfamily of
|
||
<taxonomicName id="A5ACA596743FFFEEBBC5FB484233FB61" authorityName="Latreille" authorityYear="1802" box="[413,572,1196,1222]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
|
||
(Hybosorinae). Supported by the International Code of Zoological Nomenclature (which came into effect in 1958),
|
||
<bibRefCitation id="063DA3E4743FFFEEB949FB3443C5FB4D" author="Allsopp P. G." box="[785,970,1232,1258]" pageId="5" pageNumber="6" pagination="105 - 117" refId="ref28631" refString="Allsopp P. G. 1984. Checklist of the Hybosoridae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 38 (2): 105 - 117. Available from https: // www. jstor. org / stable / 4008153 [accessed 23 Nov. 2023]." type="journal article" year="1984">Allsopp (1984)</bibRefCitation>
|
||
resumed the original (and current) spelling of
|
||
<taxonomicName id="A5ACA596743FFFEEBB66FB1741A9FAAA" authorityName="Westwood" authorityYear="1845" box="[318,422,1267,1293]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBB66FB1741A9FAAA" box="[318,422,1267,1293]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
and listed ten species in the genus: the nine cited by
|
||
<bibRefCitation id="063DA3E4743FFFEEBE52FB1744A4FAAA" author="Arrow G. J." box="[1034,1195,1267,1293]" pageId="5" pageNumber="6" pagination="1 - 65" refId="ref28783" refString="Arrow G. J. 1912. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Coleopterorum Catalogus 43: 1 - 65." type="journal article" year="1912">Arrow (1912)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="063DA3E4743FFFEEBEBAFB174102FA97" author="Blackwelder R. E." pageId="5" pageNumber="6" pagination="2" refId="ref29063" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Parts 1 and 2. Bulletin 185, United States National Museum. United States Government Printing Office, Washington DC." type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
|
||
, plus
|
||
<taxonomicName id="A5ACA596743FFFEEBB0AFAF341B5FA97" authorityName="Robinson" authorityYear="1948" box="[338,442,1302,1329]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="50D80207743FFFEEBB0AFAF341B5FA97" box="[338,442,1302,1329]" italics="true" pageId="5" pageNumber="6">C. ovalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5FABC4515FA58" blockId="5.[189,1400,1368,1535]" pageId="5" pageNumber="6">
|
||
In the molecular phylogeny of
|
||
<taxonomicName id="A5ACA596743FFFEEB86CFABC42C3FAD5" authorityName="Erichson" authorityYear="1847" box="[564,716,1368,1394]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="063DA3E4743FFFEEB8A3FABC4439FAD5" author="Ocampo F. C. & Hawks D. C." box="[763,1078,1368,1394]" pageId="5" pageNumber="6" pagination="7 - 12" refId="ref30070" refString="Ocampo F. C. & Hawks D. C. 2006. Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae 2. Molecular phylogenetics and systematic placement of the Hybosoridae. Bulletin of the University of Nebraska State Museum 19: 7 - 12." type="journal article" year="2006">Ocampo & Hawks (2006)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="A5ACA596743FFFEEBE1CFABC44A1FAD5" authorityName="Westwood" authorityYear="1845" box="[1092,1198,1368,1394]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBE1CFABC44A1FAD5" box="[1092,1198,1368,1394]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
was found to be more closely related to the Oriental genus
|
||
<taxonomicName id="A5ACA596743FFFEEB89DFA9F4423FA32" authority="Candeze, 1876" authorityName="Candeze" authorityYear="1876" box="[709,1068,1403,1429]" class="Insecta" family="Hybosoridae" genus="Phaeochroops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB89DFA9F437EFA32" box="[709,881,1403,1429]" italics="true" pageId="5" pageNumber="6">Phaeochroops</emphasis>
|
||
Candèze, 1876
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="A5ACA596743FFFEEBECBFA9F451AFA32" authorityName="MacLeay" authorityYear="1819" box="[1171,1301,1403,1429]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBECBFA9F451AFA32" box="[1171,1301,1403,1429]" italics="true" pageId="5" pageNumber="6">Hybosorus</emphasis>
|
||
</taxonomicName>
|
||
or even to
|
||
<taxonomicName id="A5ACA596743FFFEEBA85FA7A42F3FA1F" authority="Preudhomme de Borre, 1866" authorityName="Preudhomme de Borre" authorityYear="1866" box="[221,764,1438,1465]" class="Insecta" family="Hybosoridae" genus="Metachaetodus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBA85FA7A419CFA1F" box="[221,403,1438,1464]" italics="true" pageId="5" pageNumber="6">Metachaetodus</emphasis>
|
||
Preudhomme de Borre, 1866
|
||
</taxonomicName>
|
||
, although that was the only other Neotropical genus of
|
||
<taxonomicName id="A5ACA596743FFFEEBAB9FA254175FA7C" authorityName="Erichson" authorityYear="1847" box="[225,378,1473,1499]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Hybosorinae">Hybosorinae</taxonomicName>
|
||
used in the analysis. In their catalogue of
|
||
<taxonomicName id="A5ACA596743FFFEEB9DDFA254577FA7C" authority=", Ocampo & Ballerio (2006)" authorityName="Ocampo & Ballerio" authorityYear="2006" box="[901,1400,1473,1500]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">
|
||
Hybosoridae,
|
||
<bibRefCitation id="063DA3E4743FFFEEBE77FA264577FA7C" author="Ocampo F. C. & Ballerio A." box="[1071,1400,1473,1500]" pageId="5" pageNumber="6" pagination="178 - 209" refId="ref30003" refString="Ocampo F. C. & Ballerio A. 2006. Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Hybosoridae). Bulletin of the University of Nebraska State Museum 19: 178 - 209." type="journal article" year="2006">Ocampo & Ballerio (2006)</bibRefCitation>
|
||
</taxonomicName>
|
||
synonymized
|
||
<taxonomicName id="A5ACA596743FFFEEBB39FA0141CEFA59" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[353,449,1509,1535]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="50D80207743FFFEEBB39FA0141CEFA59" box="[353,449,1509,1535]" italics="true" pageId="5" pageNumber="6">C. niger</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="A5ACA596743FFFEEB85CFA014294FA58" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[516,667,1509,1535]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743FFFEEB85CFA014294FA58" box="[516,667,1509,1535]" italics="true" pageId="5" pageNumber="6">C. humeralis</emphasis>
|
||
</taxonomicName>
|
||
, differing from
|
||
<bibRefCitation id="063DA3E4743FFFEEB90DFA014408FA58" author="Allsopp P. G." box="[853,1031,1509,1535]" pageId="5" pageNumber="6" pagination="105 - 117" refId="ref28631" refString="Allsopp P. G. 1984. Checklist of the Hybosoridae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 38 (2): 105 - 117. Available from https: // www. jstor. org / stable / 4008153 [accessed 23 Nov. 2023]." type="journal article" year="1984">Allsopp (1984)</bibRefCitation>
|
||
only in this synonymy.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5F9C242AAF921" blockId="5.[187,1398,1574,1670]" pageId="5" pageNumber="6">
|
||
<bibRefCitation id="063DA3E4743FFFEEBAE5F9C24183F9E7" author="Prokofiev A. M." box="[189,396,1574,1600]" pageId="5" pageNumber="6" pagination="1 - 2" refId="ref30456" refString="Prokofiev A. M. 2013 a. A new bolboceratine beetle from Mesoamerica (Coleoptera: Geotrupidae: Bolboceratinae). Calodema 244: 1 - 2." type="journal article" year="2013">Prokofiev (2013a)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="A5ACA596743FFFEEB85DF9C242D7F9E7" authorityName="Prokofiev" authorityYear="2013" box="[517,728,1574,1600]" class="Insecta" family="Hybosoridae" genus="Gnombolbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="orosi">
|
||
<emphasis id="50D80207743FFFEEB85DF9C242D7F9E7" box="[517,728,1574,1600]" italics="true" pageId="5" pageNumber="6">Gnombolbus orosi</emphasis>
|
||
</taxonomicName>
|
||
, a new genus and species of Bolboceratinae (
|
||
<taxonomicName id="A5ACA596743FFFEEBE8EF9C24564F9E7" authorityName="Latreille" authorityYear="1802" box="[1238,1387,1574,1600]" class="Insecta" family="Geotrupidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Geotrupidae</taxonomicName>
|
||
), but in the same year, he synonymized this genus with
|
||
<taxonomicName id="A5ACA596743FFFEEB97CF9AD4383F9C4" authorityName="Westwood" authorityYear="1845" box="[804,908,1609,1635]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB97CF9AD4383F9C4" box="[804,908,1609,1635]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
, and the species became a junior synonym of
