treatments-xml/data/C0/4B/87/C04B87CFFFB9FF4530ABFF1DFB4DF897.xml

<document id="5A8866D7DC0E768EF17C1A6A74C8587E" ID="10.11646/zootaxa.4277.2.2SLASH11367" ID-CLB-Dataset="32829" ID-DOI="10.11646/zootaxa.4277.2.2" ID-GBIF-Dataset="5d4045c8-2c5a-4f75-8ca7-a2401ed0d227" ID-ISSN="1175-5326" ID-Zenodo-Dep="809634" ID-ZooBank="1551FE58-FCC9-4D45-AE2A-77655BB53147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497598594012" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2017" docId="C04B87CFFFB9FF4530ABFF1DFB4DF897" docLanguage="en" docName="zootaxa.4277.2.2SLASH11367.pdf" docOrigin="Zootaxa 4277 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheopsis paratrigona Anker 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="214" masterDocId="3C72FFB7FFB1FF4A303CFF8AFF9DFFE7" masterDocTitle="Strongly carinate species of Alpheopsis Coutière, 1897 of the tropical Atlantic and eastern Pacific, with redescription of A. trigona (Rathbun, 1901) and description of three new species (Malacostraca: Decapoda: Alpheidae)" masterLastPageNumber="227" masterPageNumber="199" pageNumber="207" updateTime="1698447107539" updateUser="ExternalLinkService">
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<mods:title id="038085842CAB885DD7E9082FEC2CA5EC">Strongly carinate species of Alpheopsis Coutière, 1897 of the tropical Atlantic and eastern Pacific, with redescription of A. trigona (Rathbun, 1901) and description of three new species (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<mods:namePart id="D0CBF99ADB9D0F8712349240E0418C99">Anker, Arthur</mods:namePart>
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<mods:date id="22E2B217800F4ECFCEB4841BD58430A7">2017</mods:date>
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<paragraph id="485D36D9FFB9FF4230ABFF1DFD8BFF55" blockId="8.[151,534,151,211]" box="[151,534,151,178]" pageId="8" pageNumber="207">
<heading id="131581B5FFB9FF4230ABFF1DFD8BFF55" bold="true" box="[151,534,151,178]" fontSize="11" level="1" pageId="8" pageNumber="207" reason="1">
<emphasis id="7A96EACBFFB9FF4230ABFF1DFD8BFF55" bold="true" box="[151,534,151,178]" pageId="8" pageNumber="207">
<taxonomicName id="8FE24D5AFFB9FF4230ABFF1DFE2CFF56" ID-CoL="85LCZ" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[151,433,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFB9FF4230ABFF1DFE2CFF56" bold="true" box="[151,433,151,177]" italics="true" pageId="8" pageNumber="207">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFB9FF423185FF12FD8BFF55" box="[441,534,152,178]" pageId="8" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="485D36D9FFB9FF4230ABFF30FE88FF34" blockId="8.[151,534,151,211]" box="[151,277,186,211]" pageId="8" pageNumber="207">
(
<figureCitation id="D0D92A5CFFB9FF4230A3FF30FE90FF34" box="[159,269,186,211]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStart-5="FIGURE 10" captionStartId-0="9.[151,250,1773,1795]" captionStartId-1="10.[151,250,1360,1382]" captionStartId-2="11.[151,250,1759,1781]" captionStartId-3="12.[151,250,1785,1807]" captionStartId-4="13.[151,250,1655,1677]" captionStartId-5="14.[151,250,1923,1945]" captionTargetBox-0="[161,1424,202,1741]" captionTargetBox-1="[158,1418,204,1325]" captionTargetBox-2="[163,1414,206,1724]" captionTargetBox-3="[175,1424,205,1748]" captionTargetBox-4="[150,1436,193,1634]" captionTargetBox-5="[150,1436,769,1901]" captionTargetId-0="figure@9.[151,1436,193,1752]" captionTargetId-1="figure@10.[151,1436,193,1339]" captionTargetId-2="figure@11.[151,1436,193,1738]" captionTargetId-3="figure@12.[151,1436,193,1764]" captionTargetId-4="figure@13.[150,1436,193,1634]" captionTargetId-5="figure@14.[150,1436,769,1901]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="14" captionText-0="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." captionText-1="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." captionText-2="FIGURE 7. Alpheopsis paratrigona sp. nov., male (cl 4.0 mm) from Barbuda, Antigua &amp; Barbuda (USNM 135846) [a – i]; ovig. female (cl 4.4 mm) from St. John Island, US Virgin Islands (USNM 134763) [j – l]; male (cl 5.0 mm) from Bermuda (USNM 107089) [m – o]; a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—left cheliped, mesial; f—same, chela, mesial; g—same, chela fingers (open), lateral; h—second pereiopod, lateral; i—third pereiopod, lateral; j—anterior region of carapace, lateral; k—left cheliped, mesial; lsame, lateral; m—carapace and anterior pleon, lateral; n—anterior region of carapace and frontal appendages, lateral; o—same, dorsal. Modified from Anker (2001), except for j, k, l (drawings prepared, but not used in Anker 2001)." captionText-3="FIGURE 8. Alpheopsis paratrigona sp. nov., male (cl 5.1 mm) [a – n] and female (cl 3.3 mm) [o, p] from Guadeloupe, French Antilles (MNHN- 2014 - IU- 19473); a—carapace, lateral; b—anterior region of carapace and frontal appendages, dorsal; csame, lateral; d—telson, dorsal; e—same, detail of posterolateral spiniform setae, dorsal; f—third maxilliped, lateral; g—left (major) cheliped, lateral; h—same, chela, mesial; i—same, chela fingers, lateral; j—right (minor) cheliped, mesial; k, same, chela, mesial; l—same, mesial; m—third pereiopod, lateral; n—uropod, dorsal; o—anterior region of carapace and frontal appendages, dorsal; p—same, lateral. Drawings prepared, but not used in Anker (2001); (e) drawn without scale." captionText-4="FIGURE 9. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222) [a]; male (cl 4.5 mm) from Carrie Bow Cay, Belize (ULLZ 8939) [b]. Photographs: A. Anker (a) and Darryl L. Felder (b)." captionText-5="FIGURE 10. Alpheopsis paratrigona sp. nov., male (cl not measured) from Guana Island, British Virgin Islands, in dorsal [a] and lateral [b] views (specimen possibly deposited in the Natural History Museum of Los Angeles Country, LACM, not available for study). Photographs: Todd Zimmerman (a, b)." httpUri-0="https://zenodo.org/record/809644/files/figure.png" httpUri-1="https://zenodo.org/record/809646/files/figure.png" httpUri-2="https://zenodo.org/record/809648/files/figure.png" httpUri-3="https://zenodo.org/record/809650/files/figure.png" httpUri-4="https://zenodo.org/record/809652/files/figure.png" httpUri-5="https://zenodo.org/record/809654/files/figure.png" pageId="8" pageNumber="207">Fig. 5–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFB9FF4230ABFF77FDE5FEB7" pageId="8" pageNumber="207" type="reference_group">
<paragraph id="485D36D9FFB9FF4230ABFF77FD1CFED5" blockId="8.[151,1338,253,336]" pageId="8" pageNumber="207">
<treatmentCitationGroup id="68F211F7FFB9FF4230ABFF77FD1CFED5" pageId="8" pageNumber="207">
<taxonomicName id="8FE24D5AFFB9FF4230ABFF77FEFDFEF4" ID-CoL="84W6C" authorityName="Rathbun" authorityYear="1901" box="[151,352,253,275]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="species" species="trigonus">
<emphasis id="7A96EACBFFB9FF4230ABFF77FEFDFEF4" box="[151,352,253,275]" italics="true" pageId="8" pageNumber="207">Alpheopsis trigonus</emphasis>
</taxonomicName>
— 
<treatmentCitation id="C94310C8FFB9FF423146FF77FD85FEF4" author="Chace" box="[378,536,253,275]" page="56" pageId="8" pageNumber="207" year="1972">
<bibRefCitation id="2C734B28FFB9FF423146FF77FE6BFEF4" author="Chace" box="[378,502,253,275]" pageId="8" pageNumber="207" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. https: // doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">Chace 1972</bibRefCitation>
: 56
</treatmentCitation>
(
<emphasis id="7A96EACBFFB9FF42321BFF74FDF7FEF4" box="[551,618,254,275]" italics="true" pageId="8" pageNumber="207">partim</emphasis>
, material from Barbuda and Yucatan) [not 
<taxonomicName id="8FE24D5AFFB9FF423423FF74FAA9FEF4" ID-CoL="84W6B" authority="Rathbun, 1901" authorityName="Rathbun" authorityYear="1901" box="[1055,1332,253,275]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFB9FF423423FF74FB1BFEF4" box="[1055,1158,254,275]" italics="true" pageId="8" pageNumber="207">A. trigona</emphasis>
(
<bibRefCitation id="2C734B28FFB9FF4234AFFF77FAB0FEF4" author="Rathbun" box="[1171,1325,253,275]" pageId="8" pageNumber="207" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 1900 (20), 1 - 137, pls. 1, 2." type="journal article" year="1901">Rathbun, 1901</bibRefCitation>
)
</taxonomicName>
] 
<taxonomicName id="8FE24D5AFFB9FF4230ABFE96FE9BFED5" ID-CoL="84LB4" authorityName="Coutiere" authorityYear="1897" box="[151,262,284,306]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="7A96EACBFFB9FF4230ABFE96FE9BFED5" box="[151,262,284,306]" italics="true" pageId="8" pageNumber="207">Alpheopsis</emphasis>
</taxonomicName>
sp. 1— 
<treatmentCitation id="C94310C8FFB9FF42316EFE96FDB0FED5" author="Rodriguez" box="[338,557,284,306]" page="121" pageId="8" pageNumber="207" year="1986">
<bibRefCitation id="2C734B28FFB9FF42316EFE96FE66FED5" author="Rodriguez" box="[338,507,284,306]" pageId="8" pageNumber="207" refString="Rodriguez, B. (1986) Los camarones (Crustacea; Decapoda; Natantia) del Parque Nacional Archipielago de los Roques. Thesis. Licenciado en Biologia, Universidad Central de Venezuela, Caracas, 350 pp." type="book" year="1986">Rodriguez 1986</bibRefCitation>
: 121
</treatmentCitation>
, fig. 29.
</treatmentCitationGroup>
</paragraph>
<paragraph id="485D36D9FFB9FF4230ABFEB0FDE5FEB7" blockId="8.[151,1338,253,336]" box="[151,632,314,336]" pageId="8" pageNumber="207">
<treatmentCitationGroup id="68F211F7FFB9FF4230ABFEB0FDE5FEB7" box="[151,632,314,336]" pageId="8" pageNumber="207">
<taxonomicName id="8FE24D5AFFB9FF4230ABFEB0FE9BFEB7" ID-CoL="84LB4" authorityName="Coutiere" authorityYear="1897" box="[151,262,314,336]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="7A96EACBFFB9FF4230ABFEB0FE9BFEB7" box="[151,262,314,336]" italics="true" pageId="8" pageNumber="207">Alpheopsis</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="8FE24D5AFFB9FF423113FEB0FE31FEB7" ID-CoL="84W6C" authorityName="Rathbun" authorityYear="1901" box="[303,428,314,336]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="species" species="trigonus">
aff. 
<emphasis id="7A96EACBFFB9FF423164FEB1FE31FEB7" box="[344,428,315,336]" italics="true" pageId="8" pageNumber="207">trigonus</emphasis>
</taxonomicName>
— 
<treatmentCitation id="C94310C8FFB9FF4231F9FEB0FDE9FEB7" author="Anker" box="[453,628,314,336]" page="163" pageId="8" pageNumber="207" year="2001">
<bibRefCitation id="2C734B28FFB9FF4231F9FEB0FDDCFEB7" author="Anker" box="[453,577,314,336]" pageId="8" pageNumber="207" refString="Anker, A. (2001) Taxonomie et evolution des Alpheidae (Crustacea, Decapoda). Fol. 1 &amp; 2. PhD thesis, Museum National d'Histoire Naturelle, Paris, 547 pp. &amp; 331 pp." type="book" year="2001">Anker 2001</bibRefCitation>
: 163
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFB9FF4230ABFE16FB67FB75" pageId="8" pageNumber="207" type="materials_examined">
<paragraph id="485D36D9FFB9FF4230ABFE16FB5FFE31" blockId="8.[151,1437,412,1970]" pageId="8" pageNumber="207">
<materialsCitation id="F88A3C84FFB9FF4230ABFE16FB5FFE31" ID-GBIF-Occurrence="1563896398" collectingDate="2007-07-21" collectionCode="MZUSP" collectorName="A. Anker &amp; D. Roche" country="Panama" county="Drake" elevation="12" location="Caribbean" municipality="Portobelo" pageId="8" pageNumber="207" specimenCode="MZUSP 34222" specimenCount="2" typeStatus="holotype">
<emphasis id="7A96EACBFFB9FF4230ABFE16FEA2FE52" bold="true" box="[151,319,412,437]" pageId="8" pageNumber="207">
<typeStatus id="9759887BFFB9FF4230ABFE16FF4FFE52" box="[151,210,412,437]" pageId="8" pageNumber="207">Type</typeStatus>
material
</emphasis>
. 
<typeStatus id="9759887BFFB9FF423173FE16FE21FE52" box="[335,444,412,437]" pageId="8" pageNumber="207" type="holotype">Holotype</typeStatus>
: male (cl 
<quantity id="8F1A9B3CFFB9FF423211FE16FD14FE53" box="[557,649,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="207" unit="mm" value="4.5">4.5 mm</quantity>
), 
<specimenCode id="18449EA2FFB9FF4232A1FE16FCD1FE53" box="[669,844,412,436]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF4232A1FE16FD67FE53" box="[669,762,412,436]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="207">MZUSP</collectionCode>
34222
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF423365FE17FC28FE53" box="[857,949,413,436]" name="Panama" pageId="8" pageNumber="207">Panama</collectingCountry>
, 
<location id="4D3D6002FFB9FF4233FDFE16FBA9FE52" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF4233FDFE16FBA9FE52" box="[961,1076,412,437]" country="Panama" county="Drake" municipality="Portobelo" name="Caribbean" pageId="8" pageNumber="207">Caribbean</location>
coast, 
<collectingMunicipality id="A839ACA3FFB9FF4234BBFE16FB6AFE52" box="[1159,1271,412,437]" pageId="8" pageNumber="207">Portobelo</collectingMunicipality>
, 
<collectingCounty id="A13C4E55FFB9FF42353FFE16FAD7FE52" box="[1283,1354,412,437]" pageId="8" pageNumber="207">Drake</collectingCounty>
, 
<elevation id="C3CFD1EAFFB9FF42356BFE16FF32FE32" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="8" pageNumber="207" unit="m" value="11.5" valueMax="12.0" valueMin="11.0">
<quantity id="8F1A9B3CFFB9FF42356BFE16FF32FE32" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="8" pageNumber="207" unit="m" value="11.5" valueMax="12.0" valueMin="11.0">11–12 m</quantity>
</elevation>
, silt-covered reef, under coral rubble, coll. 
<collectorName id="E517530FFFB9FF4232AFFE34FD66FE31" box="[659,763,445,470]" pageId="8" pageNumber="207">A. Anker</collectorName>
&amp; 
<collectorName id="E517530FFFB9FF423323FE34FC16FE31" box="[799,907,445,470]" pageId="8" pageNumber="207">D. Roche</collectorName>
, 
<collectingDate id="2C18E9F1FFB9FF4233AAFE34FBBAFE31" box="[918,1063,445,470]" pageId="8" pageNumber="207" value="2007-07-21">
<date id="3C5C1019FFB9FF4233AAFE34FBBAFE31" box="[918,1063,445,470]" pageId="8" pageNumber="207" value="2007-07-21">21 July 2007</date>
</collectingDate>
[fcn 07-204].
</materialsCitation>
</paragraph>
<paragraph id="485D36D9FFB9FF4230FBFE54FB67FB75" blockId="8.[151,1437,412,1970]" pageId="8" pageNumber="207">
<materialsCitation id="F88A3C84FFB9FF4230FBFE54FC94FDFF" ID-GBIF-Occurrence="1563896399" collectingDate="1960-08-11" collectionCode="USNM" collectorName="B. A. Hazlett &amp; H. E. Winn" country="Bermuda" location="Bermuda" pageId="8" pageNumber="207" specimenCode="USNM 107089" specimenCount="1" specimenCount-male="1">
<emphasis id="7A96EACBFFB9FF4230FBFE54FDACFE10" bold="true" box="[199,561,478,503]" pageId="8" pageNumber="207">Additional material examined</emphasis>
: 
<specimenCount id="5EE4FD50FFB9FF42327EFE55FD12FE10" box="[578,655,478,503]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 5.0 mm), 
<specimenCode id="18449EA2FFB9FF42330CFE55FC7CFE10" box="[816,993,479,503]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF42330CFE55FC1CFE10" box="[816,897,479,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">USNM</collectionCode>
107089
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF4233D3FE54FBC5FE10" box="[1007,1112,478,503]" name="Bermuda" pageId="8" pageNumber="207">Bermuda</collectingCountry>
, under rocks along southern shore, coll. 
<collectorName id="E517530FFFB9FF423127FD8AFE35FDFF" box="[283,424,511,536]" pageId="8" pageNumber="207">B.A. Hazlett</collectorName>
&amp; 
<collectorName id="E517530FFFB9FF4231F1FD8AFDD5FDFF" box="[461,584,511,536]" pageId="8" pageNumber="207">H.E. Winn</collectorName>
, 
<collectingDate id="2C18E9F1FFB9FF42326FFD8AFC94FDFF" box="[595,777,511,536]" pageId="8" pageNumber="207" value="1960-08-11">
<date id="3C5C1019FFB9FF42326FFD8AFC94FDFF" box="[595,777,511,536]" pageId="8" pageNumber="207" value="1960-08-11">11 August 1960</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF42332AFD8AFE1DFDBC" ID-GBIF-Occurrence="1563896396" collectingDate="1958-04-28" collectionCode="USNM" collectorName="Smithsonian-Bredin Caribbean Expedition" country="Antigua &amp; Barbuda, Barbuda" location="Spanish Point" pageId="8" pageNumber="207" specimenCode="USNM 135846" specimenCount="1" specimenCount-male="1">
<specimenCount id="5EE4FD50FFB9FF42332AFD8AFCFCFDFF" box="[790,865,511,536]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 4.0 mm), 
<specimenCode id="18449EA2FFB9FF4233C0FD8AFB31FDFF" box="[1020,1196,511,536]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF4233C0FD8AFBD0FDFF" box="[1020,1101,512,536]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">USNM</collectionCode>
135846
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF42348BFE75FF6BFDDD" isEnumeration="true" pageId="8" pageNumber="207">Antigua &amp; Barbuda, Barbuda</collectingCountry>
Island, west and south of 
<location id="4D3D6002FFB9FF423216FDABFD57FDDD" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF423216FDABFD57FDDD" box="[554,714,545,570]" country="Antigua &amp; Barbuda, Barbuda" name="Spanish Point" pageId="8" pageNumber="207">Spanish Point</location>
, 
<collectorName id="E517530FFFB9FF4232EBFDABFB27FDDD" box="[727,1210,545,570]" pageId="8" pageNumber="207">Smithsonian-Bredin Caribbean Expedition</collectorName>
sta. 112A-58, from coral, 
<collectingDate id="2C18E9F1FFB9FF4230E2FDC9FE1DFDBC" box="[222,384,578,603]" pageId="8" pageNumber="207" value="1958-04-28">
<date id="3C5C1019FFB9FF4230E2FDC9FE1DFDBC" box="[222,384,578,603]" pageId="8" pageNumber="207" value="1958-04-28">28 April 1958</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF4231B1FDC9FB09FD9B" ID-GBIF-Occurrence="1563896406" collectingDate="2009-02-23" collectionCode="USNM" collectorName="Carrie Bow Cay &amp; D. L. Felder" country="Belize" latitude="16.80236" location="Belize" longLatPrecision="1" longitude="-88.08175" pageId="8" pageNumber="207" specimenCode="USNM 1292510" specimenCount="1" specimenCount-male="1">
<specimenCount id="5EE4FD50FFB9FF4231B1FDC9FE44FDBC" box="[397,473,578,603]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF42323BFDC9FDFFFDBC" box="[519,610,579,603]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="207" unit="mm" value="3.1">3.1 mm</quantity>
), 
<specimenCode id="18449EA2FFB9FF42324AFDC9FCA9FDBC" box="[630,820,578,603]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF42324AFDC9FD5AFDBC" box="[630,711,579,603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">USNM</collectionCode>
1292510
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF42337CFDC8FC17FDBC" box="[832,906,578,603]" name="Belize" pageId="8" pageNumber="207">Belize</collectingCountry>
, 
<collectorName id="E517530FFFB9FF4233AAFDC8FBD2FDBC" box="[918,1103,578,603]" pageId="8" pageNumber="207">Carrie Bow Cay</collectorName>
, 
<geoCoordinate id="2DD6501EFFB9FF423467FDC8FB71FDBC" box="[1115,1260,578,603]" direction="north" orientation="latitude" pageId="8" pageNumber="207" precision="1" value="16.80236">16°48’8.5”N</geoCoordinate>
<geoCoordinate id="2DD6501EFFB9FF4234C8FDC8FA0AFDBC" box="[1268,1431,578,603]" direction="west" orientation="longitude" pageId="8" pageNumber="207" precision="1" value="-88.08175">88°04’54.3”W</geoCoordinate>
, reef flat with 
<taxonomicName id="8FE24D5AFFB9FF42310AFDEFFE1AFD9B" box="[310,391,613,636]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="207" phylum="Cnidaria" rank="genus">
<emphasis id="7A96EACBFFB9FF42310AFDEFFE1AFD9B" box="[310,391,613,636]" italics="true" pageId="8" pageNumber="207">Porites</emphasis>
</taxonomicName>
, from broken-up conchs, coll. 
<collectorName id="E517530FFFB9FF4232D1FDEFFCEDFD9A" box="[749,880,612,637]" pageId="8" pageNumber="207">D.L. Felder</collectorName>
<emphasis id="7A96EACBFFB9FF423346FDEFFC28FD9B" box="[890,949,612,636]" italics="true" pageId="8" pageNumber="207">et al.</emphasis>
, 
<collectingDate id="2C18E9F1FFB9FF4233F9FDEEFB09FD9B" box="[965,1172,612,637]" pageId="8" pageNumber="207" value="2009-02-23">
<date id="3C5C1019FFB9FF4233F9FDEEFB09FD9B" box="[965,1172,612,637]" pageId="8" pageNumber="207" value="2009-02-23">23 February 2009</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF42349EFDEEFA1CFD79" ID-GBIF-Occurrence="1563896387" collectingDate="1976-05-02" collectionCode="USNM" collectorName="K. Ruetzler" country="Belize" elevation="2" latitude="16.8" location="Carrie Bow Cay" longLatPrecision="1281" longitude="-88.083336" pageId="8" pageNumber="207" specimenCode="USNM 1171216" specimenCount="1">
1 ovig. female (cl 
<quantity id="8F1A9B3CFFB9FF423545FDEEFF5AFD79" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="207" unit="mm" value="5.5">5.5 mm</quantity>
), 
<specimenCode id="18449EA2FFB9FF4230E7FD0CFE04FD79" box="[219,409,645,670]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF4230E7FD0CFEB1FD79" box="[219,300,646,670]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">USNM</collectionCode>
1171216
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF423199FD0FFE72FD79" box="[421,495,645,670]" name="Belize" pageId="8" pageNumber="207">Belize</collectingCountry>
, 
<location id="4D3D6002FFB9FF4231C0FD0FFD28FD79" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF4231C0FD0FFD28FD79" box="[508,693,645,670]" country="Belize" latitude="16.8" longLatPrecision="1281" longitude="-88.083336" name="Carrie Bow Cay" pageId="8" pageNumber="207">Carrie Bow Cay</location>
, 
<geoCoordinate id="2DD6501EFFB9FF4232FEFD0FFCBEFD79" box="[706,803,645,670]" direction="north" orientation="latitude" pageId="8" pageNumber="207" precision="925" value="16.8">16°48’N</geoCoordinate>
<geoCoordinate id="2DD6501EFFB9FF423310FD0FFC0FFD79" box="[812,914,645,670]" direction="west" orientation="longitude" pageId="8" pageNumber="207" precision="925" value="-88.083336">88°05’W</geoCoordinate>
, depth: 
<elevation id="C3CFD1EAFFB9FF4233D2FD0CFB81FD7A" box="[1006,1052,646,670]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="207" unit="m" value="2.0">
<quantity id="8F1A9B3CFFB9FF4233D2FD0CFB81FD7A" box="[1006,1052,646,670]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="207" unit="m" value="2.0">2 m</quantity>
</elevation>
, coll. 
<collectorName id="E517530FFFB9FF42345FFD0CFB75FD79" box="[1123,1256,645,670]" pageId="8" pageNumber="207">K. Ruetzler</collectorName>
, 
<collectingDate id="2C18E9F1FFB9FF4234C8FD0CFA1CFD79" box="[1268,1409,645,670]" pageId="8" pageNumber="207" value="1976-05-02">
<date id="3C5C1019FFB9FF4234C8FD0CFA1CFD79" box="[1268,1409,645,670]" pageId="8" pageNumber="207" value="1976-05-02">2 May 1976</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF4235B2FD0CFC16FD07" ID-GBIF-Occurrence="1563896386" collectingDate="2009-02-22" collectionCode="OUMNH" collectorName="Carrie Bow Cay &amp; D. L. Felder" country="Belize" latitude="16.8023" location="Belize" longLatPrecision="1" longitude="-88.0807" pageId="8" pageNumber="207" specimenCount="1" specimenCount-male="1">
<specimenCount id="5EE4FD50FFB9FF4235B2FD0CFF50FD58" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF4230C2FD2CFEC6FD58" box="[254,347,678,703]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="207" unit="mm" value="3.6">3.6 mm</quantity>
), 
<collectionCode id="2EF3AE1CFFB9FF42314DFD2DFE7CFD58" box="[369,481,679,703]" pageId="8" pageNumber="207">OUMNH</collectionCode>
.ZC. 2009.01.0 0 68, 
<collectingCountry id="30F57649FFB9FF4232FDFD2CFC96FD58" box="[705,779,678,703]" name="Belize" pageId="8" pageNumber="207">Belize</collectingCountry>
, 
<collectorName id="E517530FFFB9FF423326FD2CFC4BFD58" box="[794,982,678,703]" pageId="8" pageNumber="207">Carrie Bow Cay</collectorName>
, 
<geoCoordinate id="2DD6501EFFB9FF4233D8FD2CFBEAFD58" box="[996,1143,678,703]" direction="north" orientation="latitude" pageId="8" pageNumber="207" precision="1" value="16.8023">16°48.138’N</geoCoordinate>
<geoCoordinate id="2DD6501EFFB9FF4234BCFD2DFA84FD58" box="[1152,1305,679,703]" direction="west" orientation="longitude" pageId="8" pageNumber="207" precision="1" value="-88.0807">88°04.842’W</geoCoordinate>
, outer reef, intertidal coral rubble, coll. 
<collectorName id="E517530FFFB9FF4231E1FD42FDFFFD07" box="[477,610,711,736]" pageId="8" pageNumber="207">D.L. Felder</collectorName>
<emphasis id="7A96EACBFFB9FF423250FD43FD3AFD07" box="[620,679,712,736]" italics="true" pageId="8" pageNumber="207">et al.</emphasis>
, 
<collectingDate id="2C18E9F1FFB9FF423285FD42FC16FD07" box="[697,907,711,736]" pageId="8" pageNumber="207" value="2009-02-22">
<date id="3C5C1019FFB9FF423285FD42FC16FD07" box="[697,907,711,736]" pageId="8" pageNumber="207" value="2009-02-22">22 February 2009</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF4233A5FD42FE61FCC4" ID-GBIF-Occurrence="1563896394" collectingDate="2009-02-20" collectionCode="OUMNH" collectorName="D. L. Felder" country="Belize" elevation="8" location="Carrie Bow Cay" pageId="8" pageNumber="207" specimenCount="1" specimenCount-male="1">
<specimenCount id="5EE4FD50FFB9FF4233A5FD42FC7AFD07" box="[921,999,711,736]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF423426FD4DFBE8FD38" box="[1050,1141,711,736]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="207" unit="mm" value="5.4">5.4 mm</quantity>
, missing both chelipeds), 
<collectionCode id="2EF3AE1CFFB9FF4230ABFD60FE9AFCE5" box="[151,263,746,770]" pageId="8" pageNumber="207">OUMNH</collectionCode>
.ZC. 2009.01.0 0 63, 
<collectingCountry id="30F57649FFB9FF4231D8FD63FDB3FCE5" box="[484,558,745,770]" name="Belize" pageId="8" pageNumber="207">Belize</collectingCountry>
, 
<location id="4D3D6002FFB9FF423206FD63FD69FCE5" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF423206FD63FD69FCE5" box="[570,756,745,770]" country="Belize" name="Carrie Bow Cay" pageId="8" pageNumber="207">Carrie Bow Cay</location>
, northeastern shore, coral rubble, depth: 
<elevation id="C3CFD1EAFFB9FF4234FFFD63FABEFCE6" box="[1219,1315,745,770]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="207" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">
<quantity id="8F1A9B3CFFB9FF4234FFFD63FABEFCE6" box="[1219,1315,745,770]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="207" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</quantity>
</elevation>
, coll. 
<collectorName id="E517530FFFB9FF423555FD60FF42FCC4" pageId="8" pageNumber="207">D.L. Felder</collectorName>
<emphasis id="7A96EACBFFB9FF4230DAFC81FEBDFCC4" box="[230,288,779,803]" italics="true" pageId="8" pageNumber="207">et al.</emphasis>
, 
<collectingDate id="2C18E9F1FFB9FF423113FC81FE61FCC4" box="[303,508,778,803]" pageId="8" pageNumber="207" value="2009-02-20">
<date id="3C5C1019FFB9FF423113FC81FE61FCC4" box="[303,508,778,803]" pageId="8" pageNumber="207" value="2009-02-20">20 February 2009</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF423234FC81FA1EFCA3" ID-GBIF-Occurrence="1563896390" collectingDate="2007-04-04" collectionCode="ULLZ" collectorName="Carrie Bow Cay &amp; D. L. Felder" country="Belize" elevation="2" location="Belize" pageId="8" pageNumber="207" specimenCode="ULLZ 8939" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="5EE4FD50FFB9FF423234FC81FDC9FCC4" box="[520,596,778,803]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF4232BFFC80FD43FCC4" box="[643,734,778,803]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="207" unit="mm" value="4.5">4.5 mm</quantity>
), 
<specimenCount id="5EE4FD50FFB9FF4232CEFC81FCC9FCC4" box="[754,852,778,803]" pageId="8" pageNumber="207" type="female">1 female</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF4233BEFC81FC40FCC4" box="[898,989,779,803]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="207" unit="mm" value="4.4">4.4 mm</quantity>
), 
<specimenCode id="18449EA2FFB9FF4233CDFC81FBE1FCC4" box="[1009,1148,779,803]" collectionCode="ULLZ" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF4233CDFC81FBA4FCC5" box="[1009,1081,779,802]" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="8" pageNumber="207">ULLZ</collectionCode>
8939
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF4234B4FC80FB4FFCC4" box="[1160,1234,778,803]" name="Belize" pageId="8" pageNumber="207">Belize</collectingCountry>
, 
<collectorName id="E517530FFFB9FF4234E3FC80FA05FCC4" box="[1247,1432,778,803]" pageId="8" pageNumber="207">Carrie Bow Cay</collectorName>
, back reef, sand flat with sea grass, depth: less than 
<elevation id="C3CFD1EAFFB9FF4232ECFCA6FD60FCA3" box="[720,765,812,836]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="207" unit="m" value="2.0">
<quantity id="8F1A9B3CFFB9FF4232ECFCA6FD60FCA3" box="[720,765,812,836]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="207" unit="m" value="2.0">2 m</quantity>
</elevation>
, in dead conch shell, coll. 
<collectorName id="E517530FFFB9FF423418FCA7FB3BFCA2" box="[1060,1190,812,837]" pageId="8" pageNumber="207">D.L. Felder</collectorName>
<emphasis id="7A96EACBFFB9FF423490FCA7FB7EFCA3" box="[1196,1251,812,836]" italics="true" pageId="8" pageNumber="207">et al.</emphasis>
, 
<collectingDate id="2C18E9F1FFB9FF4234CDFCA6FA1EFCA3" box="[1265,1411,812,837]" pageId="8" pageNumber="207" value="2007-04-04">
<date id="3C5C1019FFB9FF4234CDFCA6FA1EFCA3" box="[1265,1411,812,837]" pageId="8" pageNumber="207" value="2007-04-04">4 April 2007</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF4235B2FCA6FB6CFC60" ID-GBIF-Occurrence="1563896395" collectingDate="1960-04-06" collectorName="Espiritu Santo &amp; Smithsonian-Bredin Caribbean Expedition" country="Mexico" elevation="2" location="Bahia" pageId="8" pageNumber="207" specimenCount="1" specimenCount-male="1" stateProvince="Quintana Roo">
<specimenCount id="5EE4FD50FFB9FF4235B2FCA6FF50FC81" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 4.0 mm, missing both chelipeds), 
<collectingCountry id="30F57649FFB9FF4232BDFCC7FD44FC81" box="[641,729,845,870]" name="Mexico" pageId="8" pageNumber="207">Mexico</collectingCountry>
, 
<collectingRegion id="8A26F83BFFB9FF4232DBFCC7FC1AFC82" box="[743,903,845,870]" country="Mexico" name="Quintana Roo" pageId="8" pageNumber="207">Quintana Roo</collectingRegion>
, 
<location id="4D3D6002FFB9FF4233A8FCC7FC48FC81" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF4233A8FCC7FC48FC81" box="[916,981,845,870]" country="Mexico" name="Bahia" pageId="8" pageNumber="207" stateProvince="Quintana Roo">Bahía</location>
del 
<collectorName id="E517530FFFB9FF423436FCC7FB30FC81" box="[1034,1197,845,870]" pageId="8" pageNumber="207">Espíritu Santo</collectorName>
, 
<collectorName id="E517530FFFB9FF423486FCC7FE10FC60" pageId="8" pageNumber="207">Smithsonian-Bredin Caribbean Expedition</collectorName>
sta. 41-60, west side of reef, east of anchorage, depth: 
<elevation id="C3CFD1EAFFB9FF423439FCE5FBD2FC61" box="[1029,1103,879,903]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="207" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">
<quantity id="8F1A9B3CFFB9FF423439FCE5FBD2FC61" box="[1029,1103,879,903]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="207" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
</elevation>
, 
<collectingDate id="2C18E9F1FFB9FF423460FCE4FB6CFC60" box="[1116,1265,878,903]" pageId="8" pageNumber="207" value="1960-04-06">
<date id="3C5C1019FFB9FF423460FCE4FB6CFC60" box="[1116,1265,878,903]" pageId="8" pageNumber="207" value="1960-04-06">6 April 1960</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF4234C2FCE5FB96FC2D" ID-GBIF-Occurrence="1563896393" collectingDate="1969-03-24" collectionCode="USNM" collectorName="C. Mahnken" country="United States of America" county="Virgin Islands" elevation="20" location="Lameshur Bay" municipality="St. John Island" pageId="8" pageNumber="207" specimenCode="USNM 134762" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="5EE4FD50FFB9FF4234C2FCE5FAD7FC60" box="[1278,1354,878,903]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 6.0 mm), 
<specimenCount id="5EE4FD50FFB9FF4230E5FC1AFEA7FC4F" box="[217,314,911,936]" pageId="8" pageNumber="207" type="female">1 female</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF423159FC05FE23FC4F" box="[357,446,911,936]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="207" unit="mm" value="6.2">6.2 mm</quantity>
), 
<specimenCode id="18449EA2FFB9FF4231EDFC1AFDE2FC4F" box="[465,639,911,936]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF4231EDFC1AFDBFFC4F" box="[465,546,912,936]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="207">USNM</collectionCode>
134762
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF4232B6FC1AFD33FC4F" box="[650,686,912,936]" name="United States of America" pageId="8" pageNumber="207">US</collectingCountry>
<collectingCounty id="A13C4E55FFB9FF423288FC05FCCEFC4F" box="[692,851,911,936]" pageId="8" pageNumber="207">Virgin Islands</collectingCounty>
, 
<collectingMunicipality id="A839ACA3FFB9FF423362FC1AFB98FC4F" box="[862,1029,911,936]" pageId="8" pageNumber="207">St. John Island</collectingMunicipality>
, 
<location id="4D3D6002FFB9FF42342CFC05FB28FC4F" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF42342CFC05FB28FC4F" box="[1040,1205,911,936]" country="United States of America" county="Virgin Islands" municipality="St. John Island" name="Lameshur Bay" pageId="8" pageNumber="207">Lameshur Bay</location>
, canyon base, depth: 
<elevation id="C3CFD1EAFFB9FF4230ABFC38FF4FFC2E" box="[151,210,946,970]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="207" unit="m" value="20.0">
<quantity id="8F1A9B3CFFB9FF4230ABFC38FF4FFC2E" box="[151,210,946,970]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="207" unit="m" value="20.0">20 m</quantity>
</elevation>
, scuba diving, from empty conch shell, coll. 
<collectorName id="E517530FFFB9FF4232F6FC38FCC8FC2D" box="[714,853,945,970]" pageId="8" pageNumber="207">C. Mahnken</collectorName>
<collectingDate id="2C18E9F1FFB9FF423360FC38FB96FC2D" box="[860,1035,945,970]" pageId="8" pageNumber="207" value="1969-03-24">
<date id="3C5C1019FFB9FF423360FC38FB96FC2D" box="[860,1035,945,970]" pageId="8" pageNumber="207" value="1969-03-24">24 March 1969</date>
</collectingDate>
</materialsCitation>
; 1 male (cl 
<quantity id="8F1A9B3CFFB9FF4234B2FC3BFB75FC2D" box="[1166,1256,945,970]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="207" unit="mm" value="3.5">3.5 mm</quantity>
), 1 ovig. female (cl 
<quantity id="8F1A9B3CFFB9FF423080FC59FE8AFC0C" box="[188,279,979,1003]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="207" unit="mm" value="4.4">4.4 mm</quantity>
), USNM 134763, same collection data; 
<materialsCitation id="F88A3C84FFB9FF4232E7FC59FBA9FBEB" ID-GBIF-Occurrence="1563896401" collectorName="French Antilles &amp; Grand Cul de Sac &amp; F. Fasquel" country="Guadeloupe" location="Guadeloupe" pageId="8" pageNumber="207" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="5EE4FD50FFB9FF4232E7FC59FCAFFC0C" box="[731,818,978,1003]" pageId="8" pageNumber="207" type="male">2 males</specimenCount>
(cl 3.5, 
<quantity id="8F1A9B3CFFB9FF4233ADFC58FC71FC0C" box="[913,1004,978,1003]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="207" unit="mm" value="5.1">5.1 mm</quantity>
), 
<specimenCount id="5EE4FD50FFB9FF4233C3FC59FBFCFC0C" box="[1023,1121,978,1003]" pageId="8" pageNumber="207" type="female">1 female</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF4234B2FC59FB75FC0C" box="[1166,1256,979,1003]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="207" unit="mm" value="3.3">3.3 mm</quantity>
), MNHN-2014- IU-19473, 
<collectorName id="E517530FFFB9FF42312FFC7EFE59FBEA" box="[275,452,1012,1037]" pageId="8" pageNumber="207">French Antilles</collectorName>
, 
<collectingCountry id="30F57649FFB9FF4231F3FC7EFDC4FBEA" box="[463,601,1012,1037]" name="Guadeloupe" pageId="8" pageNumber="207">Guadeloupe</collectingCountry>
, 
<collectorName id="E517530FFFB9FF42325AFC7EFCACFBEB" box="[614,817,1012,1037]" pageId="8" pageNumber="207">Grand Cul de Sac</collectorName>
, coll. 
<collectorName id="E517530FFFB9FF42334AFC7FFC71FBEA" box="[886,1004,1012,1037]" pageId="8" pageNumber="207">F. Fasquel</collectorName>
, 2000
</materialsCitation>
; 
<materialsCitation id="F88A3C84FFB9FF42347CFC7EFD50FBA8" ID-GBIF-Occurrence="1563896397" collectingDate="2012-05-12" collectionCode="MNHN" collectorName="Baie de Baille-Argent" country="Guadeloupe" elevation="35" latitude="16.259" location="Baie de Baille-Argent" longLatPrecision="13" longitude="-61.811832" municipality="French Antilles" pageId="8" pageNumber="207" specimenCount="1" specimenCount-female="1">
<specimenCount id="5EE4FD50FFB9FF42347CFC7EFB3CFBEA" box="[1088,1185,1012,1037]" pageId="8" pageNumber="207" type="female">1 female</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF4234F3FC7EFAB7FBEB" box="[1231,1322,1012,1036]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="207" unit="mm" value="4.4">4.4 mm</quantity>
), MNHN- IU-2013-12143, 
<collectingMunicipality id="A839ACA3FFB9FF423161FB9FFD84FBC9" box="[349,537,1045,1070]" pageId="8" pageNumber="207">French Antilles</collectingMunicipality>
, KARUBENTHOS 2012, sta. GD20, 
<collectorName id="E517530FFFB9FF4233C9FB9FFA9EFBC9" box="[1013,1283,1045,1070]" pageId="8" pageNumber="207">
<location id="4D3D6002FFB9FF4233C9FB9FFA9EFBC9" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFB9FF4233C9FB9FFA9EFBC9" box="[1013,1283,1045,1070]" country="Guadeloupe" latitude="16.259" longLatPrecision="13" longitude="-61.811832" municipality="French Antilles" name="Baie de Baille-Argent" pageId="8" pageNumber="207">Baie de Baille-Argent</location>
</collectorName>
, 
<geoCoordinate id="2DD6501EFFB9FF423524FB9FFA01FBC9" box="[1304,1436,1045,1070]" direction="north" orientation="latitude" pageId="8" pageNumber="207" precision="9" value="16.259">16°15.54’N</geoCoordinate>
<geoCoordinate id="2DD6501EFFB9FF4230ABFBBCFEBDFBA8" box="[151,288,1078,1103]" direction="west" orientation="longitude" pageId="8" pageNumber="207" precision="9" value="-61.811832">61°48.71’W</geoCoordinate>
, 
<elevation id="C3CFD1EAFFB9FF423113FBBCFEF0FBA9" box="[303,365,1078,1103]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="207" unit="m" value="35.0">
<quantity id="8F1A9B3CFFB9FF423113FBBCFEF0FBA9" box="[303,365,1078,1103]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="207" unit="m" value="35.0">35 m</quantity>
</elevation>
, coll. 
<collectionCode id="2EF3AE1CFFB9FF42318AFBBDFD91FBA9" box="[438,524,1079,1102]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="207">MNHN</collectionCode>
team, 
<collectingDate id="2C18E9F1FFB9FF423260FBBDFD50FBA8" box="[604,717,1079,1103]" pageId="8" pageNumber="207" value="2012-05-12">
<date id="3C5C1019FFB9FF423260FBBDFD50FBA8" box="[604,717,1079,1103]" pageId="8" pageNumber="207" value="2012-05-12">12.v.2012</date>
</collectingDate>
</materialsCitation>
; 1 male (cl 
<quantity id="8F1A9B3CFFB9FF423366FBBCFC2AFBA8" box="[858,951,1078,1103]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="207" unit="mm" value="3.5">3.5 mm</quantity>
), MNHN-IU-2013-12150, same collection data; 
<materialsCitation id="F88A3C84FFB9FF4230EAFBD2FB67FB75" ID-GBIF-Occurrence="1563896385" collectingDate="1997-01-31" collectionCode="MZUSP" country="Brazil" location="Paraiba" pageId="8" pageNumber="207" specimenCode="MZUSP 18179" specimenCount="1" specimenCount-male="1" stateProvince="Paraiba">
<specimenCount id="5EE4FD50FFB9FF4230EAFBD2FEBEFB97" box="[214,291,1111,1136]" pageId="8" pageNumber="207" type="male">1 male</specimenCount>
(cl 
<quantity id="8F1A9B3CFFB9FF42316FFBD2FE30FB88" box="[339,429,1112,1136]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="207" unit="mm" value="2.9">2.9 mm</quantity>
, missing both chelipeds), 
<specimenCode id="18449EA2FFB9FF4232E5FBD2FC15FB97" box="[729,904,1112,1136]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="207">
<collectionCode id="2EF3AE1CFFB9FF4232E5FBD2FCABFB97" box="[729,822,1112,1136]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="207">MZUSP</collectionCode>
18179
</specimenCode>
, 
<collectingCountry id="30F57649FFB9FF4233A9FBDDFC41FB97" box="[917,988,1111,1136]" name="Brazil" pageId="8" pageNumber="207">Brazil</collectingCountry>
, continental shelf off southern 
<collectingRegion id="8A26F83BFFB9FF42357DFBDDFA05FB97" box="[1345,1432,1111,1136]" country="Brazil" name="Paraiba" pageId="8" pageNumber="207">Paraíba</collectingRegion>
, REVIZEE—Score NE II, sta. 32-2, sand-mud, 
<collectingDate id="2C18E9F1FFB9FF42329AFBF0FCC2FB75" box="[678,863,1146,1170]" pageId="8" pageNumber="207" value="1997-01-31">
<date id="3C5C1019FFB9FF42329AFBF0FCC2FB75" box="[678,863,1146,1170]" pageId="8" pageNumber="207" value="1997-01-31">31 January 1997</date>
</collectingDate>
(tentative identification, see below).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFB9FF4530FBFB10FB90FE13" lastPageId="15" lastPageNumber="214" pageId="8" pageNumber="207" type="description">
<paragraph id="485D36D9FFB9FF4230FBFB10FE43FA38" blockId="8.[151,1437,412,1970]" pageId="8" pageNumber="207">
<emphasis id="7A96EACBFFB9FF4230FBFB10FECCFB54" bold="true" box="[199,337,1178,1203]" pageId="8" pageNumber="207">Description</emphasis>
. Medium-sized species of 
<taxonomicName id="8FE24D5AFFB9FF423242FB11FD64FB54" authorityName="Coutiere" authorityYear="1897" box="[638,761,1179,1203]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="7A96EACBFFB9FF423242FB11FD64FB54" box="[638,761,1179,1203]" italics="true" pageId="8" pageNumber="207">Alpheopsis</emphasis>
</taxonomicName>
(present material: maximum cl 
<quantity id="8F1A9B3CFFB9FF423463FB10FB24FB54" box="[1119,1209,1178,1203]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="207" unit="mm" value="6.2">6.2 mm</quantity>
). Carapace with one strong MDC extending from rostral tip to almost posterior margin of carapace; each side of carapace with one strong ASC running in parallel and lateral to MDC, starting at almost frontal margin of carapace and abruptly ending at about 0.7 length of carapace, sometimes forming short high crest above eye; one strong PSC running in parallel and lateral to MDC, starting posterior and sometimes slightly ventral to ASC, not extending ventrolaterally at anterior end; one very low, short, sometimes barely noticeable or obsolete VLC, running slightly obliquely from postorbital-hepatic area to about 0.4 carapace length; and one strong ABC running along and somewhat above branchiostegial margin to about 0.1–0.2 carapace length, with stout subacute tooth at anterior end, latter protruding well beyond anterolateral margin, occasionally ABC reduced to short ridge posterior to strong anterior tooth (
<figureCitation id="D0D92A5CFFB9FF423558FA2FFF39FA38" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="8" pageNumber="207">Figs. 5</figureCitation>
a–d; 7a–d, m–o; 8a–c, o, p).
</paragraph>
<paragraph id="485D36D9FFB9FF4230FBFA62FB0BF940" blockId="8.[151,1437,412,1970]" pageId="8" pageNumber="207">
Rostrum well developed, about 1.3–1.6 times as long as wide at base, with strong carina, subacute distally, tip reaching or slightly overreaching distal margin of first article of antennular peduncle, reaching at most to 0.3 length of second article (
<figureCitation id="D0D92A5CFFB9FF42315BF9A0FE25F9A4" box="[359,440,1578,1603]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="8" pageNumber="207">Figs. 5</figureCitation>
c, d, 7c, d, n, o; 8b, c, o). Orbital hoods with distinct orbital teeth, latter ranging from broadly subtriangular and obtuse lobes to triangular and strongly projecting, distally acute teeth; tips of orbital teeth reaching to level just beyond rostral base to about 0.3 of rostral length (
<figureCitation id="D0D92A5CFFB9FF42343FF9E7FBC5F961" box="[1027,1112,1645,1670]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="8" pageNumber="207">Figs. 5</figureCitation>
c, d, 7c, d, n, o; 8b, c, o). Pterygostomial angle below strong ABC tooth broadly rounded (
<figureCitation id="D0D92A5CFFB9FF423350F904FC24F940" box="[876,953,1678,1703]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="8" pageNumber="207">Figs. 5</figureCitation>
a, c; 7c, n; 8a, c, p).
</paragraph>
<paragraph id="485D36D9FFB9FF4230FBF925FE13F855" blockId="8.[151,1437,412,1970]" pageId="8" pageNumber="207">
First and second pleomeres with well-marked, high MDC-PL1 and low MDC-PL2, continuing MDC of carapace; third to sixth pleomeres unarmed dorsally; all pleura rounded ventrally; fifth pleuron with posteroventral angle rounded; sixth somite with bluntly projecting posterior lobe and well-delimited, triangular, articulated plate (
<figureCitation id="D0D92A5CFFB9FF4230A3F89EFF73F8CA" box="[159,238,1812,1837]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="8" pageNumber="207">Figs. 5</figureCitation>
a, b, e; 7a, b). Telson moderately broad, tapering distally, about 2.3–2.5 as long as anterior width; dorsal surface with shallow median longitudinal depression and two pairs of stout spiniform setae inserted far from lateral margin, at about 0.5 and 0.75 telson length, respectively; posterior margin slightly less than half as long as anterior margin, broadly rounded; each posterolateral angle with one pair of stout spiniform setae, mesial much longer than lateral (
<figureCitation id="D0D92A5CFFB9FF4230D0F813FEB3F855" box="[236,302,1945,1970]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="8" pageNumber="207">Fig. 5</figureCitation>
f; 8d, e).
</paragraph>
<caption id="1C9D6651FFB8FF4330ABF967FF41F899" httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="9" pageNumber="208" targetBox="[161,1424,202,1741]" targetPageId="9">
<paragraph id="485D36D9FFB8FF4330ABF967FF41F899" blockId="9.[151,1436,1773,1918]" pageId="9" pageNumber="208">
<emphasis id="7A96EACBFFB8FF4330ABF967FE8FF8E4" bold="true" box="[151,274,1773,1795]" pageId="9" pageNumber="208">FIGURE 5.</emphasis>
<taxonomicName id="8FE24D5AFFB8FF433125F967FD94F8E4" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[281,521,1773,1795]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="208" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFB8FF433125F967FD94F8E4" box="[281,521,1773,1795]" italics="true" pageId="9" pageNumber="208">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFB8FF433232F964FDC2F8E4" box="[526,607,1774,1795]" pageId="9" pageNumber="208" rank="species">
<emphasis id="7A96EACBFFB8FF433232F964FDC2F8E4" bold="true" box="[526,607,1774,1795]" pageId="9" pageNumber="208">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal.
</paragraph>
</caption>
<caption id="1C9D6651FFBBFF4030ABFADAFE6DFA44" httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="10" pageNumber="209" targetBox="[158,1418,204,1325]" targetPageId="10">
<paragraph id="485D36D9FFBBFF4030ABFADAFE6DFA44" blockId="10.[151,1436,1360,1443]" pageId="10" pageNumber="209">
<emphasis id="7A96EACBFFBBFF4030ABFADAFE8FFA82" bold="true" box="[151,274,1360,1382]" pageId="10" pageNumber="209">FIGURE 6.</emphasis>
<taxonomicName id="8FE24D5AFFBBFF403125FADAFD94FA81" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[281,521,1360,1382]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="209" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBBFF403125FADAFD94FA81" box="[281,521,1360,1382]" italics="true" pageId="10" pageNumber="209">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBBFF403232FADBFDC2FA81" box="[526,607,1361,1382]" pageId="10" pageNumber="209" rank="species">
<emphasis id="7A96EACBFFBBFF403232FADBFDC2FA81" bold="true" box="[526,607,1361,1382]" pageId="10" pageNumber="209">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral.
</paragraph>
</caption>
<paragraph id="485D36D9FFBBFF4030FBFA5BFBB5F9EB" blockId="10.[151,1437,1488,2017]" pageId="10" pageNumber="209">
Eyes completely concealed by orbital hoods, not visible in dorsal and lateral views; eyestalk with wellpigmented cornea (
<figureCitation id="D0D92A5CFFBBFF403152FA7EFE2CF9EA" box="[366,433,1524,1549]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="10" pageNumber="209">Fig. 5</figureCitation>
c, d). Epistomial sclerite each with strong sharp process.
</paragraph>
<paragraph id="485D36D9FFBBFF4030FBF992FAE1F8EE" blockId="10.[151,1437,1488,2017]" pageId="10" pageNumber="209">
Antennular peduncles stout; stylocerite strong, with sharp tip, latter overreaching distal margin of second article; ventromesial carina with anteriorly directed, acute tooth; second article 1.2–1.3 times as long as wide; lateral flagellum biramous; fused portion short, with three or so joints; accessory ramus long, with several groups of long aesthetascs (
<figureCitation id="D0D92A5CFFBBFF4031B0F90EFE7DF97A" box="[396,480,1668,1693]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="10" pageNumber="209">Figs. 5</figureCitation>
c, d, g; 7c, d, n, o; 8b, c, o, p). Antenna with basicerite ending in sharp tooth distoventrally; scaphocerite ovate, with anteriorly rounded blade and sharp distolateral tooth, latter not reaching beyond anterior margin of blade; anterior margin of scaphocerite not exceeding distal margin of antennular peduncles; carpocerite reaching or slightly overreaching end of scaphocerite (
<figureCitation id="D0D92A5CFFBBFF4033C0F97AFBD7F8EE" box="[1020,1098,1776,1801]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="10" pageNumber="209">Figs. 5</figureCitation>
c, d; 7c, d, n, o; 8b, c, o, p).
</paragraph>
<paragraph id="485D36D9FFBBFF4030FBF89EFEDAF87E" blockId="10.[151,1437,1488,2017]" pageId="10" pageNumber="209">
Mouthparts typical for genus in external view. Third maxilliped slender, pediform; coxa with distally subacute lateral plate above mastigobranch; penultimate article about 2.5 times as long as wide proximally; ultimate article tapering to slender corneous tip, with one stiff subdistal spiniform seta mesiodorsally; arthrobranch well developed (
<figureCitation id="D0D92A5CFFBBFF4030A3F80AFF71F87E" box="[159,236,1920,1945]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="10" pageNumber="209">Figs. 5</figureCitation>
h, i; 8f).
</paragraph>
<paragraph id="485D36D9FFBBFF4430FBF82EFE97FEFA" blockId="10.[151,1437,1488,2017]" lastBlockId="14.[151,1437,151,717]" lastPageId="14" lastPageNumber="213" pageId="10" pageNumber="209">
Chelipeds subequal or somewhat unequal in size, subsymmetrical or slightly asymmetrical in shape and armature on finger cutting edges, carried extended when not in use (
<figureCitation id="D0D92A5CFFBBFF4033B1F842FC44F806" box="[909,985,1992,2017]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="10" pageNumber="209">Figs. 6</figureCitation>
a–c; 7e–g, k, l; 
<quantity id="8F1A9B3CFFBBFF4034B8F842FB3DF806" box="[1156,1184,1992,2017]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="10" pageNumber="209" unit="g" value="8.0">8g</quantity>
–i, k, l). Ischium stout, short, with stout spiniform seta distodorsally (
<figureCitation id="D0D92A5CFFBCFF4732A0F978FD77F8EC" box="[668,746,1778,1803]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="13" pageNumber="212">Figs. 6</figureCitation>
a; 8e). Merus stout, usually somewhat longer as wide in adult males, slenderer, noticeably longer than wide in females, broadening distally, distinctly triangular in cross-section, with three well-defined margins, dorsal margin ending in very conspicuous blunt tooth, proximodorsal margin usually with stout spiniform seta (
<figureCitation id="D0D92A5CFFBCFF47321AF8D4FDEAF890" box="[550,631,1886,1911]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="13" pageNumber="212">Figs. 6</figureCitation>
a, b; 7e; 
<quantity id="8F1A9B3CFFBCFF4732DEF8D5FC9DF890" box="[738,768,1887,1911]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="13" pageNumber="212" unit="g" value="8.0">8g</quantity>
). Carpus very stout, short, cup-shaped, widening distally, usually with strong dorsal constriction, ventral margin typically with one stiff seta (
<figureCitation id="D0D92A5CFFBCFF473469F808FB39F87C" box="[1109,1188,1922,1947]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="13" pageNumber="212">Figs. 6</figureCitation>
a–c; 7e, k; 
<quantity id="8F1A9B3CFFBCFF473519F808FAD9F87D" box="[1317,1348,1922,1946]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="13" pageNumber="212" unit="g" value="8.0">8g</quantity>
). Chela moderately robust, with palm somewhat flattened on mesial surface and fingers about 0.8–0.9 length of palm; dorsomesial-proximal surface of palm without process; cutting edges of fingers unarmed (male minor cheliped, some female chelipeds) or armed with three to five moderately developed, rounded or subtriangular, teeth (male and female major cheliped); fingers somewhat gaping, crossing distally, usually with row of short setae parallel to cutting edge; dactylus of minor cheliped not noticeably flattened; adhesive disks not distinct (
<figureCitation id="D0D92A5CFFBFFF443491FF55FB67FF1F" box="[1197,1274,223,248]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="14" pageNumber="213">Figs. 6</figureCitation>
a–c; 7f, g, k, l; 
<quantity id="8F1A9B3CFFBFFF4430ABFE8EFF2EFEFA" box="[151,179,260,285]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="14" pageNumber="213" unit="g" value="8.0">8g</quantity>
–i, k, l).
</paragraph>
<caption id="1C9D6651FFBAFF4130ABF955FB33F84A" httpUri="https://zenodo.org/record/809648/files/figure.png" pageId="11" pageNumber="210" targetBox="[163,1414,206,1724]" targetPageId="11">
<paragraph id="485D36D9FFBAFF4130ABF955FB33F84A" blockId="11.[151,1436,1759,1965]" pageId="11" pageNumber="210">
<emphasis id="7A96EACBFFBAFF4130ABF955FE92F912" bold="true" box="[151,271,1759,1781]" pageId="11" pageNumber="210">FIGURE 7</emphasis>
. 
<taxonomicName id="8FE24D5AFFBAFF413122F955FD8DF911" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[286,528,1759,1782]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="210" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBAFF413122F955FD8DF911" box="[286,528,1759,1782]" italics="true" pageId="11" pageNumber="210">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBAFF413225F96AFDF6F912" box="[537,619,1760,1781]" pageId="11" pageNumber="210" rank="species">
<emphasis id="7A96EACBFFBAFF413225F96AFDF6F912" bold="true" box="[537,619,1760,1781]" pageId="11" pageNumber="210">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (cl 4.0 mm) from Barbuda, Antigua &amp; Barbuda (USNM 135846) [a–i]; ovig. female (cl 4.4 mm) from St. John Island, US Virgin Islands (USNM 134763) [j–l]; male (cl 5.0 mm) from Bermuda (USNM 107089) [m–o]; a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—left cheliped, mesial; f—same, chela, mesial; g—same, chela fingers (open), lateral; h—second pereiopod, lateral; i—third pereiopod, lateral; j—anterior region of carapace, lateral; k—left cheliped, mesial; lsame, lateral; m—carapace and anterior pleon, lateral; n—anterior region of carapace and frontal appendages, lateral; o—same, dorsal. Modified from Anker (2001), except for j, k, l (drawings prepared, but not used in Anker 2001).
</paragraph>
</caption>
<caption id="1C9D6651FFBDFF4630ABF973FA90F84F" httpUri="https://zenodo.org/record/809650/files/figure.png" pageId="12" pageNumber="211" targetBox="[175,1424,205,1748]" targetPageId="12">
<paragraph id="485D36D9FFBDFF4630ABF973FA90F84F" blockId="12.[151,1436,1785,1960]" pageId="12" pageNumber="211">
<emphasis id="7A96EACBFFBDFF4630ABF973FF67F8E8" bold="true" box="[151,250,1785,1807]" pageId="12" pageNumber="211">FIGURE</emphasis>
8. 
<taxonomicName id="8FE24D5AFFBDFF463126F973FD96F8E8" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[282,523,1785,1807]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="211" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBDFF463126F973FD96F8E8" box="[282,523,1785,1807]" italics="true" pageId="12" pageNumber="211">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBDFF46322DF973FDFFF8E9" box="[529,610,1785,1806]" pageId="12" pageNumber="211" rank="species">
<emphasis id="7A96EACBFFBDFF46322DF973FDFFF8E9" bold="true" box="[529,610,1785,1806]" pageId="12" pageNumber="211">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (cl 5.1 mm) [a–n] and female (cl 3.3 mm) [o, p] from Guadeloupe, French Antilles (MNHN-2014-IU-19473); a—carapace, lateral; b—anterior region of carapace and frontal appendages, dorsal; csame, lateral; d—telson, dorsal; e—same, detail of posterolateral spiniform setae, dorsal; f—third maxilliped, lateral; g—left (major) cheliped, lateral; h—same, chela, mesial; i—same, chela fingers, lateral; j—right (minor) cheliped, mesial; k, same, chela, mesial; l—same, mesial; m—third pereiopod, lateral; n—uropod, dorsal; o—anterior region of carapace and frontal appendages, dorsal; p—same, lateral. Drawings prepared, but not used in Anker (2001); (e) drawn without scale.
</paragraph>
</caption>
<caption id="1C9D6651FFBCFF4730ABF9FDFF26F92D" httpUri="https://zenodo.org/record/809652/files/figure.png" pageId="13" pageNumber="212" targetBox="[150,1436,193,1634]" targetPageId="13">
<paragraph id="485D36D9FFBCFF4730ABF9FDFF26F92D" blockId="13.[151,1436,1655,1738]" pageId="13" pageNumber="212">
<emphasis id="7A96EACBFFBCFF4730ABF9FDFE8FF96B" bold="true" box="[151,274,1655,1677]" pageId="13" pageNumber="212">FIGURE 9.</emphasis>
<taxonomicName id="8FE24D5AFFBCFF473125F9FDFD94F96A" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[281,521,1655,1677]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="212" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBCFF473125F9FDFD94F96A" box="[281,521,1655,1677]" italics="true" pageId="13" pageNumber="212">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBCFF473232F9FDFDC2F96B" box="[526,607,1655,1676]" pageId="13" pageNumber="212" rank="species">
<emphasis id="7A96EACBFFBCFF473232F9FDFDC2F96B" bold="true" box="[526,607,1655,1676]" pageId="13" pageNumber="212">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222) [a]; male (cl 4.5 mm) from Carrie Bow Cay, Belize (ULLZ 8939) [b]. Photographs: A. Anker (a) and Darryl L. Felder (b).
</paragraph>
</caption>
<paragraph id="485D36D9FFBFFF4430FBFEADFDF2FD2A" blockId="14.[151,1437,151,717]" pageId="14" pageNumber="213">
Second pereiopod relatively slender; ischium subequal or slightly longer than merus; carpus with five joints; ratio of carpal joints (from proximal to distal) approximately equal to: 3.5: 1: 1: 1: 1.8; chela longer than distal carpal article, with fingers equal to palm in length, simple (
<figureCitation id="D0D92A5CFFBFFF44336CFEE5FC3FFE6F" box="[848,930,367,392]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="14" pageNumber="213">Figs. 6</figureCitation>
d; 7h). Third pereiopod moderately slender; ischium with one spiniform setae on ventrolateral surface; merus about 5.5 times as long as wide, unarmed; carpus about 0.5 length of merus, much slenderer, with slender spiniform seta on distoventral margin; propodus with three spiniform setae along ventral margin and pair of distal spiniform setae adjacent to dactylus; dactylus about 0.4–0.5 length of propodus, slender, simple, conical, gradually curving distally (
<figureCitation id="D0D92A5CFFBFFF443385FE75FB9AFDFF" box="[953,1031,511,536]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="14" pageNumber="213">Figs. 6</figureCitation>
e; 
<date id="3C5C1019FFBFFF44341DFE75FBA7FDFF" box="[1057,1082,511,536]" pageId="14" pageNumber="213">7i</date>
). Fourth pereiopod very similar to third pereiopod. Fifth pereiopod more slender than third and fourth pereiopods; ischium unarmed on ventrolateral surface; merus about 5.7 times as long as wide, unarmed; carpus about 0.8 length of merus; propodus long, slender, with four or five spiniform setae along ventral margin, one long distal spiniform setae adjacent to dactylus, and several rows of short serrulate setae on distolateral surface; dactylus about 0.4 length of propodus, similar to that of third pereiopod (
<figureCitation id="D0D92A5CFFBFFF44322FFD3EFDCBFD2A" box="[531,598,692,717]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="14" pageNumber="213">Fig. 6</figureCitation>
f).
</paragraph>
<caption id="1C9D6651FFBFFF4430ABF809FD73F831" httpUri="https://zenodo.org/record/809654/files/figure.png" pageId="14" pageNumber="213" targetBox="[150,1436,769,1901]" targetPageId="14">
<paragraph id="485D36D9FFBFFF4430ABF809FD73F831" blockId="14.[151,1435,1923,2006]" pageId="14" pageNumber="213">
<emphasis id="7A96EACBFFBFFF4430ABF809FEBDF87F" bold="true" box="[151,288,1923,1945]" pageId="14" pageNumber="213">FIGURE 10.</emphasis>
<taxonomicName id="8FE24D5AFFBFFF44311AF809FD85F87E" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[294,536,1923,1945]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="213" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBFFF44311AF809FD85F87E" box="[294,536,1923,1945]" italics="true" pageId="14" pageNumber="213">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBFFF443221F809FDF2F87F" box="[541,623,1923,1944]" pageId="14" pageNumber="213" rank="species">
<emphasis id="7A96EACBFFBFFF443221F809FDF2F87F" bold="true" box="[541,623,1923,1944]" pageId="14" pageNumber="213">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (cl not measured) from Guana Island, British Virgin Islands, in dorsal [a] and lateral [b] views (specimen possibly deposited in the Natural History Museum of Los Angeles Country, LACM, not available for study). Photographs: Todd Zimmerman (a, b).
</paragraph>
</caption>
<paragraph id="485D36D9FFBEFF4530FBFF1DFC0DFEFB" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
Male second pleopod with appendix masculina exceeding appendix interna; apex with several long stiff setae (
<figureCitation id="D0D92A5CFFBEFF4530A3FF36FF78FF32" box="[159,229,188,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="15" pageNumber="214">Fig. 5</figureCitation>
j). Uropod with lateral lobe of protopod ending in stout sharp tooth; endopod and exopod broadly ovate; exopod with distolateral tooth and adjacent spiniform seta both moderately developed; diaeresis with blunt tooth adjacent to spiniform seta and sinuous mesial portion (
<figureCitation id="D0D92A5CFFBEFF4532C7FE8EFCD5FEFA" box="[763,840,260,285]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="15" pageNumber="214">Figs. 5</figureCitation>
k, 8n).
</paragraph>
<paragraph id="485D36D9FFBEFF4530FBFEADFD6DFEA7" blockId="15.[151,1437,151,1904]" box="[199,752,295,320]" pageId="15" pageNumber="214">
Gill-exopod formula as in 
<taxonomicName id="8FE24D5AFFBEFF4531CCFEA2FDFFFED8" authorityName="Rathbun" authorityYear="1901" box="[496,610,296,319]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF4531CCFEA2FDFFFED8" box="[496,610,296,319]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
(see above).
</paragraph>
<paragraph id="485D36D9FFBEFF4530FBFEC6FB90FE13" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
<emphasis id="7A96EACBFFBEFF4530FBFEC6FE1DFE82" bold="true" box="[199,384,332,357]" pageId="15" pageNumber="214">Colour pattern</emphasis>
. Background semitransparent whitish; carapace covered with bright red chromatophores, except for anterior-most dorsal and posterior-most lateral portions; pleon typically with two broad adjacent transverse bands of red chromatophores running across posterior half of third pleomere and most of fourth pleomere, sometimes with only one band on either third or fourth pleomere; antennular and antennal flagella pale orange; chelipeds hyaline white; walking legs and tail fan whitish (
<figureCitation id="D0D92A5CFFBEFF4533B9FE56FC49FE13" box="[901,980,476,501]" captionStart="FIGURE 9" captionStartId="13.[151,250,1655,1677]" captionTargetBox="[150,1436,193,1634]" captionTargetId="figure@13.[150,1436,193,1634]" captionTargetPageId="13" captionText="FIGURE 9. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222) [a]; male (cl 4.5 mm) from Carrie Bow Cay, Belize (ULLZ 8939) [b]. Photographs: A. Anker (a) and Darryl L. Felder (b)." httpUri="https://zenodo.org/record/809652/files/figure.png" pageId="15" pageNumber="214">Figs. 9</figureCitation>
, 
<figureCitation id="D0D92A5CFFBEFF4533DCFE56FC62FE13" box="[992,1023,476,500]" captionStart="FIGURE 10" captionStartId="14.[151,250,1923,1945]" captionTargetBox="[150,1436,769,1901]" captionTargetId="figure@14.[150,1436,769,1901]" captionTargetPageId="14" captionText="FIGURE 10. Alpheopsis paratrigona sp. nov., male (cl not measured) from Guana Island, British Virgin Islands, in dorsal [a] and lateral [b] views (specimen possibly deposited in the Natural History Museum of Los Angeles Country, LACM, not available for study). Photographs: Todd Zimmerman (a, b)." httpUri="https://zenodo.org/record/809654/files/figure.png" pageId="15" pageNumber="214">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFBEFF4530FBFE75FA96FDF0" box="[199,1291,511,536]" pageId="15" pageNumber="214" type="etymology">
<paragraph id="485D36D9FFBEFF4530FBFE75FA96FDF0" blockId="15.[151,1437,151,1904]" box="[199,1291,511,536]" pageId="15" pageNumber="214">
<emphasis id="7A96EACBFFBEFF4530FBFE75FEDBFDFF" bold="true" box="[199,326,511,536]" pageId="15" pageNumber="214">Etymology</emphasis>
. The specific name refers to the general resemblance of the new species to 
<taxonomicName id="8FE24D5AFFBEFF4534AFFD8AFA99FDF0" authorityName="Rathbun" authorityYear="1901" box="[1171,1284,512,535]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF4534AFFD8AFA99FDF0" box="[1171,1284,512,535]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFBEFF4530FBFDAEFCBDFDDB" box="[199,800,548,573]" pageId="15" pageNumber="214" type="materials_examined">
<paragraph id="485D36D9FFBEFF4530FBFDAEFCBDFDDB" blockId="15.[151,1437,151,1904]" box="[199,800,548,573]" pageId="15" pageNumber="214">
<materialsCitation id="F88A3C84FFBEFF4530FBFDAEFCBDFDDB" ID-GBIF-Occurrence="1563896391" box="[199,800,548,573]" country="Panama" location="Portobelo" municipality="Caribbean" pageId="15" pageNumber="214" specimenCount="1" typeStatus="holotype">
<emphasis id="7A96EACBFFBEFF4530FBFDAEFEC0FDDA" bold="true" box="[199,349,548,573]" pageId="15" pageNumber="214">
<typeStatus id="9759887BFFBEFF4530FBFDAEFE9FFDDA" box="[199,258,548,573]" pageId="15" pageNumber="214">Type</typeStatus>
locality
</emphasis>
. 
<location id="4D3D6002FFBEFF453150FDAEFE41FDDA" LSID="urn:lsid:plazi:treatment:C04B87CFFFB9FF4530ABFF1DFB4DF897:4D3D6002FFBEFF453150FDAEFE41FDDA" box="[364,476,548,573]" country="Panama" municipality="Caribbean" name="Portobelo" pageId="15" pageNumber="214">Portobelo</location>
, 
<collectingMunicipality id="A839ACA3FFBEFF4531DAFDAEFDC4FDDA" box="[486,601,548,573]" pageId="15" pageNumber="214">Caribbean</collectingMunicipality>
coast of 
<collectingCountry id="30F57649FFBEFF453283FDAFFC86FDDB" box="[703,795,549,572]" name="Panama" pageId="15" pageNumber="214">Panama</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFBEFF4530FBFDCDFEBDFC42" pageId="15" pageNumber="214" type="distribution">
<paragraph id="485D36D9FFBEFF4530FBFDCDFEE6FCF2" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
<emphasis id="7A96EACBFFBEFF4530FBFDCDFEC4FD87" bold="true" box="[199,345,583,608]" pageId="15" pageNumber="214">Distribution</emphasis>
. Western Atlantic: 
<collectingCountry id="30F57649FFBEFF45320EFDCDFD04FD87" box="[562,665,583,608]" name="Bermuda" pageId="15" pageNumber="214">Bermuda</collectingCountry>
(
<bibRefCitation id="2C734B28FFBEFF45329BFDCDFBE3FD87" author="Chace" box="[679,1150,583,608]" pageId="15" pageNumber="214" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. https: // doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">
Chace 1972, as 
<taxonomicName id="8FE24D5AFFBEFF453366FDC2FC4BFDB8" authorityName="Rathbun" authorityYear="1901" box="[858,982,584,607]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigonus">
<emphasis id="7A96EACBFFBEFF453366FDC2FC4BFDB8" box="[858,982,584,607]" italics="true" pageId="15" pageNumber="214">A. trigonus</emphasis>
</taxonomicName>
; present study
</bibRefCitation>
); Caribbean Sea: 
<collectingRegion id="8A26F83BFFBEFF45357FFDC2FA01FDB8" box="[1347,1436,584,607]" country="Panama" name="Panama" pageId="15" pageNumber="214">
<collectingCountry id="30F57649FFBEFF45357FFDC2FA01FDB8" box="[1347,1436,584,607]" name="Panama" pageId="15" pageNumber="214">Panama</collectingCountry>
</collectingRegion>
(Portobelo, present study), 
<collectingRegion id="8A26F83BFFBEFF4531F7FDE6FDBCFD62" box="[459,545,620,645]" country="Mexico" name="Mexico" pageId="15" pageNumber="214">
<collectingCountry id="30F57649FFBEFF4531F7FDE6FDBCFD62" box="[459,545,620,645]" name="Mexico" pageId="15" pageNumber="214">Mexico</collectingCountry>
</collectingRegion>
(
<collectingRegion id="8A26F83BFFBEFF45320DFDE6FD0DFD63" box="[561,656,620,644]" country="Mexico" name="Yucatan" pageId="15" pageNumber="214">Yucatan</collectingRegion>
, 
<bibRefCitation id="2C734B28FFBEFF4532A1FDE6FCBBFD63" author="Chace" box="[669,806,620,645]" pageId="15" pageNumber="214" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. https: // doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">Chace 1972</bibRefCitation>
, as 
<taxonomicName id="8FE24D5AFFBEFF45336FFDE6FC4CFD63" authorityName="Rathbun" authorityYear="1901" box="[851,977,620,644]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigonus">
<emphasis id="7A96EACBFFBEFF45336FFDE6FC4CFD63" box="[851,977,620,644]" italics="true" pageId="15" pageNumber="214">A. trigonus</emphasis>
</taxonomicName>
; present study), 
<collectingCountry id="30F57649FFBEFF4534ACFDE6FA9EFD62" box="[1168,1283,620,645]" name="Venezuela" pageId="15" pageNumber="214">Venezuela</collectingCountry>
(Los Roques Archipelago, Rodriguez 1982, as 
<taxonomicName id="8FE24D5AFFBEFF453218FD1AFD02FD4F" authorityName="Coutiere" authorityYear="1897" box="[548,671,656,680]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="7A96EACBFFBEFF453218FD1AFD02FD4F" box="[548,671,656,680]" italics="true" pageId="15" pageNumber="214">Alpheopsis</emphasis>
</taxonomicName>
sp. 1), French Antilles (
<collectingCountry id="30F57649FFBEFF4533F5FD05FBC8FD4F" box="[969,1109,655,680]" name="Guadeloupe" pageId="15" pageNumber="214">Guadeloupe</collectingCountry>
, 
<bibRefCitation id="2C734B28FFBEFF453459FD05FB6FFD4F" author="Anker" box="[1125,1266,655,680]" pageId="15" pageNumber="214" refString="Anker, A. (2001) Taxonomie et evolution des Alpheidae (Crustacea, Decapoda). Fol. 1 &amp; 2. PhD thesis, Museum National d'Histoire Naturelle, Paris, 547 pp. &amp; 331 pp." type="book" year="2001">Anker 2001</bibRefCitation>
, as 
<emphasis id="7A96EACBFFBEFF45351AFD1AFAA2FD40" box="[1318,1343,656,679]" italics="true" pageId="15" pageNumber="214">A.</emphasis>
sp. 
<taxonomicName id="8FE24D5AFFBEFF45354AFD05FF69FD2B" authorityName="Rathbun" authorityYear="1901" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigonus">
aff. 
<emphasis id="7A96EACBFFBEFF4530ABFD3FFF69FD2B" box="[151,244,693,716]" italics="true" pageId="15" pageNumber="214">trigonus</emphasis>
</taxonomicName>
), 
<collectingCountry id="30F57649FFBEFF453133FD3EFD9AFD2A" box="[271,519,692,717]" name="British Virgin Islands" pageId="15" pageNumber="214">British Virgin Islands</collectingCountry>
(Guana Island, photographic record, see 
<figureCitation id="D0D92A5CFFBEFF4533D0FD3EFBDFFD2B" box="[1004,1090,692,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1923,1945]" captionTargetBox="[150,1436,769,1901]" captionTargetId="figure@14.[150,1436,769,1901]" captionTargetPageId="14" captionText="FIGURE 10. Alpheopsis paratrigona sp. nov., male (cl not measured) from Guana Island, British Virgin Islands, in dorsal [a] and lateral [b] views (specimen possibly deposited in the Natural History Museum of Los Angeles Country, LACM, not available for study). Photographs: Todd Zimmerman (a, b)." httpUri="https://zenodo.org/record/809654/files/figure.png" pageId="15" pageNumber="214">Fig. 10</figureCitation>
); 
<collectingCountry id="30F57649FFBEFF453467FD3EFBE2FD2B" box="[1115,1151,692,716]" name="United States of America" pageId="15" pageNumber="214">US</collectingCountry>
<collectingCountry id="30F57649FFBEFF4534B5FD3EFAB7FD2A" box="[1161,1322,692,717]" name="Virgin Islands" pageId="15" pageNumber="214">Virgin Islands</collectingCountry>
(St. John Island), 
<collectingRegion id="8A26F83BFFBEFF4530C8FD5DFECEFD17" box="[244,339,727,752]" country="Antigua and Barbuda" name="Barbuda" pageId="15" pageNumber="214">
<collectingCountry id="30F57649FFBEFF4530C8FD5DFECEFD17" box="[244,339,727,752]" name="Antigua and Barbuda" pageId="15" pageNumber="214">Barbuda</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="2C734B28FFBEFF45315FFD5DFD0AFD08" author="Chace" box="[355,663,727,752]" pageId="15" pageNumber="214" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. https: // doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">
Chace 1972, as 
<taxonomicName id="8FE24D5AFFBEFF453226FD52FD0AFD08" authorityName="Rathbun" authorityYear="1901" box="[538,663,728,751]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigonus">
<emphasis id="7A96EACBFFBEFF453226FD52FD0AFD08" box="[538,663,728,751]" italics="true" pageId="15" pageNumber="214">A. trigonus</emphasis>
</taxonomicName>
</bibRefCitation>
), 
<collectingRegion id="8A26F83BFFBEFF45328CFD5DFD65FD17" box="[688,760,727,752]" country="Belize" name="Belize" pageId="15" pageNumber="214">
<collectingCountry id="30F57649FFBEFF45328CFD5DFD65FD17" box="[688,760,727,752]" name="Belize" pageId="15" pageNumber="214">Belize</collectingCountry>
</collectingRegion>
(Carrie Bow Cay) (present study); possibly also 
<collectingCountry id="30F57649FFBEFF45351BFD5DFAEDFD17" box="[1319,1392,727,752]" name="Brazil" pageId="15" pageNumber="214">Brazil</collectingCountry>
: off 
<collectingRegion id="8A26F83BFFBEFF4530ABFD76FF76FCF2" box="[151,235,764,789]" country="Brazil" name="Paraiba" pageId="15" pageNumber="214">Paraíba</collectingRegion>
(see below).
</paragraph>
<paragraph id="485D36D9FFBEFF4530FBFC95FEBDFC42" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
<emphasis id="7A96EACBFFBEFF4530FBFC95FEB9FCDF" bold="true" box="[199,292,799,824]" pageId="15" pageNumber="214">Ecology</emphasis>
. Coral reefs and associated shallow flats with abundance of coral rubble, also on rocky-sandy mixed bottoms; in crevices of dead corals, under coral rubble and rocks, in shells of dead queen conchs, 
<taxonomicName id="8FE24D5AFFBEFF4534C0FCCEFA06FCBB" box="[1276,1435,836,860]" class="Gastropoda" family="Strombidae" genus="Lobatus" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="214" phylum="Mollusca" rank="species" species="gigas">
<emphasis id="7A96EACBFFBEFF4534C0FCCEFA06FCBB" box="[1276,1435,836,860]" italics="true" pageId="15" pageNumber="214">Lobatus gigas</emphasis>
</taxonomicName>
(L.); shallow subtidal (
<quantity id="8F1A9B3CFFBEFF4531A5FCE2FE7FFC67" box="[409,482,872,896]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="15" pageNumber="214" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
) to at least 
<quantity id="8F1A9B3CFFBEFF453259FCE2FD02FC98" box="[613,671,872,896]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="214" unit="m" value="12.0">12 m</quantity>
(
<bibRefCitation id="2C734B28FFBEFF453292FCEDFB91FC67" author="Rodriguez" box="[686,1036,871,896]" pageId="15" pageNumber="214" refString="Rodriguez, B. (1986) Los camarones (Crustacea; Decapoda; Natantia) del Parque Nacional Archipielago de los Roques. Thesis. Licenciado en Biologia, Universidad Central de Venezuela, Caracas, 350 pp." type="book" year="1986">Rodriguez 1986; present study</bibRefCitation>
), possibly in deeper water in 
<collectingCountry id="30F57649FFBEFF45356BFCEDFA06FC67" box="[1367,1435,871,896]" name="Brazil" pageId="15" pageNumber="214">Brazil</collectingCountry>
(see below).
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFBEFF4530FBFC25FB4DF897" pageId="15" pageNumber="214" type="discussion">
<paragraph id="485D36D9FFBEFF4530FBFC25FDEDFA7A" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
<emphasis id="7A96EACBFFBEFF4530FBFC25FEA9FC2F" bold="true" box="[199,308,943,968]" pageId="15" pageNumber="214">Remarks</emphasis>
. 
<taxonomicName id="8FE24D5AFFBEFF453179FC3AFDCCFC20" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[325,593,944,968]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBEFF453179FC3AFDCCFC20" box="[325,593,944,968]" italics="true" pageId="15" pageNumber="214">Alpheopsis paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBEFF453260FC25FD25FC2F" box="[604,696,943,968]" pageId="15" pageNumber="214" rank="species">
<emphasis id="7A96EACBFFBEFF453260FC25FD25FC2F" bold="true" box="[604,696,943,968]" pageId="15" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
, as its name suggests, is closely related to 
<taxonomicName id="8FE24D5AFFBEFF453488FC3AFAB4FC20" authorityName="Rathbun" authorityYear="1901" box="[1204,1321,944,967]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF453488FC3AFAB4FC20" box="[1204,1321,944,967]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
, differing from it by the presence of only one feebly developed carina (presumably VLC) on the carapace (cf. 
<figureCitation id="D0D92A5CFFBEFF4534CDFC5EFAA1FC0A" box="[1265,1340,980,1005]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="15" pageNumber="214">Figs. 5</figureCitation>
a–d; 7a– c, m–o) vs. two well developed carinae (DLC and VLC) in 
<taxonomicName id="8FE24D5AFFBEFF45330DFC72FC3EFBE8" authorityName="Rathbun" authorityYear="1901" box="[817,931,1016,1039]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF45330DFC72FC3EFBE8" box="[817,931,1016,1039]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
(
<figureCitation id="D0D92A5CFFBEFF45338EFC7DFC62FBF7" box="[946,1023,1015,1040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1739,1761]" captionTargetBox="[162,1430,203,1707]" captionTargetId="figure@3.[151,1436,193,1718]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis trigona (Rathbun, 1901), female (cl 5.5 mm) from Fort Pierce, Florida, USA (MZUSP 34221): acarapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; hthird maxilliped, lateral; i—same, tip of ultimate article, mesial; j—uropod, dorsal." httpUri="https://zenodo.org/record/809638/files/figure.png" pageId="15" pageNumber="214">Figs. 2</figureCitation>
a–d; 4a, d–g; see also 
<bibRefCitation id="2C734B28FFBEFF4534CAFC7DFA05FBF7" author="Rathbun" box="[1270,1432,1015,1040]" pageId="15" pageNumber="214" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 1900 (20), 1 - 137, pls. 1, 2." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
: fig. 21); the frontal margin of the carapace with orbital teeth, ranging from distinctly projecting and sharp teeth to rounded or blunt-angular lobes (
<figureCitation id="D0D92A5CFFBEFF4531C2FBB5FDD6FBBF" box="[510,587,1087,1112]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="15" pageNumber="214">Figs. 5</figureCitation>
d; 7d, o), vs. without orbital teeth in 
<taxonomicName id="8FE24D5AFFBEFF4533DDFBCAFBCEFBB0" authorityName="Rathbun" authorityYear="1901" box="[993,1107,1088,1111]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF4533DDFBCAFBCEFBB0" box="[993,1107,1088,1111]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
(
<figureCitation id="D0D92A5CFFBEFF45345EFBB5FB32FBBF" box="[1122,1199,1087,1112]" captionStart="FIGURE 2" captionStartId="3.[151,250,1739,1761]" captionTargetBox="[162,1430,203,1707]" captionTargetId="figure@3.[151,1436,193,1718]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis trigona (Rathbun, 1901), female (cl 5.5 mm) from Fort Pierce, Florida, USA (MZUSP 34221): acarapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; hthird maxilliped, lateral; i—same, tip of ultimate article, mesial; j—uropod, dorsal." httpUri="https://zenodo.org/record/809638/files/figure.png" pageId="15" pageNumber="214">Figs. 2</figureCitation>
b, d; 4a, e, g; see also 
<bibRefCitation id="2C734B28FFBEFF4530ABFBEEFEA4FB9B" author="Rathbun" box="[151,313,1124,1149]" pageId="15" pageNumber="214" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 1900 (20), 1 - 137, pls. 1, 2." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
: fig. 21); the dorsomesial-proximal surface of the cheliped palm unarmed (cf. 
<figureCitation id="D0D92A5CFFBEFF453494FBEEFB77FB9A" box="[1192,1258,1124,1149]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="15" pageNumber="214">Fig. 6</figureCitation>
a–c; 7e, f; 
<quantity id="8F1A9B3CFFBEFF45355DFBEEFAE6FB9A" box="[1377,1403,1124,1149]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="15" pageNumber="214" unit="g" value="8.0">8g</quantity>
–l) vs. with a strong curved process in 
<taxonomicName id="8FE24D5AFFBEFF453214FB02FD06FB78" authorityName="Rathbun" authorityYear="1901" box="[552,667,1160,1183]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF453214FB02FD06FB78" box="[552,667,1160,1183]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
(
<figureCitation id="D0D92A5CFFBEFF453296FB0DFD65FB47" box="[682,760,1159,1184]" captionStart="FIGURE 3" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[167,1421,205,1497]" captionTargetId="figure@5.[151,1436,193,1511]" captionTargetPageId="5" captionText="FIGURE 3. Alpheopsis trigona (Rathbun, 1901), female (cl 5.5 mm) from Fort Pierce, Florida, USA (MZUSP 34221): aright cheliped, lateral; b—same, mesial (proximal articles detached); c—same, chela, mesial; d—same, chela fingers, lateral; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809640/files/figure.png" pageId="15" pageNumber="214">Figs. 3</figureCitation>
b, c; 4b, c, h, k); and the fingers of the major cheliped palm moderately gaping and unarmed or armed with three to five rounded-triangular low teeth, all more or less similar in size (
<figureCitation id="D0D92A5CFFBEFF4530EDFB45FE82FB0F" box="[209,287,1231,1256]" captionStart="FIGURE 6" captionStartId="10.[151,250,1360,1382]" captionTargetBox="[158,1418,204,1325]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURE 6. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—left cheliped, lateral; b—same, mesial; c—same, chela, lateral; d—second pereiopod, lateral; e—third pereiopod, lateral; f—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809646/files/figure.png" pageId="15" pageNumber="214">Figs. 6</figureCitation>
c; 7f, g; 8h, i; see also 
<bibRefCitation id="2C734B28FFBEFF45322BFB45FD53FB0F" author="Rodriguez" box="[535,718,1231,1256]" pageId="15" pageNumber="214" refString="Rodriguez, B. (1986) Los camarones (Crustacea; Decapoda; Natantia) del Parque Nacional Archipielago de los Roques. Thesis. Licenciado en Biologia, Universidad Central de Venezuela, Caracas, 350 pp." type="book" year="1986">Rodríguez 1986</bibRefCitation>
: fig. 29e) vs. strongly gaping and armed with three much stouter teeth, especially the distal-most tooth of the pollex (
<figureCitation id="D0D92A5CFFBEFF4532E6FB7EFC80FAEA" box="[730,797,1268,1293]" captionStart="FIGURE 4" captionStartId="6.[151,250,1820,1842]" captionTargetBox="[164,1418,316,1779]" captionTargetId="figure@6.[151,1436,301,1798]" captionTargetPageId="6" captionText="FIGURE 4. Alpheopsis trigona (Rathbun, 1901), female holotype (cl 5.5 mm) from off Vieques Island, Puerto Rico (USNM 23785) [a – c]; male (cl 5.4 mm) from Peter Island, US Virgin Islands (USNM 135844); a—carapace, lateral; b—left cheliped, mesial; c—same, dorsolateral; d—carapace and anterior pleon, lateral; e—same, dorsal; f—anterior region of carapace and frontal appendages, lateral; g—same, dorsal; h—right (major) cheliped, mesial; i—same, carpus and chela, lateral; j—left (minor) cheliped, mesial; k—same, lateral; l—second pereiopod, lateral; m—third pereiopod, lateral. Modified from Anker (2001)." httpUri="https://zenodo.org/record/809642/files/figure.png" pageId="15" pageNumber="214">Fig. 4</figureCitation>
h, i). Because of the notable variation in the development of the orbital teeth and VLC and ABC in 
<taxonomicName id="8FE24D5AFFBEFF453255FA92FC92FAC8" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[617,783,1304,1327]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBEFF453255FA92FC92FAC8" box="[617,783,1304,1327]" italics="true" pageId="15" pageNumber="214">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBEFF45332AFA9DFCF2FAD7" box="[790,879,1303,1328]" pageId="15" pageNumber="214" rank="species">
<emphasis id="7A96EACBFFBEFF45332AFA9DFCF4FAD7" bold="true" box="[790,873,1303,1328]" pageId="15" pageNumber="214">sp. nov</emphasis>
.
</taxonomicNameLabel>
, the variation in the extension of DLC (including development of PLC) in 
<taxonomicName id="8FE24D5AFFBEFF453186FAB6FDB2FAB3" authorityName="Rathbun" authorityYear="1901" box="[442,559,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF453186FAB6FDB2FAB3" box="[442,559,1340,1364]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
, as well as the likely sympatry of these two taxa over at least part of their distribution ranges, a genetic analysis (e.g. comparison of the COI gene sequences) is desirable to verify the validity of the here-proposed new species.
</paragraph>
<paragraph id="485D36D9FFBEFF4530FBFA2DFC81F97F" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
The colour patterns of 
<taxonomicName id="8FE24D5AFFBEFF4531F8FA22FDABFA58" authorityName="Rathbun" authorityYear="1901" box="[452,566,1448,1471]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF4531F8FA22FDABFA58" box="[452,566,1448,1471]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
</taxonomicName>
and 
<taxonomicName id="8FE24D5AFFBEFF453257FA22FC8FFA58" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[619,786,1448,1471]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBEFF453257FA22FC8FFA58" box="[619,786,1448,1471]" italics="true" pageId="15" pageNumber="214">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBEFF453324FA2DFCECFA27" box="[792,881,1447,1472]" pageId="15" pageNumber="214" rank="species">
<emphasis id="7A96EACBFFBEFF453324FA2DFCECFA27" bold="true" box="[792,881,1447,1472]" pageId="15" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
are very similar, although the pleon of the former species appears to have an anterior transverse band across the first pleomere, extending laterally onto the second pleomere (
<bibRefCitation id="2C734B28FFBEFF453131FA65FE6AF9EF" author="Soledade" box="[269,503,1519,1544]" pageId="15" pageNumber="214" refString="Soledade, G. O., Fonseca, M. S. &amp; Almeida, A. O. (2015) Shallow-water stenopodidean and caridean shrimps from Abrolhos Archipelago, Brazil: new records and updated checklist. Zootaxa, 3905 (1), 52 - 68." type="journal article" year="2015">
Soledade 
<emphasis id="7A96EACBFFBEFF453140FA7BFE29F9EF" box="[380,436,1520,1544]" italics="true" pageId="15" pageNumber="214">et al.</emphasis>
2015
</bibRefCitation>
: fig. 3A), which is absent in the new species (
<figureCitation id="D0D92A5CFFBEFF4533C8FA65FBDEF9EF" box="[1012,1091,1519,1544]" captionStart="FIGURE 9" captionStartId="13.[151,250,1655,1677]" captionTargetBox="[150,1436,193,1634]" captionTargetId="figure@13.[150,1436,193,1634]" captionTargetPageId="13" captionText="FIGURE 9. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222) [a]; male (cl 4.5 mm) from Carrie Bow Cay, Belize (ULLZ 8939) [b]. Photographs: A. Anker (a) and Darryl L. Felder (b)." httpUri="https://zenodo.org/record/809652/files/figure.png" pageId="15" pageNumber="214">Figs. 9</figureCitation>
, 
<figureCitation id="D0D92A5CFFBEFF453473FA7AFBF3F9EF" box="[1103,1134,1520,1544]" captionStart="FIGURE 10" captionStartId="14.[151,250,1923,1945]" captionTargetBox="[150,1436,769,1901]" captionTargetId="figure@14.[150,1436,769,1901]" captionTargetPageId="14" captionText="FIGURE 10. Alpheopsis paratrigona sp. nov., male (cl not measured) from Guana Island, British Virgin Islands, in dorsal [a] and lateral [b] views (specimen possibly deposited in the Natural History Museum of Los Angeles Country, LACM, not available for study). Photographs: Todd Zimmerman (a, b)." httpUri="https://zenodo.org/record/809654/files/figure.png" pageId="15" pageNumber="214">10</figureCitation>
). However, since firstly, 
<taxonomicName id="8FE24D5AFFBEFF4535B8FA7AFE83F9CB" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBEFF4535B8FA7AFE83F9CB" italics="true" pageId="15" pageNumber="214">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBEFF453119F99EFEE3F9CA" box="[293,382,1556,1581]" pageId="15" pageNumber="214" rank="species">
<emphasis id="7A96EACBFFBEFF453119F99EFEE3F9CA" bold="true" box="[293,382,1556,1581]" pageId="15" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
displays some variation in the development of the posterior two bands (one vs. more typically two, 
<figureCitation id="D0D92A5CFFBEFF4530EFF9BDFEB8F9B7" box="[211,293,1591,1616]" captionStart="FIGURE 9" captionStartId="13.[151,250,1655,1677]" captionTargetBox="[150,1436,193,1634]" captionTargetId="figure@13.[150,1436,193,1634]" captionTargetPageId="13" captionText="FIGURE 9. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222) [a]; male (cl 4.5 mm) from Carrie Bow Cay, Belize (ULLZ 8939) [b]. Photographs: A. Anker (a) and Darryl L. Felder (b)." httpUri="https://zenodo.org/record/809652/files/figure.png" pageId="15" pageNumber="214">Figs. 9</figureCitation>
, 
<figureCitation id="D0D92A5CFFBEFF453109F9B2FEC9F9B7" box="[309,340,1592,1616]" captionStart="FIGURE 10" captionStartId="14.[151,250,1923,1945]" captionTargetBox="[150,1436,769,1901]" captionTargetId="figure@14.[150,1436,769,1901]" captionTargetPageId="14" captionText="FIGURE 10. Alpheopsis paratrigona sp. nov., male (cl not measured) from Guana Island, British Virgin Islands, in dorsal [a] and lateral [b] views (specimen possibly deposited in the Natural History Museum of Los Angeles Country, LACM, not available for study). Photographs: Todd Zimmerman (a, b)." httpUri="https://zenodo.org/record/809654/files/figure.png" pageId="15" pageNumber="214">10</figureCitation>
), and secondly, the specimen of 
<taxonomicName id="8FE24D5AFFBEFF4532E4F9B2FBE0F9B7" authority="in Soledade et al. (2015)" authorityName="in Soledade et al." authorityYear="2015" box="[728,1149,1591,1616]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFBEFF4532E4F9B2FCD2F9A8" box="[728,847,1592,1615]" italics="true" pageId="15" pageNumber="214">A. trigona</emphasis>
in 
<bibRefCitation id="2C734B28FFBEFF453346F9BDFBE0F9B7" box="[890,1149,1591,1616]" pageId="15" pageNumber="214" refString="Soledade, G. O., Fonseca, M. S. &amp; Almeida, A. O. (2015) Shallow-water stenopodidean and caridean shrimps from Abrolhos Archipelago, Brazil: new records and updated checklist. Zootaxa, 3905 (1), 52 - 68." type="journal article">
Soledade 
<emphasis id="7A96EACBFFBEFF4533D0F9B3FBB5F9B7" box="[1004,1064,1592,1616]" italics="true" pageId="15" pageNumber="214">et al.</emphasis>
(2015)
</bibRefCitation>
</taxonomicName>
was photographed postmortem (which sometimes results in fading or displacement of pigments), the importance of the colour pattern in the separation of the two species remains to be evaluated.
</paragraph>
<paragraph id="485D36D9FFBEFF4530FBF92EFB4DF897" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="214">
The Brazilian specimen tentatively identified as 
<taxonomicName id="8FE24D5AFFBEFF4532DFF92EFC15F95B" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[739,904,1700,1724]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBEFF4532DFF92EFC15F95B" box="[739,904,1700,1724]" italics="true" pageId="15" pageNumber="214">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBEFF4533B3F92EFC75F95A" box="[911,1000,1700,1725]" pageId="15" pageNumber="214" rank="species">
<emphasis id="7A96EACBFFBEFF4533B3F92EFC75F95A" bold="true" box="[911,1000,1700,1725]" pageId="15" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
is an incomplete juvenile male lacking most of its pereiopods, including both chelipeds. The orbital margins of this specimen are produced anteriorly into distally acute teeth. The carapace shows no trace of ALC or ABC. On the other hand, MDC is relatively well marked and the carapace also seems to have, albeit poorly developed, a longer ASC and a shorter PSC. Based on these features, this specimen is identifiable as 
<taxonomicName id="8FE24D5AFFBEFF45329DF8BEFCD5F8AB" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[673,840,1844,1868]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFBEFF45329DF8BEFCD5F8AB" box="[673,840,1844,1868]" italics="true" pageId="15" pageNumber="214">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFBEFF45336CF8BEFC37F8AA" box="[848,938,1844,1869]" pageId="15" pageNumber="214" rank="species">
<emphasis id="7A96EACBFFBEFF45336CF8BEFC37F8AA" bold="true" box="[848,938,1844,1869]" pageId="15" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
, however, collection of more complete and/ or adult specimens from 
<collectingCountry id="30F57649FFBEFF453191F8DDFE6CF897" box="[429,497,1879,1904]" name="Brazil" pageId="15" pageNumber="214">Brazil</collectingCountry>
is necessary to confirm its presence in the southwestern Atlantic.
</paragraph>
</subSubSection>
</treatment>
</document>