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<document id="83863405CA63AACB2F8266E5549871A5" ID-DOI="10.5852/ejt.2023.914.2377" ID-ISSN="2118-9773" ID-Zenodo-Dep="10407071" ID-ZooBank="2ECE4DA6-97CA-4917-97F1-FE506998A2D0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703002881060" checkinUser="plazi" docAuthor="Basílio, Daniel Silva, Vaz-De-Mello, Fernando Zagury, Cherman, Mariana Alejandra &amp; Almeida, Lúcia Massutti de" docDate="2023" docId="EA056F037413FFC0B85DFA0D430FF84F" docLanguage="en" docName="EJT.2023.914.1-53.pdf" docOrigin="European Journal of Taxonomy 914 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2377/10437" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Coilodes ravii Basilio" docType="treatment" docVersion="2" lastPageNumber="44" masterDocId="163C177B743AFFEBBA58FFE4400FFFA7" masterDocTitle="A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species" masterLastPageNumber="53" masterPageNumber="1" pageNumber="42" updateTime="1703003437132" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0BB59B86B6C0FA80AA12B90D390AD4F7">A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species</mods:title>
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<mods:namePart id="D540B8A78B99DD2087B9915A78A2F900">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="890BBC00BB2E9436C11D8CC45686CCB9">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
<mods:nameIdentifier id="7A7AD2BF87F85E3AAFC7EFB42E5F359D" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6B03096F95C7418564753BC875588FF1">Vaz-De-Mello, Fernando Zagury</mods:namePart>
<mods:nameIdentifier id="6541964CF3A631C6ACB19B5E1AADC5F3" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="7122F3DFD982BA77E529192D11A99D72">Laboratório de Scarabaeoidologia, Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="030EDEF7AC1F25ED0ED719DED2A63EF5" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="5E822B1EE83AD25394E434E42075C4CC">Author</mods:roleTerm>
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<mods:namePart id="184EB2DA6DB4E59EA961B619648BE18B">Cherman, Mariana Alejandra</mods:namePart>
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<mods:affiliation id="47258FBB4A4E3DEB0D92E019F694825F">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
<mods:nameIdentifier id="159C8EC43A3FC3DDDD120E822F10F23E" type="email">marianabioar@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A20E3E39554063550EE35199DF9063AF">Almeida, Lúcia Massutti de</mods:namePart>
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<mods:affiliation id="75D9AAE62D9E362E068A07D312C3737A">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
<mods:nameIdentifier id="3365D037711A6F35BE54C8B2F396739F" type="email">lalmeida51@gmail.com</mods:nameIdentifier>
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<mods:title id="889BA997B8CC3815A40AE3EA41B9FEC7">European Journal of Taxonomy</mods:title>
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<mods:date id="AADC62135FEF17A30287EC0D868BBBE9">2023</mods:date>
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<mods:identifier id="7E886437D753EFD1ABB9C19FB1DCCFE2" type="ISSN">2118-9773</mods:identifier>
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<treatment id="EA056F037413FFC0B85DFA0D430FF84F" ID-DOI="http://doi.org/10.5281/zenodo.10407087" ID-Zenodo-Dep="10407087" LSID="urn:lsid:plazi:treatment:EA056F037413FFC0B85DFA0D430FF84F" httpUri="http://treatment.plazi.org/id/EA056F037413FFC0B85DFA0D430FF84F" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
<subSubSection id="2AB68D9E7413FFC2B85DFA0D44C2F98F" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="6213DE157413FFC2B85DFA0D4420F9A3" blockId="41.[360,1229,1513,1612]" box="[517,1071,1513,1540]" pageId="41" pageNumber="42">
<heading id="395B69797413FFC2B85DFA0D4420F9A3" box="[517,1071,1513,1540]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="42" reason="2">
<taxonomicName id="A5ACA5967413FFC2B85DFA0D4309F9A4" authority="Basilio" authorityName="Basilio" box="[517,774,1513,1540]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="ravii">
<emphasis id="50D802077413FFC2B85DFA0D42A3F9A4" bold="true" box="[517,684,1513,1540]" italics="true" pageId="41" pageNumber="42">Coilodes ravii</emphasis>
Basílio
</taxonomicName>
&amp; Vaz-de-Mello sp. nov.
</heading>
</paragraph>
<paragraph id="6213DE157413FFC2BB30F9E944C2F98F" blockId="41.[360,1229,1513,1612]" box="[360,1229,1549,1576]" pageId="41" pageNumber="42">
<uri id="163DD2177413FFC2BB30F9E944C2F98F" box="[360,1229,1549,1576]" pageId="41" pageNumber="42">
urn:lsid:zoobank.org:act:
<uuid id="160AE4C07413FFC2B8D3F9E944C2F98F" box="[651,1229,1549,1576]" pageId="41" pageNumber="42">3CA8E8E5-3A5E-41A8-BF4C-404E609C4A4A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7413FFC2B896F9D54369F9EB" box="[718,870,1585,1612]" pageId="41" pageNumber="42" type="description">
<paragraph id="6213DE157413FFC2B896F9D54369F9EB" blockId="41.[360,1229,1513,1612]" box="[718,870,1585,1612]" pageId="41" pageNumber="42">
<figureCitation id="FA97C2907413FFC2B896F9D54328F9EB" box="[718,807,1585,1612]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="41" pageNumber="42">Figs 12</figureCitation>
,
<figureCitation id="FA97C2907413FFC2B96BF9D64369F9EB" box="[819,870,1586,1612]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1739]" captionTargetId="figure-12@47.[272,1315,265,1739]" captionTargetPageId="47" captionText="Fig. 14. Distribution map of species of Coilodes Westwood, 1846. A. Coilodes bezerrai Basílio &amp; Vazde-Mello sp. nov.; Coilodes castaneus Westwood, 1846; and Coilodes edeiltae Basílio &amp; Vaz-de-Mello sp. nov. B. Coilodes fumipennis Arrow, 1909; Coilodes humeralis (Mannerheim, 1829); and Coilodes lunae Basílio &amp; Vaz-de-Mello sp. nov. C. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov.;Coilodes niger (Mannerheim, 1829) stat. rev.; Coilodes ovalis Robinson, 1948; and Coilodes parvulus Westwood 1846. D. Coilodes punctipennis Arrow, 1909; Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov.; and Coilodes skelleyi Basílio &amp; Vaz-de-Mello sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10407111" httpUri="https://zenodo.org/record/10407111/files/figure.png" pageId="41" pageNumber="42">14D</figureCitation>
</paragraph>
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<subSubSection id="2AB68D9E7413FFC2BAE5F99744FCF945" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="6213DE157413FFC2BAE5F9974139F92A" blockId="41.[189,310,1651,1677]" box="[189,310,1651,1677]" pageId="41" pageNumber="42">
<heading id="395B69797413FFC2BAE5F9974139F92A" bold="true" box="[189,310,1651,1677]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="50D802077413FFC2BAE5F9974139F92A" bold="true" box="[189,310,1651,1677]" pageId="41" pageNumber="42">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157413FFC2BAE5F94744FCF945" blockId="41.[189,1398,1699,1762]" pageId="41" pageNumber="42">
Similar to
<taxonomicName id="A5ACA5967413FFC2BB61F9474230F91A" authorityName="Arrow" authorityYear="1909" box="[313,575,1699,1725]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="50D802077413FFC2BB61F9474230F91A" box="[313,575,1699,1725]" italics="true" pageId="41" pageNumber="42">Coilodes punctipennis</emphasis>
</taxonomicName>
, it is distinguished by the presence of finer punctures on elytra and by the presence of a sharp lobe on the right side of the tegmen, reaching the base of parameres.
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7413FFC2BAE5F8ED4240F8D0" pageId="41" pageNumber="42" type="etymology">
<paragraph id="6213DE157413FFC2BAE5F8ED414BF884" blockId="41.[189,324,1801,1827]" box="[189,324,1801,1827]" pageId="41" pageNumber="42">
<heading id="395B69797413FFC2BAE5F8ED414BF884" bold="true" box="[189,324,1801,1827]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="50D802077413FFC2BAE5F8ED414BF884" bold="true" box="[189,324,1801,1827]" pageId="41" pageNumber="42">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157413FFC2BAE5F8DD4240F8D0" blockId="41.[188,1399,1849,1911]" pageId="41" pageNumber="42">
Noun in the genitive singular.
<taxonomicName id="A5ACA5967413FFC2B84FF8DD431AF8F4" authority="Basilio" authorityName="Basilio" box="[535,789,1849,1875]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="ravii">
<emphasis id="50D802077413FFC2B84FF8DD42B3F8F5" box="[535,700,1849,1875]" italics="true" pageId="41" pageNumber="42">Coilodes ravii</emphasis>
Basílio
</taxonomicName>
&amp; Vaz-de-Mello sp. nov. is named after Ravi Basílio Dantas, nephew of the first author.
</paragraph>
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<paragraph id="6213DE157413FFC2BAE5F87B4163F81E" blockId="41.[189,364,1951,1977]" box="[189,364,1951,1977]" pageId="41" pageNumber="42">
<heading id="395B69797413FFC2BAE5F87B4163F81E" bold="true" box="[189,364,1951,1977]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="50D802077413FFC2BAE5F87B4163F81E" bold="true" box="[189,364,1951,1977]" pageId="41" pageNumber="42">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157413FFC2BAB2F82A4153F84F" blockId="41.[234,348,1998,2024]" box="[234,348,1998,2024]" pageId="41" pageNumber="42">
<emphasis id="50D802077413FFC2BAB2F82A4153F84F" bold="true" box="[234,348,1998,2024]" pageId="41" pageNumber="42">
<typeStatus id="BD1760B77413FFC2BAB2F82A4153F84F" box="[234,348,1998,2024]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6213DE157410FFC1BAE5FEED43FBFEE0" blockId="42.[189,1399,263,327]" pageId="42" pageNumber="43">
<materialsCitation id="D2C4D4487410FFC1BAE5FEED43FBFEE0" collectingDate="1998-09-06" collectingDateMax="1998-09-19" collectingDateMin="1998-09-06" collectionCode="CEMT" collectorName="J. E. Eger" country="Ecuador" elevation="541" latitude="-1.0345" location="Puerto Misahuali" longLatPrecision="1" longitude="-77.663666" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
<collectingCountry id="1ABB9E857410FFC1BAE5FEED4143FE84" box="[189,332,265,291]" name="Ecuador" pageId="42" pageNumber="43">ECUADOR</collectingCountry>
•
<specimenCount id="74AA159C7410FFC1BB30FEED418EFE84" box="[360,385,265,291]" count="1" pageId="42" pageNumber="43" type="male">♂</specimenCount>
; Prov.
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, vic.
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;
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,
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; alt.
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;
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</date>
;
<collectorName id="CF59BBC37410FFC1BBDCFEC941E1FEE0" box="[388,494,301,327]" pageId="42" pageNumber="43">J.E. Eger</collectorName>
leg.; fish carrion baited,
<collectingMethod id="BBEDA6027410FFC1B94BFEC8439EFEE0" box="[787,913,300,327]" pageId="42" pageNumber="43">pitfall trap</collectingMethod>
;
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.
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</paragraph>
<paragraph id="6213DE157410FFC1BAB2FE8A424FFE2F" blockId="42.[189,1399,366,533]" box="[234,576,366,392]" pageId="42" pageNumber="43">
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<typeStatus id="BD1760B77410FFC1BAB2FE8A4168FE2F" box="[234,359,366,392]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="74AA159C7410FFC1BB20FE8A41B4FE2F" box="[376,443,366,392]" count="2" pageId="42" pageNumber="43" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="74AA159C7410FFC1BBACFE8A423BFE2F" box="[500,564,366,392]" count="4" pageId="42" pageNumber="43" type="female">4 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="6213DE157410FFC1BAE5FE7542BBFDB2" blockId="42.[189,1399,366,533]" pageId="42" pageNumber="43">
<materialsCitation id="D2C4D4487410FFC1BAE5FE754401FE0C" box="[189,1038,401,427]" collectingDate="1998-09-06" collectingDateMax="1998-09-19" collectingDateMin="1998-09-06" collectionCode="CEMT" collectorName="J. E. Eger" country="Ecuador" elevation="541" latitude="-1.0345" location="Puerto Misahuali" longLatPrecision="1" longitude="-77.663666" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<collectingCountry id="1ABB9E857410FFC1BAE5FE754144FE0C" box="[189,331,401,427]" name="Ecuador" pageId="42" pageNumber="43">ECUADOR</collectingCountry>
–
<emphasis id="50D802077410FFC1BB32FE7541A2FE0C" bold="true" box="[362,429,401,427]" pageId="42" pageNumber="43">
<collectingRegion id="A06810F77410FFC1BB32FE7541A2FE0C" box="[362,429,401,427]" country="Ecuador" name="Napo" pageId="42" pageNumber="43">Napo</collectingRegion>
</emphasis>
•
<specimenCount id="74AA159C7410FFC1BB9EFE7541F9FE0C" box="[454,502,401,427]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="04BD46D07410FFC1B9E5FE754401FE0C" box="[957,1038,401,427]" pageId="42" pageNumber="43">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487410FFC1BE70FE7541BFFE69" collectingDate="1998-09-06" collectingDateMax="1998-09-19" collectingDateMin="1998-09-06" collectionCode="CEMT" collectorName="J. E. Eger" country="Ecuador" elevation="541" latitude="-1.0345" location="Puerto Misahuali" longLatPrecision="1" longitude="-77.663666" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="74AA159C7410FFC1BE70FE754459FE0C" box="[1064,1110,401,427]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="04BD46D07410FFC1BB07FE5041BFFE69" box="[351,432,436,462]" pageId="42" pageNumber="43">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487410FFC1BB94FE50441CFE68" box="[460,1043,436,463]" collectingDate="1998-09-06" collectingDateMax="1998-09-19" collectingDateMin="1998-09-06" collectionCode="DZUP" collectorName="J. E. Eger" country="Ecuador" elevation="541" latitude="-1.0345" location="Puerto Misahuali" longLatPrecision="1" longitude="-77.663666" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<specimenCount id="74AA159C7410FFC1BB94FE5041F3FE68" box="[460,508,436,463]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="04BD46D07410FFC1B99FFE51441CFE68" box="[967,1043,437,463]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="42" pageNumber="43" type="University or college">DZUP</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487410FFC1BE7FFE5041A5FE55" collectingDate="1998-09-06" collectingDateMax="1998-09-19" collectingDateMin="1998-09-06" collectionCode="DZUP" collectorName="J. E. Eger" country="Ecuador" elevation="541" latitude="-1.0345" location="Puerto Misahuali" longLatPrecision="1" longitude="-77.663666" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="74AA159C7410FFC1BE7FFE50445BFE68" box="[1063,1108,436,463]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="04BD46D07410FFC1BB06FE3C41A5FE55" box="[350,426,472,498]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="42" pageNumber="43" type="University or college">DZUP</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487410FFC1BB9CFE33440CFE56" box="[452,1027,471,498]" collectingDate="1990-03" collectionCode="CEMT" collectorName="S. Sandoval" country="Ecuador" location="Yutari" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="74AA159C7410FFC1BB9CFE3341FDFE55" box="[452,498,471,498]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
;
<location id="677388CE7410FFC1B858FE334243FE56" LSID="urn:lsid:plazi:treatment:EA056F037413FFC0B85DFA0D430FF84F:677388CE7410FFC1B858FE334243FE56" box="[512,588,471,497]" country="Ecuador" name="Yutari" pageId="42" pageNumber="43" stateProvince="Napo">Yutari</location>
;
<date id="1612F8D57410FFC1B800FE3C42DAFE56" box="[600,725,471,498]" pageId="42" pageNumber="43" value="1990-03">
<collectingDate id="0656013D7410FFC1B800FE3C42DAFE56" box="[600,725,471,498]" pageId="42" pageNumber="43" value="1990-03">Mar. 1990</collectingDate>
</date>
;
<collectorName id="CF59BBC37410FFC1B8BAFE334360FE56" box="[738,879,471,497]" pageId="42" pageNumber="43">S. Sandoval</collectorName>
leg.;
<collectionCode id="04BD46D07410FFC1B9EAFE33440CFE56" box="[946,1027,471,497]" pageId="42" pageNumber="43">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487410FFC1BE46FE3342BFFDB2" collectingDate="1990-03-08" collectionCode="CEMT" collectorName="S. Sandoval" country="Ecuador" location="Pichira" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="74AA159C7410FFC1BE46FE334443FE55" box="[1054,1100,471,498]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
;
<location id="677388CE7410FFC1BE03FE3344BDFE56" LSID="urn:lsid:plazi:treatment:EA056F037413FFC0B85DFA0D430FF84F:677388CE7410FFC1BE03FE3344BDFE56" box="[1115,1202,471,497]" country="Ecuador" name="Pichira" pageId="42" pageNumber="43" stateProvince="Napo">Pichira</location>
;
<date id="1612F8D57410FFC1BEE6FE33455DFE56" box="[1214,1362,471,498]" pageId="42" pageNumber="43" value="1990-03-08">
<collectingDate id="0656013D7410FFC1BEE6FE33455DFE56" box="[1214,1362,471,498]" pageId="42" pageNumber="43" value="1990-03-08">8 Mar. 1990</collectingDate>
</date>
;
<collectorName id="CF59BBC37410FFC1BF07FE334125FDB3" pageId="42" pageNumber="43">S. Sandoval</collectorName>
leg.; [trap], “carne baja”;
<collectionCode id="04BD46D07410FFC1B805FE1F42BFFDB2" box="[605,688,507,533]" pageId="42" pageNumber="43">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7410FFC0BAE5FDD8430FF84F" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="6213DE157410FFC1BAE5FDD84141FDF1" blockId="42.[189,334,572,598]" box="[189,334,572,598]" pageId="42" pageNumber="43">
<heading id="395B69797410FFC1BAE5FDD84141FDF1" bold="true" box="[189,334,572,598]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="50D802077410FFC1BAE5FDD84141FDF1" bold="true" box="[189,334,572,598]" pageId="42" pageNumber="43">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157410FFC1BAB2FD8E41A4FD23" blockId="42.[189,1278,618,680]" box="[234,427,618,644]" pageId="42" pageNumber="43">
<emphasis id="50D802077410FFC1BAB2FD8E4125FD23" bold="true" box="[234,298,618,644]" pageId="42" pageNumber="43">Male</emphasis>
(
<typeStatus id="BD1760B77410FFC1BB61FD8E41ACFD23" box="[313,419,618,644]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="6213DE157410FFC1BAE5FD6A44F1FD00" blockId="42.[189,1278,618,680]" box="[189,1278,653,680]" pageId="42" pageNumber="43">
MEASUREMENTS. Length
<quantity id="A55473F07410FFC1BB86FD694234FD0F" box="[478,571,653,680]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="42" pageNumber="43" unit="mm" value="6.4">6.4 mm</quantity>
. Width
<quantity id="A55473F07410FFC1B8CDFD6A42FDFD0F" box="[661,754,654,680]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="42" pageNumber="43" unit="mm" value="3.7">3.7 mm</quantity>
. Body (
<figureCitation id="FA97C2907410FFC1B915FD6943D5FD0F" box="[845,986,653,680]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="42" pageNumber="43">Fig. 12A–B</figureCitation>
) convex, oval and shiny.
</paragraph>
<caption id="36D38E9D7410FFC1BAE5F8804564F84E" ID-DOI="http://doi.org/10.5281/zenodo.10407105" ID-Zenodo-Dep="10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="42" pageNumber="43" startId="42.[189,233,1892,1918]" targetBox="[189,1398,766,1834]" targetPageId="42">
<paragraph id="6213DE157410FFC1BAE5F8804564F84E" blockId="42.[187,1399,1892,2025]" pageId="42" pageNumber="43">
<emphasis id="50D802077410FFC1BAE5F880412EF8D8" bold="true" box="[189,289,1892,1919]" pageId="42" pageNumber="43">Fig. 12.</emphasis>
<taxonomicName id="A5ACA5967410FFC1BB74F881424FF8D8" authority="Basilio" authorityName="Basilio" box="[300,576,1893,1919]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="ravii">
<emphasis id="50D802077410FFC1BB74F88141D2F8D8" box="[300,477,1893,1919]" italics="true" pageId="42" pageNumber="43">Coilodes ravii</emphasis>
Basílio
</taxonomicName>
&amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I).
<emphasis id="50D802077410FFC1BBEDF86C41C5F805" bold="true" box="[437,458,1928,1954]" pageId="42" pageNumber="43">A</emphasis>
. Body, dorsal view.
<emphasis id="50D802077410FFC1B8EBF86C42C8F805" bold="true" box="[691,711,1928,1954]" pageId="42" pageNumber="43">B</emphasis>
. Body, ventral view.
<emphasis id="50D802077410FFC1B9E1F86C43C1F805" bold="true" box="[953,974,1928,1954]" pageId="42" pageNumber="43">C</emphasis>
. Head, dorsal view.
<emphasis id="50D802077410FFC1BEEFF86C44C3F805" bold="true" box="[1207,1228,1928,1954]" pageId="42" pageNumber="43">D</emphasis>
. Head, ventral view.
<emphasis id="50D802077410FFC1BB58F84F411CF862" bold="true" box="[256,275,1963,1989]" pageId="42" pageNumber="43">E</emphasis>
. Pronotum, dorsal view.
<emphasis id="50D802077410FFC1B876F84F4268F862" bold="true" box="[558,615,1963,1990]" pageId="42" pageNumber="43">F–K</emphasis>
. Male genitalia.
<emphasis id="50D802077410FFC1B97CF84F4339F862" bold="true" box="[804,822,1963,1989]" pageId="42" pageNumber="43">F</emphasis>
. Dorsal view.
<emphasis id="50D802077410FFC1B981F84F43FFF862" bold="true" box="[985,1008,1963,1989]" pageId="42" pageNumber="43">G</emphasis>
. Ventral view.
<emphasis id="50D802077410FFC1BEC0F84F44A0F862" bold="true" box="[1176,1199,1963,1989]" pageId="42" pageNumber="43">H</emphasis>
. Right paramere.
<emphasis id="50D802077410FFC1BAE3F82A40C9F84F" bold="true" box="[187,198,1998,2024]" pageId="42" pageNumber="43">I</emphasis>
. Left paramere.
<emphasis id="50D802077410FFC1BBD3F82A4195F84F" bold="true" box="[395,410,1998,2024]" pageId="42" pageNumber="43">J</emphasis>
. Right side.
<emphasis id="50D802077410FFC1B877F82A4249F84F" bold="true" box="[559,582,1998,2024]" pageId="42" pageNumber="43">K</emphasis>
. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm.
</paragraph>
</caption>
<paragraph id="6213DE157411FFC0BAE5FEED43CBFE84" blockId="43.[189,964,265,291]" box="[189,964,265,291]" pageId="43" pageNumber="44">COLOUR. Body, yellowish brown; elytra with outer margins black.</paragraph>
<paragraph id="6213DE157411FFC0BAE5FEA8447AFD0A" blockId="43.[187,1401,332,686]" pageId="43" pageNumber="44">
HEAD (
<figureCitation id="FA97C2907411FFC0BB55FEA84193FEC1" box="[269,412,332,358]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="43" pageNumber="44">Fig. 12C–D</figureCitation>
). Surface slightly strigulate. Frons, in dorsal view, subrectangular; two small tubercles merged, in the middle; proximal border rounded; posterior margin as wide as the anterior one; lateral margin, straight and darker; surface without punctures. Eye barely visible dorsally. Canthus strong; distinct area expanded downwards. Clypeus subrectangular; lateral margins weakly rounded; anterior borders rounded; apex straight; setae thin and sparse, throughout the anterior margin dorsally; surface without punctures. Labrum semicircular; setae sparse, spreading along the entire dorsal border. Mandibles dorsally excavated, protruding beyond the apex of labrum. Labium with mentum subquadrate; inferior and lateral margins straight and superior margin slightly curved in the middle; disc strigulate, long setae on the margins. Maxilla subtriangular; long setae throughout surface. Antenna with 10 antennomeres; antennal club with three antennomeres; club with basal antennomere cupuliform.
</paragraph>
<paragraph id="6213DE157411FFC0BAE5FD32438EFC9E" blockId="43.[187,1399,726,826]" pageId="43" pageNumber="44">
PRONOTUM (
<figureCitation id="FA97C2907411FFC0BB15FD3241BAFD57" box="[333,437,726,752]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="43" pageNumber="44">Fig. 12E</figureCitation>
). Convex; strongly excavated medially; subtrapezoidal, posterior margin wider than anterior one; anterior margin straight, angles subacute; posterior margin rounded and weakly produced medially; lateral margin convex, setae and punctures absent.
</paragraph>
<paragraph id="6213DE157411FFC0BAE5FC864446FCDB" blockId="43.[189,1097,866,892]" box="[189,1097,866,892]" pageId="43" pageNumber="44">SCUTELLAR SHIELD. Subtriangular; setae and punctures absent; apex punctate.</paragraph>
<paragraph id="6213DE157411FFC0BAE5FC414544FC18" blockId="43.[189,1355,932,959]" box="[189,1355,932,959]" pageId="43" pageNumber="44">ELYTRA. Double row of longitudinal punctures, sutural stria complete, formed by thicker punctures.</paragraph>
<paragraph id="6213DE157411FFC0BAE5FC0C451CFB82" blockId="43.[187,1399,999,1061]" pageId="43" pageNumber="44">
VENTER (
<figureCitation id="FA97C2907411FFC0BB70FC03419FFBA6" box="[296,400,999,1025]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="43" pageNumber="44">Fig. 12B</figureCitation>
). Hypomeral surface strigulate, setose; prosternum (
<figureCitation id="FA97C2907411FFC0B9BBFC034464FBA6" box="[995,1131,999,1025]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="43" pageNumber="44">Fig. 12B, D</figureCitation>
) elevated at midlength, transversally grooved; anterior margin straight and flat, mesoventrite and metaventrite smooth.
</paragraph>
<paragraph id="6213DE157411FFC0BAE5FBAA44B6FAE4" blockId="43.[187,1400,1102,1347]" pageId="43" pageNumber="44">LEGS. Procoxa conical; surface strigulate. Protrochanter joint with the procoxa rounded, distally angulate. Profemur with posterior margin carinate. Protibia with inner margin convex; carinate; single spur, slightly curved at the apex; external margin with three larger teeth and series of smaller denticles along the entire margin. Protarsi with tarsal insertion beneath the protibial second tooth. Mesothoracic and metathoracic legs smooth. Mesotrochanter and metatrochanter subtriangular with bifurcated apex. Mesofemur and metafemur carinate on posterior margin. Mesotibia and metatibia with erect setae; apex weakly expanded; pair of spurs with sharp apex. Tarsi with five tarsomeres; tarsal claws toothed medially.</paragraph>
<paragraph id="6213DE157411FFC0BAE5FA8F42D2FA21" blockId="43.[189,733,1387,1414]" box="[189,733,1387,1414]" pageId="43" pageNumber="44">ABDOMEN. Six ventrites with recumbent setae.</paragraph>
<paragraph id="6213DE157411FFC0BAE5FA4A455BF9B6" blockId="43.[187,1398,1454,1553]" pageId="43" pageNumber="44">
GENITALIA (
<figureCitation id="FA97C2907411FFC0BB1CFA4A41C1FA6F" box="[324,462,1454,1480]" captionStart="Fig" captionStartId="42.[189,233,1892,1918]" captionTargetBox="[189,1398,766,1834]" captionTargetId="figure-14@42.[189,1398,765,1853]" captionTargetPageId="42" captionText="Fig. 12. Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphical representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view.F–K. Male genitalia. F. Dorsal view. G. Ventral view.H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407105" httpUri="https://zenodo.org/record/10407105/files/figure.png" pageId="43" pageNumber="44">Fig. 12F–K</figureCitation>
). Lobe of tegmen triangular on right side, reaching the paramere bases; sclerotized region next to paramere bases; parameres asymmetrical; right paramere longer than wide, straight base and apex rounded; left paramere dislocated to ventral region, straight apex strongly curved outward.
</paragraph>
<paragraph id="6213DE157411FFC0BAB2F9DE414AF9F3" blockId="43.[189,560,1594,1657]" box="[234,325,1594,1620]" pageId="43" pageNumber="44">
<emphasis id="50D802077411FFC0BAB2F9DE414AF9F3" bold="true" box="[234,325,1594,1620]" pageId="43" pageNumber="44">Female</emphasis>
</paragraph>
<paragraph id="6213DE157411FFC0BAE5F9BA423FF9DF" blockId="43.[189,560,1594,1657]" box="[189,560,1630,1657]" pageId="43" pageNumber="44">
Length
<quantity id="A55473F07411FFC0BB4FF9BA417BF9DF" box="[279,372,1630,1657]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="43" pageNumber="44" unit="mm" value="5.8">5.8 mm</quantity>
. Width
<quantity id="A55473F07411FFC0BB97F9BA4223F9DF" box="[463,556,1630,1656]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="43" pageNumber="44" unit="mm" value="3.6">3.6 mm</quantity>
.
</paragraph>
<paragraph id="6213DE157411FFC0BAB2F944416FF91D" blockId="43.[189,1399,1696,1796]" box="[234,352,1696,1722]" pageId="43" pageNumber="44">
<emphasis id="50D802077411FFC0BAB2F944416FF91D" bold="true" box="[234,352,1696,1722]" pageId="43" pageNumber="44">Variation</emphasis>
</paragraph>
<paragraph id="6213DE157411FFC0BAE5F9214567F8A4" blockId="43.[189,1399,1696,1796]" pageId="43" pageNumber="44">
Length ranging from
<quantity id="A55473F07411FFC0BB9AF921427FF947" box="[450,624,1733,1760]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" pageId="43" pageNumber="44" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">5.8 to 6.4 mm</quantity>
. Width ranging from
<quantity id="A55473F07411FFC0B92EF921442AF947" box="[886,1061,1733,1760]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="43" pageNumber="44" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.5 to 3.7 mm</quantity>
. Head and pronotum colour ranging from yellowish brown to dark brown. Pronotum in males sometimes with a slight excavation.
</paragraph>
<paragraph id="6213DE157411FFC0BAE5F8C8420DF8E1" blockId="43.[189,514,1836,1862]" box="[189,514,1836,1862]" pageId="43" pageNumber="44">
<heading id="395B69797411FFC0BAE5F8C8420DF8E1" bold="true" box="[189,514,1836,1862]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="50D802077411FFC0BAE5F8C8420DF8E1" bold="true" box="[189,514,1836,1862]" pageId="43" pageNumber="44">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157411FFC0BAE5F8B8420BF8D1" blockId="43.[189,516,1884,1910]" box="[189,516,1884,1910]" pageId="43" pageNumber="44">
<collectingCountry id="1ABB9E857411FFC0BAE5F8B84110F8D1" box="[189,287,1884,1910]" name="Ecuador" pageId="43" pageNumber="44">Ecuador</collectingCountry>
(
<collectingRegion id="A06810F77411FFC0BB76F8B8417EF8D1" box="[302,369,1884,1910]" country="Ecuador" name="Napo" pageId="43" pageNumber="44">Napo</collectingRegion>
) (
<figureCitation id="FA97C2907411FFC0BBD1F8B841F8F8D1" box="[393,503,1884,1910]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1739]" captionTargetId="figure-12@47.[272,1315,265,1739]" captionTargetPageId="47" captionText="Fig. 14. Distribution map of species of Coilodes Westwood, 1846. A. Coilodes bezerrai Basílio &amp; Vazde-Mello sp. nov.; Coilodes castaneus Westwood, 1846; and Coilodes edeiltae Basílio &amp; Vaz-de-Mello sp. nov. B. Coilodes fumipennis Arrow, 1909; Coilodes humeralis (Mannerheim, 1829); and Coilodes lunae Basílio &amp; Vaz-de-Mello sp. nov. C. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov.;Coilodes niger (Mannerheim, 1829) stat. rev.; Coilodes ovalis Robinson, 1948; and Coilodes parvulus Westwood 1846. D. Coilodes punctipennis Arrow, 1909; Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov.; and Coilodes skelleyi Basílio &amp; Vaz-de-Mello sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10407111" httpUri="https://zenodo.org/record/10407111/files/figure.png" pageId="43" pageNumber="44">Fig. 14D</figureCitation>
).
</paragraph>
<paragraph id="6213DE157411FFC0BAE5F87A4178F81E" blockId="43.[189,375,1950,1977]" box="[189,375,1950,1977]" pageId="43" pageNumber="44">
<heading id="395B69797411FFC0BAE5F87A4178F81E" bold="true" box="[189,375,1950,1977]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="50D802077411FFC0BAE5F87A4178F81E" bold="true" box="[189,375,1950,1977]" pageId="43" pageNumber="44">Biological data</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157411FFC0BAE5F82A430FF84F" blockId="43.[189,768,1998,2025]" box="[189,768,1998,2025]" pageId="43" pageNumber="44">Species collected in carcasses of fish (label data).</paragraph>
</subSubSection>
</treatment>
</document>
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