treatments-xml/data/C7/2D/87/C72D87A15349FF89FEA386F5FD1BF817.xml
2024-06-21 12:51:15 +02:00

372 lines
58 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="43D043295A776AC40056C17281A62A9F" ID-CLB-Dataset="53164" ID-DOI="10.11646/zootaxa.5124.1.1" ID-GBIF-Dataset="9666d599-2c7d-4a52-bce2-a8794f03265b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6404674" ID-ZooBank="C015F8CB-799B-4A92-90AE-02B4C576089E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648795871367" checkinUser="plazi" docAuthor="OHara, Timothy D. &amp; Thuy, Ben" docDate="2022" docId="C72D87A15349FF89FEA386F5FD1BF817" docLanguage="en" docName="zootaxa.5124.1.1.pdf" docOrigin="Zootaxa 5124 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiolebes paulensis OHara &amp; Thuy 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="26" masterDocId="3B14FFD9535EFF90FE348503FFA8FFAE" masterDocTitle="Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean" masterLastPageNumber="49" masterPageNumber="1" pageNumber="24" updateTime="1699310728982" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F7B05F8E1A52E82D4D4E0E679B5B6DFE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7D1AE90172F39E7B38BE80D35FDF72CF">
<mods:title id="63C4EB5B08B882FE780E3D23C9F25095">Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean</mods:title>
</mods:titleInfo>
<mods:name id="138A0E32E8B8521A57F5BF8E61E665FA" type="personal">
<mods:role id="E7EDBC54A6CE6EF18717138702B398C8">
<mods:roleTerm id="941CAA5BC414458A3CB1DFCC886F00E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D48AD207A00B5BE8A96DEA529E37FE65">OHara, Timothy D.</mods:namePart>
<mods:affiliation id="F7DCD5FE091B001ED643676537562160">Museums Victoria, GPO Box 666 E, Melbourne, 3001, AUSTRALIA,</mods:affiliation>
</mods:name>
<mods:name id="7F29DABCC1345CDCEF92D45E438499BC" type="personal">
<mods:role id="53D4E2E5539DC11BDCBBF3BC7A7CDC17">
<mods:roleTerm id="D787A641694E12CEEF01BD8AAF59C86D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="60FFABFC615BB7C66472BC5AF86CE633">Thuy, Ben</mods:namePart>
<mods:affiliation id="8EC22A13534FBB31760C5C401C763901">Natural History Museum of Luxembourg, 24 Rue Münster, 2160 Luxembourg</mods:affiliation>
</mods:name>
<mods:typeOfResource id="36EDCCF7C97A9961AA7C2697228AC814">text</mods:typeOfResource>
<mods:relatedItem id="A1A813CD7195E6C33119C28295EE3220" type="host">
<mods:titleInfo id="2A005EDABA3BFA271AFAD7590BC78E55">
<mods:title id="572626FCD41FDDDE624C9B96BC776A8B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FE56164AD5F80E2FD24FEFE08ED5EDF3">
<mods:date id="03E8E334D28DF73B0C8C6340C7C706C3">2022</mods:date>
<mods:detail id="E051E0B68864B74D1A3D62B3ACDE10B5" type="pubDate">
<mods:number id="240F2F8AF2E5662932F12394CCEB9164">2022-03-31</mods:number>
</mods:detail>
<mods:detail id="D3C65D566D1104E1A3E48D974DC67EBE" type="volume">
<mods:number id="9C130857461245380D99865692A90336">5124</mods:number>
</mods:detail>
<mods:detail id="8ACECEF9A0378AA07FA7859BCAE9DA5A" type="issue">
<mods:number id="31C86C573338E25954D646F57CE22FF4">1</mods:number>
</mods:detail>
<mods:extent id="D53D63AC12C3334713ECC23DED6462F9" unit="page">
<mods:start id="8E850612EABB917103CD521757FB2CC0">1</mods:start>
<mods:end id="19BCEF303AA3745193D1149D48DA7466">49</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5B704EA3FA178A6133711FB60D478552">journal article</mods:classification>
<mods:identifier id="8987754BB8600790B6B217C91BC6772C" type="CLB-Dataset">53164</mods:identifier>
<mods:identifier id="48EF26CDB973977FEEE314D5897754D7" type="DOI">10.11646/zootaxa.5124.1.1</mods:identifier>
<mods:identifier id="FFCF20E5CAF28FC6B1A387181E4BF72B" type="GBIF-Dataset">9666d599-2c7d-4a52-bce2-a8794f03265b</mods:identifier>
<mods:identifier id="123C23678BFB9989223AF9CA5E119785" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BDBE8500AADFCD2FB3C127A0E888D56E" type="Zenodo-Dep">6404674</mods:identifier>
<mods:identifier id="CE190A1B2C04260404A6D217E6E149E8" type="ZooBank">C015F8CB-799B-4A92-90AE-02B4C576089E</mods:identifier>
</mods:mods>
<treatment id="C72D87A15349FF89FEA386F5FD1BF817" ID-DOI="http://doi.org/10.5281/zenodo.6404720" ID-GBIF-Taxon="194442339" ID-Zenodo-Dep="6404720" LSID="urn:lsid:plazi:treatment:C72D87A15349FF89FEA386F5FD1BF817" httpUri="http://treatment.plazi.org/id/C72D87A15349FF89FEA386F5FD1BF817" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="079E653C5349FF87FEA386F5FE55FBBF" box="[151,509,1014,1041]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="4F3B36B75349FF87FEA386F5FE55FBBF" blockId="23.[151,1198,1014,1113]" box="[151,509,1014,1041]" pageId="23" pageNumber="24">
<heading id="147381DB5349FF87FEA386F5FE55FBBF" bold="true" box="[151,509,1014,1041]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="7DF0EAA55349FF87FEA386F5FE55FBBF" bold="true" box="[151,509,1014,1041]" pageId="23" pageNumber="24">
<taxonomicName id="88844D345349FF87FEA386F5FE30FBBF" ID-CoL="8ZDNR" authority="OHara &amp; Thuy, 2022" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[151,408,1014,1041]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="paulensis" status="sp. nov.">
<emphasis id="7DF0EAA55349FF87FEA386F5FE30FBBF" bold="true" box="[151,408,1014,1041]" italics="true" pageId="23" pageNumber="24">Ophiolebes paulensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="66C357DE5349FF87FFAB86F4FE55FBBF" box="[415,509,1015,1041]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="079E653C5349FF87FEA38118FB06FBF7" pageId="23" pageNumber="24" type="description">
