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<document id="8F3F738FCEFE6B019D394EEB0B5E0082" ID="10.11646/zootaxa.4277.2.2SLASH11367" ID-CLB-Dataset="32829" ID-DOI="10.11646/zootaxa.4277.2.2" ID-GBIF-Dataset="5d4045c8-2c5a-4f75-8ca7-a2401ed0d227" ID-ISSN="1175-5326" ID-Zenodo-Dep="809634" ID-ZooBank="1551FE58-FCC9-4D45-AE2A-77655BB53147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497598594012" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2017" docId="C04B87CFFFA1FF5C30ABFF1DFD03FAB2" docLanguage="en" docName="zootaxa.4277.2.2SLASH11367.pdf" docOrigin="Zootaxa 4277 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheopsis gotrina Anker 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="221" masterDocId="3C72FFB7FFB1FF4A303CFF8AFF9DFFE7" masterDocTitle="Strongly carinate species of Alpheopsis Coutière, 1897 of the tropical Atlantic and eastern Pacific, with redescription of A. trigona (Rathbun, 1901) and description of three new species (Malacostraca: Decapoda: Alpheidae)" masterLastPageNumber="227" masterPageNumber="199" pageNumber="215" updateTime="1698447107539" updateUser="ExternalLinkService">
<mods:mods id="ADE9069848A0300F5F445326C2D656E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BAE8A351D3B8BF08046545A2090AB10F">
<mods:title id="C02D7E17B7D633428B21FFD1434A3023">Strongly carinate species of Alpheopsis Coutière, 1897 of the tropical Atlantic and eastern Pacific, with redescription of A. trigona (Rathbun, 1901) and description of three new species (Malacostraca: Decapoda: Alpheidae)</mods:title>
</mods:titleInfo>
<mods:name id="CAAEC6166A46B26F417568EFE87F8F9F" type="personal">
<mods:role id="E8C76FCA45FD54C45418AA6EAB8301C7">
<mods:roleTerm id="5D068E703ADD160CED49FF94A0E37C04">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BBA5BBAA22F5E8ED8A680A38349E8829">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="7D12EBEFFA61171207FE29B3F6D294CF">text</mods:typeOfResource>
<mods:relatedItem id="AC42958A4CE0120383F80C8A80640C4F" type="host">
<mods:titleInfo id="BBC13F30CA54B9CA9E319865F3222F0F">
<mods:title id="38A5627519BFE7C1CF9A4F47E2DDCB13">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="06EC0DB7B068BD23888B8C39C785701F">
<mods:date id="0EE0D834A93ECE32871B4CBA7243ACD4">2017</mods:date>
<mods:detail id="C77E8A6E56B4C4195E89BFB662113685" type="volume">
<mods:number id="D618540C8F9378DC790B7DE63E5D9AC7">4277</mods:number>
</mods:detail>
<mods:detail id="0F9487DF858FE1EDF59022BE30A73EE9" type="issue">
<mods:number id="840C9DB54A915749A353F8545038FED8">2</mods:number>
</mods:detail>
<mods:extent id="E1B0A922080EE631053F8BF7D549B221" unit="page">
<mods:start id="68278B7851D467D9B78B4C4A534F3B17">199</mods:start>
<mods:end id="D9E4E67AEC2605ABCA0BC51FCC44894A">227</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="550D3C93D317ADF0A218A878090A25CA">journal article</mods:classification>
<mods:identifier id="C207BAAA4F72BD06BF189750455174FF" type="CLB-Dataset">32829</mods:identifier>
<mods:identifier id="91B2DEC06BABC8704D028821E7A8D6FD" type="DOI">10.11646/zootaxa.4277.2.2</mods:identifier>
<mods:identifier id="D9ED8E01299D0A5FC85CBC47B089349B" type="GBIF-Dataset">5d4045c8-2c5a-4f75-8ca7-a2401ed0d227</mods:identifier>
<mods:identifier id="7AF433FB7014659217A070B1859054BC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DCA23412C56F5AA06601D98AC0448A95" type="Zenodo-Dep">809634</mods:identifier>
<mods:identifier id="FEB84D7592C9910ABBF883C2B1FEEAD2" type="ZooBank">1551FE58-FCC9-4D45-AE2A-77655BB53147</mods:identifier>
</mods:mods>
<treatment id="C04B87CFFFA1FF5C30ABFF1DFD03FAB2" ID-DOI="http://doi.org/10.5281/zenodo.6039734" ID-GBIF-Taxon="131878750" ID-Zenodo-Dep="6039734" LSID="urn:lsid:plazi:treatment:C04B87CFFFA1FF5C30ABFF1DFD03FAB2" httpUri="http://treatment.plazi.org/id/C04B87CFFFA1FF5C30ABFF1DFD03FAB2" lastPageId="22" lastPageNumber="221" pageId="16" pageNumber="215">
<subSubSection id="00F86552FFA1FF5A30ABFF1DFEB1FF34" pageId="16" pageNumber="215" type="nomenclature">
<paragraph id="485D36D9FFA1FF5A30ABFF1DFE42FF55" blockId="16.[151,479,151,211]" box="[151,479,151,178]" pageId="16" pageNumber="215">
<heading id="131581B5FFA1FF5A30ABFF1DFE42FF55" bold="true" box="[151,479,151,178]" fontSize="11" level="1" pageId="16" pageNumber="215" reason="1">
<emphasis id="7A96EACBFFA1FF5A30ABFF1DFE42FF55" bold="true" box="[151,479,151,178]" pageId="16" pageNumber="215">
<taxonomicName id="8FE24D5AFFA1FF5A30ABFF1DFEE7FF56" ID-CoL="85LCS" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[151,378,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA1FF5A30ABFF1DFEE7FF56" bold="true" box="[151,378,151,177]" italics="true" pageId="16" pageNumber="215">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA1FF5A31BDFF12FE42FF55" box="[385,479,152,178]" pageId="16" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="485D36D9FFA1FF5A30ABFF30FEB1FF34" blockId="16.[151,479,151,211]" box="[151,300,186,211]" pageId="16" pageNumber="215">
(
<figureCitation id="D0D92A5CFFA1FF5A30A3FF30FEB9FF34" box="[159,292,186,211]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStartId-0="17.[151,250,1741,1763]" captionStartId-1="18.[151,250,1594,1616]" captionStartId-2="19.[151,250,1669,1691]" captionStartId-3="20.[151,250,1462,1484]" captionStartId-4="21.[151,250,1906,1928]" captionTargetBox-0="[165,1425,206,1712]" captionTargetBox-1="[167,1421,196,1558]" captionTargetBox-2="[168,1404,406,1642]" captionTargetBox-3="[158,1427,453,1428]" captionTargetBox-4="[175,1376,193,1855]" captionTargetId-0="figure@17.[151,1436,193,1720]" captionTargetId-1="figure@18.[151,1436,183,1572]" captionTargetId-2="figure@19.[157,1421,401,1658]" captionTargetId-3="figure@20.[151,1436,445,1441]" captionTargetId-4="figure@21.[175,1376,193,1855]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionTargetPageId-4="21" captionText-0="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." captionText-1="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." captionText-2="FIGURE 13. Alpheopsis gotrina sp. nov., male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021) [a – c, e – h]; male (cl 5.2 mm) from Los Negros Islands, Bahía Málaga, Pacific coast of Colombia (INV CRU 8407) [d]: a—carapace and anterior pleon, lateral; b—anterior region of carapace and frontal appendages, dorsal; c—telson, dorsal; d—antennal scaphocerite, dorsal, e—second pereiopod, lateral; f—third pereiopod, lateral; g—fifth pereiopod, lateral; h—second pleopod, lateral; i—uropod, dorsal." captionText-3="FIGURE 14. Alpheopsis gotrina sp. nov., male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021); a—left (minor) cheliped, mesial; b—same, lateral; c—same, chela, mesial; d—right (major) cheliped, lateral; e—same, chela, lateral; f—same, chela fingers, lateral." captionText-4="FIGURE 15. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507) [a]; male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021) [b]; specimen (sex and cl unknown) from Isla Tortuga, Pacific coast of Costa Rica (specimen collected but possibly lost, see text, not examined) [c]. Photographs: A. Anker (a, b) and courtesy of Ingo S. Wehrtmann (c)." httpUri-0="https://zenodo.org/record/809656/files/figure.png" httpUri-1="https://zenodo.org/record/809658/files/figure.png" httpUri-2="https://zenodo.org/record/809660/files/figure.png" httpUri-3="https://zenodo.org/record/809662/files/figure.png" httpUri-4="https://zenodo.org/record/809664/files/figure.png" pageId="16" pageNumber="215">Figs. 11–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFA1FF5A30ABFE88FE7FFDDC" pageId="16" pageNumber="215" type="materials_examined">
<paragraph id="485D36D9FFA1FF5A30ABFE88FDB2FE84" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
<materialsCitation id="F88A3C84FFA1FF5A30ABFE88FDB2FE84" ID-GBIF-Occurrence="1563896383" collectingDate="2007-04-18" collectionCode="MZUSP" collectorName="Pacific &amp; Playa Venao &amp; A. Anker &amp; I. N. Marin" country="Panama" location="Panama Canal" pageId="16" pageNumber="215" specimenCode="MZUSP 34507" specimenCount="2" typeStatus="holotype">
<emphasis id="7A96EACBFFA1FF5A30ABFE88FEDCFEFC" bold="true" box="[151,321,258,283]" pageId="16" pageNumber="215">
<typeStatus id="9759887BFFA1FF5A30ABFE88FF4FFEFC" box="[151,210,258,283]" pageId="16" pageNumber="215">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="9759887BFFA1FF5A3168FE88FE5CFEFC" box="[340,449,258,283]" pageId="16" pageNumber="215" type="holotype">Holotype</typeStatus>
: ovigerous female (cl 7.0 mm),
<specimenCode id="18449EA2FFA1FF5A3301FE89FC72FEFC" box="[829,1007,258,283]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="16" pageNumber="215">
<collectionCode id="2EF3AE1CFFA1FF5A3301FE89FC07FEFC" box="[829,922,259,283]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="16" pageNumber="215">MZUSP</collectionCode>
34507
</specimenCode>
,
<collectingCountry id="30F57649FFA1FF5A33C2FE89FBC7FEFD" box="[1022,1114,259,282]" name="Panama" pageId="16" pageNumber="215">Panama</collectingCountry>
,
<collectorName id="E517530FFFA1FF5A3454FE88FB2BFEFC" box="[1128,1206,258,283]" pageId="16" pageNumber="215">Pacific</collectorName>
coast,
<collectorName id="E517530FFFA1FF5A3531FE88FA01FEFD" box="[1293,1436,258,283]" pageId="16" pageNumber="215">Playa Venao</collectorName>
(west of
<location id="4D3D6002FFA1FF5A30C2FEADFE3BFED8" LSID="urn:lsid:plazi:treatment:C04B87CFFFA1FF5C30ABFF1DFD03FAB2:4D3D6002FFA1FF5A30C2FEADFE3BFED8" box="[254,422,294,319]" country="Panama" name="Panama Canal" pageId="16" pageNumber="215">Panama Canal</location>
), rocky intertidal exposed at low tide, under large rocks on sand, coll.
<collectorName id="E517530FFFA1FF5A34E0FEADFADAFED8" box="[1244,1351,294,319]" pageId="16" pageNumber="215">A. Anker</collectorName>
&amp;
<collectorName id="E517530FFFA1FF5A354DFEADFF43FE84" pageId="16" pageNumber="215">I.N. Marin</collectorName>
,
<collectingDate id="2C18E9F1FFA1FF5A30D5FEC1FE1BFE84" box="[233,390,330,355]" pageId="16" pageNumber="215" value="2007-04-18">
<date id="3C5C1019FFA1FF5A30D5FEC1FE1BFE84" box="[233,390,330,355]" pageId="16" pageNumber="215" value="2007-04-18">18 April 2007</date>
</collectingDate>
[fcn 07-107b].
</materialsCitation>
</paragraph>
<paragraph id="485D36D9FFA1FF5A30FBFEE4FE7FFDDC" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
<materialsCitation id="F88A3C84FFA1FF5A30FBFEE4FB6AFE28" ID-GBIF-Occurrence="1563896402" collectingDate="2009-04-25" collectionCode="INV, CRU, COL" collectorName="J. F. Lazarus" country="Colombia" elevation="2" latitude="3.9896388" location="Los Negros Islands" longLatPrecision="1" longitude="-77.29628" municipality="Pacific" pageId="16" pageNumber="215" specimenCode="CRU8407" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<emphasis id="7A96EACBFFA1FF5A30FBFEE4FDABFE60" bold="true" box="[199,566,366,391]" pageId="16" pageNumber="215">Additional material examined</emphasis>
.
<specimenCount id="5EE4FD50FFA1FF5A3275FEE5FD05FE60" box="[585,664,366,391]" pageId="16" pageNumber="215" type="male">1 male</specimenCount>
(cl
<quantity id="8F1A9B3CFFA1FF5A32F1FEE4FCB7FE61" box="[717,810,366,391]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="16" pageNumber="215" unit="mm" value="5.2">5.2 mm</quantity>
, missing major cheliped),
<collectionCode id="2EF3AE1CFFA1FF5A3459FEE5FB0AFE61" box="[1125,1175,367,390]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="16" pageNumber="215">INV</collectionCode>
<specimenCode id="18449EA2FFA1FF5A349EFEE5FA8AFE60" box="[1186,1303,367,391]" collectionCode="CRU" pageId="16" pageNumber="215">
<collectionCode id="2EF3AE1CFFA1FF5A349EFEE5FB43FE60" box="[1186,1246,367,391]" pageId="16" pageNumber="215">CRU</collectionCode>
8407
</specimenCode>
,
<collectingCountry id="30F57649FFA1FF5A351BFEE4FA05FE60" box="[1319,1432,366,391]" name="Colombia" pageId="16" pageNumber="215">Colombia</collectingCountry>
,
<collectingMunicipality id="A839ACA3FFA1FF5A30ABFE18FF78FE4C" box="[151,229,402,427]" pageId="16" pageNumber="215">Pacific</collectingMunicipality>
coast,
<location id="4D3D6002FFA1FF5A3106FE19FD85FE4C" LSID="urn:lsid:plazi:treatment:C04B87CFFFA1FF5C30ABFF1DFD03FAB2:4D3D6002FFA1FF5A3106FE19FD85FE4C" box="[314,536,402,427]" country="Colombia" latitude="3.9896388" longLatPrecision="1" longitude="-77.29628" municipality="Pacific" name="Los Negros Islands" pageId="16" pageNumber="215" stateProvince="Bahia">Los Negros Islands</location>
near entrance to
<collectingRegion id="8A26F83BFFA1FF5A32DAFE18FCBAFE4C" box="[742,807,402,427]" country="Brazil" name="Bahia" pageId="16" pageNumber="215">Bahía</collectingRegion>
Málaga,
<geoCoordinate id="2DD6501EFFA1FF5A33ABFE18FBA4FE4C" box="[919,1081,402,427]" direction="north" orientation="latitude" pageId="16" pageNumber="215" precision="1" value="3.9896388">03°59’22.7’’N</geoCoordinate>
<geoCoordinate id="2DD6501EFFA1FF5A347FFE18FB76FE4C" box="[1091,1259,402,427]" direction="west" orientation="longitude" pageId="16" pageNumber="215" precision="1" value="-77.29628">77°17’46.6’’W</geoCoordinate>
, sand and rock bottom, artificial reef substrate, depth:
<elevation id="C3CFD1EAFFA1FF5A3277FE3DFDEAFE29" box="[587,631,439,463]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="215" unit="m" value="2.0">
<quantity id="8F1A9B3CFFA1FF5A3277FE3DFDEAFE29" box="[587,631,439,463]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="215" unit="m" value="2.0">2 m</quantity>
</elevation>
, coll.
<collectorName id="E517530FFFA1FF5A3287FE3DFCDDFE29" box="[699,832,439,462]" pageId="16" pageNumber="215">J.F. Lazarus</collectorName>
<emphasis id="7A96EACBFFA1FF5A3374FE3DFCE4FE29" box="[840,889,438,462]" italics="true" pageId="16" pageNumber="215">et al</emphasis>
.,
<collectingDate id="2C18E9F1FFA1FF5A33B1FE3CFBB6FE28" box="[909,1067,438,463]" pageId="16" pageNumber="215" value="2009-04-25">
<date id="3C5C1019FFA1FF5A33B1FE3CFBB6FE28" box="[909,1067,438,463]" pageId="16" pageNumber="215" value="2009-04-25">25 April 2009</date>
</collectingDate>
[fcn
<collectionCode id="2EF3AE1CFFA1FF5A3459FE3DFB00FE28" box="[1125,1181,439,463]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="16" pageNumber="215">COL</collectionCode>
00147]
</materialsCitation>
;
<materialsCitation id="F88A3C84FFA1FF5A353EFE3DFE43FDDC" ID-GBIF-Occurrence="1563896405" collectingDate="2009-04-26" collectionCode="OUMNH, COL" collectorName="J. F. Lazarus" country="Colombia" elevation="7" latitude="3.994111" location="Curichichi" longLatPrecision="1" longitude="-77.31253" municipality="Pacific" pageId="16" pageNumber="215" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<specimenCount id="5EE4FD50FFA1FF5A353EFE3DFAD0FE28" box="[1282,1357,438,463]" pageId="16" pageNumber="215" type="male">1 male</specimenCount>
(cl
<quantity id="8F1A9B3CFFA1FF5A3545FE3DFF5AFE14" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="215" unit="mm" value="4.8">4.8 mm</quantity>
),
<collectionCode id="2EF3AE1CFFA1FF5A30D7FE51FEC6FE14" box="[235,347,475,499]" pageId="16" pageNumber="215">OUMNH</collectionCode>
.ZC. 2017.01.0 21,
<collectingCountry id="30F57649FFA1FF5A3276FE50FD26FE14" box="[586,699,474,499]" name="Colombia" pageId="16" pageNumber="215">Colombia</collectingCountry>
,
<collectingMunicipality id="A839ACA3FFA1FF5A32E4FE50FCBBFE14" box="[728,806,474,499]" pageId="16" pageNumber="215">Pacific</collectingMunicipality>
coast,
<collectingRegion id="8A26F83BFFA1FF5A33ABFE50FC45FE14" box="[919,984,474,499]" country="Brazil" name="Bahia" pageId="16" pageNumber="215">Bahía</collectingRegion>
Málaga,
<location id="4D3D6002FFA1FF5A3458FE50FB40FE14" LSID="urn:lsid:plazi:treatment:C04B87CFFFA1FF5C30ABFF1DFD03FAB2:4D3D6002FFA1FF5A3458FE50FB40FE14" box="[1124,1245,474,499]" country="Colombia" latitude="3.994111" longLatPrecision="1" longitude="-77.31253" municipality="Pacific" name="Curichichi" pageId="16" pageNumber="215" stateProvince="Bahia">Curichichi</location>
,
<geoCoordinate id="2DD6501EFFA1FF5A34C6FE50FA01FE14" box="[1274,1436,474,499]" direction="north" orientation="latitude" pageId="16" pageNumber="215" precision="1" value="3.994111">03°59’38.8’’N</geoCoordinate>
<geoCoordinate id="2DD6501EFFA1FF5A30ABFE74FEA2FDF0" box="[151,319,510,535]" direction="west" orientation="longitude" pageId="16" pageNumber="215" precision="1" value="-77.31253">77°18’45.1’’W</geoCoordinate>
, mud and silt bottom with rocks, artificial reef substrate, depth:
<elevation id="C3CFD1EAFFA1FF5A3430FE74FBF7FDF1" box="[1036,1130,510,535]" metricMagnitude="0" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="16" pageNumber="215" unit="m" value="6.75" valueMax="7.0" valueMin="6.5">
<quantity id="8F1A9B3CFFA1FF5A3430FE74FBF7FDF1" box="[1036,1130,510,535]" metricMagnitude="0" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="16" pageNumber="215" unit="m" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 m</quantity>
</elevation>
, coll.
<collectorName id="E517530FFFA1FF5A3492FE75FAA9FDF1" box="[1198,1332,511,534]" pageId="16" pageNumber="215">J.F. Lazarus</collectorName>
<emphasis id="7A96EACBFFA1FF5A3506FE75FAF6FDF1" box="[1338,1387,510,534]" italics="true" pageId="16" pageNumber="215">et al</emphasis>
.,
<collectingDate id="2C18E9F1FFA1FF5A35BCFE74FE8CFDDC" pageId="16" pageNumber="215" value="2009-04-26">
<date id="3C5C1019FFA1FF5A35BCFE74FE8CFDDC" pageId="16" pageNumber="215" value="2009-04-26">26 April 2009</date>
</collectingDate>
[fcn
<collectionCode id="2EF3AE1CFFA1FF5A3170FDA9FE19FDDC" box="[332,388,547,571]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="16" pageNumber="215">COL</collectionCode>
00188]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFA1FF5E30FBFDCCFE32F831" lastPageId="20" lastPageNumber="219" pageId="16" pageNumber="215" type="description">
<paragraph id="485D36D9FFA1FF5A30FBFDCCFAB2FC0C" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
<emphasis id="7A96EACBFFA1FF5A30FBFDCCFECCFDB8" bold="true" box="[199,337,582,607]" pageId="16" pageNumber="215">Description</emphasis>
. Medium-sized species of
<taxonomicName id="8FE24D5AFFA1FF5A3242FDCCFD64FDB9" authorityName="Coutiere" authorityYear="1897" box="[638,761,582,606]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="7A96EACBFFA1FF5A3242FDCCFD64FDB9" box="[638,761,582,606]" italics="true" pageId="16" pageNumber="215">Alpheopsis</emphasis>
</taxonomicName>
(present material: maximum cl 7.0 mm). Carapace with one strong MDC extending either from rostral tip to almost posterior margin of carapace (non-type specimens), or interrupted by wide gap between approximately 0.2 and 0.4 carapace length, this gap bearing very short low transverse crescent-shaped ridges converging to mid-dorsal line (
<typeStatus id="9759887BFFA1FF5A334AFD38FC43FD2C" box="[886,990,690,715]" pageId="16" pageNumber="215" type="holotype">holotype</typeStatus>
); each side of carapace with one strong ASC running in parallel and lateral to MDC, starting at frontal margin of carapace and abruptly ending at about 0.7 length of carapace; one PSC running in parallel and lateral to MDC, starting posterior to ASC, with anterior end extending ventrolaterally as fairly well-marked CSR; one DLC running slightly obliquely from postorbital area to about 0.6 carapace length, stopping shortly before CSR; one VLC running ventral and in parallel to DLC, starting well anterior to DLC and either reaching or falling short of carapace mid-length; one PLC ventrolateral to PSC, somewhat oblique, with posterior end adjacent to cardiac notch, sometimes very short (
<typeStatus id="9759887BFFA1FF5A3444FC00FB7DFC44" box="[1144,1248,906,931]" pageId="16" pageNumber="215" type="holotype">holotype</typeStatus>
); and one strong ABC running along and somewhat above branchiostegial margin to at most 0.3 carapace length, with stout subacute tooth at anterior end, latter protruding well beyond anterolateral margin (
<figureCitation id="D0D92A5CFFA1FF5A340CFC58FB14FC0C" box="[1072,1161,978,1003]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
a–c, e; 13a, b).
</paragraph>
<paragraph id="485D36D9FFA1FF5A30FBFC7DFEF5FB78" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
Rostrum well developed, up to twice as long as wide at base, with strong dorsal carina conspicuously sloping shortly before reaching acute tip, latter overreaching distal margin of first article of antennular peduncle, sometimes reaching mid-length of second article (
<figureCitation id="D0D92A5CFFA1FF5A32F9FBB4FC83FBB0" box="[709,798,1086,1111]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
a–e; 13a, b). Orbital hoods protruding anteriorly as broad rounded-angular orbital teeth (
<figureCitation id="D0D92A5CFFA1FF5A31D0FBE8FDDBFB9C" box="[492,582,1122,1147]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
c, e; 13b). Pterygostomial angle below strong tooth of ABC broadly rounded (
<figureCitation id="D0D92A5CFFA1FF5A30A3FB0CFF64FB78" box="[159,249,1158,1183]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
a, c; 13a).
</paragraph>
<paragraph id="485D36D9FFA1FF5A30FBFB20FE6BFA58" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
First pleomere with strong MDC-PL1 continuing MDC of carapace; second to sixth pleomeres unarmed dorsally; all pleura rounded ventrally; fifth pleuron with posteroventral angle rounded; sixth somite with bluntly projecting posterior lobe and well-delimited, triangular, articulated plate (
<figureCitation id="D0D92A5CFFA1FF5A33D1FB78FBD6FAEC" box="[1005,1099,1266,1291]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
a, f; 13a). Telson moderately broad, tapering distally, about 2.3 as long as anterior width; dorsal surface with median longitudinal depression and two pairs of stout spiniform setae inserted far from lateral margin, at about 0.5 and 0.7 telson length, respectively; posterior margin about half as long as anterior margin, broadly rounded; each posterolateral angle with one pair of stout spiniform setae, mesial much longer than lateral (
<figureCitation id="D0D92A5CFFA1FF5A331CFA08FC1DFA7C" box="[800,896,1410,1435]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">
Figs.
<quantity id="8F1A9B3CFFA1FF5A3358FA09FC1DFA7C" box="[868,896,1411,1435]" metricMagnitude="-2" metricUnit="kg" metricValue="1.1" pageId="16" pageNumber="215" unit="g" value="11.0">11</quantity>
</figureCitation>
g; 13c, note: telson of
<typeStatus id="9759887BFFA1FF5A34B2FA08FB6DFA7C" box="[1166,1264,1410,1435]" pageId="16" pageNumber="215" type="holotype">holotype</typeStatus>
with posterior margin damaged on right side).
</paragraph>
<paragraph id="485D36D9FFA1FF5A30FBFA41FBEAF9E1" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
Eyes completely concealed by orbital hoods, not visible in dorsal and lateral view; eyestalk with wellpigmented cornea (
<figureCitation id="D0D92A5CFFA1FF5A3152FA64FE23F9E0" box="[366,446,1518,1543]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Fig. 11</figureCitation>
c, e; 13b). Epistomial sclerites each with strong sharp process.
</paragraph>
<paragraph id="485D36D9FFA1FF5A30FBF998FAE7F8E4" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
Antennular peduncles stout; stylocerite strong, with sharp tip greatly overreaching distal margin of second article, sometimes reaching almost to end of third article; ventromesial carina with anteriorly directed, acute tooth; second article about 1.2 times as long as wide; lateral flagellum biramous; fused portion short, with three or four joints; accessory ramus well developed, with several groups of long aesthetascs (
<figureCitation id="D0D92A5CFFA1FF5A3425F9F4FBEEF970" box="[1049,1139,1662,1687]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
c, e, h; 13b). Antenna with basicerite ending in stout sharp tooth distoventrally; scaphocerite ovate, with anteriorly rounded blade and sharp distolateral tooth, latter reaching beyond anterior margin of blade; anterior margin of scaphocerite not or only slightly exceeding distal margin of antennular peduncles; carpocerite reaching to end of scaphocerite (
<figureCitation id="D0D92A5CFFA1FF5A34A6F960FB6FF8E4" box="[1178,1266,1770,1795]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">Figs. 11</figureCitation>
c, e; 13b, d).
</paragraph>
<paragraph id="485D36D9FFA1FF5A30FBF884FACFF888" blockId="16.[151,1437,258,1975]" pageId="16" pageNumber="215">
Mouthparts typical for genus in external view. Third maxilliped slender, pediform; coxa with distally subacute lateral plate above mastigobranch; penultimate article about 2.5 times as long as wide proximally; ultimate article tapering to slender corneous tip, with several thick subdistal setae; arthrobranch well developed (
<figureCitation id="D0D92A5CFFA1FF5A34E9F8DCFAB8F888" box="[1237,1317,1878,1903]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="16" pageNumber="215">
Fig.
<date id="3C5C1019FFA1FF5A3535F8DCFAB8F888" box="[1289,1317,1878,1903]" pageId="16" pageNumber="215" value="2011-01">11</date>
</figureCitation>
i, j).
</paragraph>
<paragraph id="485D36D9FFA1FF5930FBF8F0FC39FE82" blockId="16.[151,1437,258,1975]" lastBlockId="19.[151,1436,151,357]" lastPageId="19" lastPageNumber="218" pageId="16" pageNumber="215">
Chelipeds subequal or slightly unequal in size and asymmetrical in shape and armature on finger cutting edges (at least in adult males), carried extended when not in use (
<figureCitation id="D0D92A5CFFA1FF5A3317F814FC1BF850" box="[811,902,1950,1975]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="16" pageNumber="215">Figs. 12</figureCitation>
a–d; 14a–f; 15a, b). Ischium stout, short, with stout spiniform seta distodorsally (
<figureCitation id="D0D92A5CFFA3FF583224F93EFDEEF92A" box="[536,627,1716,1741]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="18" pageNumber="217">Figs. 12</figureCitation>
a–c; 14a, b, d). Merus almost as long as wide in males, noticeably longer than wide in females, broadening distally, triangular in cross-section, with three well-defined margins, dorsal margin ending in very conspicuous blunt tooth, proximodorsal margin with stout spiniform seta (
<figureCitation id="D0D92A5CFFA3FF5834E7F976FAABF8F2" box="[1243,1334,1788,1813]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="18" pageNumber="217">Figs. 12</figureCitation>
a–c; 14a, b, d). Carpus very stout, short, cup-shaped, widening distally, with deep dorsal constriction, ventral margin without stiff seta (
<figureCitation id="D0D92A5CFFA3FF583138F8CEFEC2F8BA" box="[260,351,1860,1885]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="18" pageNumber="217">Figs. 12</figureCitation>
a–c; 14a, b, d). Male major chela robust, with palm conspicuously flattened on mesial surface and fingers about 0.6 length of palm; dorsomesial-proximal surface of palm with strong, usually somewhat dorsally and/or anteriorly projecting, distally curved process; fingers somewhat gaping, crossing distally; cutting edges of fingers with few strong, rounded or subtriangular teeth, three on dactylus and four on pollex, distal-most teeth very large, especially on pollex; adhesive disks moderately developed (
<figureCitation id="D0D92A5CFFA3FF583391F85EFB9EF80A" box="[941,1027,2004,2029]" captionStart="FIGURE 14" captionStartId="20.[151,250,1462,1484]" captionTargetBox="[158,1427,453,1428]" captionTargetId="figure@20.[151,1436,445,1441]" captionTargetPageId="20" captionText="FIGURE 14. Alpheopsis gotrina sp. nov., male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021); a—left (minor) cheliped, mesial; b—same, lateral; c—same, chela, mesial; d—right (major) cheliped, lateral; e—same, chela, lateral; f—same, chela fingers, lateral." httpUri="https://zenodo.org/record/809662/files/figure.png" pageId="18" pageNumber="217">Fig. 14</figureCitation>
d–f). Male minor chela with palm flattened on mesial surface and fingers about 0.9 length of palm; dorsomesial-proximal surface of palm with strong, dorsally and/or anteriorly projecting, distally curved process, latter similar to that of major chela; dactylus distinctly flattened and somewhat expanded dorsally; cutting edges of fingers finely and irregularly dentate (
<figureCitation id="D0D92A5CFFA2FF593553FF55FF2EFEFB" captionStart="FIGURE 14" captionStartId="20.[151,250,1462,1484]" captionTargetBox="[158,1427,453,1428]" captionTargetId="figure@20.[151,1436,445,1441]" captionTargetPageId="20" captionText="FIGURE 14. Alpheopsis gotrina sp. nov., male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021); a—left (minor) cheliped, mesial; b—same, lateral; c—same, chela, mesial; d—right (major) cheliped, lateral; e—same, chela, lateral; f—same, chela fingers, lateral." httpUri="https://zenodo.org/record/809662/files/figure.png" pageId="19" pageNumber="218">Fig. 14</figureCitation>
a–c). Female minor chela (or one of chelae) generally similar to male minor chela, less robust, with slenderer fingers; dorsomesial-proximal surface of palm with strongly projecting, distally curved process as in males; cutting edges of fingers armed with small spaced irregular teeth (
<figureCitation id="D0D92A5CFFA2FF593326FEC6FCF6FE82" box="[794,875,332,357]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="19" pageNumber="218">Fig. 12</figureCitation>
a–d).
</paragraph>
<caption id="1C9D6651FFA0FF5B30ABF947FD1AF8B9" httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="17" pageNumber="216" targetBox="[165,1425,206,1712]" targetPageId="17">
<paragraph id="485D36D9FFA0FF5B30ABF947FD1AF8B9" blockId="17.[151,1436,1741,1886]" pageId="17" pageNumber="216">
<emphasis id="7A96EACBFFA0FF5B30ABF947FEBDF905" bold="true" box="[151,288,1741,1763]" pageId="17" pageNumber="216">FIGURE 11.</emphasis>
<taxonomicName id="8FE24D5AFFA0FF5B3114F947FE77F904" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[296,490,1741,1763]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="216" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA0FF5B3114F947FE77F904" box="[296,490,1741,1763]" italics="true" pageId="17" pageNumber="216">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA0FF5B31CEF944FDDFF904" box="[498,578,1742,1763]" pageId="17" pageNumber="216" rank="species">
<emphasis id="7A96EACBFFA0FF5B31CEF944FDDFF904" bold="true" box="[498,578,1742,1763]" pageId="17" pageNumber="216">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal.
</paragraph>
</caption>
<caption id="1C9D6651FFA3FF5830ABF9B0FAC0F96A" httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="18" pageNumber="217" targetBox="[167,1421,196,1558]" targetPageId="18">
<paragraph id="485D36D9FFA3FF5830ABF9B0FAC0F96A" blockId="18.[151,1436,1594,1677]" pageId="18" pageNumber="217">
<emphasis id="7A96EACBFFA3FF5830ABF9B0FEBCF9A8" bold="true" box="[151,289,1594,1616]" pageId="18" pageNumber="217">FIGURE 12.</emphasis>
<taxonomicName id="8FE24D5AFFA3FF583115F9B0FE76F9B7" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[297,491,1594,1616]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="217" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA3FF583115F9B0FE76F9B7" box="[297,491,1594,1616]" italics="true" pageId="18" pageNumber="217">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA3FF5831CFF9B0FDDEF9A8" box="[499,579,1594,1615]" pageId="18" pageNumber="217" rank="species">
<emphasis id="7A96EACBFFA3FF5831CFF9B0FDDEF9A8" bold="true" box="[499,579,1594,1615]" pageId="18" pageNumber="217">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral.
</paragraph>
</caption>
<caption id="1C9D6651FFA2FF5930ABF90FFCB1F8F2" httpUri="https://zenodo.org/record/809660/files/figure.png" pageId="19" pageNumber="218" targetBox="[168,1404,406,1642]" targetPageId="19">
<paragraph id="485D36D9FFA2FF5930ABF90FFCB1F8F2" blockId="19.[151,1436,1668,1813]" pageId="19" pageNumber="218">
<emphasis id="7A96EACBFFA2FF5930ABF90FFEB8F97D" bold="true" box="[151,293,1669,1691]" pageId="19" pageNumber="218">FIGURE 13.</emphasis>
<taxonomicName id="8FE24D5AFFA2FF59310DF90EFE6AF97C" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[305,503,1668,1691]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="218" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA2FF59310DF90EFE6AF97C" box="[305,503,1668,1691]" italics="true" pageId="19" pageNumber="218">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA2FF59323EF90FFDCEF97D" box="[514,595,1669,1690]" pageId="19" pageNumber="218" rank="species">
<emphasis id="7A96EACBFFA2FF59323EF90FFDCEF97D" bold="true" box="[514,595,1669,1690]" pageId="19" pageNumber="218">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH.ZC. 2017.01.021) [a–c, e–h]; male (cl 5.2 mm) from Los Negros Islands, Bahía Málaga, Pacific coast of Colombia (INV CRU8407) [d]: a—carapace and anterior pleon, lateral; b—anterior region of carapace and frontal appendages, dorsal; c—telson, dorsal; d—antennal scaphocerite, dorsal, e—second pereiopod, lateral; f—third pereiopod, lateral; g—fifth pereiopod, lateral; h—second pleopod, lateral; i—uropod, dorsal.
</paragraph>
</caption>
<paragraph id="485D36D9FFA2FF5E30FBF8B6FCACFE6F" blockId="19.[151,1437,1852,2022]" lastBlockId="20.[151,1436,151,392]" lastPageId="20" lastPageNumber="219" pageId="19" pageNumber="218">
Second pereiopod moderately slender; ischium shorter than merus; carpus five-jointed; ratio of carpal joints (from proximal to distal) approximately equal to: 3: 1.2: 1: 1: 2; chela longer than distal carpal article, with fingers equal to palm in length, simple (
<figureCitation id="D0D92A5CFFA2FF593256F80FFD57F87A" box="[618,714,1925,1950]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="19" pageNumber="218">Figs. 12</figureCitation>
e; 13e). Third pereiopod moderately slender; ischium with one stout spiniform setae on ventrolateral surface; merus about 4.8–5.1 times as long as wide, unarmed; carpus about half-length of merus, much slenderer, without or with minute seta on distoventral margin; propodus with three spiniform setae along ventral margin and pair of distal spiniform setae adjacent to dactylus; dactylus about 0.4 length of propodus, slender, simple, conical, gradually curving distally (
<figureCitation id="D0D92A5CFFA5FF5E33CDFF36FBCFFF32" box="[1009,1106,188,213]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="20" pageNumber="219">Figs. 12</figureCitation>
f, g; 13f). Fourth pereiopod generally similar to third pereiopod. Fifth pereiopod more slender than third and fourth pereiopods; ischium unarmed on ventrolateral surface; merus about five times as long as wide, unarmed; carpus about 0.7 length of merus; propodus long, slender, with at least four spiniform setae along ventral margin, one long distal spiniform setae adjacent to dactylus, and several rows of serrulate setae on distolateral surface; dactylus about 0.45 length of propodus, similar to that of third pereiopod (
<figureCitation id="D0D92A5CFFA5FF5E32B5FEE5FD46FE6F" box="[649,731,367,392]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="20" pageNumber="219">Fig. 12</figureCitation>
h;
<quantity id="8F1A9B3CFFA5FF5E32CAFEFAFCB9FE6F" box="[758,804,368,392]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="20" pageNumber="219" unit="g" value="13.0">13g</quantity>
).
</paragraph>
<caption id="1C9D6651FFA5FF5E30ABFA3CFCECF9EE" httpUri="https://zenodo.org/record/809662/files/figure.png" pageId="20" pageNumber="219" targetBox="[158,1427,453,1428]" targetPageId="20">
<paragraph id="485D36D9FFA5FF5E30ABFA3CFCECF9EE" blockId="20.[151,1436,1462,1545]" pageId="20" pageNumber="219">
<emphasis id="7A96EACBFFA5FF5E30ABFA3CFEB8FA2C" bold="true" box="[151,293,1462,1484]" pageId="20" pageNumber="219">FIGURE 14.</emphasis>
<taxonomicName id="8FE24D5AFFA5FF5E310DFA3CFE6AFA2B" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[305,503,1462,1484]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="219" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA5FF5E310DFA3CFE6AFA2B" box="[305,503,1462,1484]" italics="true" pageId="20" pageNumber="219">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA5FF5E323EFA3CFDCEFA2C" box="[514,595,1462,1483]" pageId="20" pageNumber="219" rank="species">
<emphasis id="7A96EACBFFA5FF5E323EFA3CFDCEFA2C" bold="true" box="[514,595,1462,1483]" pageId="20" pageNumber="219">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH.ZC. 2017.01.021); a—left (minor) cheliped, mesial; b—same, lateral; c—same, chela, mesial; d—right (major) cheliped, lateral; e—same, chela, lateral; f—same, chela fingers, lateral.
</paragraph>
</caption>
<paragraph id="485D36D9FFA5FF5E30FBF9BAFE9EF93E" blockId="20.[151,1437,1584,2006]" pageId="20" pageNumber="219">
Male second pleopod with appendix masculina well developed, as long as appendix interna; appendix masculina with several long stiff setae on apex and along margin. Uropod with lateral lobe of protopod ending in long sharp tooth; endopod and exopod broadly ovate; exopod with distolateral tooth and adjacent spiniform seta both strong; diaeresis with distinct subtriangular tooth adjacent to spiniform seta and sinuous mesial portion (
<figureCitation id="D0D92A5CFFA5FF5E3558F917FF2FF93E" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="20" pageNumber="219">Figs. 11</figureCitation>
k;
<date id="3C5C1019FFA5FF5E30F2F94AFF68F93E" box="[206,245,1728,1753]" pageId="20" pageNumber="219" value="2013-01">13i</date>
).
</paragraph>
<paragraph id="485D36D9FFA5FF5E30FBF96FFD6DF919" blockId="20.[151,1437,1584,2006]" box="[199,752,1765,1790]" pageId="20" pageNumber="219">
Gill-exopod formula as in
<taxonomicName id="8FE24D5AFFA5FF5E31CCF96FFDFFF91A" authorityName="in Soledade et al." authorityYear="2015" box="[496,610,1765,1789]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="219" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA5FF5E31CCF96FFDFFF91A" box="[496,610,1765,1789]" italics="true" pageId="20" pageNumber="219">A. trigona</emphasis>
</taxonomicName>
(see above).
</paragraph>
<paragraph id="485D36D9FFA5FF5E30FBF882FE32F831" blockId="20.[151,1437,1584,2006]" pageId="20" pageNumber="219">
<emphasis id="7A96EACBFFA5FF5E30FBF882FE18F8C6" bold="true" box="[199,389,1800,1825]" pageId="20" pageNumber="219">Colour pattern</emphasis>
. Background semitransparent whitish or pale yellowish; carapace covered with red chromatophores, except for anterior-most dorsal and posterior-most lateral portions; pleon with two broad adjacent transverse bands of red chromatophores running across posterior half of third pleomere and most of fourth pleomere (red bands either very intense or faded, depending on contraction state of chromatophores and shrimp’s physiological condition); antennular and antennal flagella pale orange; chelipeds hyaline white; walking legs and tail fan whitish (
<figureCitation id="D0D92A5CFFA5FF5E3172F837FE3FF831" box="[334,418,1981,2006]" captionStart="FIGURE 15" captionStartId="21.[151,250,1906,1928]" captionTargetBox="[175,1376,193,1855]" captionTargetId="figure@21.[175,1376,193,1855]" captionTargetPageId="21" captionText="FIGURE 15. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507) [a]; male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021) [b]; specimen (sex and cl unknown) from Isla Tortuga, Pacific coast of Costa Rica (specimen collected but possibly lost, see text, not examined) [c]. Photographs: A. Anker (a, b) and courtesy of Ingo S. Wehrtmann (c)." httpUri="https://zenodo.org/record/809664/files/figure.png" pageId="20" pageNumber="219">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="1C9D6651FFA4FF5F30ABF8F8FB8EF803" httpUri="https://zenodo.org/record/809664/files/figure.png" pageId="21" pageNumber="220" targetBox="[175,1376,193,1855]" targetPageId="21">
<paragraph id="485D36D9FFA4FF5F30ABF8F8FB8EF803" blockId="21.[151,1436,1906,2020]" pageId="21" pageNumber="220">
<emphasis id="7A96EACBFFA4FF5F30ABF8F8FEBCF860" bold="true" box="[151,289,1906,1928]" pageId="21" pageNumber="220">FIGURE 15.</emphasis>
<taxonomicName id="8FE24D5AFFA4FF5F3115F8F8FE76F86F" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[297,491,1906,1928]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="220" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA4FF5F3115F8F8FE76F86F" box="[297,491,1906,1928]" italics="true" pageId="21" pageNumber="220">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA4FF5F31CFF8F9FDDEF86F" box="[499,579,1907,1928]" pageId="21" pageNumber="220" rank="species">
<emphasis id="7A96EACBFFA4FF5F31CFF8F9FDDEF86F" bold="true" box="[499,579,1907,1928]" pageId="21" pageNumber="220">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507) [a]; male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH.ZC. 2017.01.021) [b]; specimen (sex and cl unknown) from Isla Tortuga, Pacific coast of Costa Rica (specimen collected but possibly lost, see text, not examined) [c]. Photographs: A. Anker (a, b) and courtesy of Ingo S. Wehrtmann (c).
</paragraph>
</caption>
<subSubSection id="00F86552FFA7FF5C30FBFF1DFEE7FF32" pageId="22" pageNumber="221" type="etymology">
<paragraph id="485D36D9FFA7FF5C30FBFF1DFEE7FF32" blockId="22.[151,1437,151,1365]" pageId="22" pageNumber="221">
<emphasis id="7A96EACBFFA7FF5C30FBFF1DFEDBFF57" bold="true" box="[199,326,151,176]" pageId="22" pageNumber="221">Etymology</emphasis>
. The new species’ name is an anagram of the specific epithet of its presumed closest relative,
<taxonomicName id="8FE24D5AFFA7FF5C35B8FF12FF77FF33" authorityName="in Soledade et al." authorityYear="2015" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C35B8FF12FF77FF33" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
(see below).
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFA7FF5C30FBFF55FC84FF10" box="[199,793,223,248]" pageId="22" pageNumber="221" type="materials_examined">
<paragraph id="485D36D9FFA7FF5C30FBFF55FC84FF10" blockId="22.[151,1437,151,1365]" box="[199,793,223,248]" pageId="22" pageNumber="221">
<materialsCitation id="F88A3C84FFA7FF5C30FBFF55FC84FF10" ID-GBIF-Occurrence="1563896392" box="[199,793,223,248]" country="Panama" location="Playa Venao" municipality="Pacific" pageId="22" pageNumber="221" specimenCount="1" typeStatus="holotype">
<emphasis id="7A96EACBFFA7FF5C30FBFF55FEC0FF1F" bold="true" box="[199,349,223,248]" pageId="22" pageNumber="221">
<typeStatus id="9759887BFFA7FF5C30FBFF55FE9FFF1F" box="[199,258,223,248]" pageId="22" pageNumber="221">Type</typeStatus>
locality
</emphasis>
.
<location id="4D3D6002FFA7FF5C3150FF55FE67FF10" LSID="urn:lsid:plazi:treatment:C04B87CFFFA1FF5C30ABFF1DFD03FAB2:4D3D6002FFA7FF5C3150FF55FE67FF10" box="[364,506,223,248]" country="Panama" municipality="Pacific" name="Playa Venao" pageId="22" pageNumber="221">Playa Venao</location>
,
<collectingMunicipality id="A839ACA3FFA7FF5C3239FF55FDCEFF1F" box="[517,595,223,248]" pageId="22" pageNumber="221">Pacific</collectingMunicipality>
coast of
<collectingCountry id="30F57649FFA7FF5C3284FF6AFC89FF10" box="[696,788,224,247]" name="Panama" pageId="22" pageNumber="221">Panama</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFA7FF5C30FBFE8EFB70FE83" pageId="22" pageNumber="221" type="distribution">
<paragraph id="485D36D9FFA7FF5C30FBFE8EFD3FFEA7" blockId="22.[151,1437,151,1365]" pageId="22" pageNumber="221">
<emphasis id="7A96EACBFFA7FF5C30FBFE8EFEC4FEFA" bold="true" box="[199,345,260,285]" pageId="22" pageNumber="221">Distribution</emphasis>
. Eastern Pacific:
<collectingCountry id="30F57649FFA7FF5C3217FE8EFD04FEFA" box="[555,665,260,285]" name="Colombia" pageId="22" pageNumber="221">Colombia</collectingCountry>
(
<collectingRegion id="8A26F83BFFA7FF5C3290FE8EFD72FEFA" box="[684,751,260,285]" country="Brazil" name="Bahia" pageId="22" pageNumber="221">Bahía</collectingRegion>
Málaga),
<collectingRegion id="8A26F83BFFA7FF5C3356FE8FFC5EFEFB" box="[874,963,261,284]" country="Panama" name="Panama" pageId="22" pageNumber="221">
<collectingCountry id="30F57649FFA7FF5C3356FE8FFC5EFEFB" box="[874,963,261,284]" name="Panama" pageId="22" pageNumber="221">Panama</collectingCountry>
</collectingRegion>
(Playa Venao), and possibly
<collectingCountry id="30F57649FFA7FF5C3522FE8EFA01FEFA" box="[1310,1436,260,285]" name="Costa Rica" pageId="22" pageNumber="221">Costa Rica</collectingCountry>
(photographic record, see below and
<figureCitation id="D0D92A5CFFA7FF5C320AFEADFD1AFEA7" box="[566,647,295,320]" captionStart="FIGURE 15" captionStartId="21.[151,250,1906,1928]" captionTargetBox="[175,1376,193,1855]" captionTargetId="figure@21.[175,1376,193,1855]" captionTargetPageId="21" captionText="FIGURE 15. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507) [a]; male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021) [b]; specimen (sex and cl unknown) from Isla Tortuga, Pacific coast of Costa Rica (specimen collected but possibly lost, see text, not examined) [c]. Photographs: A. Anker (a, b) and courtesy of Ingo S. Wehrtmann (c)." httpUri="https://zenodo.org/record/809664/files/figure.png" pageId="22" pageNumber="221">Fig. 15</figureCitation>
c).
</paragraph>
<paragraph id="485D36D9FFA7FF5C30FBFEC6FB70FE83" blockId="22.[151,1437,151,1365]" box="[199,1261,332,357]" pageId="22" pageNumber="221">
<emphasis id="7A96EACBFFA7FF5C30FBFEC6FEB9FE82" bold="true" box="[199,292,332,357]" pageId="22" pageNumber="221">Ecology</emphasis>
. Rocky-sandy shores; under rocks and rubble; intertidal and shallow subtidal (
<quantity id="8F1A9B3CFFA7FF5C34A9FEC6FB7CFE83" box="[1173,1249,332,356]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="22" pageNumber="221" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="00F86552FFA7FF5C30FBFEE5FD03FAB2" pageId="22" pageNumber="221" type="discussion">
<paragraph id="485D36D9FFA7FF5C30FBFEE5FF6DFDF0" blockId="22.[151,1437,151,1365]" pageId="22" pageNumber="221">
<emphasis id="7A96EACBFFA7FF5C30FBFEE5FEA9FE6F" bold="true" box="[199,308,367,392]" pageId="22" pageNumber="221">Remarks</emphasis>
.
<taxonomicName id="8FE24D5AFFA7FF5C317EFEFAFD8AFE60" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[322,535,368,392]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C317EFEFAFD8AFE60" box="[322,535,368,392]" italics="true" pageId="22" pageNumber="221">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C3222FEE5FDE5FE6F" box="[542,632,367,392]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C3222FEE5FDE5FE6F" bold="true" box="[542,632,367,392]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
is closely related to the western Atlantic
<taxonomicName id="8FE24D5AFFA7FF5C3470FEFAFB23FE60" authorityName="in Soledade et al." authorityYear="2015" box="[1100,1214,368,391]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C3470FEFAFB23FE60" box="[1100,1214,368,391]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
and
<taxonomicName id="8FE24D5AFFA7FF5C34C9FEFAFA06FE60" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[1269,1435,368,391]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C34C9FEFAFA06FE60" box="[1269,1435,368,391]" italics="true" pageId="22" pageNumber="221">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C30ABFE1EFF6EFE4A" box="[151,243,404,429]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C30ABFE1EFF6EFE4A" bold="true" box="[151,243,404,429]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
, especially to the former species based on the shared presence of well-developed DLC and VLC and a strong curved proximal process on the cheliped palm, both features being absent in the latter species. On the other hand,
<taxonomicName id="8FE24D5AFFA7FF5C30E0FE56FED2FE13" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[220,335,476,500]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C30E0FE56FED2FE13" box="[220,335,476,500]" italics="true" pageId="22" pageNumber="221">A. gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C316BFE56FE2FFE12" box="[343,434,476,501]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C316BFE56FE2FFE12" bold="true" box="[343,434,476,501]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
shares with
<taxonomicName id="8FE24D5AFFA7FF5C327EFE56FD74FE13" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[578,745,476,500]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C327EFE56FD74FE13" box="[578,745,476,500]" italics="true" pageId="22" pageNumber="221">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C32CEFE56FCD0FE12" box="[754,845,476,501]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C32CEFE56FCD0FE12" bold="true" box="[754,845,476,501]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
the presence of orbital teeth, which are absent in
<taxonomicName id="8FE24D5AFFA7FF5C35B8FE56FF77FDF0" authorityName="in Soledade et al." authorityYear="2015" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C35B8FE56FF77FDF0" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="485D36D9FFA7FF5C30FBFDAEFBB6FC67" blockId="22.[151,1437,151,1365]" pageId="22" pageNumber="221">
The Pacific species can be separated from
<taxonomicName id="8FE24D5AFFA7FF5C3290FDAEFC82FDDB" authorityName="in Soledade et al." authorityYear="2015" box="[684,799,548,572]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C3290FDAEFC82FDDB" box="[684,799,548,572]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
by the presence of blunt-angular orbital teeth (
<figureCitation id="D0D92A5CFFA7FF5C350BFDAEFA14FDDB" box="[1335,1417,548,573]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="22" pageNumber="221">Fig. 11</figureCitation>
e; 13b), which are absent in
<taxonomicName id="8FE24D5AFFA7FF5C3183FDC2FDAFFDB8" authorityName="in Soledade et al." authorityYear="2015" box="[447,562,584,607]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C3183FDC2FDAFFDB8" box="[447,562,584,607]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
(
<figureCitation id="D0D92A5CFFA7FF5C327EFDCDFD0FFD87" box="[578,658,583,608]" captionStart="FIGURE 2" captionStartId="3.[151,250,1739,1761]" captionTargetBox="[162,1430,203,1707]" captionTargetId="figure@3.[151,1436,193,1718]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis trigona (Rathbun, 1901), female (cl 5.5 mm) from Fort Pierce, Florida, USA (MZUSP 34221): acarapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; hthird maxilliped, lateral; i—same, tip of ultimate article, mesial; j—uropod, dorsal." httpUri="https://zenodo.org/record/809638/files/figure.png" pageId="22" pageNumber="221">Figs. 2</figureCitation>
d;
<quantity id="8F1A9B3CFFA7FF5C3293FDC2FD53FD87" box="[687,718,584,608]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="22" pageNumber="221" unit="g" value="4.0">4g</quantity>
); the absence of MDC-PL2 (
<figureCitation id="D0D92A5CFFA7FF5C3424FDCDFBF4FD87" box="[1048,1129,583,608]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="22" pageNumber="221">Fig. 11</figureCitation>
a, b; 13a), which is present albeit relatively low in
<taxonomicName id="8FE24D5AFFA7FF5C319AFDE6FD86FD63" authorityName="in Soledade et al." authorityYear="2015" box="[422,539,620,644]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C319AFDE6FD86FD63" box="[422,539,620,644]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
(cf. 2a, b; 4e); and the stronger distolateral tooth of the scaphocerite, usually reaching beyond the anterior margin of the blade (
<figureCitation id="D0D92A5CFFA7FF5C32FFFD05FC83FD4F" box="[707,798,655,680]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="22" pageNumber="221">Figs. 11</figureCitation>
e; 13b, d) vs. not reaching this margin in
<taxonomicName id="8FE24D5AFFA7FF5C34D8FD1AFACBFD40" authorityName="in Soledade et al." authorityYear="2015" box="[1252,1366,656,679]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C34D8FD1AFACBFD40" box="[1252,1366,656,679]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
(
<figureCitation id="D0D92A5CFFA7FF5C3558FD05FF3BFD2A" captionStart="FIGURE 2" captionStartId="3.[151,250,1739,1761]" captionTargetBox="[162,1430,203,1707]" captionTargetId="figure@3.[151,1436,193,1718]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis trigona (Rathbun, 1901), female (cl 5.5 mm) from Fort Pierce, Florida, USA (MZUSP 34221): acarapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; hthird maxilliped, lateral; i—same, tip of ultimate article, mesial; j—uropod, dorsal." httpUri="https://zenodo.org/record/809638/files/figure.png" pageId="22" pageNumber="221">Figs. 2</figureCitation>
d;
<quantity id="8F1A9B3CFFA7FF5C30F8FD3EFF79FD2B" box="[196,228,692,716]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="22" pageNumber="221" unit="g" value="4.0">4g</quantity>
).
<taxonomicName id="8FE24D5AFFA7FF5C30C7FD3EFE4EFD2B" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[251,467,692,716]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C30C7FD3EFE4EFD2B" box="[251,467,692,716]" italics="true" pageId="22" pageNumber="221">Alpheopsis gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C31E0FD3EFDA4FD2A" box="[476,569,692,717]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C31E0FD3EFDA4FD2A" bold="true" box="[476,569,692,717]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
may be separated from
<taxonomicName id="8FE24D5AFFA7FF5C3369FD3EFC62FD2B" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[853,1023,692,716]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C3369FD3EFC62FD2B" box="[853,1023,692,716]" italics="true" pageId="22" pageNumber="221">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C3435FD3EFBF8FD2A" box="[1033,1125,692,717]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C3435FD3EFBF8FD2A" bold="true" box="[1033,1125,692,717]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
using essentially the same criteria as for
<taxonomicName id="8FE24D5AFFA7FF5C310AFD52FE37FD08" authorityName="in Soledade et al." authorityYear="2015" box="[310,426,728,751]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C310AFD52FE37FD08" box="[310,426,728,751]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
(presence of DLC and VLC on the carapace and proximal process on the cheliped palm, see above), but also by the shape of the rostral carina, i.e. abruptly sloping in
<taxonomicName id="8FE24D5AFFA7FF5C33CCFD76FBFCFCF3" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[1008,1121,764,788]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C33CCFD76FBFCFCF3" box="[1008,1121,764,788]" italics="true" pageId="22" pageNumber="221">A. gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C345BFD76FB5DFCF2" box="[1127,1216,764,789]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C345BFD76FB5DFCF2" bold="true" box="[1127,1216,764,789]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="D0D92A5CFFA7FF5C34F2FD76FAB5FCF2" box="[1230,1320,764,789]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="22" pageNumber="221">Figs. 11</figureCitation>
d; 13a) vs. gently sloping or not sloping in
<taxonomicName id="8FE24D5AFFA7FF5C3236FCAAFD2FFCD0" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[522,690,800,823]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C3236FCAAFD2FFCD0" box="[522,690,800,823]" italics="true" pageId="22" pageNumber="221">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C3287FC95FC8AFCDF" box="[699,791,799,824]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C3287FC95FC8AFCDF" bold="true" box="[699,791,799,824]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="D0D92A5CFFA7FF5C3314FC95FCE4FCDF" box="[808,889,799,824]" captionStart="FIGURE 5" captionStartId="9.[151,250,1773,1795]" captionTargetBox="[161,1424,202,1741]" captionTargetId="figure@9.[151,1436,193,1752]" captionTargetPageId="9" captionText="FIGURE 5. Alpheopsis paratrigona sp. nov., male holotype (cl 4.5 mm) from Portobelo, Caribbean coast of Panama (MZUSP 34222): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—same, dorsal; e—posterior pleon, lateral; f—telson, dorsal; g—ventromesial tooth on first article of antennular peduncle, lateral; h—third maxilliped, lateral; i—same, tip of ultimate article, mesial; j—second pleopod, lateral; k—uropod, dorsal." httpUri="https://zenodo.org/record/809644/files/figure.png" pageId="22" pageNumber="221">Figs. 5</figureCitation>
c; 8a, c). In addition, in
<taxonomicName id="8FE24D5AFFA7FF5C34ADFCAAFA98FCD0" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[1169,1285,800,823]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C34ADFCAAFA98FCD0" box="[1169,1285,800,823]" italics="true" pageId="22" pageNumber="221">A. gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C3532FC95FAF7FCDF" box="[1294,1386,799,824]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C3532FC95FAF7FCDF" bold="true" box="[1294,1386,799,824]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
, the ventral surface of the cheliped carpus does not possess a long stiff seta (
<figureCitation id="D0D92A5CFFA7FF5C33FBFCCEFBBEFCBB" box="[967,1059,836,861]" captionStart="FIGURE 12" captionStartId="18.[151,250,1594,1616]" captionTargetBox="[167,1421,196,1558]" captionTargetId="figure@18.[151,1436,183,1572]" captionTargetPageId="18" captionText="FIGURE 12. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—left cheliped, lateral; b—same, ventrolateral; c—same, mesial; d—same, chela fingers, mesial; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809658/files/figure.png" pageId="22" pageNumber="221">Figs. 12</figureCitation>
a–c; 14a, b, d), which is typically present in both
<taxonomicName id="8FE24D5AFFA7FF5C3178FCE2FE2BFC98" authorityName="in Soledade et al." authorityYear="2015" box="[324,438,872,895]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="7A96EACBFFA7FF5C3178FCE2FE2BFC98" box="[324,438,872,895]" italics="true" pageId="22" pageNumber="221">A. trigona</emphasis>
</taxonomicName>
and
<taxonomicName id="8FE24D5AFFA7FF5C31D0FCE2FD0FFC98" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[492,658,872,895]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="paratrigona" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C31D0FCE2FD0FFC98" box="[492,658,872,895]" italics="true" pageId="22" pageNumber="221">A. paratrigona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C32A6FCEDFD6EFC67" box="[666,755,871,896]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C32A6FCEDFD6EFC67" bold="true" box="[666,755,871,896]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
(cf.
<figureCitation id="D0D92A5CFFA7FF5C331BFCEDFCEEFC67" box="[807,883,871,896]" captionStart="FIGURE 3" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[167,1421,205,1497]" captionTargetId="figure@5.[151,1436,193,1511]" captionTargetPageId="5" captionText="FIGURE 3. Alpheopsis trigona (Rathbun, 1901), female (cl 5.5 mm) from Fort Pierce, Florida, USA (MZUSP 34221): aright cheliped, lateral; b—same, mesial (proximal articles detached); c—same, chela, mesial; d—same, chela fingers, lateral; e—second pereiopod, lateral; f—third pereiopod, lateral; g—same, distal propodus and dactylus, lateral; h—fifth pereiopod, lateral." httpUri="https://zenodo.org/record/809640/files/figure.png" pageId="22" pageNumber="221">Figs. 3</figureCitation>
b; 4h; 6a, b; 7e).
</paragraph>
<paragraph id="485D36D9FFA7FF5C30FBFC06FEEFFB9A" blockId="22.[151,1437,151,1365]" pageId="22" pageNumber="221">
The rather obvious and marked differences in the configuration of the carapacial carinae between the holotype from Panama and the two non-type specimens from Colombia (cf.
<figureCitation id="D0D92A5CFFA7FF5C3343FC25FC45FC2F" box="[895,984,943,968]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="22" pageNumber="221">Figs. 11</figureCitation>
a, b; 13a) are here tentatively interpreted as intraspecific variation. The extreme condition seen in the holotype could be interpreted as (1) typical development of the carinae in very large specimens or (2) some sort of abnormal development, perhaps due to an early injury or genetic aberration. The Colombian specimens also differ from the holotype by the somewhat shorter scaphocerite and longer rostrum (cf.
<figureCitation id="D0D92A5CFFA7FF5C327DFBB5FD02FBBF" box="[577,671,1087,1112]" captionStart="FIGURE 11" captionStartId="17.[151,250,1741,1763]" captionTargetBox="[165,1425,206,1712]" captionTargetId="figure@17.[151,1436,193,1720]" captionTargetPageId="17" captionText="FIGURE 11. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507): a—carapace and anterior pleon, lateral; b—same, dorsal; c—anterior region of carapace and frontal appendages, lateral; d—detail of rostrum, lateral; e—anterior region of carapace and frontal appendages, dorsal; f—posterior pleon, lateral; g—telson, dorsal; h—ventromesial tooth on first article of antennular peduncle, lateral; i—third maxilliped, lateral; j—same, tip of ultimate article, mesial; k—uropod, dorsal." httpUri="https://zenodo.org/record/809656/files/figure.png" pageId="22" pageNumber="221">Figs. 11</figureCitation>
e; 13b). Clearly, more material of
<taxonomicName id="8FE24D5AFFA7FF5C3410FBCAFB3CFBB0" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[1068,1185,1088,1111]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C3410FBCAFB3CFBB0" box="[1068,1185,1088,1111]" italics="true" pageId="22" pageNumber="221">A. gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C3496FBB5FA9BFBBF" box="[1194,1286,1087,1112]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C3496FBB5FA9BFBBF" bold="true" box="[1194,1286,1087,1112]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
is needed to clarify these issues.
</paragraph>
<paragraph id="485D36D9FFA7FF5C30FBFB0DFD03FAB2" blockId="22.[151,1437,151,1365]" pageId="22" pageNumber="221">
The Costa Rican record is based on a photograph of a specimen from Isla Tortuga in the Nicoya Gulf (
<figureCitation id="D0D92A5CFFA7FF5C3553FB0DFF28FB22" captionStart="FIGURE 15" captionStartId="21.[151,250,1906,1928]" captionTargetBox="[175,1376,193,1855]" captionTargetId="figure@21.[175,1376,193,1855]" captionTargetPageId="21" captionText="FIGURE 15. Alpheopsis gotrina sp. nov., female holotype (cl 7.0 mm) from Playa Venao, Pacific cost of Panama (MZUSP 34507) [a]; male (cl 4.8 mm) from Curichichi, Bahía Málaga, Pacific cost of Colombia (OUMNH. ZC. 2017.01.021) [b]; specimen (sex and cl unknown) from Isla Tortuga, Pacific coast of Costa Rica (specimen collected but possibly lost, see text, not examined) [c]. Photographs: A. Anker (a, b) and courtesy of Ingo S. Wehrtmann (c)." httpUri="https://zenodo.org/record/809664/files/figure.png" pageId="22" pageNumber="221">Fig. 15</figureCitation>
c). Although the photographed shrimp has been collected and preserved, it was not located in the crustacean collection of the Universidad de Costa Rica, San José (most probably, it was lost during transportation to San José, Ingo S. Wehrtmann, pers. comm.). Although the presence of yet another species in the eastern Pacific cannot be totally excluded, this photographic record is tentatively assigned to
<taxonomicName id="8FE24D5AFFA7FF5C33B3FA92FB9FFAC8" authority="Anker, 2017" authorityName="Anker" authorityYear="2017" box="[911,1026,1304,1327]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="221" phylum="Arthropoda" rank="species" species="gotrina" status="sp. nov.">
<emphasis id="7A96EACBFFA7FF5C33B3FA92FB9FFAC8" box="[911,1026,1304,1327]" italics="true" pageId="22" pageNumber="221">A. gotrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61A557B0FFA7FF5C3435FA9DFBFEFAD7" box="[1033,1123,1303,1328]" pageId="22" pageNumber="221" rank="species">
<emphasis id="7A96EACBFFA7FF5C3435FA9DFBFEFAD7" bold="true" box="[1033,1123,1303,1328]" pageId="22" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
, awaiting collection of new material along the Pacific coast of Costa Rica.
</paragraph>
</subSubSection>
</treatment>
</document>
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