|
||
<taxonomicName id="A5ACA596743FFFEEBA85F98842AEF921" authority="(Prokofiev 2013 b)" baseAuthorityName="Prokofiev" baseAuthorityYear="2013" box="[221,673,1644,1670]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="50D80207743FFFEEBA85F98841CFF921" box="[221,448,1644,1670]" italics="true" pageId="5" pageNumber="6">Coilodes castaneus</emphasis>
|
||
(
|
||
<bibRefCitation id="063DA3E4743FFFEEBB97F9884296F921" author="Prokofiev A. M." box="[463,665,1644,1670]" pageId="5" pageNumber="6" pagination="1" refId="ref30485" refString="Prokofiev A. M. 2013 b. Gnombolbus, a synonym of Coilodes (Coleoptera: Scarabaeoidea: Hybosoridae). Calodema 252: 1." type="journal article" year="2013">Prokofiev 2013b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEEBAE5F94A43A6F8A8" blockId="5.[189,1399,1710,1807]" pageId="5" pageNumber="6">
|
||
Finally, in the most recent morphological phyllogeny proposed by
|
||
<bibRefCitation id="063DA3E4743FFFEEB9BBF94A44D9F96F" author="Basilio D. S. & Cherman M. A. & Vaz-de-Mello F. Z. & Almeida L. M." box="[995,1238,1710,1736]" pageId="5" pageNumber="6" pagination="1156 - 1170" refId="ref28944" refString="Basilio D. S., Cherman M. A., Vaz-de-Mello F. Z. & Almeida L. M. 2023. Phylogenetic relationships in Hybosoridae (Coleoptera: Scarabaeoidea). Zoological Journal of the Linnean Society 198 (4): 1156 - 1170. https: // doi. org / 10.1093 / zoolinnean / zlac 095" type="journal article" year="2023">
|
||
Basílio
|
||
<emphasis id="50D80207743FFFEEBE1BF94B448FF96F" box="[1091,1152,1710,1736]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
including all
|
||
<taxonomicName id="A5ACA596743FFFEEBAE7F935415CF94C" authorityName="Erichson" authorityYear="1847" box="[191,339,1745,1771]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Hybosorinae">Hybosorinae</taxonomicName>
|
||
genera,
|
||
<taxonomicName id="A5ACA596743FFFEEBBE8F9354218F94C" authorityName="Westwood" authorityYear="1845" box="[432,535,1745,1771]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBBE8F9354218F94C" box="[432,535,1745,1771]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
was recovered as the sister group of
|
||
<taxonomicName id="A5ACA596743FFFEEB9E1F9354426F94C" box="[953,1065,1745,1771]" class="Insecta" family="Hybosoridae" genus="Frolovius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB9E1F9354426F94C" box="[953,1065,1745,1771]" italics="true" pageId="5" pageNumber="6">Frolovius</emphasis>
|
||
</taxonomicName>
|
||
(a monospecific Neotropical genus), but distant of all other Neotropical
|
||
<taxonomicName id="A5ACA596743FFFEEB8EEF9104343F8A9" authorityName="Erichson" authorityYear="1847" box="[694,844,1780,1806]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Hybosorinae">Hybosorinae</taxonomicName>
|
||
genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E743FFFEDBAE5F8D2431EFE7F" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="6213DE15743FFFEEBAE5F8D241C2F8F7" blockId="5.[189,461,1846,1872]" box="[189,461,1846,1872]" pageId="5" pageNumber="6">
|
||
<heading id="395B6979743FFFEEBAE5F8D241C2F8F7" bold="true" box="[189,461,1846,1872]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="50D80207743FFFEEBAE5F8D241C2F8F7" bold="true" box="[189,461,1846,1872]" pageId="5" pageNumber="6">Taxonomic discussion</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743FFFEDBAE5F881431EFE7F" blockId="5.[189,1400,1893,2025]" lastBlockId="6.[187,1399,265,472]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<taxonomicName id="A5ACA596743FFFEEBAE5F8814128F8D8" authorityName="Westwood" authorityYear="1845" box="[189,295,1893,1919]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBAE5F8814128F8D8" box="[189,295,1893,1919]" italics="true" pageId="5" pageNumber="6">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from other Central and South American genera within
|
||
<taxonomicName id="A5ACA596743FFFEEBED2F881452DF8D8" authorityName="Erichson" authorityYear="1847" box="[1162,1314,1893,1919]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Hybosorinae">Hybosorinae</taxonomicName>
|
||
by the presence of an excavated mandible. This character can be observed in African and Asian genera such as
|
||
<taxonomicName id="A5ACA596743FFFEEBAE5F84F41FAF862" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[189,501,1963,1990]" class="Insecta" family="Hybosoridae" genus="Hybosorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBAE5F84F4134F862" box="[189,315,1963,1989]" italics="true" pageId="5" pageNumber="6">Hybosorus</emphasis>
|
||
MacLeay, 1819
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="50D80207743FFFEEBBA6F84F42A8F862" box="[510,679,1963,1989]" italics="true" pageId="5" pageNumber="6">Phaeochridios</emphasis>
|
||
Lansberge, 1887;
|
||
<taxonomicName id="A5ACA596743FFFEEB921F84F44DBF862" authority="Candeze, 1876" authorityName="Candeze" authorityYear="1876" box="[889,1236,1963,1989]" class="Insecta" family="Hybosoridae" genus="Phaeochroops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEB921F84F4410F862" box="[889,1055,1963,1989]" italics="true" pageId="5" pageNumber="6">Phaeochroops</emphasis>
|
||
Candèze, 1876
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A5ACA596743FFFEEBE85F84F4175F84F" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" class="Insecta" family="Hybosoridae" genus="Phaeochrous" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBE85F84F457AF862" box="[1245,1397,1963,1989]" italics="true" pageId="5" pageNumber="6">Phaeochrous</emphasis>
|
||
<bibRefCitation id="063DA3E4743FFFEEBAE5F82A4175F84F" author="Castelnau & Laporte F. D." box="[189,378,1998,2024]" pageId="5" pageNumber="6" refId="ref29111" refString="Castelnau [Laporte F. D.] 1840. Histoire naturelle des Animaux articules. Volume 2. Dumenil, Paris." type="book" year="1840">Castelnau, 1840</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743FFFEEBBECF82A42F5F84F" authority="Kuijten, 1983" authorityName="Kuijten" authorityYear="1983" box="[436,762,1998,2024]" class="Insecta" family="Hybosoridae" genus="Seleucosorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743FFFEEBBECF82A4240F84F" box="[436,591,1998,2024]" italics="true" pageId="5" pageNumber="6">Seleucosorus</emphasis>
|
||
Kuijten, 1983
|
||
</taxonomicName>
|
||
. This divergence among the Neotropical genera of the subfamily was recovered in the analysis performed with molecular data by
|
||
<bibRefCitation id="063DA3E4743CFFEDBE43FEED454AFE84" author="Ocampo F. C. & Hawks D. C." box="[1051,1349,265,291]" pageId="6" pageNumber="7" pagination="7 - 12" refId="ref30070" refString="Ocampo F. C. & Hawks D. C. 2006. Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae 2. Molecular phylogenetics and systematic placement of the Hybosoridae. Bulletin of the University of Nebraska State Museum 19: 7 - 12." type="journal article" year="2006">Ocampo & Hawks (2006)</bibRefCitation>
|
||
that shows
|
||
<taxonomicName id="A5ACA596743CFFEDBB55FEC9417AFEE0" authorityName="Westwood" authorityYear="1845" box="[269,373,301,327]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBB55FEC9417AFEE0" box="[269,373,301,327]" italics="true" pageId="6" pageNumber="7">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
as more closely related to
|
||
<taxonomicName id="A5ACA596743CFFEDB8F5FEC9435AFEE0" authorityName="Candeze" authorityYear="1876" box="[685,853,301,327]" class="Insecta" family="Hybosoridae" genus="Phaeochroops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDB8F5FEC9435AFEE0" box="[685,853,301,327]" italics="true" pageId="6" pageNumber="7">Phaeochroops</emphasis>
|
||
</taxonomicName>
|
||
than to the Neotropical genus
|
||
<taxonomicName id="A5ACA596743CFFEDBEE4FEC94560FEE0" authorityName="Preudhomme de Borre" authorityYear="1866" box="[1212,1391,301,327]" class="Insecta" family="Hybosoridae" genus="Metachaetodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBEE4FEC94560FEE0" box="[1212,1391,301,327]" italics="true" pageId="6" pageNumber="7">Metachaetodus</emphasis>
|
||
</taxonomicName>
|
||
. In the morphological hypothesis proposed by
|
||
<bibRefCitation id="063DA3E4743CFFEDB88AFEB543B2FECB" author="Basilio D. S. & Cherman M. A. & Vaz-de-Mello F. Z. & Almeida L. M." box="[722,957,337,364]" pageId="6" pageNumber="7" pagination="1156 - 1170" refId="ref28944" refString="Basilio D. S., Cherman M. A., Vaz-de-Mello F. Z. & Almeida L. M. 2023. Phylogenetic relationships in Hybosoridae (Coleoptera: Scarabaeoidea). Zoological Journal of the Linnean Society 198 (4): 1156 - 1170. https: // doi. org / 10.1093 / zoolinnean / zlac 095" type="journal article" year="2023">
