1136 lines
136 KiB
XML
1136 lines
136 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3794441" ID-GBIF-Dataset="a725007b-2d51-4c92-838a-9e8fd27484c8" ID-GBIF-Taxon="163962588" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794441" ID-ZooBank="701C1742-718D-4486-A158-AEA608BA8576" checkinTime="1588685582300" checkinUser="felipe" docAuthor="Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. & Scholtz, Clarke H." docDate="2020" docId="D474D525FF9F7C6CD7699CE1FBB0FC5D" docLanguage="en" docName="InsectSystEvol.51.1.1-61.pdf.imf" docOrigin="Insect Systematics & Evolution 51 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Sisyphini Mulsant 1842" docType="key" docVersion="6" lastPageId="8" lastPageNumber="9" masterDocId="284DAD5DFF9B7C64D7039C5BFF82FFDC" masterDocTitle="Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions" masterLastPageNumber="61" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1636469099162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Daniel, Gimo M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Davis, Adrian Lv.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sole, Catherine L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Scholtz, Clarke H.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Insect Systematics & Evolution</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2020-01-02</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>51</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>61</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.1163/1876312X-00002195</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">a725007b-2d51-4c92-838a-9e8fd27484c8</mods:identifier>
|
||
<mods:identifier type="ISSN">1876-312X</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">3786629</mods:identifier>
|
||
<mods:identifier type="ZooBank">701C1742-718D-4486-A158-AEA608BA8576</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3794441" ID-GBIF-Taxon="163962588" ID-Zenodo-Dep="3794441" LSID="urn:lsid:plazi:treatment:D474D525FF9F7C6CD7699CE1FBB0FC5D" httpUri="http://treatment.plazi.org/id/D474D525FF9F7C6CD7699CE1FBB0FC5D" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
|
||
<subSubSection box="[106,882,186,212]" pageId="4" pageNumber="5" type="nomenclature">
|
||
<paragraph blockId="4.[106,882,186,212]" box="[106,882,186,212]" pageId="4" pageNumber="5">
|
||
<heading box="[106,882,186,212]" fontSize="11" level="2" pageId="4" pageNumber="5" reason="5">
|
||
<emphasis bold="true" box="[106,882,186,212]" pageId="4" pageNumber="5">
|
||
Identification key for males of southern African
|
||
<taxonomicName authorityName="Mulsant" authorityYear="1842" box="[681,793,186,212]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Sisyphini">sisyphine</taxonomicName>
|
||
species
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="key">
|
||
<key lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
1. Lateral ridge on the pronotal disc extends from the posterior margin without reaching the lateral margin (
|
||
<figureCitation box="[471,553,272,298]" captionStart="Fig" captionStartId="8.[106,139,1511,1533]" captionTargetBox="[173,1005,995,1486]" captionTargetId="figure@8.[173,1005,995,1486]" captionTargetPageId="8" captionText="Fig. 1. A- medially narrow sternite, typical for males in Sisyphus. B- Abdominal sternite in females of Sisyphus species; C- Sisyphus; bears strong setae on the external edge of the first metatarsus; D- Neosisyphus, lacking strong setae on the external margin of the first metatarsus. E- Complete lateral ridge, which divides proepisternum from pronotal disc in Sisyphus. F- Lateral ridge does not reach antero-lateral edge of the pronotum in Neosisyphus." figureDoi="http://doi.org/10.5281/zenodo.3786632" httpUri="https://zenodo.org/record/3786632/files/figure.png" pageId="4" pageNumber="5">Fig. 1F</figureCitation>
|
||
); strong setae absent on the external margin of the first meso- and metatarsal segments (
|
||
<figureCitation box="[615,704,306,333]" captionStart="Fig" captionStartId="8.[106,139,1511,1533]" captionTargetBox="[173,1005,995,1486]" captionTargetId="figure@8.[173,1005,995,1486]" captionTargetPageId="8" captionText="Fig. 1. A- medially narrow sternite, typical for males in Sisyphus. B- Abdominal sternite in females of Sisyphus species; C- Sisyphus; bears strong setae on the external edge of the first metatarsus; D- Neosisyphus, lacking strong setae on the external margin of the first metatarsus. E- Complete lateral ridge, which divides proepisternum from pronotal disc in Sisyphus. F- Lateral ridge does not reach antero-lateral edge of the pronotum in Neosisyphus." figureDoi="http://doi.org/10.5281/zenodo.3786632" httpUri="https://zenodo.org/record/3786632/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
|
||
)........................
|
||
<taxonomicName authorityName="Muller" authorityYear="1942" box="[890,1032,306,333]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[890,1032,306,333]" italics="true" pageId="4" pageNumber="5">Neosisyphus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis bold="true" box="[1049,1064,306,332]" pageId="4" pageNumber="5">2</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Lateral prothoracic ridge between pronotal disc and prothoracic episternum complete (
|
||
<figureCitation box="[231,319,376,402]" captionStart="Fig" captionStartId="8.[106,139,1511,1533]" captionTargetBox="[173,1005,995,1486]" captionTargetId="figure@8.[173,1005,995,1486]" captionTargetPageId="8" captionText="Fig. 1. A- medially narrow sternite, typical for males in Sisyphus. B- Abdominal sternite in females of Sisyphus species; C- Sisyphus; bears strong setae on the external edge of the first metatarsus; D- Neosisyphus, lacking strong setae on the external margin of the first metatarsus. E- Complete lateral ridge, which divides proepisternum from pronotal disc in Sisyphus. F- Lateral ridge does not reach antero-lateral edge of the pronotum in Neosisyphus." figureDoi="http://doi.org/10.5281/zenodo.3786632" httpUri="https://zenodo.org/record/3786632/files/figure.png" pageId="4" pageNumber="5">Fig. 1E</figureCitation>
|
||
); first tarsal segments of the meso- and meta-thoracic legs with a row of dense setae in a straight line on the external edge (
|
||
<figureCitation box="[821,907,410,437]" captionStart="Fig" captionStartId="8.[106,139,1511,1533]" captionTargetBox="[173,1005,995,1486]" captionTargetId="figure@8.[173,1005,995,1486]" captionTargetPageId="8" captionText="Fig. 1. A- medially narrow sternite, typical for males in Sisyphus. B- Abdominal sternite in females of Sisyphus species; C- Sisyphus; bears strong setae on the external edge of the first metatarsus; D- Neosisyphus, lacking strong setae on the external margin of the first metatarsus. E- Complete lateral ridge, which divides proepisternum from pronotal disc in Sisyphus. F- Lateral ridge does not reach antero-lateral edge of the pronotum in Neosisyphus." figureDoi="http://doi.org/10.5281/zenodo.3786632" httpUri="https://zenodo.org/record/3786632/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
|
||
).................... .......................................................................................................
|
||
<taxonomicName authorityName="Latreille" authorityYear="1807" box="[915,1016,447,474]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[915,1016,447,474]" italics="true" pageId="4" pageNumber="5">Sisyphus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis bold="true" box="[1033,1064,447,473]" pageId="4" pageNumber="5">13</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
2. Mesotibia strongly modified by indentations and protrusions; a large spine distally on the posterior margin of the mesofemur. Completely black, pygidium simple and lacking a pair of distinct tubercles. Eastern coastal forests of
|
||
<collectingCountry box="[814,954,552,578]" name="South Africa" pageId="4" pageNumber="5">South Africa</collectingCountry>
|
||
and South
|
||
<collectingCountry box="[156,306,586,612]" name="Mozambique" pageId="4" pageNumber="5">Mozambique</collectingCountry>
|
||
....................................................................................................... .........(
|
||
<taxonomicName baseAuthorityName="Raffray" baseAuthorityYear="1877" box="[234,318,620,647]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tibialis">
|
||
<emphasis bold="true" box="[234,318,620,647]" italics="true" pageId="4" pageNumber="5">tibialis</emphasis>
|
||
</taxonomicName>
|
||
species-group;
|
||
<taxonomicName authority="(Arrow, 1927)" baseAuthorityName="Arrow" baseAuthorityYear="1927" box="[484,905,620,647]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis bold="true" box="[484,905,620,647]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[484,738,620,647]" italics="true" pageId="4" pageNumber="5">Neosisyphus mirabilis</emphasis>
|
||
(
|
||
<bibRefCitation author="Arrow" box="[748,895,620,646]" firstAuthor="Arrow" pageId="4" pageNumber="5" pagination="456 - 465" refId="ref28180" refString="Arrow, G. J. (1927) Notes on the Coleopterous genus Sisyphus. Annals and Magazine of natural History, 9 (19): 456 - 465." type="journal article" year="1927">Arrow,1927</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[915,1008,621,647]" captionStart="Fig" captionStartId="50.[106,139,1217,1239]" captionTargetBox="[401,777,829,1189]" captionTargetId="figure@50.[401,777,826,1192]" captionTargetPageId="50" captionText="Fig. 23. Image of the habitus of the tibialis species-group: A. N. mirabilis." figureDoi="http://doi.org/10.5281/zenodo.3786679" httpUri="https://zenodo.org/record/3786679/files/figure.png" pageId="4" pageNumber="5">Figs. 23</figureCitation>
|
||
,
|
||
<figureCitation box="[1016,1066,621,647]" captionStart="Fig" captionStartId="50.[106,139,1688,1710]" captionTargetBox="[471,707,1270,1683]" captionTargetId="figure@50.[469,710,1270,1686]" captionTargetPageId="50" captionText="Fig. 24. Image of the aedeagus of the tibialis species-group: A. N. mirabilis." figureDoi="http://doi.org/10.5281/zenodo.3786681" httpUri="https://zenodo.org/record/3786681/files/figure.png" pageId="4" pageNumber="5">24A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Mesotibia simple, without any spine projected distally on the posterior margin of the mesofemur.................................................................................................(
|
||
<emphasis bold="true" box="[1049,1064,693,719]" pageId="4" pageNumber="5">3</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
3. Femur and tibia of mesolegs without cuticular projections on the posterior margin.............................................
|
||
<taxonomicName baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[571,701,762,789]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="barbarossa">
|
||
<emphasis bold="true" box="[571,701,762,789]" italics="true" pageId="4" pageNumber="5">barbarossa</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[878,964,762,789]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
A–D) (
|
||
<emphasis bold="true" box="[1049,1064,762,788]" pageId="4" pageNumber="5">4</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[106,1074,237,1594]" box="[106,1073,800,826]" pageId="4" pageNumber="5">
|
||
<keyLead box="[106,1073,800,826]" pageId="4" pageNumber="5">
|
||
– Femur and tibia of mesolegs with cuticular projections on the posterior margin...(
|
||
<emphasis bold="true" box="[1049,1064,800,826]" pageId="4" pageNumber="5">7</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
4. Metafemur with sharp pointed spine on the mid-posterior margin (
|
||
<figureCitation box="[924,1010,834,861]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
A–B) ........................................................................................................................(
|
||
<emphasis bold="true" box="[1049,1064,869,895]" pageId="4" pageNumber="5">5</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Metafemur with angled-protuberance on the mid-posterior margin (
|
||
<figureCitation box="[917,1005,904,930]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
C–D) ........................................................................................................................(
|
||
<emphasis bold="true" box="[1049,1064,941,967]" pageId="4" pageNumber="5">6</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
5. Projecting metatrochanter longer than ¾ of the metafemur length in major males and ½ of the metafemur length in minor males. In major males the metatrochant- er is slightly inwardly-curved at the tip and straight in minor males; setae on the elytra black, recurved at the apex. Lowland dry savannas in southern Africa ........ ..........................................
|
||
<taxonomicName authority="(Klug, 1855)" baseAuthorityName="Klug" baseAuthorityYear="1855" box="[465,897,1114,1141]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="calcaratus">
|
||
<emphasis bold="true" box="[465,897,1114,1141]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[465,735,1114,1141]" italics="true" pageId="4" pageNumber="5">Neosisyphus calcaratus</emphasis>
|
||
(
|
||
<bibRefCitation author="Klug" box="[750,888,1114,1140]" firstAuthor="Klug" pageId="4" pageNumber="5" pagination="643 - 660" refId="ref29765" refString="Klug, J. C. F. (1855) Diagnosen neuer Coleoptera aus Mossambique. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 643 - 660." type="book chapter" year="1855">Klug, 1855</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[912,1004,1114,1141]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
,
|
||
<figureCitation box="[1017,1065,1115,1141]" captionStart="Fig" captionStartId="44.[106,139,640,662]" captionTargetBox="[413,766,132,615]" captionTargetId="figure@44.[413,766,131,615]" captionTargetPageId="44" captionText="Fig. 18. Images of the aedeagi of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786668" httpUri="https://zenodo.org/record/3786668/files/figure.png" pageId="4" pageNumber="5">18B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Projecting metatrochanter shorter in minor males (<¼ the length of the metafemur) and longer in major males (½ the length of the metafemur), elytral setae yellow. Moist highveld and coastal upland grassland of
|
||
<collectingCountry box="[779,921,1218,1244]" name="South Africa" pageId="4" pageNumber="5">South Africa</collectingCountry>
|
||
................... ............................
|
||
<taxonomicName authority="(Wiedemann, 1823)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[368,897,1252,1279]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="barbarossa">
|
||
<emphasis bold="true" box="[368,897,1252,1279]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[368,647,1252,1279]" italics="true" pageId="4" pageNumber="5">Neosisyphus barbarossa</emphasis>
|
||
(
|
||
<bibRefCitation author="Wiedemann" box="[662,887,1252,1278]" firstAuthor="Wiedemann" pageId="4" pageNumber="5" pagination="3 - 133" refId="ref31241" refString="Wiedemann, C. K. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin, 2 (1): 3 - 133." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[911,1003,1253,1279]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
,
|
||
<figureCitation box="[1016,1065,1253,1279]" captionStart="Fig" captionStartId="44.[106,139,640,662]" captionTargetBox="[413,766,132,615]" captionTargetId="figure@44.[413,766,131,615]" captionTargetPageId="44" captionText="Fig. 18. Images of the aedeagi of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786668" httpUri="https://zenodo.org/record/3786668/files/figure.png" pageId="4" pageNumber="5">18A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
6. Projecting metatrochanter relatively short; four times shorter than total length of metafemur. Lowland and upland moist vegetation from northeast to southern Africa .....................................
|
||
<taxonomicName authority="(Roth, 1851)" baseAuthorityName="Roth" baseAuthorityYear="1851" box="[505,892,1356,1383]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="setiger">
|
||
<emphasis bold="true" box="[505,892,1356,1383]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[505,729,1356,1383]" italics="true" pageId="4" pageNumber="5">Neosisyphus setiger</emphasis>
|
||
(Roth, 1851)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[907,999,1357,1383]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
,
|
||
<figureCitation box="[1012,1064,1357,1383]" captionStart="Fig" captionStartId="44.[106,139,640,662]" captionTargetBox="[413,766,132,615]" captionTargetId="figure@44.[413,766,131,615]" captionTargetPageId="44" captionText="Fig. 18. Images of the aedeagi of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786668" httpUri="https://zenodo.org/record/3786668/files/figure.png" pageId="4" pageNumber="5">18D</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Metatrochanter slighty inwardly-projected at the apex, metatrochanter projection is ½ the length of the metafemur. Upland woodland in southern
|
||
<collectingCountry box="[892,972,1426,1452]" name="Angola" pageId="4" pageNumber="5">Angola</collectingCountry>
|
||
............ .................................
|
||
<emphasis bold="true" box="[406,894,1463,1490]" pageId="4" pageNumber="5">
|
||
<taxonomicName authority="Daniel & Davis" authorityName="Daniel & Davis & Sole & Scholtz" authorityYear="2020" box="[406,829,1463,1490]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tembyi" status="sp. n">
|
||
<emphasis bold="true" box="[406,635,1463,1490]" italics="true" pageId="4" pageNumber="5">Neosisyphus tembyi</emphasis>
|
||
Daniel & Davis
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[837,894,1464,1490]" pageId="4" pageNumber="5" rank="species">sp. n</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[909,1001,1464,1490]" captionStart="Fig" captionStartId="42.[106,139,1599,1621]" captionTargetBox="[225,954,839,1572]" captionTargetId="figure@42.[225,954,839,1573]" captionTargetPageId="42" captionText="Fig. 17. Images of the habitus of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n, arrow showing a diagnostic feature for the new species; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786666" httpUri="https://zenodo.org/record/3786666/files/figure.png" pageId="4" pageNumber="5">Figs. 17</figureCitation>
|
||
,
|
||
<figureCitation box="[1014,1064,1464,1490]" captionStart="Fig" captionStartId="44.[106,139,640,662]" captionTargetBox="[413,766,132,615]" captionTargetId="figure@44.[413,766,131,615]" captionTargetPageId="44" captionText="Fig. 18. Images of the aedeagi of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786668" httpUri="https://zenodo.org/record/3786668/files/figure.png" pageId="4" pageNumber="5">18C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[106,1074,237,1594]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
7. Posterior margin of the mesoleg with spine or projection distally on mesofemur and basally on mesotibia; first quarter of mesotibia from femoro-tibial joint abruptly narrowed ................................
|
||
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1818" box="[608,703,1567,1593]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="spinipes">
|
||
<emphasis bold="true" box="[608,703,1567,1593]" italics="true" pageId="4" pageNumber="5">spinipes</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[878,968,1568,1594]" captionStart="Fig" captionStartId="47.[98,131,848,870]" captionTargetBox="[280,884,131,823]" captionTargetId="figure@47.[280,884,131,823]" captionTargetPageId="47" captionText="Fig. 20. Images of the habitus of the spinipes species-group: A. N. fortuitus, arrow indicating the main diagnostic feature of the species-group; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786672" httpUri="https://zenodo.org/record/3786672/files/figure.png" pageId="4" pageNumber="5">Figs. 20</figureCitation>
|
||
A–C)
|
||
<emphasis bold="true" box="[1038,1074,1567,1593]" pageId="4" pageNumber="5">(8)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" box="[98,1066,136,163]" pageId="5" pageNumber="6">
|
||
<keyLead box="[98,1066,136,163]" pageId="5" pageNumber="6">
|
||
– Mesotibial spine projecting posteriorly and gradually narrowing distally........(
|
||
<emphasis bold="true" box="[1025,1066,136,162]" pageId="5" pageNumber="6">10)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
8. Spine present on the mid-posterior margin of the metafemur; metatrochanter relatively short, ¼ of the metafemur length...........................................................
|
||
<emphasis bold="true" box="[1031,1065,205,231]" pageId="5" pageNumber="6">(9)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Spine absent on posterior margin of the metafemur; metatrochanter is ½ the length of the metafemur in major males. Shaded vegetation and thickets from east to southern Africa........
|
||
<taxonomicName authority="(Peringuey, 1901)" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[417,889,311,339]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fortuitus">
|
||
<emphasis bold="true" box="[417,889,311,339]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[417,669,312,339]" italics="true" pageId="5" pageNumber="6">Neosisyphus fortuitus</emphasis>
|
||
(
|
||
<bibRefCitation author="Peringuey" box="[683,880,311,338]" firstAuthor="Peringuey" pageId="5" pageNumber="6" pagination="1 - 563" refId="ref30622" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Transactions of the South African philosophical Society. Cape Town, 12: 1 - 563 (13 - 368)." type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[904,997,312,339]" captionStart="Fig" captionStartId="47.[98,131,848,870]" captionTargetBox="[280,884,131,823]" captionTargetId="figure@47.[280,884,131,823]" captionTargetPageId="47" captionText="Fig. 20. Images of the habitus of the spinipes species-group: A. N. fortuitus, arrow indicating the main diagnostic feature of the species-group; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786672" httpUri="https://zenodo.org/record/3786672/files/figure.png" pageId="5" pageNumber="6">Figs. 20</figureCitation>
|
||
,
|
||
<figureCitation box="[1009,1058,313,339]" captionStart="Fig" captionStartId="48.[106,139,567,589]" captionTargetBox="[315,867,131,540]" captionTargetId="figure@48.[313,867,131,541]" captionTargetPageId="48" captionText="Fig. 21. Images for the the aedeagi of the spinipes species-group: A. N. fortuitus; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786674" httpUri="https://zenodo.org/record/3786674/files/figure.png" pageId="5" pageNumber="6">21A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
9. Distinct triangular shaped projection present on the basal posterior margin of the mesotibia. Parameres not curved at the apex (
|
||
<figureCitation box="[657,762,382,408]" captionStart="Fig" captionStartId="48.[106,139,567,589]" captionTargetBox="[315,867,131,540]" captionTargetId="figure@48.[313,867,131,541]" captionTargetPageId="48" captionText="Fig. 21. Images for the the aedeagi of the spinipes species-group: A. N. fortuitus; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786674" httpUri="https://zenodo.org/record/3786674/files/figure.png" pageId="5" pageNumber="6">Fig. 21C</figureCitation>
|
||
). Moist upland and coastal grassland from east to southern Africa ................................................................. ............................................
|
||
<taxonomicName authority="(Thunberg, 1818)" baseAuthorityName="Thunberg" baseAuthorityYear="1818" box="[477,940,450,477]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinipes">
|
||
<emphasis bold="true" box="[477,940,450,477]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[477,720,450,477]" italics="true" pageId="5" pageNumber="6">Neosisyphus spinipes</emphasis>
|
||
(
|
||
<bibRefCitation author="Thunberg" box="[735,931,450,476]" firstAuthor="Thunberg" pageId="5" pageNumber="6" refId="ref31074" refString="Thunberg, C. P. (1818) Coleoptera Capensia, Antennis Lamellatis, sive, clava fissili instructa, descripta, memoires de l'Acad. T. VI." type="book" year="1818">Thunberg, 1818</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[955,1058,451,477]" captionStart="Fig" captionStartId="47.[98,131,848,870]" captionTargetBox="[280,884,131,823]" captionTargetId="figure@47.[280,884,131,823]" captionTargetPageId="47" captionText="Fig. 20. Images of the habitus of the spinipes species-group: A. N. fortuitus, arrow indicating the main diagnostic feature of the species-group; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786672" httpUri="https://zenodo.org/record/3786672/files/figure.png" pageId="5" pageNumber="6">Fig. 20C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Lacking a triangular protuberance on the basal posterior margin of the mesotibia in minor males. There is a slight sinuosity in major males. Parameres curved downward apically (
|
||
<figureCitation box="[376,478,555,581]" captionStart="Fig" captionStartId="48.[106,139,567,589]" captionTargetBox="[315,867,131,540]" captionTargetId="figure@48.[313,867,131,541]" captionTargetPageId="48" captionText="Fig. 21. Images for the the aedeagi of the spinipes species-group: A. N. fortuitus; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786674" httpUri="https://zenodo.org/record/3786674/files/figure.png" pageId="5" pageNumber="6">Fig. 21B</figureCitation>
|
||
). Lowland dry savanna from east to southern Africa .................................................
|
||
<taxonomicName authority="(Klug, 1855)" baseAuthorityName="Klug" baseAuthorityYear="1855" box="[512,943,592,619]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="infuscatus">
|
||
<emphasis bold="true" box="[512,943,592,619]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[512,781,592,619]" italics="true" pageId="5" pageNumber="6">Neosisyphus infuscatus</emphasis>
|
||
(
|
||
<bibRefCitation author="Klug" box="[796,934,592,618]" firstAuthor="Klug" pageId="5" pageNumber="6" pagination="643 - 660" refId="ref29765" refString="Klug, J. C. F. (1855) Diagnosen neuer Coleoptera aus Mossambique. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 643 - 660." type="book chapter" year="1855">Klug, 1855</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[958,1057,592,619]" captionStart="Fig" captionStartId="47.[98,131,848,870]" captionTargetBox="[280,884,131,823]" captionTargetId="figure@47.[280,884,131,823]" captionTargetPageId="47" captionText="Fig. 20. Images of the habitus of the spinipes species-group: A. N. fortuitus, arrow indicating the main diagnostic feature of the species-group; B. N. infuscatus; C. N. spinipes." figureDoi="http://doi.org/10.5281/zenodo.3786672" httpUri="https://zenodo.org/record/3786672/files/figure.png" pageId="5" pageNumber="6">Fig. 20B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
10. Basal posterior spine on mesotibia projecting perpendicularly; body uniformly brown; size of beetles 6.0–
|
||
<quantity box="[449,530,662,688]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="mm" value="10.0">10 mm</quantity>
|
||
.........
|
||
<taxonomicName baseAuthorityName="Paschalidis" baseAuthorityYear="1974" box="[602,682,661,688]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rubrus">
|
||
<emphasis bold="true" box="[602,682,661,688]" italics="true" pageId="5" pageNumber="6">rubrus</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[858,947,662,688]" captionStart="Fig" captionStartId="54.[106,139,1599,1621]" captionTargetBox="[298,881,1161,1573]" captionTargetId="figure@54.[297,881,1161,1574]" captionTargetPageId="54" captionText="Fig. 29. Images of the habitus of the rubrus species-group, arrow showing the main diagnostic feature for the species-group: A. N. macrorubrus; B. N. rubrus." figureDoi="http://doi.org/10.5281/zenodo.3786693" httpUri="https://zenodo.org/record/3786693/files/figure.png" pageId="5" pageNumber="6">Figs. 29</figureCitation>
|
||
A–B)
|
||
<emphasis bold="true" box="[1015,1065,661,687]" pageId="5" pageNumber="6">(11)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Basal posterior spine on mesotibia not projecting perpendicularly to the ventral apical margin of the femur; colour is not uniform overall, size of beetles 11.0–
|
||
<quantity box="[149,254,768,795]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="6" unit="mm" value="13.5">13.5 mm</quantity>
|
||
......................................
|
||
<taxonomicName authorityName="Gory" authorityYear="1833" box="[541,682,768,795]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis bold="true" box="[541,682,768,795]" italics="true" pageId="5" pageNumber="6">quadricollis</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[858,946,768,795]" captionStart="Fig" captionStartId="51.[98,131,471,493]" captionTargetId="figure@51.[320,843,131,445]" captionTargetPageId="51" captionText="Fig. 25. Distribution map for the tibialis species-group." figureDoi="http://doi.org/10.5281/zenodo.3786685" httpUri="https://zenodo.org/record/3786685/files/figure.png" pageId="5" pageNumber="6">Figs. 25</figureCitation>
|
||
A–B)
|
||
<emphasis bold="true" box="[1015,1066,768,794]" pageId="5" pageNumber="6">(12)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
11. Parameres with protrusion apical-laterally (
|
||
<figureCitation box="[643,749,803,829]" captionStart="Fig" captionStartId="55.[98,131,519,541]" captionTargetBox="[402,756,132,493]" captionTargetId="figure@55.[402,761,131,493]" captionTargetPageId="55" captionText="Fig. 30. Images of the aedeagi of the rubrus species-group: A. N. macrorubrus; B. N. rubrus." figureDoi="http://doi.org/10.5281/zenodo.3786695" httpUri="https://zenodo.org/record/3786695/files/figure.png" pageId="5" pageNumber="6">Fig. 30B</figureCitation>
|
||
). Upland and coastal moist grassland and savanna in southern Africa............................................................. ............................................
|
||
<taxonomicName authority="(Paschalidis, 1974)" baseAuthorityName="Paschalidis" baseAuthorityYear="1974" box="[479,943,872,899]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rubrus">
|
||
<emphasis bold="true" box="[479,943,872,899]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[479,708,872,899]" italics="true" pageId="5" pageNumber="6">Neosisyphus rubrus</emphasis>
|
||
(
|
||
<bibRefCitation author="Paschalidis, K. M." box="[722,934,872,898]" pageId="5" pageNumber="6" refId="ref30542" refString="Paschalidis, K. M. (1974 a) The genus Sisyphus Latr. (Coleoptera: Scarabaeidae) in Southern Africa. MSc. Thesis, Rhodes University, Grahamstown, South Africa, 183 pp." type="book" year="1974">Paschalidis, 1974</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[958,1057,872,899]" captionStart="Fig" captionStartId="54.[106,139,1599,1621]" captionTargetBox="[298,881,1161,1573]" captionTargetId="figure@54.[297,881,1161,1574]" captionTargetPageId="54" captionText="Fig. 29. Images of the habitus of the rubrus species-group, arrow showing the main diagnostic feature for the species-group: A. N. macrorubrus; B. N. rubrus." figureDoi="http://doi.org/10.5281/zenodo.3786693" httpUri="https://zenodo.org/record/3786693/files/figure.png" pageId="5" pageNumber="6">Fig. 29B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Parameres simple (
|
||
<figureCitation box="[366,472,907,933]" captionStart="Fig" captionStartId="55.[98,131,519,541]" captionTargetBox="[402,756,132,493]" captionTargetId="figure@55.[402,761,131,493]" captionTargetPageId="55" captionText="Fig. 30. Images of the aedeagi of the rubrus species-group: A. N. macrorubrus; B. N. rubrus." figureDoi="http://doi.org/10.5281/zenodo.3786695" httpUri="https://zenodo.org/record/3786695/files/figure.png" pageId="5" pageNumber="6">Fig. 30A</figureCitation>
|
||
). Cool, dry savanna, karoo and grassland in
|
||
<collectingCountry name="South Africa" pageId="5" pageNumber="6">South Africa</collectingCountry>
|
||
(
|
||
<collectingRegion box="[232,407,942,968]" country="South Africa" name="Northern Cape" pageId="5" pageNumber="6">Northern Cape</collectingRegion>
|
||
and
|
||
<collectingRegion box="[463,575,942,968]" country="South Africa" name="Free State" pageId="5" pageNumber="6">Free State</collectingRegion>
|
||
) and
|
||
<collectingCountry box="[639,738,942,968]" name="Namibia" pageId="5" pageNumber="6">Namibia</collectingCountry>
|
||
(Etosha Pan, Okaukuejo) ... ..................................
|
||
<taxonomicName authority="(Paschalidis, 1974)" baseAuthorityName="Paschalidis" baseAuthorityYear="1974" box="[405,943,978,1005]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macrorubrus">
|
||
<emphasis bold="true" box="[405,943,978,1005]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[405,707,978,1005]" italics="true" pageId="5" pageNumber="6">Neosisyphus macrorubrus</emphasis>
|
||
(
|
||
<bibRefCitation author="Paschalidis, K. M." box="[722,933,978,1004]" pageId="5" pageNumber="6" refId="ref30542" refString="Paschalidis, K. M. (1974 a) The genus Sisyphus Latr. (Coleoptera: Scarabaeidae) in Southern Africa. MSc. Thesis, Rhodes University, Grahamstown, South Africa, 183 pp." type="book" year="1974">Paschalidis, 1974</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[957,1058,979,1005]" captionStart="Fig" captionStartId="54.[106,139,1599,1621]" captionTargetBox="[298,881,1161,1573]" captionTargetId="figure@54.[297,881,1161,1574]" captionTargetPageId="54" captionText="Fig. 29. Images of the habitus of the rubrus species-group, arrow showing the main diagnostic feature for the species-group: A. N. macrorubrus; B. N. rubrus." figureDoi="http://doi.org/10.5281/zenodo.3786693" httpUri="https://zenodo.org/record/3786693/files/figure.png" pageId="5" pageNumber="6">Fig. 29A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
12. Lateral margin of elytra emarginate; concavity of margin distinctly deep in the middle; short projecting metatrochanter in males (1/3 of metafemur length); metafemur lacking spines. Deep sands and shrubland of southwest coast, and karoo in the
|
||
<collectingRegion box="[294,444,1118,1144]" country="South Africa" name="Eastern Cape" pageId="5" pageNumber="6">Eastern Cape</collectingRegion>
|
||
(
|
||
<collectingCountry box="[459,605,1118,1145]" name="South Africa" pageId="5" pageNumber="6">South Africa</collectingCountry>
|
||
) ............................................................ .......................................
|
||
<taxonomicName authority="(Gory, 1833)" baseAuthorityName="Gory" baseAuthorityYear="1833" box="[437,889,1152,1179]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis bold="true" box="[437,889,1152,1179]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[437,726,1152,1179]" italics="true" pageId="5" pageNumber="6">Neosisyphus quadricollis</emphasis>
|
||
(
|
||
<bibRefCitation author="Gory" box="[742,880,1152,1178]" firstAuthor="Gory" pageId="5" pageNumber="6" pagination="1 - 15" refId="ref29509" refString="Gory, M. (1833) Monographie du genre Sisyphe. Mequignon-Marvis pere et fils, Paris. 1 - 15 p." type="book chapter" year="1833">Gory, 1833</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[904,997,1152,1178]" captionStart="Fig" captionStartId="52.[106,139,1173,1195]" captionTargetBox="[287,889,762,1148]" captionTargetId="figure@52.[286,894,762,1148]" captionTargetPageId="52" captionText="Fig. 26. Image of habitus of the quadricollis species-group: A. N. quadricollis, arrow showing the main diagnostic feature of the species-group; B. N. kuehni." figureDoi="http://doi.org/10.5281/zenodo.3786687" httpUri="https://zenodo.org/record/3786687/files/figure.png" pageId="5" pageNumber="6">Figs. 26</figureCitation>
|
||
,
|
||
<figureCitation box="[1009,1058,1153,1179]" captionStart="Fig" captionStartId="52.[106,139,1645,1667]" captionTargetBox="[370,811,1262,1619]" captionTargetId="figure@52.[369,811,1262,1620]" captionTargetPageId="52" captionText="Fig. 27. Images of the aedeagi of the quadricollis species-group: A. N. quadricollis; B. N. kuehni." figureDoi="http://doi.org/10.5281/zenodo.3786689" httpUri="https://zenodo.org/record/3786689/files/figure.png" pageId="5" pageNumber="6">27A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Margin of elytra with regular shape; a protuberance distally on the posterior margin of the metafemur, the metatrochanter exceeds ½ the length of the metafemur in males. Moist highland grassland in
|
||
<collectingCountry box="[579,722,1256,1282]" name="South Africa" pageId="5" pageNumber="6">South Africa</collectingCountry>
|
||
.............................................. ...............................................
|
||
<taxonomicName authority="(Haaf, 1955)" baseAuthorityName="Haaf" baseAuthorityYear="1955" box="[497,890,1293,1320]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis bold="true" box="[497,890,1293,1320]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[497,728,1293,1320]" italics="true" pageId="5" pageNumber="6">Neosisyphus kuehni</emphasis>
|
||
(
|
||
<bibRefCitation author="Haaf" box="[743,880,1293,1319]" firstAuthor="Haaf" pageId="5" pageNumber="6" pagination="341 - 381" refId="ref29533" refString="Haaf, E. (1955) Uber die Gattung Sisyphus Latr. (Col. Scarab.). Entomologische Arbeitenaus dem Museum G. Frey, 6 (1): 341 - 381." type="journal article" year="1955">Haaf, 1955</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[904,997,1294,1320]" captionStart="Fig" captionStartId="52.[106,139,1173,1195]" captionTargetBox="[287,889,762,1148]" captionTargetId="figure@52.[286,894,762,1148]" captionTargetPageId="52" captionText="Fig. 26. Image of habitus of the quadricollis species-group: A. N. quadricollis, arrow showing the main diagnostic feature of the species-group; B. N. kuehni." figureDoi="http://doi.org/10.5281/zenodo.3786687" httpUri="https://zenodo.org/record/3786687/files/figure.png" pageId="5" pageNumber="6">Figs. 26</figureCitation>
|
||
,
|
||
<figureCitation box="[1009,1058,1294,1320]" captionStart="Fig" captionStartId="52.[106,139,1645,1667]" captionTargetBox="[370,811,1262,1619]" captionTargetId="figure@52.[369,811,1262,1620]" captionTargetPageId="52" captionText="Fig. 27. Images of the aedeagi of the quadricollis species-group: A. N. quadricollis; B. N. kuehni." figureDoi="http://doi.org/10.5281/zenodo.3786689" httpUri="https://zenodo.org/record/3786689/files/figure.png" pageId="5" pageNumber="6">27B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
13. Setae on the elytral interstriae arranged in tufts ................................................... ..........................................................
|
||
<taxonomicName authorityName="Olivier" authorityYear="1789" box="[579,699,1363,1389]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="muricatus">
|
||
<emphasis bold="true" box="[579,699,1363,1389]" italics="true" pageId="5" pageNumber="6">muricatus</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[874,948,1363,1389]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
|
||
A–F)
|
||
<emphasis bold="true" box="[1015,1066,1362,1388]" pageId="5" pageNumber="6">(14)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" box="[98,1066,1400,1427]" pageId="5" pageNumber="6">
|
||
<keyLead box="[98,1066,1400,1427]" pageId="5" pageNumber="6">
|
||
– Setae on the elytral interstriae evenly distributed, not grouped in tufts ..........
|
||
<emphasis bold="true" box="[1015,1066,1400,1426]" pageId="5" pageNumber="6">(20)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
14. Setae arranged in tufts on the pronotum, which alternate with depressed bare patches ..........................................................................................................
|
||
<emphasis bold="true" box="[1015,1066,1469,1495]" pageId="5" pageNumber="6">(15)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[98,1066,136,1638]" box="[98,1066,1506,1534]" pageId="5" pageNumber="6">
|
||
<keyLead box="[98,1066,1506,1534]" pageId="5" pageNumber="6">
|
||
– Setae on the pronotum not arranged in tufts .................................................
|
||
<emphasis bold="true" box="[1015,1066,1506,1532]" pageId="5" pageNumber="6">(17)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[98,1066,136,1638]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
15. Concave and upwardly curved margin between the anterior medial teeth of the clypeus; macropterous (
|
||
<figureCitation box="[424,507,1576,1603]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="5" pageNumber="6">Fig. 2I</figureCitation>
|
||
). Woodland thickets and forest in north
|
||
<collectingCountry name="South Africa" pageId="5" pageNumber="6">South Africa</collectingCountry>
|
||
and
|
||
<collectingCountry box="[273,386,1611,1637]" name="Swaziland" pageId="5" pageNumber="6">Swaziland</collectingCountry>
|
||
..........
|
||
<taxonomicName authority="Boheman, 1857" authorityName="Boheman" authorityYear="1857" box="[471,916,1610,1637]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fasciculatus">
|
||
<emphasis bold="true" box="[471,916,1610,1637]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[471,718,1610,1637]" italics="true" pageId="5" pageNumber="6">Sisyphus fasciculatus</emphasis>
|
||
<bibRefCitation author="Boheman" box="[725,916,1611,1637]" firstAuthor="Boheman" pageId="5" pageNumber="6" refId="ref28273" refString="Boheman, C. H. (1857) Insecta Caffrariae annis 1838 - 1845 a JA Wahlberg collecta. Pars. II. Coleoptera (Scarabaeides). Holmiae: Offieina Nordstedtiana. - 395 pp, 1." type="book" year="1857">Boheman, 1857</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[931,1009,1611,1637]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation box="[1021,1057,1611,1637]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="5" pageNumber="6">3C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Anterior margin between medial clypeal teeth not upcurved; hind wings reduced or completely brachypterous (
|
||
<figureCitation box="[494,569,172,198]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
|
||
H–J) .....................................................(
|
||
<emphasis bold="true" box="[1033,1064,171,197]" pageId="6" pageNumber="7">16</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
16. Anterior margin between medial clypeal teeth concave; pronotum wider anteriorly than posteriorly. Parameres (
|
||
<figureCitation box="[477,563,241,267]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="6" pageNumber="7">Fig. 3F</figureCitation>
|
||
). Only known from
|
||
<collectingCountry box="[797,910,241,267]" name="Swaziland" pageId="6" pageNumber="7">Swaziland</collectingCountry>
|
||
..................... ...................................................
|
||
<emphasis bold="true" box="[533,967,275,302]" pageId="6" pageNumber="7">
|
||
<taxonomicName authority="Daniel & Davis" authorityName="Daniel & Davis & Sole & Scholtz" authorityYear="2020" box="[533,902,275,302]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="swazi" status="sp. n">
|
||
<emphasis bold="true" box="[533,708,275,302]" italics="true" pageId="6" pageNumber="7">Sisyphus swazi</emphasis>
|
||
Daniel & Davis
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[910,967,276,302]" pageId="6" pageNumber="7" rank="species">sp. n</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[982,1065,276,302]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Fig. 2F</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Straight edge between medial teeth of clypeus; width of pronotum uniform from anterior to posterior margins. Parameres (
|
||
<figureCitation box="[634,727,345,371]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="6" pageNumber="7">Fig. 3G</figureCitation>
|
||
). Popa Falls forest (
|
||
<collectingCountry box="[956,1061,345,372]" name="Namibia" pageId="6" pageNumber="7">Namibia</collectingCountry>
|
||
), Central
|
||
<collectingCountry box="[249,399,380,406]" name="Mozambique" pageId="6" pageNumber="7">Mozambique</collectingCountry>
|
||
and East Africa.................................................................. ..................................................
|
||
<taxonomicName authority="Boucomont, 1935" authorityName="Boucomont" authorityYear="1935" box="[529,960,416,443]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alveatus">
|
||
<emphasis bold="true" box="[529,960,416,443]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[529,736,416,443]" italics="true" pageId="6" pageNumber="7">Sisyphus alveatus</emphasis>
|
||
<bibRefCitation author="Boucomont" box="[743,960,417,443]" firstAuthor="Boucomont" pageId="6" pageNumber="7" pagination="279 - 290" refId="ref28313" refString="Boucomont, A. (1935) Coleoptera. VI. Scarabaeidae Scarabaeini. In: Jeannel, R. (Ed.), Mission Scientifique de l'Omo. Tome 2 Zoologie, Fascicule 16. Paul Chevalier & Fils, Paris, France, pp. 279 - 290." type="book chapter" year="1935">Boucomont, 1935</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[975,1065,417,443]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Fig. 2G</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
17. Very strong lateral emargination and distinct shiny punctation on pronotum. Cooler southern forest and montane grassland in
|
||
<collectingCountry box="[716,859,486,512]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
|
||
............................ ................................................
|
||
<taxonomicName authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[511,926,520,547]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="muricatus">
|
||
<emphasis bold="true" box="[511,926,520,547]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[511,740,520,547]" italics="true" pageId="6" pageNumber="7">Sisyphus muricatus</emphasis>
|
||
(
|
||
<bibRefCitation author="Olivier" box="[755,916,520,546]" firstAuthor="Olivier" pageId="6" pageNumber="7" pagination="1 - 455" refId="ref30408" refString="Olivier, M. (1789) Entomologie, Histoire Naturelle des Insectes, Avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres, Tome premier, Paris, 1 - 455 pp." type="book chapter" year="1789">Olivier, 1789</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[941,1019,521,547]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation box="[1031,1065,521,547]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="6" pageNumber="7">3A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" box="[106,1073,558,585]" pageId="6" pageNumber="7">
|
||
<keyLead box="[106,1073,558,585]" pageId="6" pageNumber="7">
|
||
– No distinct lateral emargination on the pronotum.........................................(
|
||
<emphasis bold="true" box="[1033,1064,558,584]" pageId="6" pageNumber="7">18</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
18. Punctation on the pronotum unclear; interstriae 1, 3, 5, and 7 with rows of black setae arranged in tufts, interstriae 2, 4, and 6 with sparsely arranged single setae. Coastal sand forest of northeast
|
||
<collectingCountry box="[520,662,662,688]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
|
||
and southeast
|
||
<collectingCountry box="[831,981,662,688]" name="Mozambique" pageId="6" pageNumber="7">Mozambique</collectingCountry>
|
||
........... .....................
|
||
<taxonomicName authority="Daniel & Davis, 2016" authorityName="Daniel & Davis" authorityYear="2016" box="[317,926,696,723]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="neobornemisszanus">
|
||
<emphasis bold="true" box="[317,926,696,723]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[317,656,696,723]" italics="true" pageId="6" pageNumber="7">Sisyphus neobornemisszanus</emphasis>
|
||
<bibRefCitation author="Daniel & Davis" box="[663,926,697,723]" firstAuthor="Daniel" pageId="6" pageNumber="7" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, & Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel & Davis, 2016</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[941,1019,697,723]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation box="[1032,1065,697,723]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="6" pageNumber="7">3B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Punctation on the pronotum conspicuous; setae on pronotal disc arise from the centre of ocellate punctures (
|
||
<figureCitation box="[482,571,769,796]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="6" pageNumber="7">Fig. 5H</figureCitation>
|
||
)............................................................(
|
||
<emphasis bold="true" box="[1033,1073,768,794]" pageId="6" pageNumber="7">19)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
19. Proximal elytral setae evenly arranged; mid-basal elytral setae arranged in tufts of scattered individual setae. Pronotal disc with three prominent round depressions. Parameres (
|
||
<figureCitation box="[287,375,873,899]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="6" pageNumber="7">Fig. 3E</figureCitation>
|
||
). Primarily upland and highland grassland in
|
||
<collectingRegion box="[901,1002,873,899]" country="South Africa" name="Gauteng" pageId="6" pageNumber="7">Gauteng</collectingRegion>
|
||
, Kwa- Zulu Natal,
|
||
<collectingRegion box="[296,404,908,934]" country="South Africa" name="Limpopo" pageId="6" pageNumber="7">Limpopo</collectingRegion>
|
||
,
|
||
<collectingRegion box="[416,567,908,934]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="7">Mpumalanga</collectingRegion>
|
||
and
|
||
<collectingRegion box="[624,849,908,934]" country="Botswana" name="North West" pageId="6" pageNumber="7">Northwest Province</collectingRegion>
|
||
............................. ........................................................
|
||
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[574,965,942,969]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="manni">
|
||
<emphasis bold="true" box="[574,965,942,969]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[574,760,942,969]" italics="true" pageId="6" pageNumber="7">Sisyphus manni</emphasis>
|
||
<bibRefCitation author="Montreuil, O." box="[768,965,942,968]" pageId="6" pageNumber="7" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[981,1065,942,969]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Fig. 2E</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Elytra entirely covered in tufts of setae, each tuft dense and compact; depressions on the pronotal disc absent; body larger than
|
||
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[740,841,1012,1038]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="manni">
|
||
<emphasis box="[740,841,1012,1038]" italics="true" pageId="6" pageNumber="7">S. manni</emphasis>
|
||
</taxonomicName>
|
||
and oval. Parameres (
|
||
<figureCitation box="[165,256,1046,1073]" captionStart="Fig" captionStartId="12.[106,139,1637,1659]" captionTargetBox="[334,848,1087,1611]" captionTargetId="figure@12.[332,848,1087,1612]" captionTargetPageId="12" captionText="Fig. 3. Images of the aedeagi of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n; G. S. alveatus." figureDoi="http://doi.org/10.5281/zenodo.3786636" httpUri="https://zenodo.org/record/3786636/files/figure.png" pageId="6" pageNumber="7">Fig. 3D</figureCitation>
|
||
). Unshaded vegetation in the
|
||
<collectingRegion box="[596,746,1047,1073]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="7">Eastern Cape</collectingRegion>
|
||
........................................... ..............................................
|
||
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[500,960,1083,1110]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="perissinottoi">
|
||
<emphasis bold="true" box="[500,960,1083,1110]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[500,755,1083,1110]" italics="true" pageId="6" pageNumber="7">Sisyphus perissinottoi</emphasis>
|
||
<bibRefCitation author="Montreuil, O." box="[762,960,1083,1109]" pageId="6" pageNumber="7" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[975,1065,1084,1110]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="6" pageNumber="7">Fig. 2D</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
20. Well-developed tufts of epipleural setae (
|
||
<figureCitation box="[620,712,1118,1145]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="6" pageNumber="7">Fig. 5H</figureCitation>
|
||
); edge between medial teeth almost straight; setae on the pronotum evenly distributed ...................................... ..................................................................
|
||
<taxonomicName authorityName="Harold" authorityYear="1859" box="[647,707,1187,1213]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="goryi">
|
||
<emphasis bold="true" box="[647,707,1187,1213]" italics="true" pageId="6" pageNumber="7">goryi</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[882,955,1188,1215]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="6" pageNumber="7">Figs. 5</figureCitation>
|
||
A–E) (
|
||
<emphasis bold="true" box="[1033,1064,1187,1213]" pageId="6" pageNumber="7">22</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Weak tufts of epipleural setae; margin between medial teeth V-shaped)............... ..................................................................................
|
||
<taxonomicName authorityName="Daniel & Davis" authorityYear="2016" box="[761,913,1259,1286]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="umbraphilus">
|
||
<emphasis bold="true" box="[761,913,1259,1286]" italics="true" pageId="6" pageNumber="7">umbraphilus</emphasis>
|
||
</taxonomicName>
|
||
species-group
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
21. Pronotal disc with setae interspersed with bald patches. Dense woodland and thicket in KwaZulu Natal,
|
||
<collectingRegion box="[462,560,1329,1355]" country="South Africa" name="Gauteng" pageId="6" pageNumber="7">Gauteng</collectingRegion>
|
||
and
|
||
<collectingRegion box="[616,750,1329,1356]" country="South Africa" name="North West" pageId="6" pageNumber="7">North West</collectingRegion>
|
||
Provinces (
|
||
<collectingCountry box="[879,1025,1329,1356]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
|
||
) .... .....................................
|
||
<taxonomicName authority="Daniel & Davis, 2016" authorityName="Daniel & Davis" authorityYear="2016" box="[433,964,1363,1389]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="umbraphilus">
|
||
<emphasis bold="true" box="[433,964,1363,1389]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[433,694,1363,1389]" italics="true" pageId="6" pageNumber="7">Sisyphus umbraphilus</emphasis>
|
||
<bibRefCitation author="Daniel & Davis" box="[701,964,1363,1389]" firstAuthor="Daniel" pageId="6" pageNumber="7" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, & Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel & Davis, 2016</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[980,1065,1364,1390]" captionStart="Fig" captionStartId="25.[98,131,1628,1650]" captionTargetBox="[362,802,1074,1603]" captionTargetId="figure@25.[362,802,1074,1603]" captionTargetPageId="25" captionText="Fig. 8. Images of the habitus of the umbraphilus species-group: A. S. umbraphilus." figureDoi="http://doi.org/10.5281/zenodo.3786646" httpUri="https://zenodo.org/record/3786646/files/figure.png" pageId="6" pageNumber="7">Fig. 8A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" box="[106,1074,1400,1428]" pageId="6" pageNumber="7">
|
||
<keyLead box="[106,1074,1400,1428]" pageId="6" pageNumber="7">
|
||
– Pronotal disc without clear setae interspersed with bald patches ....................
|
||
<emphasis bold="true" box="[1023,1074,1400,1426]" pageId="6" pageNumber="7">(26)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
22. Setae on the pronotum inserted between or on the posterior margin of ocellate punctures (
|
||
<figureCitation box="[283,366,1470,1497]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="6" pageNumber="7">Fig. 5F</figureCitation>
|
||
)........................................................................................
|
||
<emphasis bold="true" box="[1023,1073,1470,1496]" pageId="6" pageNumber="7">(23)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[106,1074,134,1638]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Setae on the pronotum inserted centrally on ocellate punctures ............................ ...................................................................................................... (
|
||
<figureCitation box="[919,1008,1542,1569]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="6" pageNumber="7">Fig. 5G</figureCitation>
|
||
) (
|
||
<emphasis bold="true" box="[1033,1064,1542,1568]" pageId="6" pageNumber="7">25</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[106,1074,134,1638]" lastBlockId="7.[98,1067,134,1670]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<keyLead lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
23. Fine setae on elytra arranged in single rows, epipleural tufts containing few setae (<10); larger-bodied species. Highland grassland under cooler conditions in
|
||
<collectingCountry box="[149,295,134,160]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
|
||
,
|
||
<collectingCountry box="[306,418,135,161]" name="Botswana" pageId="7" pageNumber="8">Botswana</collectingCountry>
|
||
, and
|
||
<collectingCountry box="[479,578,134,160]" name="Namibia" pageId="7" pageNumber="8">Namibia</collectingCountry>
|
||
................................................................. ......................................................
|
||
<taxonomicName authority="Boheman, 1857" authorityName="Boheman" authorityYear="1857" box="[546,918,168,195]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="caffer">
|
||
<emphasis bold="true" box="[546,918,168,195]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[546,721,168,195]" italics="true" pageId="7" pageNumber="8">Sisyphus caffer</emphasis>
|
||
<bibRefCitation author="Boheman" box="[728,918,169,195]" firstAuthor="Boheman" pageId="7" pageNumber="8" refId="ref28273" refString="Boheman, C. H. (1857) Insecta Caffrariae annis 1838 - 1845 a JA Wahlberg collecta. Pars. II. Coleoptera (Scarabaeides). Holmiae: Offieina Nordstedtiana. - 395 pp, 1." type="book" year="1857">Boheman, 1857</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[935,1011,169,196]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="7" pageNumber="8">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation box="[1024,1057,169,196]" captionStart="Fig" captionStartId="21.[98,131,1632,1654]" captionTargetBox="[349,815,1116,1606]" captionTargetId="figure@21.[348,815,1116,1607]" captionTargetPageId="21" captionText="Fig. 6. Images of the aedagi of the goryi species-group: A. S. bicuariensis sp. n; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus." figureDoi="http://doi.org/10.5281/zenodo.3786642" httpUri="https://zenodo.org/record/3786642/files/figure.png" pageId="7" pageNumber="8">6B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Much finer setae on elytra, distributed in single rows; epipleural tufts containing many setae (>10); elongate body shape ..........................................................(
|
||
<emphasis bold="true" box="[1026,1057,241,267]" pageId="7" pageNumber="8">24</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
24. Setae on the pronotum and elytra of uniform colour and regularly arranged. Parameres (
|
||
<figureCitation box="[230,322,310,337]" captionStart="Fig" captionStartId="21.[98,131,1632,1654]" captionTargetBox="[349,815,1116,1606]" captionTargetId="figure@21.[348,815,1116,1607]" captionTargetPageId="21" captionText="Fig. 6. Images of the aedagi of the goryi species-group: A. S. bicuariensis sp. n; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus." figureDoi="http://doi.org/10.5281/zenodo.3786642" httpUri="https://zenodo.org/record/3786642/files/figure.png" pageId="7" pageNumber="8">Fig. 6D</figureCitation>
|
||
). Widespread woodland species in sub-Saharan Africa................. ...............................................................
|
||
<taxonomicName authority="Harold, 1859" authorityName="Harold" authorityYear="1859" box="[615,952,344,371]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="goryi">
|
||
<emphasis bold="true" box="[615,952,344,371]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[615,783,344,371]" italics="true" pageId="7" pageNumber="8">Sisyphus goryi</emphasis>
|
||
<bibRefCitation author="Harold" box="[790,952,345,371]" firstAuthor="Harold" pageId="7" pageNumber="8" pagination="193 - 224" refId="ref29570" refString="Harold, E. von (1859) Beitrage zur Kenntniss einiger coprophagen Lamellicornien. Deutsche Entomologische Zeitschrift, 3 (2 - 3), 193 - 224." type="journal article" year="1859">Harold, 1859</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[968,1057,345,372]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Setae on pronotum and elytra bicoloured (black and gray). Parameres (
|
||
<figureCitation box="[963,1054,380,407]" captionStart="Fig" captionStartId="21.[98,131,1632,1654]" captionTargetBox="[349,815,1116,1606]" captionTargetId="figure@21.[348,815,1116,1607]" captionTargetPageId="21" captionText="Fig. 6. Images of the aedagi of the goryi species-group: A. S. bicuariensis sp. n; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus." figureDoi="http://doi.org/10.5281/zenodo.3786642" httpUri="https://zenodo.org/record/3786642/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
|
||
). Open woodland in southern
|
||
<collectingCountry box="[478,558,414,440]" name="Angola" pageId="7" pageNumber="8">Angola</collectingCountry>
|
||
.................................................................... .......................................
|
||
<emphasis bold="true" box="[439,957,451,478]" pageId="7" pageNumber="8">
|
||
<taxonomicName authority="Daniel & Davis" authorityName="Daniel & Davis & Sole & Scholtz" authorityYear="2020" box="[439,886,451,478]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bicuariensis" status="sp. n">
|
||
<emphasis bold="true" box="[439,690,451,478]" italics="true" pageId="7" pageNumber="8">Sisyphus bicuariensis</emphasis>
|
||
Daniel & Davis
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[899,957,452,478]" pageId="7" pageNumber="8" rank="species">sp. n</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[972,1058,452,479]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="7" pageNumber="8">Fig. 5A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
25. Elytral setae thick, arranged in non-single rows and forming bunches; body size: 6.6–
|
||
<quantity box="[200,295,521,548]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="8" unit="mm" value="4.4">4.4 mm</quantity>
|
||
. Open lowland vegetation in
|
||
<collectingCountry box="[633,780,521,547]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
|
||
,
|
||
<collectingCountry box="[792,945,521,547]" name="Mozambique" pageId="7" pageNumber="8">Mozambique</collectingCountry>
|
||
, and
|
||
<collectingCountry name="Zimbabwe" pageId="7" pageNumber="8">Zimbabwe</collectingCountry>
|
||
.......................................
|
||
<taxonomicName authority="Boheman, 1857" authorityName="Boheman" authorityYear="1857" box="[513,919,555,582]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sordidus">
|
||
<emphasis bold="true" box="[513,919,555,582]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[513,721,555,582]" italics="true" pageId="7" pageNumber="8">Sisyphus sordidus</emphasis>
|
||
<bibRefCitation author="Boheman" box="[728,919,555,582]" firstAuthor="Boheman" pageId="7" pageNumber="8" refId="ref28273" refString="Boheman, C. H. (1857) Insecta Caffrariae annis 1838 - 1845 a JA Wahlberg collecta. Pars. II. Coleoptera (Scarabaeides). Holmiae: Offieina Nordstedtiana. - 395 pp, 1." type="book" year="1857">Boheman, 1857</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[934,1011,556,583]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="7" pageNumber="8">
|
||
Figs.
|
||
<geoCoordinate box="[995,1011,556,583]" degrees="5, 6" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5555" value="5.6">5</geoCoordinate>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation box="[1025,1058,556,583]" captionStart="Fig" captionStartId="21.[98,131,1632,1654]" captionTargetBox="[349,815,1116,1606]" captionTargetId="figure@21.[348,815,1116,1607]" captionTargetPageId="21" captionText="Fig. 6. Images of the aedagi of the goryi species-group: A. S. bicuariensis sp. n; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus." figureDoi="http://doi.org/10.5281/zenodo.3786642" httpUri="https://zenodo.org/record/3786642/files/figure.png" pageId="7" pageNumber="8">6E</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Elytral setae fine, regularly arranged on interstriae; body size: 10–8.0 mm. East African miombo woodland; known only from Quirimbas National Park (Northern
|
||
<collectingCountry box="[149,301,662,689]" name="Mozambique" pageId="7" pageNumber="8">Mozambique</collectingCountry>
|
||
).....................
|
||
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[466,916,662,689]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="genierorum">
|
||
<emphasis bold="true" box="[466,916,662,689]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[466,711,662,689]" italics="true" pageId="7" pageNumber="8">Sisyphus genierorum</emphasis>
|
||
<bibRefCitation author="Montreuil, O." box="[718,916,662,688]" pageId="7" pageNumber="8" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[931,1008,662,689]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="7" pageNumber="8">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation box="[1021,1057,662,689]" captionStart="Fig" captionStartId="21.[98,131,1632,1654]" captionTargetBox="[349,815,1116,1606]" captionTargetId="figure@21.[348,815,1116,1607]" captionTargetPageId="21" captionText="Fig. 6. Images of the aedagi of the goryi species-group: A. S. bicuariensis sp. n; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus." figureDoi="http://doi.org/10.5281/zenodo.3786642" httpUri="https://zenodo.org/record/3786642/files/figure.png" pageId="7" pageNumber="8">6C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
26. Setae on outer margin of elytra not arranged in tufts; margin between medial teeth concave or straight; very large ocellate punctures on dorsal surface of pronotal disc......................................................
|
||
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1818" box="[590,683,766,792]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis bold="true" box="[590,683,766,792]" italics="true" pageId="7" pageNumber="8">costatus</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[858,943,766,793]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Figs. 11</figureCitation>
|
||
A–E)
|
||
<emphasis bold="true" box="[1015,1066,766,792]" pageId="7" pageNumber="8">(27)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Setae on outer margin of elytra not arranged in tufts; margin between medial teeth narrowed (almost V-shaped); very fine ocellate punctures on dorsal surface of pronotal disc.........................................
|
||
<taxonomicName authorityName="Gerstaecker" authorityYear="1871" box="[563,695,872,899]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="seminulum">
|
||
<emphasis bold="true" box="[563,695,872,899]" italics="true" pageId="7" pageNumber="8">seminulum</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation box="[870,944,873,900]" captionStart="Fig" captionStartId="21.[98,131,1632,1654]" captionTargetBox="[349,815,1116,1606]" captionTargetId="figure@21.[348,815,1116,1607]" captionTargetPageId="21" captionText="Fig. 6. Images of the aedagi of the goryi species-group: A. S. bicuariensis sp. n; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus." figureDoi="http://doi.org/10.5281/zenodo.3786642" httpUri="https://zenodo.org/record/3786642/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
|
||
A–C)
|
||
<emphasis bold="true" box="[1015,1066,872,898]" pageId="7" pageNumber="8">(30)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
27. Dorsal surface of the clypeus and frons with inconspicuous ocellate punctation, but more visible on the vertex (
|
||
<figureCitation box="[501,588,942,969]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Figs. 11</figureCitation>
|
||
G–I)..................................................
|
||
<emphasis bold="true" box="[1015,1066,942,968]" pageId="7" pageNumber="8">(28)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Dorsal surface of the head (clypeus, frons and vertex) with distinct ocellate punctation (
|
||
<figureCitation box="[231,345,1014,1041]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Figure11J</figureCitation>
|
||
)..........................................................................................
|
||
<emphasis bold="true" box="[1015,1066,1014,1040]" pageId="7" pageNumber="8">(30)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
28. Vertex with dense ocellate punctation, margin between medial teeth straight; genae convex anteriorly (
|
||
<figureCitation box="[408,501,1084,1110]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11I</figureCitation>
|
||
); elytral setae fine and uniformly arranged. Moist savanna or dense woodland and riverine vegetation in the interior of
|
||
<collectingCountry box="[917,1063,1118,1144]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
|
||
. Shaded vegetation on coastline from
|
||
<collectingRegion box="[576,644,1153,1179]" country="Mozambique" name="Sofala" pageId="7" pageNumber="8">Sofala</collectingRegion>
|
||
Bay (
|
||
<collectingCountry box="[711,864,1153,1180]" name="Mozambique" pageId="7" pageNumber="8">Mozambique</collectingCountry>
|
||
) to
|
||
<collectingRegion box="[914,1066,1153,1180]" country="South Africa" name="Eastern Cape" pageId="7" pageNumber="8">Eastern Cape</collectingRegion>
|
||
(
|
||
<collectingCountry box="[157,303,1188,1215]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
|
||
)................
|
||
<emphasis bold="true" box="[430,957,1187,1214]" pageId="7" pageNumber="8">
|
||
<taxonomicName authority="Daniel & Davis" authorityName="Daniel & Davis & Sole & Scholtz" authorityYear="2020" box="[430,887,1187,1214]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="inconspicuus" status="sp. n">
|
||
<emphasis bold="true" box="[430,691,1187,1214]" italics="true" pageId="7" pageNumber="8">Sisyphus inconspicuus</emphasis>
|
||
Daniel & Davis
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[900,957,1188,1214]" pageId="7" pageNumber="8" rank="species">sp. n</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[973,1057,1188,1215]" captionStart="Fig" captionStartId="20.[106,139,1467,1489]" captionTargetBox="[107,1071,525,1442]" captionTargetId="figure@20.[106,1074,525,1442]" captionTargetPageId="20" captionText="Fig. 5. Images of the habitus of the goryi species-group: A. S. bicuariensis sp. n; with enphasis of the black and gray setae on pronotum and elytra; B. S. caffer; C. S. genierorum; D. S. goryi; E. S. sordidus.F. Pronotal setae originate from the posterior edge or between ocellate points. G. Pronotal setae arise from the centre of ocellate punctures, commonly seen in S. genierorum, S. sordidus; S. perissinottoi; S. manni and S. swazi sp. n. H. Tufts of setae on the epipleurae visible from above." figureDoi="http://doi.org/10.5281/zenodo.3786640" httpUri="https://zenodo.org/record/3786640/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Vertex with scattered ocellate punctation (
|
||
<figureCitation box="[618,695,1222,1249]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
|
||
G–H), elytral setae thicker, alternating on the interstriae between rows of well-developed and less developed setae (
|
||
<figureCitation box="[157,254,1294,1321]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11F</figureCitation>
|
||
) ......................................................................................................(
|
||
<emphasis bold="true" box="[1025,1066,1294,1320]" pageId="7" pageNumber="8">29)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
29. Margin between medial teeth distinctly concave and upcurved; lacking carinae on the frons; genae virtually straight laterally (
|
||
<figureCitation box="[685,793,1364,1390]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11G</figureCitation>
|
||
). Parameres (
|
||
<figureCitation box="[948,1053,1364,1390]" captionStart="Fig" captionStartId="29.[98,131,1599,1621]" captionTargetBox="[346,816,1085,1574]" captionTargetId="figure@29.[346,817,1081,1574]" captionTargetPageId="29" captionText="Fig. 12. Images of the aedeagi of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786654" httpUri="https://zenodo.org/record/3786654/files/figure.png" pageId="7" pageNumber="8">Fig. 12B</figureCitation>
|
||
). Highland grassland in
|
||
<collectingCountry box="[405,548,1398,1424]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
|
||
..................................................................... ..................................................
|
||
<taxonomicName authority="(Thunberg, 1818)" baseAuthorityName="Thunberg" baseAuthorityYear="1818" box="[521,943,1432,1459]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis bold="true" box="[521,943,1432,1459]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[521,722,1432,1459]" italics="true" pageId="7" pageNumber="8">Sisyphus costatus</emphasis>
|
||
(
|
||
<bibRefCitation author="Thunberg" box="[737,934,1432,1458]" firstAuthor="Thunberg" pageId="7" pageNumber="8" refId="ref31074" refString="Thunberg, C. P. (1818) Coleoptera Capensia, Antennis Lamellatis, sive, clava fissili instructa, descripta, memoires de l'Acad. T. VI." type="book" year="1818">Thunberg, 1818</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[958,1057,1433,1459]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[98,1067,134,1670]" pageId="7" pageNumber="8">
|
||
<keyLead pageId="7" pageNumber="8">
|
||
– Edge between medial teeth straight; well-defined clypeo-frontal carina; genal margin virtually convex (
|
||
<figureCitation box="[396,506,1502,1529]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11H</figureCitation>
|
||
). Parameres (
|
||
<figureCitation box="[663,768,1502,1529]" captionStart="Fig" captionStartId="29.[98,131,1599,1621]" captionTargetBox="[346,816,1085,1574]" captionTargetId="figure@29.[346,817,1081,1574]" captionTargetPageId="29" captionText="Fig. 12. Images of the aedeagi of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786654" httpUri="https://zenodo.org/record/3786654/files/figure.png" pageId="7" pageNumber="8">Fig. 12E</figureCitation>
|
||
). Shaded vegetation from Western and
|
||
<collectingRegion box="[299,449,1537,1564]" country="South Africa" name="Eastern Cape" pageId="7" pageNumber="8">Eastern Cape</collectingRegion>
|
||
Provinces,
|
||
<collectingCountry box="[578,721,1537,1563]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
|
||
.............................................. ............................................
|
||
<emphasis bold="true" box="[473,943,1574,1601]" pageId="7" pageNumber="8">
|
||
<taxonomicName authority="Daniel & Davis" authorityName="Daniel & Davis & Sole & Scholtz" authorityYear="2020" box="[473,879,1574,1601]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="australis" status="sp. n">
|
||
<emphasis bold="true" box="[473,684,1574,1601]" italics="true" pageId="7" pageNumber="8">Sisyphus australis</emphasis>
|
||
Daniel & Davis
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[886,943,1574,1600]" pageId="7" pageNumber="8" rank="species">sp. n</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[959,1057,1574,1601]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="7" pageNumber="8">Fig. 11E</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
<paragraph blockId="7.[98,1067,134,1670]" lastBlockId="8.[106,1074,134,897]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
<keyLead lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
30. Small-bodied; genae with shallow depression dorsally; pronotal disc showing distinct pattern of bare patches amongst sparse short setae; metafemur simple. Highland forest under cooler conditions in
|
||
<collectingCountry box="[649,770,134,160]" name="Zimbabwe" pageId="8" pageNumber="9">Zimbabwe</collectingCountry>
|
||
and
|
||
<collectingCountry box="[827,911,134,160]" name="Malawi" pageId="8" pageNumber="9">Malawi</collectingCountry>
|
||
..................... ...................................................
|
||
<taxonomicName authority="Arrow, 1907" authorityName="Arrow" authorityYear="1907" box="[534,894,168,195]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gazanus">
|
||
<emphasis bold="true" box="[534,894,168,195]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[534,739,168,195]" italics="true" pageId="8" pageNumber="9">Sisyphus gazanus</emphasis>
|
||
Arrow, 1907
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[909,1001,169,195]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="8" pageNumber="9">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation box="[1014,1065,169,195]" captionStart="Fig" captionStartId="29.[98,131,1599,1621]" captionTargetBox="[346,816,1085,1574]" captionTargetId="figure@29.[346,817,1081,1574]" captionTargetPageId="29" captionText="Fig. 12. Images of the aedeagi of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786654" httpUri="https://zenodo.org/record/3786654/files/figure.png" pageId="8" pageNumber="9">12C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[106,1074,134,897]" pageId="8" pageNumber="9">
|
||
<keyLead pageId="8" pageNumber="9">
|
||
– Large-bodied; genal depression absent; setae arranged regularly on pronotal disc; anvil- shaped projection on mid-posterior margin of metafemur. Lowland shaded Savanna from east to southern Africa................................................................... ..................................
|
||
<taxonomicName authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[408,896,310,337]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="impressipennis">
|
||
<emphasis bold="true" box="[408,896,310,337]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[408,692,310,337]" italics="true" pageId="8" pageNumber="9">Sisyphus impressipennis</emphasis>
|
||
<bibRefCitation author="Lansberge" box="[698,896,310,336]" firstAuthor="Lansberge" pageId="8" pageNumber="9" pagination="69 - 120" refId="ref29861" refString="Lansberge, J. W. (1886) Scarabaeides, Buprestides et Cerambycides de l'afrique occidentale. Notes from Leyden Museum, 8 (2): 69 - 120." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[911,1003,310,337]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="8" pageNumber="9">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation box="[1016,1066,311,337]" captionStart="Fig" captionStartId="29.[98,131,1599,1621]" captionTargetBox="[346,816,1085,1574]" captionTargetId="figure@29.[346,817,1081,1574]" captionTargetPageId="29" captionText="Fig. 12. Images of the aedeagi of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786654" httpUri="https://zenodo.org/record/3786654/files/figure.png" pageId="8" pageNumber="9">12A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="9">
|
||
<paragraph blockId="8.[106,1074,134,897]" pageId="8" pageNumber="9">
|
||
<keyLead pageId="8" pageNumber="9">
|
||
31. Relatively large-bodied; pronotal disc with conspicuous metallic sheen; open woodland in north of
|
||
<collectingCountry box="[407,506,380,406]" name="Namibia" pageId="8" pageNumber="9">Namibia</collectingCountry>
|
||
and south of
|
||
<collectingCountry box="[663,743,380,406]" name="Angola" pageId="8" pageNumber="9">Angola</collectingCountry>
|
||
............................................ ................................................
|
||
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[511,950,414,441]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="splendidus">
|
||
<emphasis bold="true" box="[511,950,414,441]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[511,745,414,441]" italics="true" pageId="8" pageNumber="9">Sisyphus splendidus</emphasis>
|
||
<bibRefCitation author="Montreuil, O." box="[752,950,414,440]" pageId="8" pageNumber="9" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[966,1066,414,441]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="8" pageNumber="9">Fig. 14A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[106,1074,134,897]" box="[106,1074,451,478]" pageId="8" pageNumber="9">
|
||
<keyLead box="[106,1074,451,478]" pageId="8" pageNumber="9">
|
||
– Relatively small to medium-bodied................................................................
|
||
<emphasis bold="true" box="[1023,1074,451,477]" pageId="8" pageNumber="9">(32)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="9">
|
||
<paragraph blockId="8.[106,1074,134,897]" pageId="8" pageNumber="9">
|
||
<keyLead pageId="8" pageNumber="9">
|
||
32. Very small-sized body (3.0–
|
||
<quantity box="[468,560,487,513]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="9" unit="mm" value="3.8">3.8 mm</quantity>
|
||
); pronotal disc dull bearing some depressions. Shaded vegetation in eastern and southern Africa ................................................ .......................................
|
||
<taxonomicName authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[450,876,555,582]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nanniscus">
|
||
<emphasis bold="true" box="[450,876,555,582]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[450,677,555,582]" italics="true" pageId="8" pageNumber="9">Sisyphus nanniscus</emphasis>
|
||
<bibRefCitation author="Peringuey" box="[684,876,555,582]" firstAuthor="Peringuey" pageId="8" pageNumber="9" pagination="1 - 563" refId="ref30622" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Transactions of the South African philosophical Society. Cape Town, 12: 1 - 563 (13 - 368)." type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[891,1005,556,582]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="8" pageNumber="9">Figs. 14C</figureCitation>
|
||
,
|
||
<figureCitation box="[1017,1065,556,583]" captionStart="Fig" captionStartId="38.[106,139,502,524]" captionTargetBox="[279,902,131,477]" captionTargetId="figure@38.[278,902,131,477]" captionTargetPageId="38" captionText="Fig. 15. Images of the aedeagi of the seminulum species-group: A. S. oralensis; B. S. nanniscus; C. S. auricomus sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786662" httpUri="https://zenodo.org/record/3786662/files/figure.png" pageId="8" pageNumber="9">15B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[106,1074,134,897]" box="[106,1073,592,620]" pageId="8" pageNumber="9">
|
||
<keyLead box="[106,1073,592,620]" pageId="8" pageNumber="9">
|
||
– Medium-sized body (3.9–5.0 mm)................................................................
|
||
<emphasis bold="true" box="[1023,1073,592,618]" pageId="8" pageNumber="9">(33)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="9">
|
||
<paragraph blockId="8.[106,1074,134,897]" pageId="8" pageNumber="9">
|
||
<keyLead pageId="8" pageNumber="9">
|
||
33. Pronotal disc bears sparse, long and fine yellow setae, separated by linear bare patches (
|
||
<figureCitation box="[257,360,662,689]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="8" pageNumber="9">Fig. 14E</figureCitation>
|
||
); parameres notched dorsally (
|
||
<figureCitation box="[711,816,662,689]" captionStart="Fig" captionStartId="38.[106,139,502,524]" captionTargetBox="[279,902,131,477]" captionTargetId="figure@38.[278,902,131,477]" captionTargetPageId="38" captionText="Fig. 15. Images of the aedeagi of the seminulum species-group: A. S. oralensis; B. S. nanniscus; C. S. auricomus sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786662" httpUri="https://zenodo.org/record/3786662/files/figure.png" pageId="8" pageNumber="9">Fig. 15A</figureCitation>
|
||
). Dense coastal woodlands and forest from northeast
|
||
<collectingCountry box="[518,660,697,723]" name="South Africa" pageId="8" pageNumber="9">South Africa</collectingCountry>
|
||
to southern
|
||
<collectingCountry box="[805,955,697,723]" name="Mozambique" pageId="8" pageNumber="9">Mozambique</collectingCountry>
|
||
.............. ..........................................
|
||
<taxonomicName authority="Daniel & Davis, 2016" authorityName="Daniel & Davis" authorityYear="2016" box="[467,950,731,758]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oralensis">
|
||
<emphasis bold="true" box="[467,950,731,758]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[467,680,731,758]" italics="true" pageId="8" pageNumber="9">Sisyphus oralensis</emphasis>
|
||
<bibRefCitation author="Daniel & Davis" box="[687,950,731,757]" firstAuthor="Daniel" pageId="8" pageNumber="9" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, & Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel & Davis, 2016</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[966,1065,732,758]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="8" pageNumber="9">Fig. 14B</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[106,1074,134,897]" pageId="8" pageNumber="9">
|
||
<keyLead pageId="8" pageNumber="9">
|
||
– Pronotum bears thick and short golden setae; directional projection of the setae well-defined (
|
||
<figureCitation box="[358,461,801,827]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="8" pageNumber="9">Fig. 14F</figureCitation>
|
||
); parameres without excavation dorsally, attenuating sharply at the apex (
|
||
<figureCitation box="[403,511,836,863]" captionStart="Fig" captionStartId="38.[106,139,502,524]" captionTargetBox="[279,902,131,477]" captionTargetId="figure@38.[278,902,131,477]" captionTargetPageId="38" captionText="Fig. 15. Images of the aedeagi of the seminulum species-group: A. S. oralensis; B. S. nanniscus; C. S. auricomus sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786662" httpUri="https://zenodo.org/record/3786662/files/figure.png" pageId="8" pageNumber="9">Fig. 15C</figureCitation>
|
||
). Forest and dense woodland species in central
|
||
<collectingCountry box="[156,306,870,896]" name="Mozambique" pageId="8" pageNumber="9">Mozambique</collectingCountry>
|
||
...................
|
||
<emphasis bold="true" box="[453,945,870,897]" pageId="8" pageNumber="9">
|
||
<taxonomicName authority="Daniel & Davis" authorityName="Daniel & Davis & Sole & Scholtz" authorityYear="2020" box="[453,880,870,897]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="auricomus" status="sp. n">
|
||
<emphasis bold="true" box="[453,686,870,897]" italics="true" pageId="8" pageNumber="9">Sisyphus auricomus</emphasis>
|
||
Daniel & Davis
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[888,945,870,896]" pageId="8" pageNumber="9" rank="species">sp. n</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[960,1064,870,897]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="8" pageNumber="9">Fig. 14D</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |