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<document id="34C9D6FD6DCDC56C75DC3DDA1463DD9F" ID-CLB-Dataset="8672" ID-DOI="10.5281/zenodo.4468383" ID-GBIF-Dataset="5c5c8aad-73f1-4a26-bdca-d1d02ba3fec1" ID-Zenodo-Dep="4468383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611044739228" checkinUser="plazi" docAuthor="Martínez, Leonel, Brescovit, Antonio D., Villarreal, Eduardo &amp; Oliveira, Luiz Fernando M." docDate="2021" docId="A86D27653D5FFFB8FF1DFC8AFE49FB7F" docLanguage="en" docName="zootaxa.4914.1.1.pdf" docOrigin="Zootaxa 4914 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Patrera philipi Duperre &amp; Tapia 2016" docType="treatment" docVersion="11" lastPageNumber="45" masterDocId="54545F1D3D73FF94FF8AFFC9FFEBFF96" masterDocTitle="An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia" masterLastPageNumber="64" masterPageNumber="1" pageNumber="45" updateTime="1698924792971" updateUser="ExternalLinkService">
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<mods:title id="30A7D5D50C28B33B4F5CFDCE92C83628">An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia</mods:title>
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<mods:namePart id="286B2E7D427D9AFAE92CA23A4E3BDF53">Martínez, Leonel</mods:namePart>
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<mods:nameIdentifier id="024B89145BC5075C4846BF2E7563CDDE" type="email">leonelmarbio@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7750EBF8471D930222440E333BC71AD8">Brescovit, Antonio D.</mods:namePart>
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<mods:nameIdentifier id="BCB5BA37E03174D8D742C27D6251D4A2" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:namePart id="41218D5ADFB1548C74BA97741225E658">Villarreal, Eduardo</mods:namePart>
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<mods:nameIdentifier id="BA098078878775888B17E3A5396900C8" type="email">leonelmarbio@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A1F4A771C5B12FF2BBA8419407FE931B">Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:title id="CEEB3DD8F87FF4D4F20C6B10DCA0993B">Zootaxa</mods:title>
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<mods:date id="3F89F2A0C797ACD1AB1457DE6F1119FE">2021</mods:date>
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<mods:number id="812DA9AB345CDADA0343F36D7EAEABCF">4914</mods:number>
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<treatment id="A86D27653D5FFFB8FF1DFC8AFE49FB7F" ID-DOI="http://doi.org/10.5281/zenodo.4469617" ID-GBIF-Taxon="177121404" ID-Zenodo-Dep="4469617" LSID="urn:lsid:plazi:treatment:A86D27653D5FFFB8FF1DFC8AFE49FB7F" httpUri="http://treatment.plazi.org/id/A86D27653D5FFFB8FF1DFC8AFE49FB7F" lastPageNumber="45" pageId="44" pageNumber="45">
<subSubSection id="68DEC5F83D5FFFB8FF1DFC8AFE1CFCCB" box="[151,503,834,861]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="207B96733D5FFFB8FF1DFC8AFE1CFCCB" blockId="44.[151,503,834,861]" box="[151,503,834,861]" pageId="44" pageNumber="45">
<heading id="7B33211F3D5FFFB8FF1DFC8AFE1CFCCB" bold="true" box="[151,503,834,861]" fontSize="11" level="2" pageId="44" pageNumber="45" reason="4">
<emphasis id="12B04A613D5FFFB8FF1DFC8AFE1CFCCB" bold="true" box="[151,503,834,861]" pageId="44" pageNumber="45">
<taxonomicName id="E7C4EDF03D5FFFB8FF1DFC8AFEA3FCCB" ID-CoL="6TTNK" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[151,328,834,861]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="philipi">
<emphasis id="12B04A613D5FFFB8FF1DFC8AFEA3FCCB" bold="true" box="[151,328,834,861]" italics="true" pageId="44" pageNumber="45">Patrera philipi</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="68DEC5F83D5FFFB8FF1DFC43FE49FB7F" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="207B96733D5FFFB8FF1DFC43FAE1FBCD" blockId="44.[151,1437,906,1257]" pageId="44" pageNumber="45">
<emphasis id="12B04A613D5FFFB8FF1DFC43FEFFFC32" bold="true" box="[151,276,906,932]" pageId="44" pageNumber="45">Diagnosis:</emphasis>
males can be diagnosed by following character combination: usually with queliceral projections on anterior region of the distal edge; a prominent, conspicuous and strongly sclerotized ventral tegular process on the tegulum, closely related to the embolus; embolus short, proximally inserted on the tegulum, retrolateral tibial apophysis bifid from base (
<figureCitation id="B8FF8AF63D5FFFB8FEC4FC3EFE52FB85" box="[334,441,1015,1043]" captionStart="FIGURE 22" captionStartId="42.[151,250,1809,1833]" captionTargetBox="[272,1303,181,1785]" captionTargetId="figure-27@42.[272,1316,181,1785]" captionTargetPageId="42" captionText="FIGURE 22AD. Patrera armata (Chickering, 1940). Male (ICN-Ar-10608): A habitus, dorsal view; B Chelicerae ventral view; C left palp, retroventral view (Arrow: indicates projection basal of the tibia); D palp, ventral view; E palp, retrolateral view. Female (ICN-Ar-5481): B habitus, dorsal view; G epigynum, ventral view; F epigynum, dorsal view. Abbreviations: BE, base of the embolus; Cvp, cheliceral ventral projection; dps, dorsal projection of the subtegulum; E, embolus; FD, fertilization ducts; H, hood; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; ST, subtegulum; T, tegulum. Scale bars: AB: 2mm. C: 1mm; DF: 0.5mm; GH: 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.4468433" httpUri="https://zenodo.org/record/4468433/files/figure.png" pageId="44" pageNumber="45">Figs 22D</figureCitation>
̅F; 23C̅E; 24E̅G; 25C̅E; 26D̅F; 27D̅F; 28C̅E; 29C̅E). Females are distinguished by having the hood with undulations on the base (absent in
<taxonomicName id="E7C4EDF03D5FFFB8FCA4FBD2FC46FBA3" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[814,941,1051,1077]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="chucurui">
<emphasis id="12B04A613D5FFFB8FCA4FBD2FC46FBA3" box="[814,941,1051,1077]" italics="true" pageId="44" pageNumber="45">P. chucurui</emphasis>
</taxonomicName>
); seminal receptacles absent and posteriorly positioned spermathecae (except in
<taxonomicName id="E7C4EDF03D5FFFB8FDA2FBF6FD43FBCF" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[552,680,1087,1113]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="chucurui">
<emphasis id="12B04A613D5FFFB8FDA2FBF6FD43FBCF" box="[552,680,1087,1113]" italics="true" pageId="44" pageNumber="45">P. chucurui</emphasis>
</taxonomicName>
, anteriorly positioned) (
<figureCitation id="B8FF8AF63D5FFFB8FC3CFBF6FBC9FBCD" box="[950,1058,1087,1115]" captionStart="FIGURE 22" captionStartId="42.[151,250,1809,1833]" captionTargetBox="[272,1303,181,1785]" captionTargetId="figure-27@42.[272,1316,181,1785]" captionTargetPageId="42" captionText="FIGURE 22AD. Patrera armata (Chickering, 1940). Male (ICN-Ar-10608): A habitus, dorsal view; B Chelicerae ventral view; C left palp, retroventral view (Arrow: indicates projection basal of the tibia); D palp, ventral view; E palp, retrolateral view. Female (ICN-Ar-5481): B habitus, dorsal view; G epigynum, ventral view; F epigynum, dorsal view. Abbreviations: BE, base of the embolus; Cvp, cheliceral ventral projection; dps, dorsal projection of the subtegulum; E, embolus; FD, fertilization ducts; H, hood; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; ST, subtegulum; T, tegulum. Scale bars: AB: 2mm. C: 1mm; DF: 0.5mm; GH: 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.4468433" httpUri="https://zenodo.org/record/4468433/files/figure.png" pageId="44" pageNumber="45">Figs 22G</figureCitation>
̅H; 24H̅I; 28F̅G).
</paragraph>
<paragraph id="207B96733D5FFFB8FF4DFBABFE49FB7F" blockId="44.[151,1437,906,1257]" pageId="44" pageNumber="45">
<emphasis id="12B04A613D5FFFB8FF4DFBABFE82FBEA" bold="true" box="[199,361,1122,1148]" pageId="44" pageNumber="45">Composition:</emphasis>
Thirteen species:
<taxonomicName id="E7C4EDF03D5FFFB8FDB1FBAAFC67FBEB" authority="(Chickering, 1940)" baseAuthorityName="Chickering" baseAuthorityYear="1940" box="[571,908,1123,1149]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="12B04A613D5FFFB8FDB1FBAAFD41FBEA" box="[571,682,1123,1148]" italics="true" pageId="44" pageNumber="45">P. armata</emphasis>
(
<bibRefCitation id="4455EB823D5FFFB8FD30FBAAFC6EFBEB" author="Chickering, A. M." box="[698,901,1123,1149]" pageId="44" pageNumber="45" pagination="78" refId="ref40903" refString="Chickering, A. M. (1940) New Anyphaenidae from Panama with notes on known species. Transactions of the American Microscopical Society, 59, 78 ‾ 122." type="journal article" year="1940">Chickering, 1940</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="E7C4EDF03D5FFFB8FC10FBAAFC0DFBEA" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[922,998,1123,1148]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="sutu" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FC10FBAAFC0DFBEA" box="[922,998,1123,1148]" italics="true" pageId="44" pageNumber="45">P. sutu</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FC65FBAAFBDAFBEB" bold="true" box="[1007,1073,1123,1149]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FC65FBAAFBDAFBEB" box="[1007,1073,1123,1149]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FBCAFBAAFB29FBEB" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[1088,1218,1123,1149]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="chucurui" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FBCAFBAAFB29FBEB" box="[1088,1218,1123,1149]" italics="true" pageId="44" pageNumber="45">P. chucurui</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FB40FBAAFAE7FBEB" bold="true" box="[1226,1292,1123,1149]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FB40FBAAFAE7FBEB" box="[1226,1292,1123,1149]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FA91FBAAFA76FBEB" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[1307,1437,1123,1149]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="dawkinsi" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FA91FBAAFA76FBEB" box="[1307,1437,1123,1149]" italics="true" pageId="44" pageNumber="45">P. dawkinsi</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FF1DFB4EFF3CFB37" bold="true" box="[151,215,1159,1185]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FF1DFB4EFF3CFB37" box="[151,215,1159,1185]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FF6DFB4EFEB3FB37" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[231,344,1159,1185]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="dentata" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FF6DFB4EFEB3FB37" box="[231,344,1159,1185]" italics="true" pageId="44" pageNumber="45">P. dentata</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FEEAFB4EFE4BFB37" bold="true" box="[352,416,1159,1185]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FEEAFB4EFE4BFB37" box="[352,416,1159,1185]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FE3AFB4EFDC1FB37" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[432,554,1159,1185]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="4" phylum="Arthropoda" rank="species" species="dracula" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FE3AFB4EFDC1FB37" box="[432,554,1159,1185]" italics="true" pageId="44" pageNumber="45">P. Dracula</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FDB8FB4EFD98FB37" bold="true" box="[562,627,1159,1185]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FDB8FB4EFD98FB37" box="[562,627,1159,1185]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FD08FB4EFBF5FB37" authority="Duperre &amp; Tapia, 2016" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[642,1054,1159,1185]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="hatunkiru">
<emphasis id="12B04A613D5FFFB8FD08FB4EFCE7FB37" box="[642,780,1159,1185]" italics="true" pageId="44" pageNumber="45">P. hatunkiru</emphasis>
<bibRefCitation id="4455EB823D5FFFB8FC9EFB4EFBF5FB37" author="Duperre, N. &amp; Tapia, E." box="[788,1054,1159,1185]" pageId="44" pageNumber="45" pagination="1 - 50" refId="ref40935" refString="Duperre, N. &amp; Tapia, E. (2016) Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Choco forest of Ecuador, with the description of thirteen new species. European Journal of Taxonomy, 255, 1 - 50. https: // doi. org / 10.5852 / ejt. 2016.255" type="journal article" year="2016">Dupérré &amp; Tapia, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7C4EDF03D5FFFB8FBA1FB4EFB6AFB37" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[1067,1153,1159,1185]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="kuryi" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FBA1FB4EFB6AFB37" box="[1067,1153,1159,1185]" italics="true" pageId="44" pageNumber="45">P. kuryi</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FB03FB4EFB22FB37" bold="true" box="[1161,1225,1159,1185]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FB03FB4EFB22FB37" box="[1161,1225,1159,1185]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FB53FB4EFA96FB37" authority="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira, 2021" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[1241,1405,1159,1185]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="longitibialis" status="sp. nov.">
<emphasis id="12B04A613D5FFFB8FB53FB4EFA96FB37" box="[1241,1405,1159,1185]" italics="true" pageId="44" pageNumber="45">P. longitibialis</emphasis>
</taxonomicName>
<emphasis id="12B04A613D5FFFB8FA0CFB4EFF53FB53" bold="true" pageId="44" pageNumber="45">
<taxonomicNameLabel id="0983F71A3D5FFFB8FA0CFB4EFF53FB53" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E7C4EDF03D5FFFB8FF4DFB62FDD3FB53" authority="Duperre &amp; Tapia, 2016" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[199,568,1195,1221]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="philipi">
<emphasis id="12B04A613D5FFFB8FF4DFB62FEC0FB53" box="[199,299,1195,1221]" italics="true" pageId="44" pageNumber="45">P. philipi</emphasis>
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<taxonomicName id="E7C4EDF03D5FFFB8FDC9FB62FC42FB53" authority="Duperre &amp; Tapia, 2016" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[579,937,1195,1221]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="shida">
<emphasis id="12B04A613D5FFFB8FDC9FB62FD77FB53" box="[579,668,1195,1221]" italics="true" pageId="44" pageNumber="45">P. shida</emphasis>
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<taxonomicName id="E7C4EDF03D5FFFB8FC3EFB62FAE0FB53" authority="Duperre &amp; Tapia, 2016" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[948,1291,1195,1221]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="suni">
<emphasis id="12B04A613D5FFFB8FC3EFB62FC15FB52" box="[948,1022,1195,1220]" italics="true" pageId="44" pageNumber="45">P. suni</emphasis>
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<taxonomicName id="E7C4EDF03D5FFFB8FACCFB62FE75FB7F" authority="Duperre &amp; Tapia, 2016" authorityName="Duperre &amp; Tapia" authorityYear="2016" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="witsu">
<emphasis id="12B04A613D5FFFB8FACCFB62FA77FB52" box="[1350,1436,1195,1220]" italics="true" pageId="44" pageNumber="45">P. witsu</emphasis>
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.
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