|
||
Basílio
|
||
<emphasis id="50D80207743CFFEDB973FEB6436AFECC" box="[811,869,337,363]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
, except for the close relationship with
|
||
<taxonomicName id="A5ACA596743CFFEDBAE4FE924122FE37" box="[188,301,374,400]" class="Insecta" family="Hybosoridae" genus="Frolovius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBAE4FE924122FE37" box="[188,301,374,400]" italics="true" pageId="6" pageNumber="7">Frolovius</emphasis>
|
||
</taxonomicName>
|
||
, the divergence from the other Neotropical genera was also recovered. In that study,
|
||
<taxonomicName id="A5ACA596743CFFEDBF56FE924579FE37" authorityName="Westwood" authorityYear="1845" box="[1294,1398,374,400]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBF56FE924579FE37" box="[1294,1398,374,400]" italics="true" pageId="6" pageNumber="7">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A5ACA596743CFFEDBAA9FE7E416BFE13" box="[241,356,410,436]" class="Insecta" family="Hybosoridae" genus="Frolovius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBAA9FE7E416BFE13" box="[241,356,410,436]" italics="true" pageId="6" pageNumber="7">Frolovius</emphasis>
|
||
</taxonomicName>
|
||
are considered sister groups, closely related to
|
||
<taxonomicName id="A5ACA596743CFFEDB9F2FE7E4447FE13" authorityName="Kuijten" authorityYear="1983" box="[938,1096,410,436]" class="Insecta" family="Hybosoridae" genus="Seleucosorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDB9F2FE7E4447FE13" box="[938,1096,410,436]" italics="true" pageId="6" pageNumber="7">Seleucosorus</emphasis>
|
||
</taxonomicName>
|
||
and the Palearctic genus
|
||
<taxonomicName id="A5ACA596743CFFEDBAE7FE5A4217FE7F" authority="Faimaire, 1893" authorityName="Faimaire" authorityYear="1893" box="[191,536,446,472]" class="Insecta" family="Hybosoridae" genus="Hypseloderus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBAE7FE5A416FFE7F" box="[191,352,446,472]" italics="true" pageId="6" pageNumber="7">Hypseloderus</emphasis>
|
||
Faimaire, 1893
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="063DA3E4743CFFEDB87FFE5A430AFE7F" author="Basilio D. S. & Cherman M. A. & Vaz-de-Mello F. Z. & Almeida L. M." box="[551,773,446,472]" pageId="6" pageNumber="7" pagination="1156 - 1170" refId="ref28944" refString="Basilio D. S., Cherman M. A., Vaz-de-Mello F. Z. & Almeida L. M. 2023. Phylogenetic relationships in Hybosoridae (Coleoptera: Scarabaeoidea). Zoological Journal of the Linnean Society 198 (4): 1156 - 1170. https: // doi. org / 10.1093 / zoolinnean / zlac 095" type="journal article" year="2023">
|
||
Basílio
|
||
<emphasis id="50D80207743CFFEDB8DBFE5B42B1FE7F" box="[643,702,446,472]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2023
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E743CFFEDBAE5FDE542EEFD10" pageId="6" pageNumber="7" type="distribution">
|
||
<paragraph id="6213DE15743CFFEDBAE5FDE5420DFDBC" blockId="6.[189,514,513,539]" box="[189,514,513,539]" pageId="6" pageNumber="7">
|
||
<heading id="395B6979743CFFEDBAE5FDE5420DFDBC" bold="true" box="[189,514,513,539]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
|
||
<emphasis id="50D80207743CFFEDBAE5FDE5420DFDBC" bold="true" box="[189,514,513,539]" pageId="6" pageNumber="7">Geographical distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBAE5FDD442EEFD10" blockId="6.[188,1400,560,696]" pageId="6" pageNumber="7">
|
||
<taxonomicName id="A5ACA596743CFFEDBAE5FDD4412AFDED" authorityName="Westwood" authorityYear="1845" box="[189,293,560,586]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBAE5FDD4412AFDED" box="[189,293,560,586]" italics="true" pageId="6" pageNumber="7">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
occurs in the Neotropical Region, from Central America (
|
||
<collectingCountry id="1ABB9E85743CFFEDB988FDD44442FDED" box="[976,1101,560,586]" name="Nicaragua" pageId="6" pageNumber="7">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBE01FDD544D3FDED" box="[1113,1244,560,587]" name="Costa Rica" pageId="6" pageNumber="7">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBEB0FDD5454AFDEC" box="[1256,1349,561,587]" name="Panama" pageId="6" pageNumber="7">Panama</collectingCountry>
|
||
and the Antilles) to South America (French Guiana,
|
||
<collectingCountry id="1ABB9E85743CFFEDB8A7FDB14373FDC8" box="[767,892,597,623]" name="Venezuela" pageId="6" pageNumber="7">Venezuela</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDB9D1FDB1440EFDC8" box="[905,1025,597,623]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBE57FDB1447BFDC8" box="[1039,1140,597,623]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBED9FDB144CBFDC8" box="[1153,1220,597,623]" name="Chile" pageId="6" pageNumber="7">Chile</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBE89FDB14505FDC8" box="[1233,1290,597,623]" name="Peru" pageId="6" pageNumber="7">Peru</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBF40FDB1457DFDC8" box="[1304,1394,597,623]" name="Bolivia" pageId="6" pageNumber="7">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBAE7FD9D4105FD34" box="[191,266,633,659]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBB4CFD9E418EFD33" box="[276,385,634,660]" name="Paraguay" pageId="6" pageNumber="7">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743CFFEDBBE0FD9D423DFD34" box="[440,562,633,659]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
|
||
). However, the record from
|
||
<collectingCountry id="1ABB9E85743CFFEDB921FD9D43B6FD34" box="[889,953,633,659]" name="Chile" pageId="6" pageNumber="7">Chile</collectingCountry>
|
||
is doubtful. No records are yet known from
|
||
<collectingCountry id="1ABB9E85743CFFEDBAA5FD794154FD10" box="[253,347,669,695]" name="Guyana" pageId="6" pageNumber="7">Guyana</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBB3FFD7941D5FD10" box="[359,474,669,695]" name="Suriname" pageId="6" pageNumber="7">Suriname</collectingCountry>
|
||
, or
|
||
<collectingCountry id="1ABB9E85743CFFEDB85EFD7A4262FD1F" box="[518,621,670,696]" name="Uruguay" pageId="6" pageNumber="7">Uruguay</collectingCountry>
|
||
(
|
||
<figureCitation id="FA97C290743CFFEDB824FD7942DBFD10" box="[636,724,669,695]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1739]" captionTargetId="figure-12@47.[272,1315,265,1739]" captionTargetPageId="47" captionText="Fig. 14. Distribution map of species of Coilodes Westwood, 1846. A. Coilodes bezerrai Basílio & Vazde-Mello sp. nov.; Coilodes castaneus Westwood, 1846; and Coilodes edeiltae Basílio & Vaz-de-Mello sp. nov. B. Coilodes fumipennis Arrow, 1909; Coilodes humeralis (Mannerheim, 1829); and Coilodes lunae Basílio & Vaz-de-Mello sp. nov. C. Coilodes mayae Basílio & Vaz-de-Mello sp. nov.;Coilodes niger (Mannerheim, 1829) stat. rev.; Coilodes ovalis Robinson, 1948; and Coilodes parvulus Westwood 1846. D. Coilodes punctipennis Arrow, 1909; Coilodes ravii Basílio & Vaz-de-Mello sp. nov.; and Coilodes skelleyi Basílio & Vaz-de-Mello sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10407111" httpUri="https://zenodo.org/record/10407111/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E743CFFEDBAE5FD0443DAFB33" pageId="6" pageNumber="7" type="biology_ecology">
|
||
<paragraph id="6213DE15743CFFEDBAE5FD044178FD5D" blockId="6.[189,375,736,762]" box="[189,375,736,762]" pageId="6" pageNumber="7">
|
||
<heading id="395B6979743CFFEDBAE5FD044178FD5D" bold="true" box="[189,375,736,762]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
|
||
<emphasis id="50D80207743CFFEDBAE5FD044178FD5D" bold="true" box="[189,375,736,762]" pageId="6" pageNumber="7">Biological data</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBAE5FCEB43DAFB33" blockId="6.[188,1401,783,1173]" pageId="6" pageNumber="7">
|
||
<taxonomicName id="A5ACA596743CFFEDBAE5FCEB412AFC8E" authorityName="Westwood" authorityYear="1845" box="[189,293,783,809]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBAE5FCEB412AFC8E" box="[189,293,783,809]" italics="true" pageId="6" pageNumber="7">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
usually presents copronecrophagous habits. It is commonly collected using vertebrate faeces or meat in bait traps. Some authors reported the association of adults with carcasses of many kinds of vertebrate, including mammals (monkey, sloth, pig, hedgehog, opossum, rabbit, and mouse), birds (chicken), reptiles (lizard), amphibians (frog), and fishes (
|
||
<bibRefCitation id="063DA3E4743CFFEDB914FC9843FBFC30" author="Cornaby B. W." box="[844,1012,892,919]" pageId="6" pageNumber="7" pagination="51 - 63" refId="ref29134" refString="Cornaby B. W. 1974. Carrion reduction by animals in contrasting tropical habitats. Biotropica 6 (1): 51 - 63. https: // doi. org / 10.2307 / 2989697" type="journal article" year="1974">Cornaby 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="063DA3E4743CFFEDB9A6FC98452CFC30" author="Wehncke E. V. & Dalling J. W." box="[1022,1315,892,919]" pageId="6" pageNumber="7" pagination="73 - 80" refId="ref30980" refString="Wehncke E. V. & Dalling J. W. 2005. Post-dispersal seed removal and germination selected tree species dispersed by Cebus capucinus on Barro Colorado Island, Panama. Biotropica 37 (1): 73 - 80. https: // doi. org / 10.1111 / j. 1744 - 7429.2005.03037. x" type="journal article" year="2005">Wehncke & Dalling 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="063DA3E4743CFFEDBF75FC9940F5FC1C" author="Young O. P." pageId="6" pageNumber="7" pagination="247 - 253" refId="ref31099" refString="Young O. P. 1983. The distribution and ecology of Coilodes castanea (Coleoptera: Scarabaeidae: Hybosorinae). The Coleopterists Bulletin 37 (3): 247 - 253. Available from https: // www. jstor. org / stable / 4008026 [accessed 23 Nov. 2023]." type="journal article" year="1983">Young 1983</bibRefCitation>
|
||
; label data, personal observation). This association mostly occurs in the first stages of decomposition. In addition, specimens were collected in carcasses of invertebrates (ant, shrimp, squid, and earthworm), in fungi, and a few times in decomposing fruit (
|
||
<bibRefCitation id="063DA3E4743CFFEDB887FC0D437DFBA4" author="Young O. P." box="[735,882,1001,1027]" pageId="6" pageNumber="7" pagination="247 - 253" refId="ref31099" refString="Young O. P. 1983. The distribution and ecology of Coilodes castanea (Coleoptera: Scarabaeidae: Hybosorinae). The Coleopterists Bulletin 37 (3): 247 - 253. Available from https: // www. jstor. org / stable / 4008026 [accessed 23 Nov. 2023]." type="journal article" year="1983">Young 1983</bibRefCitation>
|
||
; label data). They are considered good flyers (
|
||
<bibRefCitation id="063DA3E4743CFFEDBA9DFBE94184FB8F" author="Lacordaire J. T." box="[197,395,1037,1064]" pageId="6" pageNumber="7" refId="ref29690" refString="Lacordaire J. T. 1856. Histoire naturelle des Insectes: Genera des Coleopteres ou Expose methodique et critique de tous les Genres proposes jusqu'ici dans cet Ordre d'Insectes. Tome 3. Contenant les Familles des Pectinicornes et Lamellicornes. Librairie encyclopedique de Roret, Paris." type="book" year="1856">Lacordaire 1856</bibRefCitation>
|
||
), with diffuse distribution and diurnal habits (
|
||
<bibRefCitation id="063DA3E4743CFFEDB9F8FBEA443AFB8F" author="Young O. P." box="[928,1077,1038,1064]" pageId="6" pageNumber="7" pagination="247 - 253" refId="ref31099" refString="Young O. P. 1983. The distribution and ecology of Coilodes castanea (Coleoptera: Scarabaeidae: Hybosorinae). The Coleopterists Bulletin 37 (3): 247 - 253. Available from https: // www. jstor. org / stable / 4008026 [accessed 23 Nov. 2023]." type="journal article" year="1983">Young 1983</bibRefCitation>
|
||
). There are records of adult specimens of
|
||
<taxonomicName id="A5ACA596743CFFEDBB3AFBD641C5FBEB" authorityName="Westwood" authorityYear="1845" box="[354,458,1074,1100]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="50D80207743CFFEDBB3AFBD641C5FBEB" box="[354,458,1074,1100]" italics="true" pageId="6" pageNumber="7">Coilodes</emphasis>
|
||
</taxonomicName>
|
||
collected using light traps, which suggests attraction to light (label data). They also bury themselves quickly when threatened (personal observation). In addition, stridulating behaviour has been observed in adults of
|
||
<taxonomicName id="A5ACA596743CFFEDB87FFB9F42B1FB33" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[551,702,1146,1173]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743CFFEDB87FFB9F42B1FB33" box="[551,702,1146,1173]" italics="true" pageId="6" pageNumber="7">C. humeralis</emphasis>
|
||
</taxonomicName>
|
||
(personal observation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2AB68D9E743CFFECBAE5FB594578FABE" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="key">
|
||
<paragraph id="6213DE15743CFFEDBAE5FB5942FBFB70" blockId="6.[189,756,1212,1239]" box="[189,756,1212,1239]" pageId="6" pageNumber="7">
|
||
<heading id="395B6979743CFFEDBAE5FB5942FBFB70" bold="true" box="[189,756,1212,1239]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
|
||
<emphasis id="50D80207743CFFEDBAE5FB5942FBFB70" bold="true" box="[189,756,1212,1239]" pageId="6" pageNumber="7">
|
||
Identification key to males of
|
||
<taxonomicName id="A5ACA596743CFFEDB876FB5842FBFB70" box="[558,756,1212,1239]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undefined-1">
|
||
<emphasis id="50D80207743CFFEDB876FB584299FB70" bold="true" box="[558,662,1212,1239]" italics="true" pageId="6" pageNumber="7">Coilodes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="163D8924743CFFECBAE5FB094578FABE" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<keyStep id="D958C6B0743CFFEDBAE5FB094578FA8C" pageId="6" pageNumber="7">
|
||
<paragraph id="6213DE15743CFFEDBAE5FB094578FAA0" blockId="6.[189,1399,1260,1323]" box="[189,1399,1260,1287]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBAE5FB094578FAA0" box="[189,1399,1260,1287]" pageId="6" pageNumber="7">
|
||
1. Aedeagus with dorsal lobe of tegmen (
|
||
<figureCitation id="FA97C290743CFFEDB8E8FB08434CFAA0" box="[688,835,1260,1287]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1397,698,1750]" captionTargetId="figure-14@24.[189,1398,697,1784]" captionTargetPageId="24" captionText="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407089" httpUri="https://zenodo.org/record/10407089/files/figure.png" pageId="6" pageNumber="7">Fig. 5F, J–K</figureCitation>
|
||
) ........................................................................ 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBA99FAF54578FA8C" blockId="6.[189,1399,1260,1323]" box="[193,1399,1297,1323]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBA99FAF54578FA8C" box="[193,1399,1297,1323]" pageId="6" pageNumber="7">
|
||
– Aedeagus without dorsal lobe of tegmen (
|
||
<figureCitation id="FA97C290743CFFEDB88DFAF54366FA8C" box="[725,873,1297,1323]" captionStart="Fig" captionStartId="18.[189,232,1787,1813]" captionTargetBox="[189,1398,661,1744]" captionTargetId="figure-15@18.[189,1398,660,1748]" captionTargetPageId="18" captionText="Fig. 4. Coilodes fumipennis Arrow, 1909, ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (A, C, E), ♂ at CERPE from Rondônia, Brazil (B), ♂ at CERPE from Alta Floresta, Mato Grosso, Brazil (D); ♂ at CEMT from Rondônia, Brazil (F–G), graphical representation (H–I), and another ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407085" httpUri="https://zenodo.org/record/10407085/files/figure.png" pageId="6" pageNumber="7">Fig. 4F, J–K</figureCitation>
|
||
) ................................................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743CFFEDBAE5FAB74578FA35" pageId="6" pageNumber="7">
|
||
<paragraph id="6213DE15743CFFEDBAE5FAB74578FACA" blockId="6.[189,1399,1363,1426]" box="[189,1399,1363,1390]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBAE5FAB74578FACA" box="[189,1399,1363,1390]" pageId="6" pageNumber="7">
|
||
2. Aedeagus with the dorsal lobe base in the middle of tegmen (
|
||
<figureCitation id="FA97C290743CFFEDB9E1FAB74428FACA" box="[953,1063,1363,1390]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1397,698,1750]" captionTargetId="figure-14@24.[189,1398,697,1784]" captionTargetPageId="24" captionText="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407089" httpUri="https://zenodo.org/record/10407089/files/figure.png" pageId="6" pageNumber="7">Fig. 5F, J</figureCitation>
|
||
