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<mods:title id="DCD042593EA3AB176BBC90E94AF1CA0C">Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean</mods:title>
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<heading id="147381DB534EFF80FEA38611FE55FC82" bold="true" box="[151,509,786,813]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="7DF0EAA5534EFF80FEA38611FE55FC82" bold="true" box="[151,509,786,813]" pageId="16" pageNumber="17">
Genus
<taxonomicName id="88844D34534EFF80FEDA8611FE2FFC83" ID-CoL="8VSMC" authority="OHara &amp; Thuy, 2022" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[238,391,786,813]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="7DF0EAA5534EFF80FEDA8611FE2FFC83" bold="true" box="[238,391,786,813]" italics="true" pageId="16" pageNumber="17">Ophiosabine</emphasis>
</taxonomicName>
<taxonomicNameLabel id="66C357DE534EFF80FFBB8611FE55FC82" box="[399,509,786,812]" pageId="16" pageNumber="17" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="4F3B36B7534EFF80FEA38635FB05FCFF" blockId="16.[151,1197,786,849]" box="[151,1197,822,849]" pageId="16" pageNumber="17">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="079E653C534EFF80FEA3867DFE2EFB17" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="4F3B36B7534EFF80FEA3867DFE2EFB17" blockId="16.[151,1437,894,2037]" pageId="16" pageNumber="17">
<emphasis id="7DF0EAA5534EFF80FEA3867DFEBAFC36" bold="true" box="[151,274,894,920]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Disc covered in thin scales and armoured with stout thorny spines, smooth spines or granules; radial shields mostly obscured, only evident by a shallow furrow and small exposed distal section on the disc; oral shields 13 times longer than wide, pentagonal to square, with an obtuse proximal angle or straight proximal margin and rounded to square distal margin; adoral shields small, usually situated proximal to the oral shields; 35 spiniform oral papillae, the outer one usually widened, suboral papillae can be present around the second oral tentacle pore; DAPs triangular to fan-shaped, separate, sometimes edged with small thorns; VAPs with an obtuse angle proximally, and an uplifted rounded distal edge, can be glassy and striated; 610 long hollow cylindrical arm spines, rounded at tip, finely denticulate; tentacle pores small, one large oval to pointed tentacle scale on most pores, sometimes two on first arm segment.
</paragraph>
</subSubSection>
<subSubSection id="079E653C534EFF80FEF381C1FC5EFB73" box="[199,1014,1218,1245]" pageId="16" pageNumber="17" type="type_taxon">
<paragraph id="4F3B36B7534EFF80FEF381C1FC5EFB73" blockId="16.[151,1437,894,2037]" box="[199,1014,1218,1245]" pageId="16" pageNumber="17">
<emphasis id="7DF0EAA5534EFF80FEF381C1FE3AFB72" bold="true" box="[199,402,1218,1244]" pageId="16" pageNumber="17">Included species</emphasis>
.
<taxonomicName id="88844D34534EFF80FF9C81C1FC96FB72" ID-CoL="49V66" authority="Lyman, 1878 b" authorityName="Lyman" authorityYear="1878" box="[424,830,1218,1245]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534EFF80FF9C81C1FD2EFB72" box="[424,646,1218,1244]" italics="true" pageId="16" pageNumber="17">Ophiacantha rosea</emphasis>
<bibRefCitation id="2B154B46534EFF80FCA081C0FC96FB72" author="Lyman, T." box="[660,830,1218,1245]" pageId="16" pageNumber="17" pagination="65 - 168" refId="ref33150" refString="Lyman, T. (1878 b) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard University, 5, 65 - 168, 10 pls." type="journal article" year="1878">Lyman, 1878b</bibRefCitation>
</taxonomicName>
(
<typeStatus id="903F8815534EFF80FD6081C1FC2EFB72" box="[852,902,1218,1244]" pageId="16" pageNumber="17">type</typeStatus>
species);
</paragraph>
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<paragraph id="4F3B36B7534EFF80FA3181C1FE1BFAC2" blockId="16.[151,1437,894,2037]" pageId="16" pageNumber="17">
<taxonomicName id="88844D34534EFF80FA3181C1FA3FFB72" authority="Sars, 1872" authorityName="Sars" authorityYear="1872" box="[1029,1431,1218,1244]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="anomala">
<emphasis id="7DF0EAA5534EFF80FA3181C1FAA0FB72" box="[1029,1288,1218,1244]" italics="true" pageId="16" pageNumber="17">Ophiacantha anomala</emphasis>
<bibRefCitation id="2B154B46534EFF80FB2381C1FA3FFB72" author="Sars, G. O." box="[1303,1431,1218,1244]" pageId="16" pageNumber="17" pagination="1 - 31" refId="ref35602" refString="Sars, G. O. (1872) Nye Echinodermer fra den norske kyst. Videnskabelige Selskabets Forhandlinger, 1871, 1 - 31." type="journal article" year="1872">Sars, 1872</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FEA381E5FD97FAAE" authority="Koehler, 1896 b" authorityName="Koehler" authorityYear="1896" box="[151,575,1254,1280]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="aristata">
<emphasis id="7DF0EAA5534EFF80FEA381E5FE2FFAAE" box="[151,391,1254,1280]" italics="true" pageId="16" pageNumber="17">Ophiacantha aristata</emphasis>
<bibRefCitation id="2B154B46534EFF80FFBA81E5FD97FAAE" author="Koehler, R." box="[398,575,1254,1280]" pageId="16" pageNumber="17" pagination="33 - 122" refId="ref32068" refString="Koehler, R. (1896 b) Resultats scientifiques de la compagne du Caudan dans de Golfe de Gascogne. Echinodermes. Annales de l'Universite de Lyon, 26, 33 - 122. https: // doi. org / 10.5962 / bhl. title. 65730" type="journal article" year="1896">Koehler, 1896b</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FC7D81E5FC53FAAE" authority="Lyman, 1878 b" authorityName="Lyman" authorityYear="1878" box="[585,1019,1254,1281]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="cuspidata">
<emphasis id="7DF0EAA5534EFF80FC7D81E5FCE7FAAE" box="[585,847,1254,1280]" italics="true" pageId="16" pageNumber="17">Ophiacantha cuspidata</emphasis>
<bibRefCitation id="2B154B46534EFF80FD6281E4FC53FAAE" author="Lyman, T." box="[854,1019,1254,1281]" pageId="16" pageNumber="17" pagination="65 - 168" refId="ref33150" refString="Lyman, T. (1878 b) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard University, 5, 65 - 168, 10 pls." type="journal article" year="1878">Lyman, 1878b</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FA3281E5FF7AFA8A" authority="Mortensen, 1936" authorityName="Mortensen" authorityYear="1936" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="densispina">
<emphasis id="7DF0EAA5534EFF80FA3281E5FABFFAAE" box="[1030,1303,1254,1280]" italics="true" pageId="16" pageNumber="17">Ophiacantha densispina</emphasis>
<bibRefCitation id="2B154B46534EFF80FB2981E4FF7AFA8A" author="Mortensen, T." pageId="16" pageNumber="17" pagination="199 - 348" refId="ref34461" refString="Mortensen, T. (1936) Echinoidea and Ophiuroidea. Discovery Reports, 12, 199 - 348, 9 pls. https: // doi. org / 10.5962 / bhl. part. 8051" type="journal article" year="1936">Mortensen, 1936</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FED68009FD2CFA8A" authority="Lyman, 1878 b" authorityName="Lyman" authorityYear="1878" box="[226,644,1290,1317]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="nodosa">
<emphasis id="7DF0EAA5534EFF80FED68009FE78FA8A" box="[226,464,1290,1316]" italics="true" pageId="16" pageNumber="17">Ophiacantha nodosa</emphasis>
<bibRefCitation id="2B154B46534EFF80FFEF8008FD2CFA8A" author="Lyman, T." box="[475,644,1290,1317]" pageId="16" pageNumber="17" pagination="65 - 168" refId="ref33150" refString="Lyman, T. (1878 b) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard University, 5, 65 - 168, 10 pls." type="journal article" year="1878">Lyman, 1878b</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FCA78009FB9DFA8A" authority="Koehler, 1906" authorityName="Koehler" authorityYear="1906" box="[659,1077,1290,1317]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="notata">
<emphasis id="7DF0EAA5534EFF80FCA78009FCD0FA8A" box="[659,888,1290,1316]" italics="true" pageId="16" pageNumber="17">Ophiacantha notata</emphasis>
<bibRefCitation id="2B154B46534EFF80FDB78009FB9DFA8A" author="Koehler, R." box="[899,1077,1290,1317]" pageId="16" pageNumber="17" pagination="5 - 34" refId="ref32177" refString="Koehler, R. (1906) Description des ophiures nouvelles recueillies par le Travailleur et le Talisman. Memoires de la Societe Zoologique de France, 19, 5 - 34 pls. 1 - 3." type="journal article" year="1906">Koehler, 1906</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FA708009FF7AFAE6" authority="Koehler, 1906" authorityName="Koehler" authorityYear="1906" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="parcita">
<emphasis id="7DF0EAA5534EFF80FA708009FA99FA8A" box="[1092,1329,1290,1316]" italics="true" pageId="16" pageNumber="17">Ophiacantha parcita</emphasis>
<bibRefCitation id="2B154B46534EFF80FB088009FF7AFAE6" author="Koehler, R." pageId="16" pageNumber="17" pagination="5 - 34" refId="ref32177" refString="Koehler, R. (1906) Description des ophiures nouvelles recueillies par le Travailleur et le Talisman. Memoires de la Societe Zoologique de France, 19, 5 - 34 pls. 1 - 3." type="journal article" year="1906">Koehler, 1906</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FED4802DFD04FAE6" authority="Mortensen, 1936" authorityName="Mortensen" authorityYear="1936" box="[224,684,1326,1353]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="pentactis">
<emphasis id="7DF0EAA5534EFF80FED4802DFE48FAE6" box="[224,480,1326,1352]" italics="true" pageId="16" pageNumber="17">Ophiacantha pentactis</emphasis>
<bibRefCitation id="2B154B46534EFF80FFDD802CFD04FAE6" author="Mortensen, T." box="[489,684,1326,1353]" pageId="16" pageNumber="17" pagination="199 - 348" refId="ref34461" refString="Mortensen, T. (1936) Echinoidea and Ophiuroidea. Discovery Reports, 12, 199 - 348, 9 pls. https: // doi. org / 10.5962 / bhl. part. 8051" type="journal article" year="1936">Mortensen, 1936</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FC8D802DFB2AFAE6" authority="Ljungman, 1871" authorityName="Ljungman" authorityYear="1871" box="[697,1154,1326,1353]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="vivipara">
<emphasis id="7DF0EAA5534EFF80FC8D802DFC1AFAE6" box="[697,946,1326,1352]" italics="true" pageId="16" pageNumber="17">Ophiacantha vivipara</emphasis>
Ljungman, 1871
</taxonomicName>
;
<taxonomicName id="88844D34534EFF80FABB802DFE07FAC2" authority="Martin-Ledo et al., 2013" authorityName="Martin-Ledo" authorityYear="2013" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="wolfarntzi">
<emphasis id="7DF0EAA5534EFF80FABB802DFA35FAE6" box="[1167,1437,1326,1352]" italics="true" pageId="16" pageNumber="17">Ophiacantha wolfarntzi</emphasis>
<bibRefCitation id="2B154B46534EFF80FEA38051FE07FAC2" author="Martin-Ledo, R. &amp; Sands, C. J. &amp; Lopez-Gonzalez, P. J." box="[151,431,1362,1389]" pageId="16" pageNumber="17" pagination="115 - 126" refId="ref33745" refString="Martin-Ledo, R., Sands, C. J. &amp; Lopez-Gonzalez, P. J. (2013) A new brooding species of brittle star (Echinodermata: Ophiuroidea) from Antarctic waters Polar Biology, 36, 115 - 126. https: // doi. org / 10.1007 / s 00300 - 012 - 1242 - z" type="journal article" year="2013">
Martín-Ledo
<emphasis id="7DF0EAA5534EFF80FF198050FECEFAC2" box="[301,358,1362,1388]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
, 2013
</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="079E653C534EFF81FEF38075FCCBFBC1" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="4F3B36B7534EFF80FEF38075FF45F922" blockId="16.[151,1437,894,2037]" pageId="16" pageNumber="17">
<emphasis id="7DF0EAA5534EFF80FEF38075FE9CFA3E" bold="true" box="[199,308,1398,1424]" pageId="16" pageNumber="17">Remarks</emphasis>
. The molecular phylogenies presented in
<bibRefCitation id="2B154B46534EFF80FD3F8075FC59FA3E" author="O'Hara, T. D. &amp; Hugall, A. F. &amp; Thuy, B. &amp; Stohr, S. &amp; Martynov, A. V." box="[779,1009,1398,1424]" pageId="16" pageNumber="17" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. &amp; Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
OHara
<emphasis id="7DF0EAA5534EFF80FD528074FC37FA3E" box="[870,927,1398,1424]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
(2017)
</bibRefCitation>
and
<bibRefCitation id="2B154B46534EFF80FA1C8075FAF2FA3E" author="Christodoulou, M. &amp; O'Hara, T. D. &amp; Hugall, A. F. &amp; Arbizu, P. M." box="[1064,1370,1398,1424]" pageId="16" pageNumber="17" pagination="3909 - 3912" refId="ref29723" refString="Christodoulou, M., O'Hara, T. D., Hugall, A. F. &amp; Arbizu, P. M. (2019) Dark Ophiuroid Biodiversity in a Prospective Abyssal Mine Field. Current Biology, 29 (e 3), 3909 - 3912." type="journal article" year="2019">
Christodoulou
<emphasis id="7DF0EAA5534EFF80FAFB8074FAA0FA3E" box="[1231,1288,1398,1424]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
(2019)
</bibRefCitation>
show that the genus
<taxonomicName id="88844D34534EFF80FF778099FCAAFA1A" authority="Muller &amp; Troschel, 1842" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[323,770,1434,1460]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA5534EFF80FF778099FE7DFA1A" box="[323,469,1434,1460]" italics="true" pageId="16" pageNumber="17">Ophiacantha</emphasis>
<bibRefCitation id="2B154B46534EFF80FFEB8099FCAAFA1A" author="Muller, J. &amp; Troschel, F. H." box="[479,770,1434,1460]" pageId="16" pageNumber="17" refId="ref34576" refString="Muller, J. &amp; Troschel, F. H. (1842) System der Asteriden. Braunschweig: Papier, Druck und Verlag." type="book" year="1842">Müller &amp; Troschel, 1842</bibRefCitation>
</taxonomicName>
, is polyphyletic, with some species even occurring in the
<taxonomicName id="88844D34534EFF80FEA380BDFE9FFA76" authorityName="Paterson" authorityYear="1985" box="[151,311,1470,1496]" class="Ophiuroidea" family="Ophiotomidae" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="family">Ophiotomidae</taxonomicName>
(see
<taxonomicName id="88844D34534EFF80FF4D80BCFDBEFA76" box="[377,534,1470,1497]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiotreta" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="spectabilis">
<emphasis id="7DF0EAA5534EFF80FF4D80BCFDBEFA76" box="[377,534,1470,1497]" italics="true" pageId="16" pageNumber="17">O. spectabilis</emphasis>
</taxonomicName>
below). Amongst the many nominal
<taxonomicName id="88844D34534EFF80FDFC80BDFB1DFA76" box="[968,1205,1470,1496]" pageId="16" pageNumber="17">
<emphasis id="7DF0EAA5534EFF80FDFC80BDFBF2FA76" box="[968,1114,1470,1496]" italics="true" pageId="16" pageNumber="17">Ophiacantha</emphasis>
species
</taxonomicName>
that are not closely related to the
<typeStatus id="903F8815534EFF80FF1A80E0FEF6FA53" box="[302,350,1507,1533]" pageId="16" pageNumber="17">type</typeStatus>
species
<taxonomicName id="88844D34534EFF80FF8880E0FCA3FA53" authority="(Bruzelius 1805)" baseAuthorityName="Bruzelius" baseAuthorityYear="1805" box="[444,779,1506,1533]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="bidentata">
<emphasis id="7DF0EAA5534EFF80FF8880E0FDEFFA52" box="[444,583,1506,1533]" italics="true" pageId="16" pageNumber="17">O. bidentata</emphasis>
(
<bibRefCitation id="2B154B46534EFF80FC6180E1FCABFA53" author="Bruzelius, N." box="[597,771,1506,1533]" pageId="16" pageNumber="17" refId="ref29608" refString="Bruzelius, N. (1805) Dissertatio sistens species cognitas asteriarum, quamr. sub praesidio A. J. Retzii. exhibet N. Bruzelius. s. n., Lundae, 37 pp." type="book" year="1805">Bruzelius 1805</bibRefCitation>
)
</taxonomicName>
, there is a monophyletic clade with a coherent morphology that contains the species
<taxonomicName id="88844D34534EFF80FF898304FD88F98E" authorityName="Lyman" authorityYear="1878" box="[445,544,1543,1569]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534EFF80FF898304FD88F98E" box="[445,544,1543,1569]" italics="true" pageId="16" pageNumber="17">O. rosea</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534EFF80FC6A8304FD4AF98E" authorityName="Ljungman" authorityYear="1871" box="[606,738,1543,1569]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="vivipara">
<emphasis id="7DF0EAA5534EFF80FC6A8304FD4AF98E" box="[606,738,1543,1569]" italics="true" pageId="16" pageNumber="17">O. vivipara</emphasis>
</taxonomicName>
that is here named as a new genus
<taxonomicName id="88844D34534EFF80FABB8305FAB6F98E" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[1167,1310,1542,1568]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA5534EFF80FABB8305FAB6F98E" box="[1167,1310,1542,1568]" italics="true" pageId="16" pageNumber="17">Ophiosabine</emphasis>
</taxonomicName>
. The most prominent feature is the unusual oral shield that usually is longer than wide and can be almost rectangular in some species (e.g.,
<taxonomicName id="88844D34534EFF80FF07834CFE3DF9C6" authorityName="Lyman" authorityYear="1878" box="[307,405,1615,1641]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534EFF80FF07834CFE3DF9C6" box="[307,405,1615,1641]" italics="true" pageId="16" pageNumber="17">O. rosea</emphasis>
</taxonomicName>
) and the small interradially-contiguous adoral shields that are placed proximal to the oral shields.
</paragraph>
<paragraph id="4F3B36B7534EFF80FEF38395FCC7F87F" blockId="16.[151,1437,894,2037]" pageId="16" pageNumber="17">
There is DNA evidence that, as well as
<taxonomicName id="88844D34534EFF80FCB98394FD46F91E" authorityName="Lyman" authorityYear="1878" box="[653,750,1687,1713]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534EFF80FCB98394FD46F91E" box="[653,750,1687,1713]" italics="true" pageId="16" pageNumber="17">O. rosea</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534EFF80FD1C8394FC00F91E" authorityName="Ljungman" authorityYear="1871" box="[808,936,1687,1713]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="vivipara">
<emphasis id="7DF0EAA5534EFF80FD1C8394FC00F91E" box="[808,936,1687,1713]" italics="true" pageId="16" pageNumber="17">O. vivipara</emphasis>
</taxonomicName>
, this genus includes the Southern Ocean
<taxonomicName id="88844D34534EFF80FBB58394FF54F97A" authorityName="Mortensen" authorityYear="1936" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="pentactis">
<emphasis id="7DF0EAA5534EFF80FBB58394FF54F97A" italics="true" pageId="16" pageNumber="17">O. pentactis</emphasis>
</taxonomicName>
,
<taxonomicName id="88844D34534EFF80FF3883B8FE01F97A" authorityName="Mortensen" authorityYear="1936" box="[268,425,1722,1749]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="densispina">
<emphasis id="7DF0EAA5534EFF80FF3883B8FE01F97A" box="[268,425,1722,1749]" italics="true" pageId="16" pageNumber="17">O. densispina</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534EFF80FFD783B8FDD1F97A" box="[483,633,1722,1749]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="wolfarntzi">
<emphasis id="7DF0EAA5534EFF80FFD783B8FDD1F97A" box="[483,633,1722,1749]" italics="true" pageId="16" pageNumber="17">O. wolfarntzi</emphasis>
</taxonomicName>
, as well as the Atlantic species
<taxonomicName id="88844D34534EFF80FDDD83B8FBD8F97A" authorityName="Sars" authorityYear="1872" box="[1001,1136,1722,1749]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="anomala">
<emphasis id="7DF0EAA5534EFF80FDDD83B8FBD8F97A" box="[1001,1136,1722,1749]" italics="true" pageId="16" pageNumber="17">O. anomala</emphasis>
</taxonomicName>
,
<taxonomicName id="88844D34534EFF80FAB483B8FAB9F97A" authorityName="Lyman" authorityYear="1878" box="[1152,1297,1722,1749]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="cuspidata">
<emphasis id="7DF0EAA5534EFF80FAB483B8FAB9F97A" box="[1152,1297,1722,1749]" italics="true" pageId="16" pageNumber="17">O. cuspidata</emphasis>
</taxonomicName>
,
<taxonomicName id="88844D34534EFF80FB1583B8FA34F97A" authorityName="Koehler" authorityYear="1896" box="[1313,1436,1723,1749]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="aristata">
<emphasis id="7DF0EAA5534EFF80FB1583B8FA34F97A" box="[1313,1436,1723,1749]" italics="true" pageId="16" pageNumber="17">O. aristata</emphasis>
</taxonomicName>
(
<bibRefCitation id="2B154B46534EFF80FEAB83DDFE66F956" author="Christodoulou, M. &amp; O'Hara, T. D. &amp; Hugall, A. F. &amp; Arbizu, P. M." box="[159,462,1758,1784]" pageId="16" pageNumber="17" pagination="3909 - 3912" refId="ref29723" refString="Christodoulou, M., O'Hara, T. D., Hugall, A. F. &amp; Arbizu, P. M. (2019) Dark Ophiuroid Biodiversity in a Prospective Abyssal Mine Field. Current Biology, 29 (e 3), 3909 - 3912." type="journal article" year="2019">
Christodoulou
<emphasis id="7DF0EAA5534EFF80FF7883DCFE20F956" box="[332,392,1758,1784]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="2B154B46534EFF80FFEA83DDFD52F956" author="Martin-Ledo, R. &amp; Sands, C. J. &amp; Lopez-Gonzalez, P. J." box="[478,762,1758,1784]" pageId="16" pageNumber="17" pagination="115 - 126" refId="ref33745" refString="Martin-Ledo, R., Sands, C. J. &amp; Lopez-Gonzalez, P. J. (2013) A new brooding species of brittle star (Echinodermata: Ophiuroidea) from Antarctic waters Polar Biology, 36, 115 - 126. https: // doi. org / 10.1007 / s 00300 - 012 - 1242 - z" type="journal article" year="2013">
Martín-Ledo
<emphasis id="7DF0EAA5534EFF80FC4C83DCFD1CF956" box="[632,692,1758,1784]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="2B154B46534EFF80FD3E83DDFC45F956" author="O'Hara, T. D. &amp; Smith, P. J. &amp; Mills, V. S. &amp; Smirnov, A. V. &amp; Steinke, D." box="[778,1005,1758,1784]" pageId="16" pageNumber="17" pagination="321 - 333" refId="ref35087" refString="O'Hara, T. D., Smith, P. J., Mills, V. S., Smirnov, A. V. &amp; Steinke, D. (2013) Biogeographical and phylogeographical relationships of the bathyal ophiuroid fauna of the Macquarie Ridge, Southern Ocean. Polar Biology, 36, 321 - 333. https: // doi. org / 10.1007 / s 00300 - 012 - 1261 - 9" type="journal article" year="2013">
OHara
<emphasis id="7DF0EAA5534EFF80FD5D83DCFC0EF956" box="[873,934,1758,1784]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2013
</bibRefCitation>
). In addition, the Atlantic species
<taxonomicName id="88844D34534EFF80FBB583DCFF40F8B2" authorityName="Lyman" authorityYear="1878" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="nodosa">
<emphasis id="7DF0EAA5534EFF80FBB583DCFF40F8B2" italics="true" pageId="16" pageNumber="17">O. nodosa</emphasis>
</taxonomicName>
,
<taxonomicName id="88844D34534EFF80FEC28200FEC8F8B2" authorityName="Koehler" authorityYear="1906" box="[246,352,1795,1821]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="notata">
<emphasis id="7DF0EAA5534EFF80FEC28200FEC8F8B2" box="[246,352,1795,1821]" italics="true" pageId="16" pageNumber="17">O. notata</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534EFF80FFA38200FDA1F8B2" authorityName="Koehler" authorityYear="1906" box="[407,521,1795,1821]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="parcita">
<emphasis id="7DF0EAA5534EFF80FFA38200FDA1F8B2" box="[407,521,1795,1821]" italics="true" pageId="16" pageNumber="17">O. parcita</emphasis>
</taxonomicName>
share many morphological features of the group. Several taxonomists have noted the similarity of the six-armed
<taxonomicName id="88844D34534EFF80FFC38224FDC3F8EE" authorityName="Lyman" authorityYear="1878" box="[503,619,1830,1857]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="nodosa">
<emphasis id="7DF0EAA5534EFF80FFC38224FDC3F8EE" box="[503,619,1830,1857]" italics="true" pageId="16" pageNumber="17">O. nodosa</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534EFF80FC978224FC81F8EE" authorityName="Sars" authorityYear="1872" box="[675,809,1830,1857]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="anomala">
<emphasis id="7DF0EAA5534EFF80FC978224FC81F8EE" box="[675,809,1830,1857]" italics="true" pageId="16" pageNumber="17">O. anomala</emphasis>
</taxonomicName>
(
<bibRefCitation id="2B154B46534EFF80FD0C8225FBAAF8EE" author="Mortensen, T." box="[824,1026,1830,1856]" pageId="16" pageNumber="17" pagination="1 - 120" refId="ref34437" refString="Mortensen, T. (1933 c) Ophiuroidea. The Danish Ingolf Expedition, 4, 1 - 120, 3 pls." type="journal article" year="1933">Mortensen 1933c</bibRefCitation>
;
<bibRefCitation id="2B154B46534EFF80FA3A8224FB1BF8EF" author="Paterson, G. L. J." box="[1038,1203,1830,1857]" pageId="16" pageNumber="17" pagination="1 - 162" refId="ref35334" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History) Zoology, 49, 1 - 162." type="journal article" year="1985">Paterson 1985</bibRefCitation>
). However,
<bibRefCitation id="2B154B46534EFF80FB088224FF4AF8CB" author="Paterson, G. L. J." pageId="16" pageNumber="17" pagination="1 - 162" refId="ref35334" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History) Zoology, 49, 1 - 162." type="journal article" year="1985">Paterson (1985)</bibRefCitation>
found morphological characters that distinguished the two species and the two species differ in depth distribution, with the
<typeStatus id="903F8815534EFF80FFB3826DFE41F826" box="[391,489,1902,1928]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
and only known specimen of
<taxonomicName id="88844D34534EFF80FD0C826CFC03F826" authorityName="Lyman" authorityYear="1878" box="[824,939,1902,1929]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="nodosa">
<emphasis id="7DF0EAA5534EFF80FD0C826CFC03F826" box="[824,939,1902,1929]" italics="true" pageId="16" pageNumber="17">O. nodosa</emphasis>
</taxonomicName>
collected from much lower depths (
<quantity id="887C9B52534EFF80FB09826DFA3CF827" box="[1341,1428,1902,1929]" metricMagnitude="3" metricUnit="m" metricValue="2.7889999999999997" pageId="16" pageNumber="17" unit="m" value="2789.0">2789 m</quantity>
) than is characteristic for
<taxonomicName id="88844D34534EFF80FF938290FD83F802" authorityName="Sars" authorityYear="1872" box="[423,555,1938,1965]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="anomala">
<emphasis id="7DF0EAA5534EFF80FF938290FD83F802" box="[423,555,1938,1965]" italics="true" pageId="16" pageNumber="17">O. anomala</emphasis>
</taxonomicName>
(
<quantity id="887C9B52534EFF80FC0C8291FD62F803" box="[568,714,1938,1965]" metricMagnitude="2" metricUnit="m" metricValue="6.705" metricValueMax="12.0" metricValueMin="1.41" pageId="16" pageNumber="17" unit="m" value="670.5" valueMax="1200.0" valueMin="141.0">1411200 m</quantity>
). As well as the typical oral shield,
<taxonomicName id="88844D34534EFF80FA7A8290FB1FF802" authorityName="Koehler" authorityYear="1906" box="[1102,1207,1939,1965]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="notata">
<emphasis id="7DF0EAA5534EFF80FA7A8290FB1FF802" box="[1102,1207,1939,1965]" italics="true" pageId="16" pageNumber="17">O. notata</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534EFF80FADE8290FAF3F802" authorityName="Koehler" authorityYear="1906" box="[1258,1371,1939,1965]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="parcita">
<emphasis id="7DF0EAA5534EFF80FADE8290FAF3F802" box="[1258,1371,1939,1965]" italics="true" pageId="16" pageNumber="17">O. parcita</emphasis>
</taxonomicName>
share the glassy striated nature of the VAPs that occur in
<taxonomicName id="88844D34534EFF80FCE082B4FCC0F87E" box="[724,872,1974,2001]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="wolfarntzi">
<emphasis id="7DF0EAA5534EFF80FCE082B4FCC0F87E" box="[724,872,1974,2001]" italics="true" pageId="16" pageNumber="17">O. wolfarntzi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4F3B36B7534EFF81FEF382D9FD43FECD" blockId="16.[151,1437,894,2037]" lastBlockId="17.[151,1437,151,1208]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Several other little-known species are also similar but have been described or figured with certain morphological features that make their assignment to
<taxonomicName id="88844D34534FFF81FC798594FD74FF1F" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[589,732,151,177]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA5534FFF81FC798594FD74FF1F" box="[589,732,151,177]" italics="true" pageId="17" pageNumber="18">Ophiosabine</emphasis>
</taxonomicName>
uncertain. The Atlantic species
<taxonomicName id="88844D34534FFF81FA738594FAC4FF1F" authority="Koehler, 1896 a" authorityName="Koehler" authorityYear="1896" box="[1095,1388,151,177]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="lineata">
<emphasis id="7DF0EAA5534FFF81FA738594FB10FF1F" box="[1095,1208,151,177]" italics="true" pageId="17" pageNumber="18">O. lineata</emphasis>
<bibRefCitation id="2B154B46534FFF81FAF48594FAC4FF1F" author="Koehler, R." box="[1216,1388,151,177]" pageId="17" pageNumber="18" pagination="241 - 253" refId="ref32032" refString="Koehler, R. (1896 a) Note preliminaire sur les ophiures recueillies pendant les campagnes de la ' Princesse Alice' Memoires de la Societe Zoologique de France, 9, 241 - 253." type="journal article" year="1896">Koehler, 1896a</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="88844D34534FFF81FEA385B8FDACFF7B" authority="H. L. Clark, 1915" authorityName="H. L. Clark" authorityYear="1915" box="[151,516,186,213]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="metallacta">
<emphasis id="7DF0EAA5534FFF81FEA385B8FE9BFF7B" box="[151,307,187,213]" italics="true" pageId="17" pageNumber="18">O. metallacta</emphasis>
H.L.
<bibRefCitation id="2B154B46534FFF81FF4885B8FDACFF7B" author="Clark, H. L." box="[380,516,186,213]" pageId="17" pageNumber="18" pagination="165 - 376" refId="ref29982" refString="Clark, H. L. (1915) Catalogue of Recent ophiurans: based on the collection of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard University, 25, 165 - 376, pls. 1 - 20. https: // doi. org / 10.5962 / bhl. title. 48598" type="journal article" year="1915">Clark, 1915</bibRefCitation>
</taxonomicName>
have disc spines in the form of short cylindrical stumps with a rim of thorns.
<taxonomicName id="88844D34534FFF81FEA385DCFDE9FF57" authority="Koehler, 1914" authorityName="Koehler" authorityYear="1914" box="[151,577,223,249]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomitrella" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="7DF0EAA5534FFF81FEA385DCFE33FF56" box="[151,411,223,249]" italics="true" pageId="17" pageNumber="18">Ophiomitrella porrecta</emphasis>
<bibRefCitation id="2B154B46534FFF81FF9685DCFDE9FF57" author="Koehler, R." box="[418,577,223,249]" pageId="17" pageNumber="18" pagination="1 - 173" refId="ref32434" refString="Koehler, R. (1914) A contribution to the study of ophiurans of the United States National Museum. Bulletin of the United States National Museum, 84, 1 - 173 pls. 1 - 18. https: // doi. org / 10.5962 / bhl. title. 27414" type="journal article" year="1914">Koehler, 1914</bibRefCitation>
</taxonomicName>
from the Straits of Magellan (
<quantity id="887C9B52534FFF81FDA685DCFC73FF57" box="[914,987,223,249]" metricMagnitude="2" metricUnit="m" metricValue="6.36" pageId="17" pageNumber="18" unit="m" value="636.0">636 m</quantity>
) has a very elongated jaw,
<taxonomicName id="88844D34534FFF81FB3E85DCFE4BFEB2" authority="H. L. Clark, 1911" authorityName="H. L. Clark" authorityYear="1911" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="atopostoma">
<emphasis id="7DF0EAA5534FFF81FB3E85DCFEB2FEB2" italics="true" pageId="17" pageNumber="18">Ophiacantha atopostoma</emphasis>
H.L.
<bibRefCitation id="2B154B46534FFF81FF6A8401FE4BFEB2" author="Clark, H. L." box="[350,483,258,284]" pageId="17" pageNumber="18" pagination="1 - 302" refId="ref29926" refString="Clark, H. L. (1911) North Pacific ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum, 75, 1 - 302, figs. 1 - 144. https: // doi. org / 10.5962 / bhl. title. 32556" type="journal article" year="1911">Clark, 1911</bibRefCitation>
</taxonomicName>
from the Bering Sea (
<quantity id="887C9B52534FFF81FCD78401FCCEFEB3" box="[739,870,258,285]" metricMagnitude="2" metricUnit="m" metricValue="6.55" metricValueMax="6.81" metricValueMin="6.29" pageId="17" pageNumber="18" unit="m" value="655.0" valueMax="681.0" valueMin="629.0">629681 m</quantity>
) has round exposed patches on the radial shields, and
<taxonomicName id="88844D34534FFF81FEF38425FDE5FEEE" authority="H. L. Clark, 1911" authorityName="H. L. Clark" authorityYear="1911" box="[199,589,294,320]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="acanthinotata">
<emphasis id="7DF0EAA5534FFF81FEF38425FE2EFEEE" box="[199,390,294,320]" italics="true" pageId="17" pageNumber="18">O. acanthinotata</emphasis>
H.L.
<bibRefCitation id="2B154B46534FFF81FFFD8425FDE5FEEE" author="Clark, H. L." box="[457,589,294,320]" pageId="17" pageNumber="18" pagination="1 - 302" refId="ref29926" refString="Clark, H. L. (1911) North Pacific ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum, 75, 1 - 302, figs. 1 - 144. https: // doi. org / 10.5962 / bhl. title. 32556" type="journal article" year="1911">Clark, 1911</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="37937627534FFF81FCA68425FD7AFEEE" box="[658,722,294,320]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
(
<quantity id="887C9B52534FFF81FCD68425FCCCFEEE" box="[738,868,294,320]" metricMagnitude="2" metricUnit="m" metricValue="3.2" metricValueMax="3.31" metricValueMin="3.09" pageId="17" pageNumber="18" unit="m" value="320.0" valueMax="331.0" valueMin="309.0">309331 m</quantity>
) has oral shields that “tend to become triangular”. All these species require further critical examination.
</paragraph>
<paragraph id="4F3B36B7534FFF81FEF3846FFA22FD96" blockId="17.[151,1437,151,1208]" pageId="17" pageNumber="18">
There are also numerous cryptic species in this group which complicate the nomenclature.
<taxonomicName id="88844D34534FFF81FA86846FFA35FE28" box="[1202,1437,364,390]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="vivpara">
<emphasis id="7DF0EAA5534FFF81FA86846FFA35FE28" box="[1202,1437,364,390]" italics="true" pageId="17" pageNumber="18">Ophiosabine vivpara</emphasis>
</taxonomicName>
consists of three clades in the Southern Ocean (
<bibRefCitation id="2B154B46534FFF81FC868493FC65FE05" author="Brogger, M. I. &amp; O'Hara, T. D." box="[690,973,400,427]" pageId="17" pageNumber="18" pagination="432 - 440" refId="ref29561" refString="Brogger, M. I. &amp; O'Hara, T. D. (2015) Revision of some ophiuroid records from Argentina. Zootaxa, 3972 (3), 432 - 440. https: // doi. org / 10.11646 / zootaxa. 3972.3.8" type="journal article" year="2015">Brogger &amp; OHara 2015</bibRefCitation>
;
<bibRefCitation id="2B154B46534FFF81FDEE8493FB1CFE04" author="O'Hara, T. D. &amp; Smith, P. J. &amp; Mills, V. S. &amp; Smirnov, A. V. &amp; Steinke, D." box="[986,1204,400,426]" pageId="17" pageNumber="18" pagination="321 - 333" refId="ref35087" refString="O'Hara, T. D., Smith, P. J., Mills, V. S., Smirnov, A. V. &amp; Steinke, D. (2013) Biogeographical and phylogeographical relationships of the bathyal ophiuroid fauna of the Macquarie Ridge, Southern Ocean. Polar Biology, 36, 321 - 333. https: // doi. org / 10.1007 / s 00300 - 012 - 1261 - 9" type="journal article" year="2013">
OHara
<emphasis id="7DF0EAA5534FFF81FA028492FBC7FE04" box="[1078,1135,400,426]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2013
</bibRefCitation>
).
<taxonomicName id="88844D34534FFF81FAFC8493FA34FE04" box="[1224,1436,400,426]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534FFF81FAFC8493FA34FE04" box="[1224,1436,400,426]" italics="true" pageId="17" pageNumber="18">Ophiosabine rosea</emphasis>
</taxonomicName>
consists of numerous clades around
<collectingCountry id="37937627534FFF81FC1C84B0FD27FE63" box="[552,655,435,461]" name="Australia" pageId="17" pageNumber="18">Australia</collectingCountry>
/
<collectingCountry id="37937627534FFF81FCA384B0FC84FE63" box="[663,812,435,461]" name="New Zealand" pageId="17" pageNumber="18">New Zealand</collectingCountry>
and
<collectingCountry id="37937627534FFF81FD5584B7FBA6FE63" box="[865,1038,435,462]" name="New Caledonia" pageId="17" pageNumber="18">New Caledonia</collectingCountry>
(
<bibRefCitation id="2B154B46534FFF81FA2884B0FA95FE60" author="Christodoulou, M. &amp; O'Hara, T. D. &amp; Hugall, A. F. &amp; Arbizu, P. M." box="[1052,1341,435,462]" pageId="17" pageNumber="18" pagination="3909 - 3912" refId="ref29723" refString="Christodoulou, M., O'Hara, T. D., Hugall, A. F. &amp; Arbizu, P. M. (2019) Dark Ophiuroid Biodiversity in a Prospective Abyssal Mine Field. Current Biology, 29 (e 3), 3909 - 3912." type="journal article" year="2019">
Christodoulou
<emphasis id="7DF0EAA5534FFF81FAF084B7FB54FE63" box="[1220,1276,435,461]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="2B154B46534FFF81FB7C84B7FEB8FE5F" author="O'Hara, T. D. &amp; Smith, P. J. &amp; Mills, V. S. &amp; Smirnov, A. V. &amp; Steinke, D." pageId="17" pageNumber="18" pagination="321 - 333" refId="ref35087" refString="O'Hara, T. D., Smith, P. J., Mills, V. S., Smirnov, A. V. &amp; Steinke, D. (2013) Biogeographical and phylogeographical relationships of the bathyal ophiuroid fauna of the Macquarie Ridge, Southern Ocean. Polar Biology, 36, 321 - 333. https: // doi. org / 10.1007 / s 00300 - 012 - 1261 - 9" type="journal article" year="2013">
OHara
<emphasis id="7DF0EAA5534FFF81FEA384DBFF66FE5F" box="[151,206,471,497]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2013
</bibRefCitation>
), which may not be the same species as the
<typeStatus id="903F8815534FFF81FCDA84D4FCB6FE5F" box="[750,798,471,497]" pageId="17" pageNumber="18">type</typeStatus>
collected from off
<collectingCountry id="37937627534FFF81FDC484D4FB98FE5F" box="[1008,1072,471,497]" name="Chile" pageId="17" pageNumber="18">Chile</collectingCountry>
. COI sequences of
<taxonomicName id="88844D34534FFF81FB3084D4FA34FE5F" authorityName="Mortensen" authorityYear="1936" box="[1284,1436,471,497]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="densispina">
<emphasis id="7DF0EAA5534FFF81FB3084D4FA34FE5F" box="[1284,1436,471,497]" italics="true" pageId="17" pageNumber="18">O. densispina</emphasis>
</taxonomicName>
from
<collectingCountry id="37937627534FFF81FEEC84F9FED5FDBA" box="[216,381,506,532]" name="South Georgia and the South Sandwich Islands" pageId="17" pageNumber="18">South Georgia</collectingCountry>
(
<bibRefCitation id="2B154B46534FFF81FFBA84F9FD00FDBB" author="Martin-Ledo, R. &amp; Sands, C. J. &amp; Lopez-Gonzalez, P. J." box="[398,680,506,533]" pageId="17" pageNumber="18" pagination="115 - 126" refId="ref33745" refString="Martin-Ledo, R., Sands, C. J. &amp; Lopez-Gonzalez, P. J. (2013) A new brooding species of brittle star (Echinodermata: Ophiuroidea) from Antarctic waters Polar Biology, 36, 115 - 126. https: // doi. org / 10.1007 / s 00300 - 012 - 1242 - z" type="journal article" year="2013">
Martín-Ledo
<emphasis id="7DF0EAA5534FFF81FC1D84F8FDCDFDBA" box="[553,613,506,532]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2013
</bibRefCitation>
) are distinct from those around southern
<collectingCountry id="37937627534FFF81FABD84F9FB58FDBA" box="[1161,1264,506,532]" name="Australia" pageId="17" pageNumber="18">Australia</collectingCountry>
(
<bibRefCitation id="2B154B46534FFF81FB3684F8FF7CFD96" author="O'Hara, T. D. &amp; Smith, P. J. &amp; Mills, V. S. &amp; Smirnov, A. V. &amp; Steinke, D." pageId="17" pageNumber="18" pagination="321 - 333" refId="ref35087" refString="O'Hara, T. D., Smith, P. J., Mills, V. S., Smirnov, A. V. &amp; Steinke, D. (2013) Biogeographical and phylogeographical relationships of the bathyal ophiuroid fauna of the Macquarie Ridge, Southern Ocean. Polar Biology, 36, 321 - 333. https: // doi. org / 10.1007 / s 00300 - 012 - 1261 - 9" type="journal article" year="2013">
OHara
<emphasis id="7DF0EAA5534FFF81FB5584F8FA35FDBA" box="[1377,1437,506,532]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2013
</bibRefCitation>
). There is a second unnamed species at
<collectingCountry id="37937627534FFF81FCA5871DFC9BFD96" box="[657,819,542,568]" name="South Georgia and the South Sandwich Islands" pageId="17" pageNumber="18">South Georgia</collectingCountry>
closely related to
<taxonomicName id="88844D34534FFF81FA34871DFA2EFD96" authority="(Martin-Ledo, 2013)" baseAuthorityName="Martin-Ledo" baseAuthorityYear="2013" box="[1024,1414,542,568]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="wolfarntzi">
<emphasis id="7DF0EAA5534FFF81FA34871DFB3CFD96" box="[1024,1172,542,568]" italics="true" pageId="17" pageNumber="18">O. wolfarntzi</emphasis>
(
<bibRefCitation id="2B154B46534FFF81FA96871DFAD6FD96" author="Martin-Ledo, R. &amp; Sands, C. J. &amp; Lopez-Gonzalez, P. J." box="[1186,1406,542,568]" pageId="17" pageNumber="18" pagination="115 - 126" refId="ref33745" refString="Martin-Ledo, R., Sands, C. J. &amp; Lopez-Gonzalez, P. J. (2013) A new brooding species of brittle star (Echinodermata: Ophiuroidea) from Antarctic waters Polar Biology, 36, 115 - 126. https: // doi. org / 10.1007 / s 00300 - 012 - 1242 - z" type="journal article" year="2013">Martin-Ledo, 2013</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="4F3B36B7534FFF81FEF38742FC95FCD9" blockId="17.[151,1437,151,1208]" pageId="17" pageNumber="18">
The micromorphology of ophiuroid lateral arm plates has been shown to yield a wealth of characters potentially relevant for taxon diagnoses and phylogenetic analyses (
<bibRefCitation id="2B154B46534FFF81FD388766FC17FDD1" author="Martynov, A. V." box="[780,959,613,639]" pageId="17" pageNumber="18" pagination="1 - 154" refId="ref33807" refString="Martynov, A. V. (2010) Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae. Zootaxa, 2697, 1 - 154. https: // doi. org / 10.11646 / zootaxa. 2697.1.1" type="journal article" year="2010">Martynov 2010</bibRefCitation>
;
<bibRefCitation id="2B154B46534FFF81FDFC8766FB0CFDD1" author="Thuy, B. &amp; Stohr, S." box="[968,1188,613,639]" pageId="17" pageNumber="18" pagination="1 - 47" refId="ref36265" refString="Thuy, B. &amp; Stohr, S. (2011) Lateral arm plate morphology in brittle stars (Echinodermata: Ophiuroidea): new perspectives for ophiuroid micropalaeontology and classification. Zootaxa, 3013, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 3013.1.1" type="journal article" year="2011">Thuy &amp; Stöhr 2011</bibRefCitation>
,
<bibRefCitation id="2B154B46534FFF81FA9A8766FB43FD2E" author="Thuy, B. &amp; Stohr, S." box="[1198,1259,613,640]" pageId="17" pageNumber="18" pagination="0156140" refId="ref36318" refString="Thuy, B. &amp; Stohr, S. (2016) A New Morphological Phylogeny of the Ophiuroidea (Echinodermata) Accords with Molecular Evidence and Renders Microfossils Accessible for Cladistics. PLoSONE, 11, e 0156140. https: // doi. org / 10.1371 / journal. pone. 0156140" type="journal article" year="2016">2016</bibRefCitation>
). A recent study suggested that lateral arm plate morphologies among species of the
<taxonomicName id="88844D34534FFF81FD82878BFBCEFD0C" authorityName="Ljungman" authorityYear="1867" box="[950,1126,648,674]" class="Ophiuroidea" family="Ophiacanthidae" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Ophiacanthidae</taxonomicName>
are consistent with clades resolved by molecular evidence (
<bibRefCitation id="2B154B46534FFF81FC2187AFFC2CFD68" author="Numberger-Thuy, L. &amp; Thuy, B." box="[533,900,684,710]" pageId="17" pageNumber="18" pagination="19 - 30" refId="ref34675" refString="Numberger-Thuy, L. &amp; Thuy, B. (2020) A new bathyal ophiacanthid brittle star (Ophiuroidea: Ophiacanthidae) with Caribbean affinities from the Plio-Pleistocene of the Mediterranean. Zootaxa, 4820, 19 - 30. https: // doi. org / 10.11646 / zootaxa. 4820.1.2" type="journal article" year="2020">Numberger-Thuy &amp; Thuy 2020</bibRefCitation>
;
<bibRefCitation id="2B154B46534FFF81FDA687AFFBDAFD68" author="O'Hara, T. D. &amp; Hugall, A. F. &amp; Thuy, B. &amp; Stohr, S. &amp; Martynov, A. V." box="[914,1138,684,710]" pageId="17" pageNumber="18" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. &amp; Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
OHara
<emphasis id="7DF0EAA5534FFF81FDC487AEFB83FD68" box="[1008,1067,684,710]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2017
</bibRefCitation>
). Although the number of species analysed by
<bibRefCitation id="2B154B46534FFF81FF4887CCFCAAFD47" author="Numberger-Thuy, L. &amp; Thuy, B." box="[380,770,719,745]" pageId="17" pageNumber="18" pagination="19 - 30" refId="ref34675" refString="Numberger-Thuy, L. &amp; Thuy, B. (2020) A new bathyal ophiacanthid brittle star (Ophiuroidea: Ophiacanthidae) with Caribbean affinities from the Plio-Pleistocene of the Mediterranean. Zootaxa, 4820, 19 - 30. https: // doi. org / 10.11646 / zootaxa. 4820.1.2" type="journal article" year="2020">Numberger-Thuy and Thuy (2020)</bibRefCitation>
only reflects a small part of the ophiacanthid spectrum, the study suggests that the lateral arm plates of
<taxonomicName id="88844D34534FFF81FCB787F0FCFEFCA2" box="[643,854,755,781]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534FFF81FCB787F0FCFEFCA2" box="[643,854,755,781]" italics="true" pageId="17" pageNumber="18">Ophiosabine rosea</emphasis>
</taxonomicName>
have a uniquely shaped vertebral articular structure on their inner side. In order to corroborate this preliminary observation and potentially establish the shape of the vertebral articular structure as a diagnosing character of the genus
<taxonomicName id="88844D34534FFF81FD9D8639FB90FCFA" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[937,1080,826,852]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA5534FFF81FD9D8639FB90FCFA" box="[937,1080,826,852]" italics="true" pageId="17" pageNumber="18">Ophiosabine</emphasis>
</taxonomicName>
, more species will have to be examined with respect to their lateral arm plate morphology.
</paragraph>
<paragraph id="4F3B36B7534FFF81FEF38683FCCBFBC1" blockId="17.[151,1437,151,1208]" pageId="17" pageNumber="18">
There is no available genus-level name for the group.
<bibRefCitation id="2B154B46534FFF81FD128683FC65FC35" author="Verrill, A. E." box="[806,973,896,923]" pageId="17" pageNumber="18" pagination="1 - 88" refId="ref36700" refString="Verrill, A. E. (1899 b) Report on the Ophiuroidea collected by the Bahama expedition in 1893. Bulletin of the Laboratories of Natural History of the State of Iowa, 5, 1 - 88 pls. 1 - 8." type="journal article" year="1899">Verrill (1899b)</bibRefCitation>
created the subgenus
<taxonomicName id="88844D34534FFF81FAF28683FAFAFC34" authorityName="Verrill" authorityYear="1899" box="[1222,1362,896,922]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophientodia" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA5534FFF81FAF28683FAFAFC34" box="[1222,1362,896,922]" italics="true" pageId="17" pageNumber="18">Ophientodia</emphasis>
</taxonomicName>
for
<taxonomicName id="88844D34534FFF81FBB58682FEACFC10" authorityName="Lyman" authorityYear="1878" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cuspidata">
<emphasis id="7DF0EAA5534FFF81FBB58682FEACFC10" italics="true" pageId="17" pageNumber="18">O. cuspidata</emphasis>
</taxonomicName>
,
<taxonomicName id="88844D34534FFF81FF2186A7FD07FC10" authority="Lyman, 1878 a" authorityName="Lyman" authorityYear="1878" box="[277,687,932,959]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="scutata">
<emphasis id="7DF0EAA5534FFF81FF2186A7FDAAFC10" box="[277,514,932,958]" italics="true" pageId="17" pageNumber="18">Ophiacantha scutata</emphasis>
<bibRefCitation id="2B154B46534FFF81FC3886A6FD07FC10" author="Lyman, T." box="[524,687,932,959]" pageId="17" pageNumber="18" pagination="217 - 238" refId="ref33100" refString="Lyman, T. (1878 a) Ophiurans and Astrophytons. Reports on the dredging operations of the U. S. coast survey Str. ' Blake'. Bulletin of the Museum of Comparative Zoology, Harvard University, 5, 217 - 238, 3 pls." type="journal article" year="1878">Lyman, 1878a</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="88844D34534FFF81FCDF86A7FBB7FC10" authority="Verrill, 1899 a" authorityName="Verrill" authorityYear="1899" box="[747,1055,932,958]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="pectinula">
<emphasis id="7DF0EAA5534FFF81FCDF86A7FCD0FC10" box="[747,888,932,958]" italics="true" pageId="17" pageNumber="18">O. pectinula</emphasis>
<bibRefCitation id="2B154B46534FFF81FDB686A7FBB7FC10" author="Verrill, A. E." box="[898,1055,932,958]" pageId="17" pageNumber="18" pagination="301 - 396" refId="ref36643" refString="Verrill, A. E. (1899 a) North American Ophiuroidea. 1. Revision of certain families and genera of West Indian ophiurans. Transactions of the Connecticut Academy of Arts and Sciences, 10, 301 - 396. https: // doi. org / 10.5962 / bhl. part. 7032" type="journal article" year="1899">Verrill, 1899a</bibRefCitation>
</taxonomicName>
(=
<emphasis id="7DF0EAA5534FFF81FA7686A7FB10FC10" box="[1090,1208,932,958]" italics="true" pageId="17" pageNumber="18">echinulata</emphasis>
<bibRefCitation id="2B154B46534FFF81FAF686A6FACFFC10" author="Lyman, T." box="[1218,1383,932,959]" pageId="17" pageNumber="18" pagination="217 - 238" refId="ref33100" refString="Lyman, T. (1878 a) Ophiurans and Astrophytons. Reports on the dredging operations of the U. S. coast survey Str. ' Blake'. Bulletin of the Museum of Comparative Zoology, Harvard University, 5, 217 - 238, 3 pls." type="journal article" year="1878">Lyman, 1878a</bibRefCitation>
) but didnt designate a
<typeStatus id="903F8815534FFF81FF5E86CBFE32FC4C" box="[362,410,968,994]" pageId="17" pageNumber="18">type</typeStatus>
species. However, as he only reported examining specimens of
<taxonomicName id="88844D34534FFF81FA4C86CBFB44FC4F" authorityName="Lyman" authorityYear="1878" box="[1144,1260,968,994]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="scutata">
<emphasis id="7DF0EAA5534FFF81FA4C86CBFB44FC4F" box="[1144,1260,968,994]" italics="true" pageId="17" pageNumber="18">O. scutata</emphasis>
</taxonomicName>
and
<emphasis id="7DF0EAA5534FFF81FB1286C4FA34FC4F" box="[1318,1436,967,993]" italics="true" pageId="17" pageNumber="18">echinulata</emphasis>
it seems improbable that he would have intended
<taxonomicName id="88844D34534FFF81FCEE86E8FCC5FBAB" authorityName="Lyman" authorityYear="1878" box="[730,877,1003,1029]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cuspidata">
<emphasis id="7DF0EAA5534FFF81FCEE86E8FCC5FBAB" box="[730,877,1003,1029]" italics="true" pageId="17" pageNumber="18">O. cuspidata</emphasis>
</taxonomicName>
to be the
<typeStatus id="903F8815534FFF81FDD386E8FBBFFBAB" box="[999,1047,1003,1029]" pageId="17" pageNumber="18">type</typeStatus>
species. Likewise, he created the subgenus
<taxonomicName id="88844D34534FFF81FF33810DFD9FFB86" authority="Verrill, 1899 b" authorityName="Verrill" authorityYear="1899" box="[263,567,1038,1064]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="subGenus" subGenus="Ophiectodia">
<emphasis id="7DF0EAA5534FFF81FF33810DFE39FB86" box="[263,401,1038,1064]" italics="true" pageId="17" pageNumber="18">Ophiectodia</emphasis>
<bibRefCitation id="2B154B46534FFF81FFAC810DFD9FFB86" author="Verrill, A. E." box="[408,567,1038,1064]" pageId="17" pageNumber="18" pagination="1 - 88" refId="ref36700" refString="Verrill, A. E. (1899 b) Report on the Ophiuroidea collected by the Bahama expedition in 1893. Bulletin of the Laboratories of Natural History of the State of Iowa, 5, 1 - 88 pls. 1 - 8." type="journal article" year="1899">Verrill, 1899b</bibRefCitation>
</taxonomicName>
, without designating a
<typeStatus id="903F8815534FFF81FD0F810CFCC3FB87" box="[827,875,1039,1065]" pageId="17" pageNumber="18">type</typeStatus>
species, which contained
<taxonomicName id="88844D34534FFF81FAA6810CFA30FB87" authority="Verrill, 1885" authorityName="Verrill" authorityYear="1885" box="[1170,1432,1038,1065]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="enopla">
<emphasis id="7DF0EAA5534FFF81FAA6810CFAA8FB86" box="[1170,1280,1038,1065]" italics="true" pageId="17" pageNumber="18">O. enopla</emphasis>
<bibRefCitation id="2B154B46534FFF81FB33810DFA30FB87" author="Verrill, A. E." box="[1287,1432,1038,1065]" pageId="17" pageNumber="18" pagination="149 - 157" refId="ref36586" refString="Verrill, A. E. (1885) Notice of the remarkable marine fauna occupying the outer banks off the southern coast of New England, No. 11. American Journal of Science, 29, 149 - 157. https: // doi. org / 10.2475 / ajs. s 3 - 29.170.149" type="journal article" year="1885">Verrill, 1885</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="88844D34534FFF81FEA38131FF52FBE2" box="[151,250,1074,1100]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosabine" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="rosea">
<emphasis id="7DF0EAA5534FFF81FEA38131FF52FBE2" box="[151,250,1074,1100]" italics="true" pageId="17" pageNumber="18">O. rosea</emphasis>
</taxonomicName>
and
<taxonomicName id="88844D34534FFF81FF0C8131FE7EFBE2" box="[312,470,1074,1100]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiotreta" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="spectabilis">
<emphasis id="7DF0EAA5534FFF81FF0C8131FE7EFBE2" box="[312,470,1074,1100]" italics="true" pageId="17" pageNumber="18">O. spectabilis</emphasis>
</taxonomicName>
but only listed material he had examined of
<taxonomicName id="88844D34534FFF81FDDE8131FBF4FBE2" authorityName="Verrill" authorityYear="1885" box="[1002,1116,1074,1100]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="enopla">
<emphasis id="7DF0EAA5534FFF81FDDE8131FBF4FBE2" box="[1002,1116,1074,1100]" italics="true" pageId="17" pageNumber="18">O. enopla</emphasis>
</taxonomicName>
. Both of Verrills proposed subgenera would now be regarded as polyphyletic assemblages.
</paragraph>
</subSubSection>
<subSubSection id="079E653C534FFF81FEF3817AFB51FB19" pageId="17" pageNumber="18" type="etymology">
<paragraph id="4F3B36B7534FFF81FEF3817AFB51FB19" blockId="17.[151,1437,151,1208]" pageId="17" pageNumber="18">
<emphasis id="7DF0EAA5534FFF81FEF3817AFEE0FB3D" bold="true" box="[199,328,1145,1171]" pageId="17" pageNumber="18">Etymology</emphasis>
. Named using the typical “Ophio” prefix of brittle star genera and after our colleague, ophiuroid taxonomist and friend Sabine Stöhr of the Swedish Museum of Natural History. Gender is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>