<paragraph id="4F3B36B75349FF87FEA38118FEC1FB9B" blockId="23.[151,1198,1014,1113]" box="[151,361,1051,1077]" pageId="23" pageNumber="24">
<figureCitation id="D7BF2A325349FF87FEA38118FEB8FB9B" box="[151,272,1051,1077]" captionStart="FIGURE 9" captionStartId="24.[151,250,1678,1702]" captionTargetBox="[151,1436,309,1653]" captionTargetId="figure-88@24.[151,1436,309,1654]" captionTargetPageId="24" captionText="FIGURE 9. Ophiolebes paulensis sp. nov., holotype, MNHN IE.2009.1566, A: dorsal and B: ventral views, C: inset showing hook-shaped distal arm spines." figureDoi="http://doi.org/10.5281/zenodo.6404700" httpUri="https://zenodo.org/record/6404700/files/figure.png" pageId="23" pageNumber="24">Fig. 9AC</figureCitation>
,
<figureCitation id="D7BF2A325349FF87FF2F8118FEC1FB9B" box="[283,361,1051,1077]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="23" pageNumber="24">10AP</figureCitation>
</paragraph>
<paragraph id="4F3B36B75349FF87FEA3813DFB06FBF7" blockId="23.[151,1198,1014,1113]" box="[151,1198,1086,1113]" pageId="23" pageNumber="24">
http://zoobank.org/
<uri id="3B153AB55349FF87FF45813DFB06FBF7" box="[369,1198,1086,1113]" pageId="23" pageNumber="24">
urn:lsid:zoobank.org:act:
<uuid id="3B220C625349FF87FC4B813DFB06FBF7" box="[639,1198,1086,1113]" pageId="23" pageNumber="24">F57E6CCA-DD72-4FED-80BB-439479B0C13D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="079E653C5349FF89FEA38184FEDEF983" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="4F3B36B75349FF87FEA38184FBB6FB6B" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
<emphasis id="7DF0EAA55349FF87FEA38184FEEAFB0E" bold="true" box="[151,322,1158,1185]" pageId="23" pageNumber="24">Type material</emphasis>
.
<materialsCitation id="FFEC3CEA5349FF87FF618185FF55FB6B" ID-GBIF-Occurrence="3746276322" collectingDate="1986-07-09" collectionCode="MD" elevation="940" latitude="-37.786667" longLatPrecision="92" longitude="77.649666" pageId="23" pageNumber="24" specimenCode="MD50" specimenCount="1">
<specimenCode id="1F229ECC5349FF87FF618185FE37FB0F" box="[341,415,1158,1185]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="23" pageNumber="24">MD50</specimenCode>
CP7, north-east of Île Amsterdam,
<geoCoordinate id="2AB050705349FF87FD7C8184FC6FFB0F" box="[840,967,1159,1185]" degrees="37" direction="south" minutes="47.2" orientation="latitude" pageId="23" pageNumber="24" precision="92" value="-37.786667">37° 47.2´S</geoCoordinate>
,
<geoCoordinate id="2AB050705349FF87FDE38184FBCEFB0F" box="[983,1126,1159,1185]" degrees="77" direction="east" minutes="38.98" orientation="longitude" pageId="23" pageNumber="24" precision="9" value="77.649666">77° 38.98´E</geoCoordinate>
, 1680
<quantity id="887C9B525349FF87FA898184FB57FB0F" box="[1213,1279,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="23" pageNumber="24" unit="m" value="940.0">
<elevation id="C4A9D1845349FF87FA898184FB57FB0F" box="[1213,1279,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="23" pageNumber="24" unit="m" value="940.0">940m</elevation>
</quantity>
, Beam trawl,
<date id="3B3A10775349FF87FEA381A8FF55FB6B" box="[151,253,1195,1221]" pageId="23" pageNumber="24" value="1986-07-09">
<collectingDate id="2B7EE99F5349FF87FEA381A8FF55FB6B" box="[151,253,1195,1221]" pageId="23" pageNumber="24" value="1986-07-09">9/7/1986</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="FFEC3CEA5349FF87FF3D81A8FDD4FB68" ID-GBIF-Occurrence="3746276304" box="[265,636,1194,1222]" collectionCode="MNHN, IE" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
<collectionCode id="2995AE725349FF87FF3D81A8FEF7FB6B" box="[265,351,1195,1221]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
<collectionCode id="2995AE725349FF87FF5181A9FE28FB6B" box="[357,384,1194,1221]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="23" pageNumber="24">IE</collectionCode>
.2009.1566 (
<typeStatus id="903F88155349FF87FC3981A8FDDDFB68" box="[525,629,1195,1222]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
)
</materialsCitation>
,
<materialsCitation id="FFEC3CEA5349FF87FCB381A8FBB2FB6B" ID-GBIF-Occurrence="3746276324" box="[647,1050,1194,1221]" collectionCode="MNHN, IE" pageId="23" pageNumber="24" specimenCount="5" typeStatus="paratype">
<collectionCode id="2995AE725349FF87FCB381A8FD75FB6B" box="[647,733,1195,1221]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
<collectionCode id="2995AE725349FF87FCD081A9FD57FB6B" box="[740,767,1194,1221]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="23" pageNumber="24">IE</collectionCode>
.2009.1567 (
<specimenCount id="5982FD3E5349FF87FDB881A9FBBBFB6B" box="[908,1043,1194,1221]" pageId="23" pageNumber="24" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4F3B36B75349FF87FEF381CDFC6FFB47" blockId="23.[151,1437,1158,2013]" box="[199,967,1230,1257]" pageId="23" pageNumber="24">
<emphasis id="7DF0EAA55349FF87FEF381CDFE54FB46" bold="true" box="[199,508,1230,1256]" pageId="23" pageNumber="24">Other material examined.</emphasis>
<collectionCode id="2995AE725349FF87FC3681CDFD87FB47" box="[514,559,1230,1257]" country="Germany" name="Museum Donaueschingen" pageId="23" pageNumber="24">MD</collectionCode>
50 DC91,
<collectionCode id="2995AE725349FF87FC9781CCFD51FB47" box="[675,761,1231,1257]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
<collectionCode id="2995AE725349FF87FD3481CDFCB3FB47" box="[768,795,1230,1257]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="23" pageNumber="24">IE</collectionCode>
.2009.1568 (2).
</paragraph>
<paragraph id="4F3B36B75349FF87FEF381F1FB44FA33" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
<emphasis id="7DF0EAA55349FF87FEF381F1FE7EFAA2" bold="true" box="[199,470,1266,1292]" pageId="23" pageNumber="24">Comparative Material</emphasis>
.
<taxonomicName id="88844D345349FF87FFD381F1FC17FAA3" authority="McKnight, 2003" authorityName="McKnight" authorityYear="2003" box="[487,959,1266,1293]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="comatulina">
<emphasis id="7DF0EAA55349FF87FFD381F1FD5DFAA3" bold="true" box="[487,757,1266,1293]" italics="true" pageId="23" pageNumber="24">Ophiolebes comatulina</emphasis>
<materialsCitation id="FFEC3CEA5349FF87FCCB81F0FC17FAA3" ID-GBIF-Occurrence="3746276321" box="[767,959,1267,1293]" collectionCode="NZOI" pageId="23" pageNumber="24" specimenCode="E728" specimenCount="1">
<bibRefCitation id="2B154B465349FF87FCCB81F0FC17FAA3" author="McKnight, D. G." box="[767,959,1267,1293]" pageId="23" pageNumber="24" pagination="1 - 36" refId="ref34183" refString="McKnight, D. G. (2003) New brittle-stars (Echinodermata: Ophiuroidea) from New Zealand waters. Zootaxa, 352, 1 - 36. https: // doi. org / 10.11646 / zootaxa. 352.1.1" type="journal article" year="2003">McKnight, 2003</bibRefCitation>
</materialsCitation>
</taxonomicName>
:
<collectionCode id="2995AE725349FF87FDF881F0FBB9FAA3" box="[972,1041,1267,1293]" country="New Zealand" httpUri="http://grbio.org/cool/chf3-57df" name="New Zealand Oceanographic Institute" pageId="23" pageNumber="24">NZOI</collectionCode>
/
<specimenCode id="1F229ECC5349FF87FA2E81F0FBFDFAA3" box="[1050,1109,1267,1293]" collectionCode="NZOI" country="New Zealand" httpUri="http://grbio.org/cool/chf3-57df" name="New Zealand Oceanographic Institute" pageId="23" pageNumber="24">E728</specimenCode>
,
<materialsCitation id="FFEC3CEA5349FF87FA5781F0FD7FFA9F" ID-GBIF-Occurrence="3746276303" collectingDate="1967-03-25" collectionCode="NIWA" elevation="432" latitude="-37.625" longLatPrecision="92" longitude="177.2033" pageId="23" pageNumber="24" specimenCode="NIWA 3343" specimenCount="1" typeStatus="holotype">
<geoCoordinate id="2AB050705349FF87FA5781F0FB48FAA3" box="[1123,1248,1266,1293]" degrees="37" direction="south" minutes="37.5" orientation="latitude" pageId="23" pageNumber="24" precision="92" value="-37.625">37° 37.5´S</geoCoordinate>
,
<geoCoordinate id="2AB050705349FF87FADB81F0FA30FAA3" box="[1263,1432,1267,1293]" degrees="177" direction="east" minutes="12.198" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="177.2033">177° 12.198´E</geoCoordinate>
, 688
<quantity id="887C9B525349FF87FEFB8014FEB3FA9F" box="[207,283,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="23" pageNumber="24" unit="m" value="432.0">
<elevation id="C4A9D1845349FF87FEFB8014FEB3FA9F" box="[207,283,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="23" pageNumber="24" unit="m" value="432.0">432 m</elevation>
</quantity>
,
<date id="3B3A10775349FF87FF1E8015FE36FA9F" box="[298,414,1302,1329]" pageId="23" pageNumber="24" value="1967-03-25">
<collectingDate id="2B7EE99F5349FF87FF1E8015FE36FA9F" box="[298,414,1302,1329]" pageId="23" pageNumber="24" value="1967-03-25">25/3/1967</collectingDate>
</date>
,
<typeStatus id="903F88155349FF87FF998014FDBAFA9F" box="[429,530,1303,1329]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
,
<specimenCode id="1F229ECC5349FF87FC148014FD02FA9F" box="[544,682,1303,1329]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="23" pageNumber="24" type="Other institutional type">NIWA 3343</specimenCode>
(1)
</materialsCitation>
.
<materialsCitation id="FFEC3CEA5349FF87FCD28014FD10FAFB" ID-GBIF-Occurrence="3746276323" collectingDate="2005-10-18" collectionCode="MNHN, IE" elevation="569" latitude="-19.6" location="Plateau des Chesterfield" longLatPrecision="925" longitude="158.71667" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
EBISCO/DW2603,
<location id="4A5B606C5349FF87FDFD8014FB49FA9F" LSID="urn:lsid:plazi:treatment:C72D87A15349FF89FEA386F5FD1BF817:4A5B606C5349FF87FDFD8014FB49FA9F" box="[969,1249,1303,1329]" latitude="-19.6" longLatPrecision="925" longitude="158.71667" name="Plateau des Chesterfield" pageId="23" pageNumber="24">Plateau des Chesterfield</location>
,
<geoCoordinate id="2AB050705349FF87FADB8014FAFFFA9F" box="[1263,1367,1303,1329]" degrees="19" direction="south" minutes="36" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="-19.6">19° 36´S</geoCoordinate>
,
<geoCoordinate id="2AB050705349FF87FB538015FEB8FAFB" degrees="158" direction="east" minutes="43.0005" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="158.71667">158° 43.0005´E</geoCoordinate>
,
<quantity id="887C9B525349FF87FF298039FE08FAFB" box="[285,416,1338,1365]" metricMagnitude="2" metricUnit="m" metricValue="5.6899999999999995" metricValueMax="5.7" metricValueMin="5.68" pageId="23" pageNumber="24" unit="m" value="569.0" valueMax="570.0" valueMin="568.0">
<elevation id="C4A9D1845349FF87FF298039FE08FAFB" box="[285,416,1338,1365]" metricMagnitude="2" metricUnit="m" metricValue="5.6899999999999995" metricValueMax="5.7" metricValueMin="5.68" pageId="23" pageNumber="24" unit="m" value="569.0" valueMax="570.0" valueMin="568.0">568570 m</elevation>
</quantity>
,
<date id="3B3A10775349FF87FF9A8039FD99FAFB" box="[430,561,1338,1365]" pageId="23" pageNumber="24" value="2005-10-18">
<collectingDate id="2B7EE99F5349FF87FF9A8039FD99FAFB" box="[430,561,1338,1365]" pageId="23" pageNumber="24" value="2005-10-18">18/10/2005</collectingDate>
</date>
,
<collectionCode id="2995AE725349FF87FC0A8038FD3CFAFB" box="[574,660,1339,1365]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
<collectionCode id="2995AE725349FF87FCA98038FD10FAFB" box="[669,696,1339,1365]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="23" pageNumber="24">IE</collectionCode>
</materialsCitation>
.
<materialsCitation id="FFEC3CEA5349FF87FC8F8038FC90FAD7" ID-GBIF-Occurrence="3746276301" collectingDate="2005-10-19" collectionCode="MNHN, IE" elevation="521" latitude="-19.616667" location="Plateau des Chesterfield" longLatPrecision="925" longitude="158.7" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
2013.6311 (1). EBISCO/DW2613,
<location id="4A5B606C5349FF87FA658038FAC0FAFB" LSID="urn:lsid:plazi:treatment:C72D87A15349FF89FEA386F5FD1BF817:4A5B606C5349FF87FA658038FAC0FAFB" box="[1105,1384,1339,1365]" latitude="-19.616667" longLatPrecision="925" longitude="158.7" name="Plateau des Chesterfield" pageId="23" pageNumber="24">Plateau des Chesterfield</location>
,
<geoCoordinate id="2AB050705349FF87FB418038FF67FAD7" degrees="19" direction="south" minutes="37" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="-19.616667">19° 37´S</geoCoordinate>
,
<geoCoordinate id="2AB050705349FF87FEE8805DFE3BFAD7" box="[220,403,1374,1401]" degrees="158" direction="east" minutes="41.9998" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="158.7">158° 41.9998´E</geoCoordinate>
,
<quantity id="887C9B525349FF87FFAB805DFD8AFAD7" box="[415,546,1374,1401]" metricMagnitude="2" metricUnit="m" metricValue="5.205" metricValueMax="5.22" metricValueMin="5.1899999999999995" pageId="23" pageNumber="24" unit="m" value="520.5" valueMax="522.0" valueMin="519.0">
<elevation id="C4A9D1845349FF87FFAB805DFD8AFAD7" box="[415,546,1374,1401]" metricMagnitude="2" metricUnit="m" metricValue="5.205" metricValueMax="5.22" metricValueMin="5.1899999999999995" pageId="23" pageNumber="24" unit="m" value="520.5" valueMax="522.0" valueMin="519.0">519522 m</elevation>
</quantity>
,
<date id="3B3A10775349FF87FC1B805DFD1AFAD7" box="[559,690,1374,1401]" pageId="23" pageNumber="24" value="2005-10-19">
<collectingDate id="2B7EE99F5349FF87FC1B805DFD1AFAD7" box="[559,690,1374,1401]" pageId="23" pageNumber="24" value="2005-10-19">19/10/2005</collectingDate>
</date>
,
<collectionCode id="2995AE725349FF87FC8B805CFCBDFAD7" box="[703,789,1375,1401]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
<collectionCode id="2995AE725349FF87FD29805CFC90FAD7" box="[797,824,1375,1401]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="23" pageNumber="24">IE</collectionCode>
</materialsCitation>
.
<materialsCitation id="FFEC3CEA5349FF87FD0F805CFB15FA33" ID-GBIF-Occurrence="3746276317" collectingDate="1999-06-03" collectionCode="NIWA" elevation="980" latitude="-39.475166" location="Ritchie Hill Seamount" longLatPrecision="11" longitude="178.42467" pageId="23" pageNumber="24" specimenCode="NIWA 41434" specimenCount="1" typeStatus="holotype">
2013.6287 (2). KAH9907/37,
<location id="4A5B606C5349FF87FAA3805CFA30FAD7" LSID="urn:lsid:plazi:treatment:C72D87A15349FF89FEA386F5FD1BF817:4A5B606C5349FF87FAA3805CFA30FAD7" box="[1175,1432,1375,1401]" latitude="-39.475166" longLatPrecision="11" longitude="178.42467" name="Ritchie Hill Seamount" pageId="23" pageNumber="24">Ritchie Hill Seamount</location>
,
<geoCoordinate id="2AB050705349FF87FEA38080FE88FA33" box="[151,288,1411,1437]" degrees="39" direction="south" minutes="29.44" orientation="latitude" pageId="23" pageNumber="24" precision="9" value="-39.490665">39° 29.44´S</geoCoordinate>
,
<geoCoordinate id="2AB050705349FF87FF1F8080FE69FA33" box="[299,449,1410,1437]" degrees="178" direction="east" minutes="25.05" orientation="longitude" pageId="23" pageNumber="24" precision="9" value="178.4175">178° 25.05´E</geoCoordinate>
to 39° 28.51´S, 178° 25.48´E, 1000
<quantity id="887C9B525349FF87FD568080FC03FA33" box="[866,939,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="23" pageNumber="24" unit="m" value="980.0">
<elevation id="C4A9D1845349FF87FD568080FC03FA33" box="[866,939,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="23" pageNumber="24" unit="m" value="980.0">980 m</elevation>
</quantity>
,
<date id="3B3A10775349FF87FD838080FBB6FA33" box="[951,1054,1411,1437]" pageId="23" pageNumber="24" value="1999-06-03">
<collectingDate id="2B7EE99F5349FF87FD838080FBB6FA33" box="[951,1054,1411,1437]" pageId="23" pageNumber="24" value="1999-06-03">3/6/1999</collectingDate>
</date>
,
<specimenCode id="1F229ECC5349FF87FA1D8080FB15FA33" box="[1065,1213,1411,1437]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="23" pageNumber="24" type="Other institutional type">NIWA 41434</specimenCode>
</materialsCitation>
(1).
</paragraph>
<paragraph id="4F3B36B75349FF87FEF380A4FBD1F8AB" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
<emphasis id="7DF0EAA55349FF87FEF380A4FE68FA6E" bold="true" box="[199,448,1446,1473]" pageId="23" pageNumber="24">
<typeStatus id="903F88155349FF87FEF380A4FE9CFA6F" box="[199,308,1447,1473]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
description
</emphasis>
. Disc is
<quantity id="887C9B525349FF87FC2B80A5FDDCFA6F" box="[543,628,1446,1473]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="23" pageNumber="24" unit="mm" value="5.5">5.5 mm</quantity>
dd, the longest attached arm piece is ~
<quantity id="887C9B525349FF87FA1480A4FBC6FA6F" box="[1056,1134,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="mm" value="10.0">10 mm</quantity>
long; disc is tumid radially and sunken interradially, covered in rounded non-imbricating scales, embedded in thin skin, ~
<quantity id="887C9B525349FF87FA9880C9FAABFA4B" box="[1196,1283,1482,1509]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="24" unit="mm" value="0.5">0.5 mm</quantity>
dia, minutely perforated, fully contiguous and more polygonal over the radial shields, slightly separated in the centre of the disc (
<figureCitation id="D7BF2A325349FF87FE948310FF53F983" box="[160,251,1555,1581]" captionStart="FIGURE 9" captionStartId="24.[151,250,1678,1702]" captionTargetBox="[151,1436,309,1653]" captionTargetId="figure-88@24.[151,1436,309,1654]" captionTargetPageId="24" captionText="FIGURE 9. Ophiolebes paulensis sp. nov., holotype, MNHN IE.2009.1566, A: dorsal and B: ventral views, C: inset showing hook-shaped distal arm spines." figureDoi="http://doi.org/10.5281/zenodo.6404700" httpUri="https://zenodo.org/record/6404700/files/figure.png" pageId="23" pageNumber="24">Fig. 9A</figureCitation>
). Scales support one, rarely 2, disc spines, tapered conical to cylindrical, thorny on blunt tip, 3 times as high as wide, ~
<quantity id="887C9B525349FF87FF768334FE32F9FF" box="[322,410,1591,1617]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="24" unit="mm" value="0.3">0.3 mm</quantity>
high in centre, smaller near margin, base smooth, upper half with more sculptured stereom and some small thorns. Radial shields are evidently bar-like but covered by skin and scales. Ventral disc (
<figureCitation id="D7BF2A325349FF87FB098358FA3CF9DB" box="[1341,1428,1627,1653]" captionStart="FIGURE 9" captionStartId="24.[151,250,1678,1702]" captionTargetBox="[151,1436,309,1653]" captionTargetId="figure-88@24.[151,1436,309,1654]" captionTargetPageId="24" captionText="FIGURE 9. Ophiolebes paulensis sp. nov., holotype, MNHN IE.2009.1566, A: dorsal and B: ventral views, C: inset showing hook-shaped distal arm spines." figureDoi="http://doi.org/10.5281/zenodo.6404700" httpUri="https://zenodo.org/record/6404700/files/figure.png" pageId="23" pageNumber="24">Fig. 9B</figureCitation>
) surfaces are covered in similar scales, 45 rows from oral shield to margin, marginal scales with short conical stumps, 2 times as long as wide, but extend only half the way to the oral shield. Genital slits extend from oral shield for 2.5 arm segments, stopping short of the margin. Genital plates visible externally, adradial plates with a short base and broadly curved, abradial ones abut it closely, extending for half the length of the slit.
</paragraph>
<paragraph id="4F3B36B75349FF88FEF3820CFCDBFEB3" blockId="23.[151,1437,1158,2013]" lastBlockId="24.[151,1436,151,285]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Jaws are as long as wide, supporting a tapered ventralmost tooth and three spiniform oral papillae, all arising from the oral plate, the proximally pointed inner one representing the infradental papilla, the obliquely pointed middle one the lateral oral papilla and the distal one the adoral shield spine (following
<bibRefCitation id="2B154B465349FF87FA258254FB06F8DF" author="Hendler, G." box="[1041,1198,1879,1905]" pageId="23" pageNumber="24" pagination="189 - 311" refId="ref31642" refString="Hendler, G. (2018) Armed to the teeth: a new paradigm for the buccal skeleton of brittle stars (Echinodermata: Ophiuroidea). Contributions in Science, 526, 189 - 311. https: // doi. org / 10.5962 / p. 324539" type="journal article" year="2018">Hendler 2018</bibRefCitation>
, fig. 16). All papillae are pitted with tiny thorns near the tip, hollow. Four are teeth visible from the external ventral surface, similar shape to the ventralmost one. Oral plates are raised near the papillae and sunken slightly behind. Second oral podia arise from a naked patch between the adoral shields and first VAP. Adoral shields are 3 times as wide as long, sausage shaped, with concave inner and convex outer margins, rounded outer ends, fully contiguous interradially, separated radially by the first VAP. Oral shields are pentagonal, with concave proximolateral margins, obtuse proximal angle, slightly tapering distolateral sides and a truncate distal margin, the distal section of the plate is sunken down to the level of the ventral disc, central section tumid in the madreporite.
</paragraph>
<caption id="1BFB663F5346FF88FEA3838DFE78F964" ID-DOI="http://doi.org/10.5281/zenodo.6404700" ID-Zenodo-Dep="6404700" httpUri="https://zenodo.org/record/6404700/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1678,1702]" targetBox="[151,1436,309,1653]" targetPageId="24">
<paragraph id="4F3B36B75346FF88FEA3838DFE78F964" blockId="24.[151,1436,1678,1738]" pageId="24" pageNumber="25">
<emphasis id="7DF0EAA55346FF88FEA3838DFEBCF908" bold="true" box="[151,276,1678,1702]" pageId="24" pageNumber="25">FIGURE 9.</emphasis>
<taxonomicName id="88844D345346FF88FF2F838DFE5DF908" authority="OHara &amp; Thuy, 2022" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[283,501,1678,1702]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="25" phylum="Echinodermata" rank="species" species="paulensis" status="sp. nov.">
<emphasis id="7DF0EAA55346FF88FF2F838DFE5DF908" box="[283,501,1678,1702]" italics="true" pageId="24" pageNumber="25">Ophiolebes paulensis</emphasis>
</taxonomicName>
<emphasis id="7DF0EAA55346FF88FFC8838DFDE5F908" bold="true" box="[508,589,1678,1702]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="66C357DE5346FF88FFC8838DFDE5F908" box="[508,589,1678,1702]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, MNHN IE.2009.1566, A: dorsal and B: ventral views, C: inset showing hook-shaped distal arm spines.
</paragraph>
</caption>
<paragraph id="4F3B36B75346FF89FEF383FBFA34FEB3" blockId="24.[151,1436,1784,2027]" lastBlockId="25.[151,1437,151,1979]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">First 12 DAPs are triangular, wide as long, contiguous. Succeeding DAPs are rounded triangular, with convex distal margin, and long tapered proximolateral sides, with a small truncate proximal margin, sometimes contiguous, or with a small skin-covered bare patch or a small supplementary plate between plates, typically 1.5 to 2 times as long as wide. LAPs are roughly quadrangular, contiguous, arm spine ridge raised slightly, contiguous ventrally but not dorsally. VAPs on the first segment are 1.5 times as long as wide, widest proximally, with straight proximolateral edges and a straight to concave proximal edge, concave sides (to accommodate the adoral shields), straight distolateral edges and a truncate to rounded distal margin, just touching the proximal angle of the 2nd VAPs, 2nd VAPs are fanshaped with straight proximolateral edges, an acute proximal angle and a rounded distal edge, widest at the distal 1/3 of the plate. Tube foot openings are very small, rounded, the proximolateral edges of the VAPs are not concave around the pore. Succeeding plates become as wide as long, well separated from their neighbours, widest in the middle of the plate, and notched around the pore, the distal edge can become centrally concave on distal segments.</paragraph>
<paragraph id="4F3B36B75347FF89FEF38424FA2CFD93" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
Arm spines are rounded, tapering to a blunt tip, hollow, with sculptured stereom but generally no thorns, do not meet basally even on basal segments just free of the disc. Three arm spines occur on the first segments, four on second and third, four on the fourth segment, then five on segment five and six, just free of the disc, an occasional segment has six spines in the middle of the arm. Upper arm spines are generally longest, particularly on the 5th and 6th segment, to
<quantity id="887C9B525347FF89FF7984B4FE0EFE7F" box="[333,422,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="mm" value="1.7">1.7 mm</quantity>
, 2nd spines (from dorsal) about half its length, lower three 1/4 as short and more slender. From about the 10th segment, the upper spine reduces in size becoming subequal with the others, and the lower arm spines become shorter and stouter with proximally directed thorns (
<figureCitation id="D7BF2A325347FF89FDBA84FCFC4FFDB7" box="[910,999,511,537]" captionStart="FIGURE 9" captionStartId="24.[151,250,1678,1702]" captionTargetBox="[151,1436,309,1653]" captionTargetId="figure-88@24.[151,1436,309,1654]" captionTargetPageId="24" captionText="FIGURE 9. Ophiolebes paulensis sp. nov., holotype, MNHN IE.2009.1566, A: dorsal and B: ventral views, C: inset showing hook-shaped distal arm spines." figureDoi="http://doi.org/10.5281/zenodo.6404700" httpUri="https://zenodo.org/record/6404700/files/figure.png" pageId="25" pageNumber="26">Fig. 9C</figureCitation>
). Tentacle scales absent proximally (or rubbed off), a tiny pointed scale on the ventral edge of LAP in the middle of the arm, 1/10 the length of the VAP.
</paragraph>
<paragraph id="4F3B36B75347FF89FEF38744FAC4FDCF" blockId="25.[151,1437,151,1979]" box="[199,1388,583,610]" pageId="25" pageNumber="26">Colour (preserved). light brown, the tube feet and tips of the arm spines (skin covered?) are darker brown.</paragraph>
<paragraph id="4F3B36B75347FF89FEF38768FA96FD63" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
<emphasis id="7DF0EAA55347FF89FEF38768FE10FD2A" bold="true" box="[199,440,618,645]" pageId="25" pageNumber="26">
<typeStatus id="903F88155347FF89FEF38768FE9CFD2B" box="[199,308,619,645]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
variations.
</emphasis>
Discs vary from 4.0
<quantity id="887C9B525347FF89FC918769FD50FD2B" box="[677,760,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="25" pageNumber="26" unit="mm" value="5.5">5.5 mm</quantity>
dd, longest arm fragment is
<quantity id="887C9B525347FF89FA198769FB27FD2B" box="[1069,1167,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="25" pageNumber="26" unit="mm" value="12.5">12.5 mm</quantity>
on a
<quantity id="887C9B525347FF89FAF38769FAAEFD2B" box="[1223,1286,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="5.0">5 mm</quantity>
dd specimen. The distal section of the bar-like radial shields are exposed on some radii. Some specimens have long disc stumps (to five times height/dia) with stereom extending into short thorns around the spine stem as well as the tip.
</paragraph>
<paragraph id="4F3B36B75347FF89FEF387D4FCBEFBF7" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
<emphasis id="7DF0EAA55347FF89FEF387D4FDB4FD5E" bold="true" box="[199,540,726,753]" pageId="25" pageNumber="26">
<typeStatus id="903F88155347FF89FEF387D4FE9CFD5F" box="[199,308,727,753]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
ossicle description.
</emphasis>
Proximal LAPs (
<figureCitation id="D7BF2A325347FF89FCD687D4FCC1FD5F" box="[738,873,727,753]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10AB</figureCitation>
) are slightly higher than long, of rounded outline, relatively thin, with strongly convex distal edge; ventral edge oblique, straight, slightly protruding; dorsal edge slightly concave as a result of a weak constriction; outer surface with coarsely meshed stereom, devoid of tubercles or a vertical striation; outer proximal edge with a proximalwards-pointed spur, entirely lined by a band of more finely meshed stereom. Spine articulations (
<figureCitation id="D7BF2A325347FF89FCBF8664FD59FC2F" box="[651,753,871,897]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10C</figureCitation>
) are freestanding in a continuous vertical row on a strongly raised distal portion of the LAP, ear-shaped, moderately large, composed of proximally merged dorsal and ventral lobes enclosing nearly equal-sized muscle and nerve openings and with a moderately well-developed sigmoidal fold. Inner side of LAPs have a large, prominent, well-defined, slender digit-1-shaped vertebral articular ridge; poorly-defined spur in the dorsal half of the inner distal edge corresponding to the protruding spur on the outer distal edge of the LAPs; tentacle notches are small, oblique, shallow. Very small, inconspicuous perforations occur close to the ventro-distal tip of the vertebral articular structure.
</paragraph>
<paragraph id="4F3B36B75347FF89FEF38160FCBFFA9F" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
Vertebrae (
<figureCitation id="D7BF2A325347FF89FF708160FE68FBD3" box="[324,448,1123,1149]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10EI</figureCitation>
) have wide triangular dorsal fossae and with relatively large ventral muscle fossae on both distal and proximal faces; distal articulations are relatively large, hour-glass-shaped (streptospondylous) with a very small ventral projection of the middle saddle; hourglass-shaped proximal articulation with a vertical pore in the ventral half; oblique L-shaped lateral articulation structure corresponding to articular structure on the inner side of the LAP; dorso-distal muscle fossae are deeply carved into the lateral side of the vertebra; relatively large dorsal and ventral furrows, with moderately large podial basins.
</paragraph>
<paragraph id="4F3B36B75347FF89FEF38038FEDEF983" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
Oral plates (Fig. L) are split in distal and proximal halves, approximately as high as long, with adradial muscle fossa lining the ventro-distal edge of articulation area. Dental plates (
<figureCitation id="D7BF2A325347FF89FDA5805CFC5DFAD7" box="[913,1013,1375,1401]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10K</figureCitation>
) are undivided, very narrow, rounded, with single row of small, shallow tooth sockets surrounded by weakly-defined protruding ring. Abradial genital plates (
<figureCitation id="D7BF2A325347FF89FEDE80A4FEF9FA6F" box="[234,337,1447,1473]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10N</figureCitation>
) shorter than the adradial ones (
<figureCitation id="D7BF2A325347FF89FCF180A4FC87FA6F" box="[709,815,1447,1473]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10O</figureCitation>
), slender, bar-like without ridges or groves and with a straight adradio-distal tip. Radial shields (
<figureCitation id="D7BF2A325347FF89FC4580C8FD7CFA4B" box="[625,724,1483,1509]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10P</figureCitation>
) are strongly elongated, bar-like, slightly bent, devoid of outer surface ornamentation. A proximal dorsal arm spine (
<figureCitation id="D7BF2A325347FF89FCC480ECFCF0F9A7" box="[752,856,1519,1545]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10D</figureCitation>
), disc spine (
<figureCitation id="D7BF2A325347FF89FDDA80ECFBF3F9A7" box="[1006,1115,1519,1545]" captionStart="FIGURE 10" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[172,1415,181,1813]" captionTargetId="figure-26@26.[172,1415,181,1813]" captionTargetPageId="26" captionText="FIGURE 10. (AP) Ophiolebes paulensis sp. nov., paratype, MNHN IE.2009.1567.AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D: arm spine. EI: proximal vertebra in distal (E), proximal (F), lateral (G), dorsal (H) and ventral (I) views. J: ventral arm plate. K: dental plate. L: oral plate split in proximal and distal halves. M: disc spine. N: abradial genital plate. O: adradial genital plate. P: radial shield.Abbreviations:AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404704" httpUri="https://zenodo.org/record/6404704/files/figure.png" pageId="25" pageNumber="26">Fig. 10M</figureCitation>
), and ventral arm plate (
<figureCitation id="D7BF2A325347FF89FB5B80ECFF18F983" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (AS) Ophiolebes felli sp. nov. paratype, IE.2009.1569. AB: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. DE: arm spines. F: tentacle scale. GK: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="25" pageNumber="26">Fig. 8J</figureCitation>
) are also figured.
</paragraph>
</subSubSection>
<subSubSection id="079E653C5347FF89FEF38335FDECF9FF" box="[199,580,1590,1617]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="4F3B36B75347FF89FEF38335FDECF9FF" blockId="25.[151,1437,151,1979]" box="[199,580,1590,1617]" pageId="25" pageNumber="26">
<emphasis id="7DF0EAA55347FF89FEF38335FEF1F9FE" bold="true" box="[199,345,1590,1616]" pageId="25" pageNumber="26">Distribution</emphasis>
. SPA (
<quantity id="887C9B525347FF89FF918334FD90F9FF" box="[421,568,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.31" metricValueMax="1.68" metricValueMin="0.94" pageId="25" pageNumber="26" unit="m" value="1310.0" valueMax="1680.0" valueMin="940.0">9401680 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="079E653C5347FF89FEF38358FCBEF83B" pageId="25" pageNumber="26" type="discussion">
<paragraph id="4F3B36B75347FF89FEF38358FCBEF83B" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
<emphasis id="7DF0EAA55347FF89FEF38358FE9CF9DB" bold="true" box="[199,308,1627,1653]" pageId="25" pageNumber="26">Remarks</emphasis>
. This species is similar to
<taxonomicName id="88844D345347FF89FC5F8358FB9FF9DB" authority="McKnight, 2003" authorityName="McKnight" authorityYear="2003" box="[619,1079,1627,1653]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="comatulina">
<emphasis id="7DF0EAA55347FF89FC5F8358FCDAF9DB" box="[619,882,1627,1653]" italics="true" pageId="25" pageNumber="26">Ophiolebes comatulina</emphasis>
<bibRefCitation id="2B154B465347FF89FD4F8358FB9FF9DB" author="McKnight, D. G." box="[891,1079,1627,1653]" pageId="25" pageNumber="26" pagination="1 - 36" refId="ref34183" refString="McKnight, D. G. (2003) New brittle-stars (Echinodermata: Ophiuroidea) from New Zealand waters. Zootaxa, 352, 1 - 36. https: // doi. org / 10.11646 / zootaxa. 352.1.1" type="journal article" year="2003">McKnight, 2003</bibRefCitation>
</taxonomicName>
known from NE
<collectingCountry id="379376275347FF89FB308358FA34F9DB" box="[1284,1436,1627,1653]" name="New Zealand" pageId="25" pageNumber="26">New Zealand</collectingCountry>
(
<quantity id="887C9B525347FF89FEAB837CFE99F937" box="[159,305,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" metricValueMax="10.0" metricValueMin="4.32" pageId="25" pageNumber="26" unit="m" value="716.0" valueMax="1000.0" valueMin="432.0">4321000 m</quantity>
) and the Chesterfield Bank,
<collectingCountry id="379376275347FF89FCB5837CFC9AF937" box="[641,818,1663,1689]" name="New Caledonia" pageId="25" pageNumber="26">New Caledonia</collectingCountry>
(
<quantity id="887C9B525347FF89FD77837DFC60F937" box="[835,968,1662,1689]" metricMagnitude="2" metricUnit="m" metricValue="5.445" metricValueMax="5.7" metricValueMin="5.1899999999999995" pageId="25" pageNumber="26" unit="m" value="544.5" valueMax="570.0" valueMin="519.0">519570 m</quantity>
) (OHara unpublished data).
<taxonomicName id="88844D345347FF89FB29837CFEBDF913" authorityName="McKnight" authorityYear="2003" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="comatulina">
<emphasis id="7DF0EAA55347FF89FB29837CFEBDF913" italics="true" pageId="25" pageNumber="26">Ophiolebes comatulina</emphasis>
</taxonomicName>
differs in having diamond shaped oral shields (tapering to a distal angle) and a stout tentacle scale on proximal segments, 1/4 the length of the VAP.All the
<taxonomicName id="88844D345347FF89FCDF83C4FC22F94F" authorityName="McKnight" authorityYear="2003" box="[747,906,1735,1761]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="comatulina">
<emphasis id="7DF0EAA55347FF89FCDF83C4FC22F94F" box="[747,906,1735,1761]" italics="true" pageId="25" pageNumber="26">O. comatulina</emphasis>
</taxonomicName>
specimens to date have been found on crinoids. Although there is now no evidence of crinoid material with the MD50
<taxonomicName id="88844D345347FF89FDF683E8FB1CF8AB" authority="OHara &amp; Thuy, 2022" authorityName="OHara &amp; Thuy" authorityYear="2022" box="[962,1204,1771,1797]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="paulensis" status="sp. nov.">
<emphasis id="7DF0EAA55347FF89FDF683E8FB1CF8AB" box="[962,1204,1771,1797]" italics="true" pageId="25" pageNumber="26">Ophiolebes paulensis</emphasis>
</taxonomicName>
<emphasis id="7DF0EAA55347FF89FA8983E8FAB0F8AB" bold="true" box="[1213,1304,1771,1797]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="66C357DE5347FF89FA8983E8FAB0F8AB" box="[1213,1304,1771,1797]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens, the
<typeStatus id="903F88155347FF89FEF0820CFF5CF887" box="[196,244,1807,1833]" pageId="25" pageNumber="26">type</typeStatus>
locality (stn CP7) was the only sample from the survey recorded with crinoids (
<bibRefCitation id="2B154B465347FF89FA9D820CFAEDF887" author="Arnaud, P. M." box="[1193,1349,1807,1833]" pageId="25" pageNumber="26" pagination="1 - 130" refId="ref29101" refString="Arnaud, P. M. (1990) Les rapports des campagnes a la mer aux iles Saint-Paul et Amsterdam a bord du Marion-Dufresne 3 juillet - 1 aout 1986. Publications de la Mission de Recherche T. A. A. F., 86 - 04, pp. 1 - 130." type="book chapter" year="1990">Arnaud 1990</bibRefCitation>
). Other
<taxonomicName id="88844D345347FF89FEA38230FEC6F8E3" box="[151,366,1843,1869]" pageId="25" pageNumber="26">
<emphasis id="7DF0EAA55347FF89FEA38230FEBFF8E3" box="[151,279,1843,1869]" italics="true" pageId="25" pageNumber="26">Ophiolebes</emphasis>
species
</taxonomicName>
have disc granules and/or short spines on the disc (see above) rather than tall stumps. The South African species
<taxonomicName id="88844D345347FF89FF658254FCDEF8DF" authority="(H. L. Clark, 1923)" baseAuthorityName="H. L. Clark" baseAuthorityYear="1923" box="[337,886,1879,1905]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosemnotes" kingdom="Animalia" order="Ophiacanthida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="corynephora">
<emphasis id="7DF0EAA55347FF89FF658254FD3EF8DF" box="[337,662,1879,1905]" italics="true" pageId="25" pageNumber="26">Ophiosemnotes corynephora</emphasis>
(H.L.
<bibRefCitation id="2B154B465347FF89FCD28254FCC6F8DF" author="Clark, H. L." box="[742,878,1879,1905]" pageId="25" pageNumber="26" pagination="221 - 435" refId="ref30081" refString="Clark, H. L. (1923) The echinoderm fauna of South Africa. Annals of the South African Museum, 13, 221 - 435, pls. 8 - 23." type="journal article" year="1923">Clark, 1923</bibRefCitation>
)
</taxonomicName>
has superficially similar disc spines to the new species but differs in having wide exposed radial shields.
</paragraph>
</subSubSection>
<subSubSection id="079E653C5347FF89FEF3829CFD1BF817" box="[199,691,1951,1979]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="4F3B36B75347FF89FEF3829CFD1BF817" blockId="25.[151,1437,151,1979]" box="[199,691,1951,1979]" pageId="25" pageNumber="26">
<emphasis id="7DF0EAA55347FF89FEF3829CFEE0F817" bold="true" box="[199,328,1951,1977]" pageId="25" pageNumber="26">Etymology</emphasis>
. Named after the Île Saint-Paul.
</paragraph>
</subSubSection>
</treatment>
</document>