) ......................................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBA99FA9C4578FA35" blockId="6.[189,1399,1363,1426]" box="[193,1399,1399,1426]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBA99FA9C4578FA35" box="[193,1399,1399,1426]" pageId="6" pageNumber="7">
|
||
– Aedeagus with the dorsal lobe base on the right side of tegmen (
|
||
<figureCitation id="FA97C290743CFFEDB984FA9C4457FA35" box="[988,1112,1400,1426]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="6" pageNumber="7">Fig. 12F, J</figureCitation>
|
||
) .................................. 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743CFFEDBAE5FA5E4578F9C1" pageId="6" pageNumber="7">
|
||
<paragraph id="6213DE15743CFFEDBAE5FA5E4578FA5E" blockId="6.[189,1399,1466,1638]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBAE5FA5E4578FA5E" pageId="6" pageNumber="7">
|
||
3. Clypeus subtrapezoidal in shape with angulate ends (
|
||
<figureCitation id="FA97C290743CFFEDB914FA5E43A9FA72" box="[844,934,1466,1493]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1397,698,1750]" captionTargetId="figure-14@24.[189,1398,697,1784]" captionTargetPageId="24" captionText="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407089" httpUri="https://zenodo.org/record/10407089/files/figure.png" pageId="6" pageNumber="7">Fig. 5C</figureCitation>
|
||
); lobe of tegmen long, reaching the base of the parameres (
|
||
<figureCitation id="FA97C290743CFFEDBBE4FA3A425FFA5E" box="[444,592,1502,1529]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1397,698,1750]" captionTargetId="figure-14@24.[189,1398,697,1784]" captionTargetPageId="24" captionText="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407089" httpUri="https://zenodo.org/record/10407089/files/figure.png" pageId="6" pageNumber="7">Fig. 5F, J–K</figureCitation>
|
||
) ......................................................................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBA99F9E74578F9C1" blockId="6.[189,1399,1466,1638]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBA99F9E74578F9C1" pageId="6" pageNumber="7">
|
||
– Clypeus subrectangular in shape with rounded ends (
|
||
<figureCitation id="FA97C290743CFFEDB924F9E743D1F9BA" box="[892,990,1539,1565]" captionStart="Fig" captionStartId="32.[189,232,1822,1848]" captionTargetBox="[189,1398,698,1768]" captionTargetId="figure-14@32.[189,1398,697,1784]" captionTargetPageId="32" captionText="Fig. 8. Coilodes niger (Mannerheim, 1829) stat. rev., two ♂♂ from Macaé de Cima, Nova Friburgo, Rio de Janeiro, Brazil, at CERPE (A–E) and at CEMT (F–G); another ♂ at CEMT from Nova Friburgo, Rio de Janeiro, Brazil (J–K); and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407097" httpUri="https://zenodo.org/record/10407097/files/figure.png" pageId="6" pageNumber="7">Fig. 8C</figureCitation>
|
||
); lobe of tegmen short, less than one half the distance between the base of the lobe and the base of the parameres (
|
||
<figureCitation id="FA97C290743CFFEDBE9EF9C34566F9E6" box="[1222,1385,1575,1602]" captionStart="Fig" captionStartId="32.[189,232,1822,1848]" captionTargetBox="[189,1398,698,1768]" captionTargetId="figure-14@32.[189,1398,697,1784]" captionTargetPageId="32" captionText="Fig. 8. Coilodes niger (Mannerheim, 1829) stat. rev., two ♂♂ from Macaé de Cima, Nova Friburgo, Rio de Janeiro, Brazil, at CERPE (A–E) and at CEMT (F–G); another ♂ at CEMT from Nova Friburgo, Rio de Janeiro, Brazil (J–K); and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407097" httpUri="https://zenodo.org/record/10407097/files/figure.png" pageId="6" pageNumber="7">Fig. 8F, J– K</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743CFFEDBAB4F9AF413DF9C2" box="[236,306,1611,1637]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
|
||
............................................................................
|
||
<taxonomicName id="A5ACA596743CFFEDB93CF9AF4502F9C2" ID-CoL="3MYST" authority="(Mannerheim, 1929)" authorityName="Basílio & Vaz-De-Mello & Cherman & Almeida" authorityYear="2023" baseAuthorityName="Mannerheim" baseAuthorityYear="1929" box="[868,1293,1611,1638]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="niger" status="stat. nov.">
|
||
<emphasis id="50D80207743CFFEDB93CF9AF441CF9C2" bold="true" box="[868,1043,1611,1638]" italics="true" pageId="6" pageNumber="7">Coilodes niger</emphasis>
|
||
(Mannerheim, 1929)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4BEBBF7C743CFFEDBF4CF9A84578F9C1" box="[1300,1399,1612,1638]" pageId="6" pageNumber="7">stat. rev.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743CFFEDBAE5F96A4578F8F9" pageId="6" pageNumber="7">
|
||
<paragraph id="6213DE15743CFFEDBAE5F96A4579F957" blockId="6.[189,1399,1678,1886]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBAE5F96A4579F957" pageId="6" pageNumber="7">
|
||
4. Pronotum subelliptical with lateral margins strongly rounded and posterior margin as wide as the anterior one (
|
||
<figureCitation id="FA97C290743CFFEDBBD6F95641E3F96A" box="[398,492,1714,1741]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1397,698,1750]" captionTargetId="figure-14@24.[189,1398,697,1784]" captionTargetPageId="24" captionText="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407089" httpUri="https://zenodo.org/record/10407089/files/figure.png" pageId="6" pageNumber="7">Fig. 5E</figureCitation>
|
||
); colour ranging from dark brown to black (
|
||
<figureCitation id="FA97C290743CFFEDBE5CF956446AF96A" box="[1028,1125,1714,1741]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1397,698,1750]" captionTargetId="figure-14@24.[189,1398,697,1784]" captionTargetPageId="24" captionText="Fig. 5. Coilodes humeralis (Mannerheim, 1829), ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (A, D), four different ♂♂ at DZUP from Curitiba, Brazil (B–C, E–G); graphical representation (H–I), and another ♂ at DZUP from Baixo Gandu, Espírito Santo, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407089" httpUri="https://zenodo.org/record/10407089/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743CFFEDBE24F95644C7F96B" box="[1148,1224,1714,1740]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743CFFEDBE8DF957454BF96A" box="[1237,1348,1715,1741]" name="Paraguay" pageId="6" pageNumber="7">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743CFFEDBAB4F932416EF957" box="[236,353,1750,1776]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
|
||
...........................................................................
|
||
<taxonomicName id="A5ACA596743CFFEDB9CAF9324579F957" ID-CoL="WWJ2" authority="(Mannerheim, 1929)" baseAuthorityName="Mannerheim" baseAuthorityYear="1929" box="[914,1398,1750,1777]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="50D80207743CFFEDB9CAF9324473F956" bold="true" box="[914,1148,1750,1777]" italics="true" pageId="6" pageNumber="7">Coilodes humeralis</emphasis>
|
||
(Mannerheim, 1929)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBA99F91F4578F8F9" blockId="6.[189,1399,1678,1886]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBA99F91F4578F8F9" pageId="6" pageNumber="7">
|
||
– Pronotum subtrapezoidal in shape, with lateral margins straight and posterior margin wider than anterior one (
|
||
<figureCitation id="FA97C290743CFFEDBBD1F8FB41EBF89E" box="[393,484,1823,1849]" captionStart="Fig" captionStartId="15.[189,233,1892,1918]" captionTargetBox="[190,1398,774,1820]" captionTargetId="figure-11@15.[189,1398,768,1855]" captionTargetPageId="15" captionText="Fig. 3. Coilodes edeiltae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407081" httpUri="https://zenodo.org/record/10407081/files/figure.png" pageId="6" pageNumber="7">Fig. 3E</figureCitation>
|
||
); yellowish brown in colour (
|
||
<figureCitation id="FA97C290743CFFEDB966F8FB4392F89E" box="[830,925,1823,1849]" captionStart="Fig" captionStartId="15.[189,233,1892,1918]" captionTargetBox="[190,1398,774,1820]" captionTargetId="figure-11@15.[189,1398,768,1855]" captionTargetPageId="15" captionText="Fig. 3. Coilodes edeiltae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407081" httpUri="https://zenodo.org/record/10407081/files/figure.png" pageId="6" pageNumber="7">Fig. 3A</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743CFFEDB9EAF8FB43F5F89E" box="[946,1018,1823,1849]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
|
||
................................................... .............................................................................
|
||
<taxonomicName id="A5ACA596743CFFEDB970F8A74440F8FA" authority="Basilio" authorityName="Basilio" box="[808,1103,1859,1886]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="edeiltae">
|
||
<emphasis id="50D80207743CFFEDB970F8A743FBF8F9" bold="true" box="[808,1012,1859,1886]" italics="true" pageId="6" pageNumber="7">Coilodes edeiltae</emphasis>
|
||
Basílio
|
||
</taxonomicName>
|
||
& Vaz-de-Mello sp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743CFFEDBAE5F8624578F84E" pageId="6" pageNumber="7">
|
||
<paragraph id="6213DE15743CFFEDBAE5F8624578F807" blockId="6.[189,1406,1926,2025]" box="[189,1399,1926,1953]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBAE5F8624578F807" box="[189,1399,1926,1953]" pageId="6" pageNumber="7">
|
||
5. Lobe of tegmen with apex sharp (
|
||
<figureCitation id="FA97C290743CFFEDB82DF86242D9F807" box="[629,726,1926,1952]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="6" pageNumber="7">Fig. 12J</figureCitation>
|
||
) ....................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743CFFEDBA99F84F4578F84E" blockId="6.[189,1406,1926,2025]" pageId="6" pageNumber="7">
|
||
<keyLead id="D95D7D20743CFFEDBA99F84F4578F84E" pageId="6" pageNumber="7">
|
||
– Lobe of tegmen with apex rounded (
|
||
<figureCitation id="FA97C290743CFFEDB8C7F84E430CF863" box="[671,771,1962,1988]" captionStart="Fig" captionStartId="45.[189,232,1822,1848]" captionTargetBox="[189,1398,698,1763]" captionTargetId="figure-11@45.[189,1398,697,1784]" captionTargetPageId="45" captionText="Fig. 13. Coilodes skelleyi Basílio & Vaz-de-Mello sp. nov., paratype ♂ at CEMT from Pico da Neblina, Amazonas, Brazil (A, F–G), paratype ♀ at DZUP from Loreto, Peru (B), paratype ♂ at DZUP from Benjamin Constant, Amazonas, Brazil (C, E), holotype ♂ (D, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407109" httpUri="https://zenodo.org/record/10407109/files/figure.png" pageId="6" pageNumber="7">Fig. 13J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743CFFEDB942F84E436DF863" box="[794,866,1962,1988]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743CFFEDB9C5F84F43DCF862" box="[925,979,1963,1989]" name="Peru" pageId="6" pageNumber="7">Peru</collectingCountry>
|
||
.......................................................... .............................................................................
|
||
<taxonomicName id="A5ACA596743CFFEDB973F82A4440F84F" authority="Basilio" authorityName="Basilio" box="[811,1103,1998,2025]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="skelleyi">
|
||
<emphasis id="50D80207743CFFEDB973F82A43FAF84E" bold="true" box="[811,1013,1998,2025]" italics="true" pageId="6" pageNumber="7">Coilodes skelleyi</emphasis>
|
||
Basílio
|
||
</taxonomicName>
|
||
& Vaz-de-Mello sp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FEED4578FE37" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FEED4578FEEF" blockId="7.[189,1414,265,400]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FEED4578FEEF" pageId="7" pageNumber="8">
|
||
6. Lobe of tegmen long, reaching the base of the parameres (
|
||
<figureCitation id="FA97C290743DFFECB9FBFEED4407FE84" box="[931,1032,265,291]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="7" pageNumber="8">Fig. 12J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECBE46FEED448FFE84" box="[1054,1152,265,291]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
|
||
................................... ..................................................................................
|
||
<taxonomicName id="A5ACA596743DFFECB915FEC94440FEE0" authority="Basilio" authorityName="Basilio" box="[845,1103,301,328]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ravii">
|
||
<emphasis id="50D80207743DFFECB915FEC943FBFEE0" bold="true" box="[845,1012,301,328]" italics="true" pageId="7" pageNumber="8">Coilodes ravii</emphasis>
|
||
Basílio
|
||
</taxonomicName>
|
||
& Vaz-de-Mello sp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FEB64578FE37" blockId="7.[189,1414,265,400]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FEB64578FE37" pageId="7" pageNumber="8">
|
||
– Lobe of tegmen short, less than one half the distance between the base of the lobe and the base of the parameres (
|
||
<figureCitation id="FA97C290743DFFECBB2AFE9141CBFE37" box="[370,452,373,400]" captionStart="Fig" captionStartId="8.[189,232,1857,1883]" captionTargetBox="[189,1398,730,1795]" captionTargetId="figure-14@8.[189,1398,729,1816]" captionTargetPageId="8" captionText="Fig. 1. Coilodes bezerrai Basílio & Vaz-de-Mello sp. nov., paratype ♂ at DZUP (A, B, D), holotype ♂ (C, E–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407073" httpUri="https://zenodo.org/record/10407073/files/figure.png" pageId="7" pageNumber="8">Fig. 1J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECBB83FE914240FE28" box="[475,591,373,399]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
|
||
............................
|
||
<taxonomicName id="A5ACA596743DFFECB97BFE914440FE28" authority="Basilio" authorityName="Basilio & Vazde-Mello" authorityYear="2023" box="[803,1103,373,400]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bezerrai">
|
||
<emphasis id="50D80207743DFFECB97BFE9143FBFE37" bold="true" box="[803,1012,373,400]" italics="true" pageId="7" pageNumber="8">Coilodes bezerrai</emphasis>
|
||
Basílio
|
||
</taxonomicName>
|
||
& Vaz-de-Mello sp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FE5C4579FE51" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FE5C4578FE75" blockId="7.[189,1399,440,503]" box="[189,1399,440,466]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FE5C4578FE75" box="[189,1399,440,466]" pageId="7" pageNumber="8">
|
||
7. Right paramere with distinctly rounded apex (
|
||
<figureCitation id="FA97C290743DFFECB95EFE5C4373FE75" box="[774,892,440,466]" captionStart="Fig" captionStartId="18.[189,232,1787,1813]" captionTargetBox="[189,1398,661,1744]" captionTargetId="figure-15@18.[189,1398,660,1748]" captionTargetPageId="18" captionText="Fig. 4. Coilodes fumipennis Arrow, 1909, ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (A, C, E), ♂ at CERPE from Rondônia, Brazil (B), ♂ at CERPE from Alta Floresta, Mato Grosso, Brazil (D); ♂ at CEMT from Rondônia, Brazil (F–G), graphical representation (H–I), and another ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407085" httpUri="https://zenodo.org/record/10407085/files/figure.png" pageId="7" pageNumber="8">Fig. 4H, J</figureCitation>
|
||
) ................................................................ 8
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FE394579FE51" blockId="7.[189,1399,440,503]" box="[193,1398,476,503]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FE394579FE51" box="[193,1398,476,503]" pageId="7" pageNumber="8">
|
||
– Right paramere with straight or almost straight apex (
|
||
<figureCitation id="FA97C290743DFFECB906FE3843DBFE51" box="[862,980,476,502]" captionStart="Fig" captionStartId="27.[189,232,1857,1883]" captionTargetBox="[189,1398,731,1793]" captionTargetId="figure-196@27.[189,1398,731,1819]" captionTargetPageId="27" captionText="Fig. 6. Coilodes lunae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–C, E–G, J–K), paratype ♀ at DZUP from Itatiaia, Rio de Janeiro (D), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407093" httpUri="https://zenodo.org/record/10407093/files/figure.png" pageId="7" pageNumber="8">Fig. 6H, J</figureCitation>
|
||
) .................................................. 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FDFB4578FD26" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FDFB4578FD9E" blockId="7.[189,1399,542,641]" box="[189,1399,542,569]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FDFB4578FD9E" box="[189,1399,542,569]" pageId="7" pageNumber="8">
|
||
8. Left paramere placed ventrally (
|
||
<figureCitation id="FA97C290743DFFECB83BFDFA42FEFD9E" box="[611,753,542,569]" captionStart="Fig" captionStartId="40.[189,232,1822,1848]" captionTargetBox="[189,1398,703,1775]" captionTargetId="figure-14@40.[189,1398,695,1784]" captionTargetPageId="40" captionText="Fig. 11. Coilodes punctipennis Arrow, 1909, ♂ at UCCC from Tropica Region Chapare, Bolivia (A, C–E), ♂ at CEMT from Cotococha, Napo, Ecuador (B), ♂ at DZUP from Tugurahua, Ecuador (F–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407103" httpUri="https://zenodo.org/record/10407103/files/figure.png" pageId="7" pageNumber="8">Fig. 11G, K</figureCitation>
|
||
).................................................................................... 9
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FDA74578FD26" blockId="7.[189,1399,542,641]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FDA74578FD26" pageId="7" pageNumber="8">
|
||
– Left paramere fully located on the left side (
|
||
<figureCitation id="FA97C290743DFFECB94DFDA643AEFDFA" box="[789,929,578,605]" captionStart="Fig" captionStartId="35.[189,232,1857,1883]" captionTargetBox="[197,1398,733,1807]" captionTargetId="figure-231@35.[189,1398,732,1819]" captionTargetPageId="35" captionText="Fig. 9. Coilodes ovalis Robinson, 1948, three ♂♂ from Jaji, Sucre, Venezuela, two at CEMT (first A, C–D; and second B, E), and one at DZUP (F–G, J–K); and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view.C. Head, dorsal view. D. Head, ventral view.E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407099" httpUri="https://zenodo.org/record/10407099/files/figure.png" pageId="7" pageNumber="8">Fig. 9G, K</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECB9E0FDA6443DFDFB" box="[952,1074,578,604]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECBE19FDA6454EFDFB" box="[1089,1345,578,604]" name="Trinidad and Tobago" pageId="7" pageNumber="8">Trinidad and Tobago</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743DFFECBAB2FD83416CFD26" box="[234,355,615,641]" name="Venezuela" pageId="7" pageNumber="8">Venezuela</collectingCountry>
|
||
..........................................................................................
|
||
<taxonomicName id="A5ACA596743DFFECBE58FD824578FD26" ID-CoL="WWJ4" authority="Robinson, 1948" authorityName="Robinson" authorityYear="1948" box="[1024,1399,614,641]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="50D80207743DFFECBE58FD8244BAFD26" bold="true" box="[1024,1205,614,641]" italics="true" pageId="7" pageNumber="8">Coilodes ovalis</emphasis>
|
||
<bibRefCitation id="063DA3E4743DFFECBEE5FD834578FD26" author="Robinson M." box="[1213,1399,615,641]" pageId="7" pageNumber="8" pagination="29 - 36" refId="ref30632" refString="Robinson M. 1948. Studies in the Scarabaeidae IV (Coleoptera). Transactions of the American Entomological Society 74 (1): 29 - 36. Available from https: // www. jstor. org / stable / 25077572 [accessed 23 Nov. 2023]." type="journal article" year="1948">Robinson, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FD4D4578FC97" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FD4D4578FD40" blockId="7.[189,1399,681,816]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FD4D4578FD40" pageId="7" pageNumber="8">
|
||
9. Tegmen with small, sharp projection next to the base of the right paramere (
|
||
<figureCitation id="FA97C290743DFFECBE1AFD4D449BFD64" box="[1090,1172,681,707]" captionStart="Fig" captionStartId="18.[189,232,1787,1813]" captionTargetBox="[189,1398,661,1744]" captionTargetId="figure-15@18.[189,1398,660,1748]" captionTargetPageId="18" captionText="Fig. 4. Coilodes fumipennis Arrow, 1909, ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (A, C, E), ♂ at CERPE from Rondônia, Brazil (B), ♂ at CERPE from Alta Floresta, Mato Grosso, Brazil (D); ♂ at CEMT from Rondônia, Brazil (F–G), graphical representation (H–I), and another ♂ at CEMT from Alta Floresta, Mato Grosso, Brazil (J–K). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407085" httpUri="https://zenodo.org/record/10407085/files/figure.png" pageId="7" pageNumber="8">Fig. 4J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECBEFFFD4D4513FD64" box="[1191,1308,681,707]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECBF7DFD4D4131FD40" name="French Guiana" pageId="7" pageNumber="8">French Guiana</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743DFFECBB2EFD2941B1FD40" box="[374,446,717,743]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
|
||
.........................................................................
|
||
<taxonomicName id="A5ACA596743DFFECB9BBFD294578FD40" ID-CoL="WWHY" authority="Arrow, 1909" authorityName="Arrow" authorityYear="1909" box="[995,1399,717,744]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fumipennis">
|
||
<emphasis id="50D80207743DFFECB9BBFD2944D4FD40" bold="true" box="[995,1243,717,744]" italics="true" pageId="7" pageNumber="8">Coilodes fumipennis</emphasis>
|
||
<bibRefCitation id="063DA3E4743DFFECBEBAFD2A4578FD40" author="Arrow G. J." box="[1250,1399,717,744]" pageId="7" pageNumber="8" pagination="479 - 507" refId="ref28738" refString="Arrow G. J. 1909. On the characters and relationships of the less know groups of lamellicorn Coleoptera, with description of a new species of Hybosorinae, etc. Transactions of the Entomological Society of London 2: 479 - 507." type="journal article" year="1909">Arrow, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FD164578FC97" blockId="7.[189,1399,681,816]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FD164578FC97" pageId="7" pageNumber="8">
|
||
– Tegmen without a sharp projection and with a more sclerotized region next to the right paramere base (
|
||
<figureCitation id="FA97C290743DFFECBAAAFCF2415BFC97" box="[242,340,790,816]" captionStart="Fig" captionStartId="40.[189,232,1822,1848]" captionTargetBox="[189,1398,703,1775]" captionTargetId="figure-14@40.[189,1398,695,1784]" captionTargetPageId="40" captionText="Fig. 11. Coilodes punctipennis Arrow, 1909, ♂ at UCCC from Tropica Region Chapare, Bolivia (A, C–E), ♂ at CEMT from Cotococha, Napo, Ecuador (B), ♂ at DZUP from Tugurahua, Ecuador (F–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407103" httpUri="https://zenodo.org/record/10407103/files/figure.png" pageId="7" pageNumber="8">Fig. 11J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECBB30FCF241C3FC97" box="[360,460,790,816]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECBB80FCF24201FC97" box="[472,526,790,816]" name="Peru" pageId="7" pageNumber="8">Peru</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743DFFECB81FFCF24292FC97" box="[583,669,790,816]" name="Bolivia" pageId="7" pageNumber="8">Bolivia</collectingCountry>
|
||
.........................................
|
||
<taxonomicName id="A5ACA596743DFFECB997FCF14578FC97" ID-CoL="WWJ6" authority="Arrow, 1909" authorityName="Arrow" authorityYear="1909" box="[975,1399,789,816]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctipennis">
|
||
<emphasis id="50D80207743DFFECB997FCF144D4FC97" bold="true" box="[975,1243,789,816]" italics="true" pageId="7" pageNumber="8">Coilodes punctipennis</emphasis>
|
||
<bibRefCitation id="063DA3E4743DFFECBEBAFCF24578FC97" author="Arrow G. J." box="[1250,1399,790,816]" pageId="7" pageNumber="8" pagination="479 - 507" refId="ref28738" refString="Arrow G. J. 1909. On the characters and relationships of the less know groups of lamellicorn Coleoptera, with description of a new species of Hybosorinae, etc. Transactions of the Entomological Society of London 2: 479 - 507." type="journal article" year="1909">Arrow, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FCBC4578FC1D" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FCBC4579FCD5" blockId="7.[189,1399,856,955]" box="[189,1398,856,883]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FCBC4579FCD5" box="[189,1398,856,883]" pageId="7" pageNumber="8">
|
||
10. Right paramere with at least one of the lateral margins arched (
|
||
<figureCitation id="FA97C290743DFFECB995FCBC444DFCD5" box="[973,1090,856,882]" captionStart="Fig" captionStartId="27.[189,232,1857,1883]" captionTargetBox="[189,1398,731,1793]" captionTargetId="figure-196@27.[189,1398,731,1819]" captionTargetPageId="27" captionText="Fig. 6. Coilodes lunae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–C, E–G, J–K), paratype ♀ at DZUP from Itatiaia, Rio de Janeiro (D), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407093" httpUri="https://zenodo.org/record/10407093/files/figure.png" pageId="7" pageNumber="8">Fig. 6H, J</figureCitation>
|
||
) ....................................11
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FC994578FC1D" blockId="7.[189,1399,856,955]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FC994578FC1D" pageId="7" pageNumber="8">
|
||
– Right paramere with both lateral margins straight (
|
||
<figureCitation id="FA97C290743DFFECB96CFC9843A6FC31" box="[820,937,892,918]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[194,1397,736,1817]" captionTargetId="figure-92@10.[189,1398,731,1819]" captionTargetPageId="10" captionText="Fig. 2. Coilodes castaneus Westwood, 1846, ♂ at CEMT from Cerro Campana, Panama (A, C–E), ♂ at DZUP from Puntarenas, Costa Rica (B), ♂ at DZUP from Canal Zone, Panamá (F–G, J–K), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407075" httpUri="https://zenodo.org/record/10407075/files/figure.png" pageId="7" pageNumber="8">Fig. 2H, J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECB9E6FC984435FC31" box="[958,1082,892,918]" name="Nicaragua" pageId="7" pageNumber="8">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECBE1CFC9844C9FC31" box="[1092,1222,892,918]" name="Costa Rica" pageId="7" pageNumber="8">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECBE88FC99453FFC30" box="[1232,1328,893,919]" name="Panama" pageId="7" pageNumber="8">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECBF62FC98423EFC1D" name="Saint Vincent and the Grenadines" pageId="7" pageNumber="8">Saint Vincent and the Grenadines</collectingCountry>
|
||
,
|
||
<collectingCountry id="1ABB9E85743DFFECB865FC4442BEFC1D" box="[573,689,928,954]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="1ABB9E85743DFFECB8B1FC444344FC1D" box="[745,843,928,954]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
|
||
...............
|
||
<taxonomicName id="A5ACA596743DFFECB99CFC444578FC1D" ID-CoL="WWHW" authority="Westwood, 1846" authorityName="Westwood" authorityYear="1846" box="[964,1399,928,955]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="50D80207743DFFECB99CFC4444A4FC1C" bold="true" box="[964,1195,928,955]" italics="true" pageId="7" pageNumber="8">Coilodes castaneus</emphasis>
|
||
<bibRefCitation id="063DA3E4743DFFECBEEAFC444578FC1D" author="Westwood J. O." box="[1202,1399,928,954]" pageId="7" pageNumber="8" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FC074578FBCE" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FC074579FB86" blockId="7.[189,1399,995,1130]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FC074579FB86" pageId="7" pageNumber="8">
|
||
11. Clypeus subrectangular, wider than long (
|
||
<figureCitation id="FA97C290743DFFECB88BFC07433FFC5A" box="[723,816,995,1021]" captionStart="Fig" captionStartId="27.[189,232,1857,1883]" captionTargetBox="[189,1398,731,1793]" captionTargetId="figure-196@27.[189,1398,731,1819]" captionTargetPageId="27" captionText="Fig. 6. Coilodes lunae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–C, E–G, J–K), paratype ♀ at DZUP from Itatiaia, Rio de Janeiro (D), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407093" httpUri="https://zenodo.org/record/10407093/files/figure.png" pageId="7" pageNumber="8">Fig. 6C</figureCitation>
|
||
); left paramere curved at the apex, with curvature area rounded (
|
||
<figureCitation id="FA97C290743DFFECBBCBFBE34206FB86" box="[403,521,1031,1057]" captionStart="Fig" captionStartId="27.[189,232,1857,1883]" captionTargetBox="[189,1398,731,1793]" captionTargetId="figure-196@27.[189,1398,731,1819]" captionTargetPageId="27" captionText="Fig. 6. Coilodes lunae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–C, E–G, J–K), paratype ♀ at DZUP from Itatiaia, Rio de Janeiro (D), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407093" httpUri="https://zenodo.org/record/10407093/files/figure.png" pageId="7" pageNumber="8">Fig. 6I, K</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECB846FBE34269FB86" box="[542,614,1031,1057]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
|
||
...................................................................................................... 12
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FBC84578FBCE" blockId="7.[189,1399,995,1130]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FBC84578FBCE" pageId="7" pageNumber="8">
|
||
– Clypeus subtrapezoidal, as wide as long (
|
||
<figureCitation id="FA97C290743DFFECB893FBCF4339FBE2" box="[715,822,1067,1093]" captionStart="Fig" captionStartId="37.[191,235,1857,1883]" captionTargetBox="[189,1398,731,1779]" captionTargetId="figure-80@37.[189,1398,730,1817]" captionTargetPageId="37" captionText="Fig. 10. Coilodes parvulus Westwood, 1846, Three ♂♂ at CEMT, two from Ubajara, Ceará, Brazil (A–B, E) and (C), and one from Igrapiuna, Bahia, Brazil (D, F–G, J–K); graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407101" httpUri="https://zenodo.org/record/10407101/files/figure.png" pageId="7" pageNumber="8">Fig. 10C</figureCitation>
|
||
); left paramere curved at the apex, with curvature area angulated (
|
||
<figureCitation id="FA97C290743DFFECBBFCFBAB4226FBCE" box="[420,553,1103,1129]" captionStart="Fig" captionStartId="37.[191,235,1857,1883]" captionTargetBox="[189,1398,731,1779]" captionTargetId="figure-80@37.[189,1398,730,1817]" captionTargetPageId="37" captionText="Fig. 10. Coilodes parvulus Westwood, 1846, Three ♂♂ at CEMT, two from Ubajara, Ceará, Brazil (A–B, E) and (C), and one from Igrapiuna, Bahia, Brazil (D, F–G, J–K); graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407101" httpUri="https://zenodo.org/record/10407101/files/figure.png" pageId="7" pageNumber="8">Fig. 10I, K</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECB866FBAB4289FBCE" box="[574,646,1103,1129]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
|
||
............................................
|
||
<taxonomicName id="A5ACA596743DFFECB989FBAB4578FBCE" ID-CoL="WWJ5" authority="Westwood, 1846" authorityName="Westwood" authorityYear="1846" box="[977,1399,1103,1130]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="50D80207743DFFECB989FBAB44A4FBCD" bold="true" box="[977,1195,1103,1130]" italics="true" pageId="7" pageNumber="8">Coilodes parvulus</emphasis>
|
||
<bibRefCitation id="063DA3E4743DFFECBEEAFBAB4578FBCE" author="Westwood J. O." box="[1202,1399,1103,1129]" pageId="7" pageNumber="8" pagination="155 - 180" refId="ref31038" refString="Westwood J. O. 1846. XXVI. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4 (3): 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1846">Westwood, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="D958C6B0743DFFECBAE5FB764578FABE" pageId="7" pageNumber="8">
|
||
<paragraph id="6213DE15743DFFECBAE5FB764578FB77" blockId="7.[189,1414,1170,1305]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBAE5FB764578FB77" pageId="7" pageNumber="8">
|
||
12. Right paramere with transparent region on its distal half (
|
||
<figureCitation id="FA97C290743DFFECB9C0FB76441DFB0B" box="[920,1042,1170,1196]" captionStart="Fig" captionStartId="27.[189,232,1857,1883]" captionTargetBox="[189,1398,731,1793]" captionTargetId="figure-196@27.[189,1398,731,1819]" captionTargetPageId="27" captionText="Fig. 6. Coilodes lunae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–C, E–G, J–K), paratype ♀ at DZUP from Itatiaia, Rio de Janeiro (D), and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407093" httpUri="https://zenodo.org/record/10407093/files/figure.png" pageId="7" pageNumber="8">Fig. 6H, J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECBE72FB76447DFB0B" box="[1066,1138,1170,1196]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
|
||
..................................... ................................................................................
|
||
<taxonomicName id="A5ACA596743DFFECB918FB514440FB77" authority="Basilio" authorityName="Basilio" box="[832,1103,1205,1232]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lunae">
|
||
<emphasis id="50D80207743DFFECB918FB5143FBFB77" bold="true" box="[832,1012,1205,1232]" italics="true" pageId="7" pageNumber="8">Coilodes lunae</emphasis>
|
||
Basílio
|
||
</taxonomicName>
|
||
& Vaz-de-Mello sp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6213DE15743DFFECBA99FB3F4578FABE" blockId="7.[189,1414,1170,1305]" pageId="7" pageNumber="8">
|
||
<keyLead id="D95D7D20743DFFECBA99FB3F4578FABE" pageId="7" pageNumber="8">
|
||
– Right paramere without transparent region on its distal half (
|
||
<figureCitation id="FA97C290743DFFECB9E6FB3E4437FB53" box="[958,1080,1242,1269]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio & Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="7" pageNumber="8">Fig. 7H, J</figureCitation>
|
||
).
|
||
<collectingCountry id="1ABB9E85743DFFECBE17FB3E4498FB53" box="[1103,1175,1242,1268]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
|
||
................................ ...............................................................................
|
||
<taxonomicName id="A5ACA596743DFFECB96EFB1A4440FABF" authority="Basilio" authorityName="Basilio" box="[822,1103,1278,1305]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mayae">
|
||
<emphasis id="50D80207743DFFECB96EFB1A43FBFABE" bold="true" box="[822,1012,1278,1305]" italics="true" pageId="7" pageNumber="8">Coilodes mayae</emphasis>
|
||
Basílio
|
||
</taxonomicName>
|
||
& Vaz-de-Mello sp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |