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<document id="A4266E40A45615F8A56CB1B82979691B" ID-CLB-Dataset="32732" ID-DOI="10.11646/zootaxa.4286.4.2" ID-GBIF-Dataset="85257999-58b3-4ea2-9b44-d86f686122ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="828729" ID-ZooBank="24BA3710-2416-47D2-8E29-B13AE35F5589" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500109527176" checkinUser="plazi" docAuthor="Botero-Trujillo, Ricardo &amp; Flórez, Eduardo" docDate="2017" docId="864D67014D6AFFF1FF1AFF49310F38C8" docLanguage="en" docName="zootaxa.4286.4.2.pdf" docOrigin="Zootaxa 4286 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cryptocellus chiruisla Botero-Trujillo &amp; Flórez, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="492" masterDocId="7A741F794D68FFF8FF8DFFDE356A3B6D" masterDocTitle="Two new ricinuleid species from Ecuador and Colombia belonging to the peckorum species-group of Cryptocellus Westwood (Arachnida, Ricinulei)" masterLastPageNumber="498" masterPageNumber="483" pageNumber="485" updateTime="1698465282705" updateUser="ExternalLinkService">
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<mods:title id="A215ABA3FDB0E0A4E57F96161119690E">Two new ricinuleid species from Ecuador and Colombia belonging to the peckorum species-group of Cryptocellus Westwood (Arachnida, Ricinulei)</mods:title>
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<mods:namePart id="DBF7F3CE73A932C7E7B8AFE5323DCC99">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:namePart id="0C886C0EC1F37FE8BC04C404FF5FD7C1">Flórez, Eduardo</mods:namePart>
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<mods:date id="9D6D6240BFFB5ECA7659A196E2AEA7B0">2017</mods:date>
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<treatment id="864D67014D6AFFF1FF1AFF49310F38C8" ID-DOI="http://doi.org/10.5281/zenodo.6046879" ID-GBIF-Taxon="132549025" ID-Zenodo-Dep="6046879" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8" httpUri="http://treatment.plazi.org/id/864D67014D6AFFF1FF1AFF49310F38C8" lastPageId="9" lastPageNumber="492" pageId="2" pageNumber="485">
<subSubSection id="46FE859C4D6AFFFAFF1AFF4934DF3BBE" pageId="2" pageNumber="485" type="nomenclature">
<paragraph id="0E5BD6174D6AFFFAFF1AFF49376C3BDF" blockId="2.[151,518,151,211]" box="[151,518,151,178]" pageId="2" pageNumber="485">
<heading id="5513617B4D6AFFFAFF1AFF49376C3BDF" bold="true" box="[151,518,151,178]" fontSize="11" level="1" pageId="2" pageNumber="485" reason="1">
<emphasis id="3C900A054D6AFFFAFF1AFF49376C3BDF" bold="true" box="[151,518,151,178]" pageId="2" pageNumber="485">
<taxonomicName id="C9E4AD944D6AFFFAFF1AFF4934CB3BDC" ID-CoL="6BMCN" box="[151,417,151,177]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFF1AFF4934CB3BDC" bold="true" box="[151,417,151,177]" italics="true" pageId="2" pageNumber="485">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6AFFFAFE25FF46376C3BDF" box="[424,518,152,178]" pageId="2" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0E5BD6174D6AFFFAFF1AFF6434DF3BBE" blockId="2.[151,518,151,211]" box="[151,437,186,211]" pageId="2" pageNumber="485">
<figureCitation id="96DFCA924D6AFFFAFF1AFF6434463BBE" box="[151,300,186,211]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 10" captionStart-2="FIGURES 11 17" captionStart-3="FIGURES 18 21" captionStart-4="FIGURES 22 24" captionStartId-0="3.[151,269,1660,1683]" captionStartId-1="4.[151,264,1719,1741]" captionStartId-2="5.[151,264,1809,1831]" captionStartId-3="6.[151,264,1452,1474]" captionStartId-4="7.[151,264,1191,1213]" captionTargetBox-0="[176,1420,194,1621]" captionTargetBox-1="[151,1436,193,1665]" captionTargetBox-2="[151,1436,337,1748]" captionTargetBox-3="[157,1421,193,1422]" captionTargetBox-4="[158,1426,243,1090]" captionTargetId-0="figure@3.[176,1420,194,1642]" captionTargetId-1="figure@4.[151,1436,193,1698]" captionTargetId-2="figure@5.[151,1436,337,1788]" captionTargetId-3="figure@6.[157,1421,193,1422]" captionTargetId-4="figure@7.[151,1436,221,1126]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." captionText-1="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." captionText-2="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." captionText-3="FIGURES 18 21. Cryptocellus chiruisla sp. nov. Female paratype (QCAZI 3441). 18. Carapace, dorsal aspect. 19. Prosoma, ventral aspect (sternal region). 20. Opisthosoma, dorsal aspect. 21. Opisthosoma, pygidium. Scale bars: 500 µm." captionText-4="FIGURES 22 24. Cryptocellus chiruisla sp. nov. Female paratype (QCAZI 3441). Spermathecae. 22 23. Posterior and anterior aspects (apically trilobated apex of the posterior genital lip indicated by arrow). 24. Frontal aspect. Scale bars: 200 µm." httpUri-0="https://zenodo.org/record/828731/files/figure.png" httpUri-1="https://zenodo.org/record/828733/files/figure.png" httpUri-2="https://zenodo.org/record/828735/files/figure.png" httpUri-3="https://zenodo.org/record/828737/files/figure.png" httpUri-4="https://zenodo.org/record/828739/files/figure.png" pageId="2" pageNumber="485">Figures 124</figureCitation>
,
<figureCitation id="96DFCA924D6AFFFAFEB5FF65343D3BBE" box="[312,343,187,211]" captionStart="FIGURE 41" captionStartId="14.[151,250,863,885]" captionTargetBox="[151,1436,193,842]" captionTargetId="figure@14.[151,1436,193,842]" captionTargetPageId="14" captionText="FIGURE 41. Map of Northwestern South America, plotting the known distribution of Cryptocellus chiruisla sp. nov. and Cryptocellus guaviarensis sp. nov." httpUri="https://zenodo.org/record/828747/files/figure.png" pageId="2" pageNumber="485">41</figureCitation>
;
<tableCitation id="4366E3AC4D6AFFFAFEEEFF6434DF3BBE" box="[355,437,186,211]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Measurements of Cryptocellus chiruisla sp. nov. and Cryptocellus guaviarensis sp. nov. Measurements in millimeters; L = length; W = width; H = height. 1 It excludes pygidium and cucullus lenght. 2 Greatest width. 3 Greatest height. 4 Sum of individual segment lengths, excluding coxa. * Measured at level between coxae II and III, where widest. ** Measured at level of tergite XII, where widest." pageId="2" pageNumber="485">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="46FE859C4D6AFFFAFF1AFF23361F3A30" pageId="2" pageNumber="485" type="materials_examined">
<paragraph id="0E5BD6174D6AFFFAFF1AFF23361F3A30" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<materialsCitation id="BE8CDC4A4D6AFFFAFF1AFF2331253A54" ID-GBIF-Occurrence="1572325258" collectingDate="2005-12-08" collectingDateMax="2005-12-13" collectingDateMin="2005-12-08" collectionCode="QCAZI" collectorName="J. Viera" country="Ecuador" elevation="218" latitude="-0.6138889" location="Orellana" longLatPrecision="21" longitude="-75.876114" pageId="2" pageNumber="485" specimenCode="QCAZI 3440" specimenCount="2" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="3C900A054D6AFFFAFF1AFF2334A03A78" bold="true" box="[151,458,252,278]" pageId="2" pageNumber="485">
<typeStatus id="D15F68B54D6AFFFAFF1AFF2335B83A7B" box="[151,210,253,278]" pageId="2" pageNumber="485">Type</typeStatus>
material.
<emphasis id="3C900A054D6AFFFAFED5FF2234A03A78" bold="true" box="[344,458,252,277]" italics="true" pageId="2" pageNumber="485">
<typeStatus id="D15F68B54D6AFFFAFED5FF2234A93A78" box="[344,451,252,277]" pageId="2" pageNumber="485" type="holotype">Holotype</typeStatus>
:
</emphasis>
</emphasis>
male from
<collectingCountry id="76F396874D6AFFFAFDD3FF2037803A7B" box="[606,746,254,278]" name="Ecuador" pageId="2" pageNumber="485">ECUADOR</collectingCountry>
:
<collectingRegion id="CC2018F54D6AFFFAFD76FF23360A3A7B" box="[763,864,253,278]" country="Ecuador" name="Orellana" pageId="2" pageNumber="485">Orellana</collectingRegion>
:
<taxonomicName id="C9E4AD944D6AFFFAFCFDFF2336BD3A7B" box="[880,983,253,278]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">Chiruisla</taxonomicName>
km 0.2, primary forest,
<elevation id="85C931244D6AFFFAFA8AFF20303E3A78" box="[1287,1364,254,278]" metricMagnitude="2" metricUnit="m" metricValue="2.18" pageId="2" pageNumber="485" unit="m" value="218.0">
<quantity id="C91C7BF24D6AFFFAFA8AFF20303E3A78" box="[1287,1364,254,278]" metricMagnitude="2" metricUnit="m" metricValue="2.18" pageId="2" pageNumber="485" unit="m" value="218.0">218 m</quantity>
</elevation>
elev.,
<geoCoordinate id="6BD0B0D04D6AFFFAFF1AFEFF34483A54" box="[151,290,289,314]" direction="south" orientation="latitude" pageId="2" pageNumber="485" precision="15" value="-0.6138889">00°3650″ S</geoCoordinate>
<geoCoordinate id="6BD0B0D04D6AFFFAFEA3FEFF34AE3A54" box="[302,452,289,314]" direction="west" orientation="longitude" pageId="2" pageNumber="485" precision="15" value="-75.876114">75°5234″ W</geoCoordinate>
,
<collectingDate id="6A1E093F4D6AFFFAFE59FEFF37EA3A57" box="[468,640,289,314]" pageId="2" pageNumber="485" value="2005-12-08" valueMax="2005-12-13" valueMin="2005-12-08">
<date id="7A5AF0D74D6AFFFAFE59FEFF37EA3A57" box="[468,640,289,314]" pageId="2" pageNumber="485" value="2005-12-08" valueMax="2005-12-13" valueMin="2005-12-08">0813.xii.2005</date>
</collectingDate>
,
<collectingMethod id="D7A5AE004D6AFFFAFD1CFEFF364C3A54" box="[657,806,289,314]" pageId="2" pageNumber="485">Winkler trap</collectingMethod>
,
<collectorName id="A311B3C14D6AFFFAFCBAFEFC36F93A57" box="[823,915,289,314]" pageId="2" pageNumber="485">J. Viera</collectorName>
(
<specimenCode id="5E427E6C4D6AFFFAFC2BFEFF312D3A54" box="[934,1095,289,313]" collectionCode="QCAZI" pageId="2" pageNumber="485">
<collectionCode id="68F54ED24D6AFFFAFC2BFEFF36943A54" box="[934,1022,289,313]" pageId="2" pageNumber="485">QCAZI</collectionCode>
3440
</specimenCode>
)
</materialsCitation>
.
<emphasis id="3C900A054D6AFFFAFBD3FEFF31B33A55" bold="true" box="[1118,1241,289,312]" italics="true" pageId="2" pageNumber="485">Paratypes:</emphasis>
1 female and 1 tritonymph with same collection data of holotype (QCAZI 3441).
</paragraph>
</subSubSection>
<subSubSection id="46FE859C4D6AFFFAFF4AFEB631983AED" box="[199,1266,360,385]" pageId="2" pageNumber="485" type="etymology">
<paragraph id="0E5BD6174D6AFFFAFF4AFEB631983AED" blockId="2.[151,1437,252,2029]" box="[199,1266,360,385]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AFEB634243AEC" bold="true" box="[199,334,360,385]" pageId="2" pageNumber="485">Etymology.</emphasis>
The specific epithet is a noun in apposition taken from the
<typeStatus id="D15F68B54D6AFFFAFC6AFEB7317D3AEC" box="[999,1047,361,385]" pageId="2" pageNumber="485">type</typeStatus>
locality:
<taxonomicName id="C9E4AD944D6AFFFAFB0CFEB631813AED" box="[1153,1259,360,384]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFB0CFEB631813AED" box="[1153,1259,360,384]" italics="true" pageId="2" pageNumber="485">Chiruisla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="46FE859C4D6AFFFAFF4AFE5236E93F69" pageId="2" pageNumber="485" type="diagnosis">
<paragraph id="0E5BD6174D6AFFFAFF4AFE52362E398B" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AFE52342B3AC8" bold="true" box="[199,321,396,421]" pageId="2" pageNumber="485">Diagnosis.</emphasis>
<taxonomicName id="C9E4AD944D6AFFFAFEC4FE52372F3AC9" box="[329,581,396,420]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFEC4FE52372F3AC9" box="[329,581,396,420]" italics="true" pageId="2" pageNumber="485">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6AFFFAFDC0FE5237C23AC8" box="[589,680,396,421]" pageId="2" pageNumber="485" rank="species">
<emphasis id="3C900A054D6AFFFAFDC0FE5237C23AC8" bold="true" box="[589,680,396,421]" pageId="2" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
most closely resembles
<taxonomicName id="C9E4AD944D6AFFFAFC33FE5331043AC9" box="[958,1134,397,420]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="guaviarensis" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFC33FE5331043AC9" box="[958,1134,397,420]" italics="true" pageId="2" pageNumber="485">C. guaviarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6AFFFAFBFBFE5231BB3AC8" box="[1142,1233,396,421]" pageId="2" pageNumber="485" rank="species">
<emphasis id="3C900A054D6AFFFAFBFBFE5231BB3AC8" bold="true" box="[1142,1233,396,421]" pageId="2" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
, among members of the
<taxonomicName id="C9E4AD944D6AFFFAFF6EFE6E343B3AA5" box="[227,337,432,456]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="peckorum">
<emphasis id="3C900A054D6AFFFAFF6EFE6E343B3AA5" box="[227,337,432,456]" italics="true" pageId="2" pageNumber="485">peckorum</emphasis>
</taxonomicName>
species-group to which both belong, in having the following morphological features of male: a large and distinct, flat, triangle-like region with tubercles on the anterior section of cucullus (
<figureCitation id="96DFCA924D6AFFFAFB59FE0D30763A81" box="[1236,1308,467,492]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="2" pageNumber="485">Fig. 7</figureCitation>
), absent in female, on top of which a median elevation is visible on dorsal and ventral aspects (
<figureCitation id="96DFCA924D6AFFFAFBEFFE2931BA397D" box="[1122,1232,503,528]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="2" pageNumber="485">Figs. 12</figureCitation>
); the anterior and anterolateral margins of carapace smooth (
<figureCitation id="96DFCA924D6AFFFAFD0FFDC537AD395E" box="[642,711,539,564]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="2" pageNumber="485">Fig. 1</figureCitation>
); the movable finger of the chelicera noticeably widened with respect to the fixed finger (
<figureCitation id="96DFCA924D6AFFFAFE50FDE0374D393A" box="[477,551,574,599]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="2" pageNumber="485">Fig. 7</figureCitation>
); leg I tibia with a pronounced ventral “notch” formed by well-developed, ventrally protruding pro- (large) and retrolateral (small) distal lobes, a sub-basal tubercle-like protrusion, and a distinct concavity in the middle (
<figureCitation id="96DFCA924D6AFFFAFD81FD5837FC39F2" box="[524,662,646,671]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 1314</figureCitation>
); leg I metatarsus laterally-compressed, prolaterally sunken on basal two thirds, and with a well-developed, proventral distal “elbow-like” expansion (
<figureCitation id="96DFCA924D6AFFFAFBE1FD74319739AF" box="[1132,1277,682,707]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 1314</figureCitation>
); and leg III metatarsus with disto-ventral tuft of long setae (
<figureCitation id="96DFCA924D6AFFFAFD22FD13365D398B" box="[687,823,717,742]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 1112</figureCitation>
).
</paragraph>
<paragraph id="0E5BD6174D6AFFFAFF4AFD2F36E93F69" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
Males of
<taxonomicName id="C9E4AD944D6AFFFAFEB9FD2C34D73864" box="[308,445,753,777]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFEB9FD2C34D73864" box="[308,445,753,777]" italics="true" pageId="2" pageNumber="485">C. chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6AFFFAFE45FD2F374F3867" box="[456,549,753,778]" pageId="2" pageNumber="485" rank="species">
<emphasis id="3C900A054D6AFFFAFE45FD2F374F3867" bold="true" box="[456,549,753,778]" pageId="2" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
can be readily recognized by having the cheliceral movable finger apically truncated (
<figureCitation id="96DFCA924D6AFFFAFE9DFCCB343C3843" box="[272,342,789,814]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="2" pageNumber="485">Fig. 7</figureCitation>
); suture line of coxae II about half the length of that of coxae III (
<figureCitation id="96DFCA924D6AFFFAFBD8FCCB31F13843" box="[1109,1179,789,814]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
); leg I tibia sub-basal, tubercle-like protrusion part of the “notch”, placed pro-ventrally on basal half of segment, such that the “notch” is visible dorsally and prolaterally (
<figureCitation id="96DFCA924D6AFFFAFDBDFC8237A83818" box="[560,706,860,885]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 1314</figureCitation>
); median concavity of the “notch” of leg I tibia is short, approximately one third of the segments length (
<figureCitation id="96DFCA924D6AFFFAFD42FC5E364F38F4" box="[719,805,896,921]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Fig. 14</figureCitation>
); apex of the lamina cyathiformis is distinctly rounded (
<figureCitation id="96DFCA924D6AFFFAFF12FC7A344438D0" box="[159,302,932,957]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 1112</figureCitation>
); the tarsal process of the copulatory apparatus is U-shaped, with apex directed dorsally; the retrolateral edge of the tarsal process ventral opening lacks spiniform projections (
<figureCitation id="96DFCA924D6AFFFAFBB2FC193199388D" box="[1087,1267,967,992]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 11, 1517</figureCitation>
); and femur of legs, particularly that of leg IV, remarkably bulky (
<figureCitation id="96DFCA924D6AFFFAFD5DFC35367F3F69" box="[720,789,1003,1028]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="2" pageNumber="485">Fig. 9</figureCitation>
,
<tableCitation id="4366E3AC4D6AFFFAFCADFC3536183F69" box="[800,882,1003,1028]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Measurements of Cryptocellus chiruisla sp. nov. and Cryptocellus guaviarensis sp. nov. Measurements in millimeters; L = length; W = width; H = height. 1 It excludes pygidium and cucullus lenght. 2 Greatest width. 3 Greatest height. 4 Sum of individual segment lengths, excluding coxa. * Measured at level between coxae II and III, where widest. ** Measured at level of tergite XII, where widest." pageId="2" pageNumber="485">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="46FE859C4D6AFFFAFF4AFBD134663E04" pageId="2" pageNumber="485" type="discussion">
<paragraph id="0E5BD6174D6AFFFAFF4AFBD134663E04" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AFBD134513F45" bold="true" box="[199,315,1039,1064]" pageId="2" pageNumber="485">Remarks.</emphasis>
<taxonomicName id="C9E4AD944D6AFFFAFEC9FBD1372B3F4A" box="[324,577,1039,1063]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFEC9FBD1372B3F4A" box="[324,577,1039,1063]" italics="true" pageId="2" pageNumber="485">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6AFFFAFDC6FBD137CD3F45" box="[587,679,1039,1064]" pageId="2" pageNumber="485" rank="species">
<emphasis id="3C900A054D6AFFFAFDC6FBD137CD3F45" bold="true" box="[587,679,1039,1064]" pageId="2" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
most closely resembles
<taxonomicName id="C9E4AD944D6AFFFAFC4FFBCE31393F4A" box="[962,1107,1039,1063]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="peckorum">
<emphasis id="3C900A054D6AFFFAFC4FFBCE31393F4A" box="[962,1107,1039,1063]" italics="true" pageId="2" pageNumber="485">C. peckorum</emphasis>
</taxonomicName>
among previously described members of the species-group by having the tegument densely covered with conspicuous tubercles, and similar shape of the male copulatory apparatus and male leg III metatarsus. Unlike the male of the new species referred above, in males of
<taxonomicName id="C9E4AD944D6AFFFAFEE1FBA534903FFE" box="[364,506,1147,1171]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="peckorum">
<emphasis id="3C900A054D6AFFFAFEE1FBA534903FFE" box="[364,506,1147,1171]" italics="true" pageId="2" pageNumber="485">C. peckorum</emphasis>
</taxonomicName>
there is no flat region on cucullus, hence it is undifferentiated from that of female; the carapace is uniformly densely granular on its entire surface; the width of the cheliceral movable finger is similar to that of the fixed finger; leg I tibia bears similarly-developed pro- and retroventral distal lobes but lacks a sub-basal protrusion, making the ventral concavity less evident and the notch is absent; leg I metatarsus is not laterally-compressed and lacks the “elbow-like” expansion; and leg III metatarsal tuft is absent. Furthermore, the posterior genital lip of the female of
<taxonomicName id="C9E4AD944D6AFFFAFDBDFAF037DE3E28" box="[560,692,1325,1349]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6AFFFAFDBDFAF037DE3E28" box="[560,692,1325,1349]" italics="true" pageId="2" pageNumber="485">C. chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6AFFFAFD36FAF3367E3E2B" box="[699,788,1325,1350]" pageId="2" pageNumber="485" rank="species">
<emphasis id="3C900A054D6AFFFAFD36FAF3367E3E2B" bold="true" box="[699,788,1325,1350]" pageId="2" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
is apically trilobated (
<figureCitation id="96DFCA924D6AFFFAFB80FAF331FC3E2B" box="[1037,1174,1325,1350]" captionStart="FIGURES 22 24" captionStartId="7.[151,264,1191,1213]" captionTargetBox="[158,1426,243,1090]" captionTargetId="figure@7.[151,1436,221,1126]" captionTargetPageId="7" captionText="FIGURES 22 24. Cryptocellus chiruisla sp. nov. Female paratype (QCAZI 3441). Spermathecae. 22 23. Posterior and anterior aspects (apically trilobated apex of the posterior genital lip indicated by arrow). 24. Frontal aspect. Scale bars: 200 µm." httpUri="https://zenodo.org/record/828739/files/figure.png" pageId="2" pageNumber="485">Figs. 2223</figureCitation>
), whereas it is not in
<taxonomicName id="C9E4AD944D6AFFFAFA0FFAF0346F3E04" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="peckorum">
<emphasis id="3C900A054D6AFFFAFA0FFAF0346F3E04" italics="true" pageId="2" pageNumber="485">C. peckorum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="46FE859C4D6AFFFFFF4AFAAA37693DE5" lastPageId="7" lastPageNumber="490" pageId="2" pageNumber="485" type="description">
<paragraph id="0E5BD6174D6AFFFAFF4AFAAA37773EE0" blockId="2.[151,1437,252,2029]" box="[199,541,1396,1421]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AFAAA34333EE0" bold="true" box="[199,345,1396,1421]" pageId="2" pageNumber="485">Description.</emphasis>
Male (
<typeStatus id="D15F68B54D6AFFFAFE25FAAA377B3EE0" box="[424,529,1396,1421]" pageId="2" pageNumber="485" type="holotype">holotype</typeStatus>
):
</paragraph>
<paragraph id="0E5BD6174D6AFFFAFF4AFA46345E3E95" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AFA4634213EDD" box="[199,331,1432,1456]" italics="true" pageId="2" pageNumber="485">Coloration:</emphasis>
<figureCitation id="96DFCA924D6AFFFAFED9FA4634A53EDC" box="[340,463,1432,1457]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 10" captionStart-2="FIGURES 11 17" captionStartId-0="3.[151,269,1660,1683]" captionStartId-1="4.[151,264,1719,1741]" captionStartId-2="5.[151,264,1809,1831]" captionTargetBox-0="[176,1420,194,1621]" captionTargetBox-1="[151,1436,193,1665]" captionTargetBox-2="[151,1436,337,1748]" captionTargetId-0="figure@3.[176,1420,194,1642]" captionTargetId-1="figure@4.[151,1436,193,1698]" captionTargetId-2="figure@5.[151,1436,337,1788]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." captionText-1="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." captionText-2="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri-0="https://zenodo.org/record/828731/files/figure.png" httpUri-1="https://zenodo.org/record/828733/files/figure.png" httpUri-2="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 114</figureCitation>
. Body and appendages immaculately reddish orange, except for, pedipalps, metatarsi and tarsi of legs, and central area of opisthosomal Sternites XIXII yellowish red. Carapacial translucent areas yellow inside.
</paragraph>
<paragraph id="0E5BD6174D6AFFFAFF4AF9DA367C3D2D" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AF9DA34413D76" box="[199,299,1540,1563]" italics="true" pageId="2" pageNumber="485">Setation:</emphasis>
<figureCitation id="96DFCA924D6AFFFAFEB9F9DD34DA3D71" box="[308,432,1539,1564]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 10" captionStart-2="FIGURES 11 17" captionStartId-0="3.[151,269,1660,1683]" captionStartId-1="4.[151,264,1719,1741]" captionStartId-2="5.[151,264,1809,1831]" captionTargetBox-0="[176,1420,194,1621]" captionTargetBox-1="[151,1436,193,1665]" captionTargetBox-2="[151,1436,337,1748]" captionTargetId-0="figure@3.[176,1420,194,1642]" captionTargetId-1="figure@4.[151,1436,193,1698]" captionTargetId-2="figure@5.[151,1436,337,1788]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." captionText-1="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." captionText-2="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri-0="https://zenodo.org/record/828731/files/figure.png" httpUri-1="https://zenodo.org/record/828733/files/figure.png" httpUri-2="https://zenodo.org/record/828735/files/figure.png" pageId="2" pageNumber="485">Figs. 114</figureCitation>
. Body and appendages covered with fine bristle-like, predominantly straight to eyelashlike, translucent setae, which are sparse in sternal region.
</paragraph>
<paragraph id="0E5BD6174D6AFFFAFF4AF99531BE3C57" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AF99534543D0F" box="[199,318,1611,1634]" italics="true" pageId="2" pageNumber="485">Carapace:</emphasis>
<figureCitation id="96DFCA924D6AFFFAFEC8F99534FB3D0E" box="[325,401,1611,1636]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="2" pageNumber="485">Figs. 1</figureCitation>
,
<figureCitation id="96DFCA924D6AFFFAFE13F99534C53D09" box="[414,431,1611,1636]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="2" pageNumber="485">6</figureCitation>
. Trapezoidal in shape, with lateral margins curved and not parallel (narrowing anteriorly); anterior margin straight in dorsal aspect, re-curved in frontal aspect; posterior margin gently re-curved; carapace as long as wide; laterally with moderate protrusion at margins at level between coxae II and III, where widest. Longitudinal translucent areas placed at the level between coxae I and II, glabrous with well-defined borders, elevated, clearly visible on either lateral or dorsal aspects. Carapace densely covered with conspicuous, rounded, iridescent tubercles throughout, except on triangle-like area on anterior third of carapace, and on lateral margins anterior to the translucent areas, where predominantly smooth. Carapace devoid of cuticular pits.
</paragraph>
<paragraph id="0E5BD6174D6AFFFAFF4AF89B36393C80" blockId="2.[151,1437,252,2029]" pageId="2" pageNumber="485">
<emphasis id="3C900A054D6AFFFAFF4AF89B34593C30" box="[199,307,1861,1885]" italics="true" pageId="2" pageNumber="485">Cucullus:</emphasis>
<figureCitation id="96DFCA924D6AFFFAFEB0F89B34C33C33" box="[317,425,1861,1886]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="2" pageNumber="485">Figs. 12</figureCitation>
,
<figureCitation id="96DFCA924D6AFFFAFE35F89834A33C30" box="[440,457,1862,1885]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="2" pageNumber="485">7</figureCitation>
. Trapezoidal, with lateral margins notably diverging anteriorly, noticeably wider than long; anterior margin straight on ventral view, with shallow median concavity on frontal view; cucullus with pronounced median elevation (visible on dorsal and ventral aspects), and anterior triangle-like flat section densely covered with tubercles similar to those of carapace; tubercles also in two patches on each side of the midline; front corners smooth. Cucullus devoid of cuticular pits and furrows.
</paragraph>
<caption id="5A9B869F4D6BFFFBFF1AF9A231CF3DDE" httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="3" pageNumber="486" targetBox="[176,1420,194,1621]" targetPageId="3">
<paragraph id="0E5BD6174D6BFFFBFF1AF9A231CF3DDE" blockId="3.[151,1436,1659,1715]" pageId="3" pageNumber="486">
<emphasis id="3C900A054D6BFFFBFF1AF9A2342E3DFE" bold="true" box="[151,324,1660,1683]" pageId="3" pageNumber="486">FIGURES 14.</emphasis>
<taxonomicName id="C9E4AD944D6BFFFBFEC1F9A537513DFE" box="[332,571,1659,1683]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6BFFFBFEC1F9A537513DFE" box="[332,571,1659,1683]" italics="true" pageId="3" pageNumber="486">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6BFFFBFDCEF9A237F03DFE" box="[579,666,1660,1683]" pageId="3" pageNumber="486" rank="species">
<emphasis id="3C900A054D6BFFFBFDCEF9A237F03DFE" bold="true" box="[579,666,1660,1683]" pageId="3" pageNumber="486">sp. nov.</emphasis>
</taxonomicNameLabel>
Male holotype (QCAZI 3440).
<emphasis id="3C900A054D6BFFFBFC79F9A231623DFE" bold="true" box="[1012,1032,1660,1683]" pageId="3" pageNumber="486">1.</emphasis>
Carapace, dorsal aspect.
<emphasis id="3C900A054D6BFFFBFA91F9A2305A3DFE" bold="true" box="[1308,1328,1660,1683]" pageId="3" pageNumber="486">2.</emphasis>
Prosoma, ventral aspect (sternal region).
<emphasis id="3C900A054D6BFFFBFE6FF942377A3DDE" bold="true" box="[482,528,1692,1715]" pageId="3" pageNumber="486">34.</emphasis>
Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm.
</paragraph>
</caption>
<paragraph id="0E5BD6174D6BFFFBFF4AF90130F63C2D" blockId="3.[151,1437,1759,2036]" pageId="3" pageNumber="486">
<emphasis id="3C900A054D6BFFFBFF4AF90134213D9A" box="[199,331,1759,1783]" italics="true" pageId="3" pageNumber="486">Chelicerae:</emphasis>
<figureCitation id="96DFCA924D6BFFFBFED9F90134F63D9A" box="[340,412,1759,1784]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="3" pageNumber="486">Fig. 7</figureCitation>
. Movable finger almost twice longer than fixed finger and more robust; movable fingers apically truncated, armed with seven/nine small and sharp teeth which progressively increase in size distally; fixed fingers with row of five/six teeth, the distalmost greatly enlarged as compared to the others which small and sharp.
</paragraph>
<paragraph id="0E5BD6174D6BFFFBFF4AF892370A3C99" blockId="3.[151,1437,1759,2036]" pageId="3" pageNumber="486">
<emphasis id="3C900A054D6BFFFBFF4AF892341B3C0E" box="[199,369,1868,1892]" italics="true" pageId="3" pageNumber="486">Sternal region:</emphasis>
<figureCitation id="96DFCA924D6BFFFBFEF6F89534AB3C09" box="[379,449,1867,1892]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="3" pageNumber="486">Fig. 2</figureCitation>
. Coxae I not meeting tritosternum; coxae II meeting along their posterior half, IIIIV meeting entirely; IIIV progressively decreasing in length; coxae II with anterior and posterior margins subparallel, not perpendicular to the median axis but inclined anteriorly; suture line of coxae II about half the length of that of coxae III and IV (coxae III are larger). Cuticle with scarce tubercles, similar to those of carapace, along coxal margins and without cuticular pits.
</paragraph>
<caption id="5A9B869F4D6CFFFCFF1AF96936223C44" httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="4" pageNumber="487" targetBox="[151,1436,193,1665]" targetPageId="4">
<paragraph id="0E5BD6174D6CFFFCFF1AF96936223C44" blockId="4.[151,1436,1719,1833]" pageId="4" pageNumber="487">
<emphasis id="3C900A054D6CFFFCFF1AF96934223DA0" bold="true" box="[151,328,1719,1741]" pageId="4" pageNumber="487">FIGURES 510.</emphasis>
<taxonomicName id="C9E4AD944D6CFFFCFEDDF96937593DA0" box="[336,563,1719,1741]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6CFFFCFEDDF96937593DA0" box="[336,563,1719,1741]" italics="true" pageId="4" pageNumber="487">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6CFFFCFDB6F96937E73DA0" box="[571,653,1719,1741]" pageId="4" pageNumber="487" rank="species">
<emphasis id="3C900A054D6CFFFCFDB6F96937E73DA0" bold="true" box="[571,653,1719,1741]" pageId="4" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
Male holotype (QCAZI 3440).
<emphasis id="3C900A054D6CFFFCFC54F96936863DA0" bold="true" box="[985,1004,1719,1741]" pageId="4" pageNumber="487">5.</emphasis>
Opisthosoma, lateral aspect.
<emphasis id="3C900A054D6CFFFCFA91F96930453DA0" bold="true" box="[1308,1327,1719,1741]" pageId="4" pageNumber="487">6.</emphasis>
Carapace, detail of translucent area.
<emphasis id="3C900A054D6CFFFCFE13F90834DB3D81" bold="true" box="[414,433,1750,1772]" pageId="4" pageNumber="487">7.</emphasis>
Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow).
<emphasis id="3C900A054D6CFFFCFF1AF92A35C03C67" bold="true" box="[151,170,1780,1802]" pageId="4" pageNumber="487">8.</emphasis>
Opisthosoma, pygidium, posterior view.
<emphasis id="3C900A054D6CFFFCFDDEF92A370F3C67" bold="true" box="[595,613,1780,1802]" pageId="4" pageNumber="487">9.</emphasis>
Right leg IV, retrolateral aspect.
<emphasis id="3C900A054D6CFFFCFC30F92A36B73C67" bold="true" box="[957,989,1780,1802]" pageId="4" pageNumber="487">10.</emphasis>
Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10).
</paragraph>
</caption>
<paragraph id="0E5BD6174D6CFFFDFF4AF88E301C3A70" blockId="4.[151,1436,1872,2005]" lastBlockId="5.[151,1437,151,285]" lastPageId="5" lastPageNumber="488" pageId="4" pageNumber="487">
<emphasis id="3C900A054D6CFFFCFF4AF88E340E3C05" box="[199,356,1872,1896]" italics="true" pageId="4" pageNumber="487">Opisthosoma:</emphasis>
<figureCitation id="96DFCA924D6CFFFCFEE1F88E34BD3C04" box="[364,471,1872,1897]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1660,1683]" captionTargetBox="[176,1420,194,1621]" captionTargetId="figure@3.[176,1420,194,1642]" captionTargetPageId="3" captionText="FIGURES 1 4. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 1. Carapace, dorsal aspect. 2. Prosoma, ventral aspect (sternal region). 3 4. Opisthosoma, dorsal and ventral aspects. Scale bars: 500 µm." httpUri="https://zenodo.org/record/828731/files/figure.png" pageId="4" pageNumber="487">Figs. 34</figureCitation>
,
<figureCitation id="96DFCA924D6CFFFCFE69F88E377A3C04" box="[484,528,1872,1897]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="4" pageNumber="487">5, 8</figureCitation>
. Oblong truncate, longer than wide, widest at level of tergite XII. Median plates of tergites XIXIII with paired anterolateral depressions; lateral margins converging posteriorly on XI, approximately parallel on XII, converging anteriorly on XIII; median plate of tergites XI and XII wider than long, that of XIII approximately as wide as long, that of X slit-like trapezoidal. Central region of tergite XI median plate with pronounced, elevated rounded bulge; tergite XIII median plate with rear corners in 90° angle (not protruding laterally). Dorsal and ventral surfaces with tubercles similar to those of carapace and without cuticular pits. Sternites XIXIII with paired anterolateral depressions similar to those of tergites and containing densely packed tubercles. Basal segment of pygidium with deep notch on dorsal posterior border; ventral border without notch.
</paragraph>
<caption id="5A9B869F4D6DFFFDFF1AF8CF301C3CCF" httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="5" pageNumber="488" targetBox="[151,1436,337,1748]" targetPageId="5">
<paragraph id="0E5BD6174D6DFFFDFF1AF8CF301C3CCF" blockId="5.[151,1436,1809,1954]" pageId="5" pageNumber="488">
<emphasis id="3C900A054D6DFFFDFF1AF8CF34383C4A" bold="true" box="[151,338,1809,1831]" pageId="5" pageNumber="488">FIGURES 1117.</emphasis>
<taxonomicName id="C9E4AD944D6DFFFDFED5F8CF37503C4A" box="[344,570,1809,1831]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6DFFFDFED5F8CF37503C4A" box="[344,570,1809,1831]" italics="true" pageId="5" pageNumber="488">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6DFFFDFDCDF8CF37FA3C4A" box="[576,656,1809,1831]" pageId="5" pageNumber="488" rank="species">
<emphasis id="3C900A054D6DFFFDFDCDF8CF37FA3C4A" bold="true" box="[576,656,1809,1831]" pageId="5" pageNumber="488">sp. nov.</emphasis>
</taxonomicNameLabel>
Male holotype (QCAZI 3440).
<emphasis id="3C900A054D6DFFFDFC59F8CF31733C4A" bold="true" box="[980,1049,1809,1831]" pageId="5" pageNumber="488">1112.</emphasis>
Left leg III, prolateral and retrolateral aspects.
<emphasis id="3C900A054D6DFFFDFF62F8EE345E3C2B" bold="true" box="[239,308,1840,1862]" pageId="5" pageNumber="488">1314.</emphasis>
Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “elbow-like” expansion indicated by arrow).
<emphasis id="3C900A054D6DFFFDFDE0F89037D93C09" bold="true" box="[621,691,1870,1892]" pageId="5" pageNumber="488">1517.</emphasis>
Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations:
<emphasis id="3C900A054D6DFFFDFC5EF8B036813CEE" box="[979,1003,1902,1923]" italics="true" pageId="5" pageNumber="488">ac</emphasis>
, accessory piece;
<emphasis id="3C900A054D6DFFFDFB2AF8B331B93CEE" box="[1191,1235,1901,1923]" italics="true" pageId="5" pageNumber="488">M-tf</emphasis>
, metatarsal tuft;
<emphasis id="3C900A054D6DFFFDFA0FF8B030FC3CEE" box="[1410,1430,1902,1923]" italics="true" pageId="5" pageNumber="488">tP</emphasis>
, tarsal process;
<emphasis id="3C900A054D6DFFFDFEA0F85334323CCF" box="[301,344,1933,1954]" italics="true" pageId="5" pageNumber="488">T-vn</emphasis>
, tibial ventral “notch”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 1517).
</paragraph>
</caption>
<caption id="5A9B869F4D6EFFFEFF1AFA7230633E8D" httpUri="https://zenodo.org/record/828737/files/figure.png" pageId="6" pageNumber="489" targetBox="[157,1421,193,1422]" targetPageId="6">
<paragraph id="0E5BD6174D6EFFFEFF1AFA7230633E8D" blockId="6.[151,1436,1452,1504]" pageId="6" pageNumber="489">
<emphasis id="3C900A054D6EFFFEFF1AFA72343E3EAF" bold="true" box="[151,340,1452,1474]" pageId="6" pageNumber="489">FIGURES 1821.</emphasis>
<taxonomicName id="C9E4AD944D6EFFFEFED6FA7237543EAF" box="[347,574,1452,1474]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="489" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6EFFFEFED6FA7237543EAF" box="[347,574,1452,1474]" italics="true" pageId="6" pageNumber="489">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6EFFFEFDC9FA7237FC3EAF" box="[580,662,1452,1474]" pageId="6" pageNumber="489" rank="species">
<emphasis id="3C900A054D6EFFFEFDC9FA7237FC3EAF" bold="true" box="[580,662,1452,1474]" pageId="6" pageNumber="489">sp. nov.</emphasis>
</taxonomicNameLabel>
Female paratype (QCAZI 3441).
<emphasis id="3C900A054D6EFFFEFC7FFA7231783EAF" bold="true" box="[1010,1042,1452,1474]" pageId="6" pageNumber="489">18.</emphasis>
Carapace, dorsal aspect.
<emphasis id="3C900A054D6EFFFEFA9BFA72305C3EAF" bold="true" box="[1302,1334,1452,1474]" pageId="6" pageNumber="489">19.</emphasis>
Prosoma, ventral aspect (sternal region).
<emphasis id="3C900A054D6EFFFEFE5CFA14349B3E8D" bold="true" box="[465,497,1482,1504]" pageId="6" pageNumber="489">20.</emphasis>
Opisthosoma, dorsal aspect.
<emphasis id="3C900A054D6EFFFEFC94FA1436533E8D" bold="true" box="[793,825,1482,1504]" pageId="6" pageNumber="489">21.</emphasis>
Opisthosoma, pygidium. Scale bars: 500 µm.
</paragraph>
</caption>
<paragraph id="0E5BD6174D6EFFFEFF4AF9D6361A3DE0" blockId="6.[151,1436,1544,2037]" pageId="6" pageNumber="489">
<emphasis id="3C900A054D6EFFFEFF4AF9D634553D4D" box="[199,319,1544,1568]" italics="true" pageId="6" pageNumber="489">Pedipalps:</emphasis>
<figureCitation id="96DFCA924D6EFFFEFEC5F9D634F13D4D" box="[328,411,1544,1569]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="6" pageNumber="489">Fig. 10</figureCitation>
. Without cuticular pits; with few tubercles restricted to trochanters I, II, and base of femur on retrolateral aspect. Femur dorsally convex, widened in basal half. Tibia longer than femur, with dorsal surface straight, slightly widened ventrally in basal third and without tubercles. Movable claw about twice the length of fixed claw and more robust; both claws armed with minute teeth.
</paragraph>
<paragraph id="0E5BD6174D6EFFFEFF4AF947374D3C99" blockId="6.[151,1436,1544,2037]" pageId="6" pageNumber="489">
<emphasis id="3C900A054D6EFFFEFF4AF947346C3DDD" box="[199,262,1689,1712]" italics="true" pageId="6" pageNumber="489">Legs:</emphasis>
<figureCitation id="96DFCA924D6EFFFEFE80F94634373DDD" box="[269,349,1688,1713]" captionStart="FIGURES 5 10" captionStartId="4.[151,264,1719,1741]" captionTargetBox="[151,1436,193,1665]" captionTargetId="figure@4.[151,1436,193,1698]" captionTargetPageId="4" captionText="FIGURES 5 10. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 5. Opisthosoma, lateral aspect. 6. Carapace, detail of translucent area. 7. Cucullus and chelicerae, frontal aspect (truncated apex of the movable finger indicated by arrow). 8. Opisthosoma, pygidium, posterior view. 9. Right leg IV, retrolateral aspect. 10. Left pedipalp tibia, prolateral aspect. Scale bars: 500 µm (Figs. 5, 9); 300 µm (Figs. 7, 8); 200 µm (Figs. 6, 10)." httpUri="https://zenodo.org/record/828733/files/figure.png" pageId="6" pageNumber="489">Figs. 9</figureCitation>
,
<figureCitation id="96DFCA924D6EFFFEFEE6F94634D93DDD" box="[363,435,1688,1712]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="6" pageNumber="489">1114</figureCitation>
. Without cuticular pits; leg segments with tubercles similar to those of carapace. Leg II longest; legs progressively increasing in width (i.e., at femur) in the order II &lt;III &lt;I &lt;IV. Tibia of leg I with pronounced ventral “notch” formed by well-developed, ventrally protruding pro- (large) and retrolateral (small) distal lobes, a tubercle-like protrusion pro-ventrally on basal half of segment, and a deep concavity in the middle; leg I metatarsus laterally-compressed, prolaterally sunken on basal two thirds, and with a well-developed, proventral distal “elbow-like” expansion. Legs I and II, femur ventrally with flat, basal apophyses, absent on legs III and IV; apophyses are projected proximally and fully (leg I) or partially (leg II) cover the articular membranes between femur and trochanter. Leg III metatarsus moderately inflated, deeply excavated, with disto-ventral tuft of long setae and metatarsal process as in
<figureCitation id="96DFCA924D6EFFFEFDE9F86637873CBD" box="[612,749,1976,2001]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="6" pageNumber="489">Figs. 1112</figureCitation>
; lamina cyathiformis higher than long, with apex distinctly rounded. Leg length formula: 2341.
</paragraph>
<caption id="5A9B869F4D6FFFFFFF1AFB7930F63FB6" httpUri="https://zenodo.org/record/828739/files/figure.png" pageId="7" pageNumber="490" targetBox="[158,1426,243,1090]" targetPageId="7">
<paragraph id="0E5BD6174D6FFFFFFF1AFB7930F63FB6" blockId="7.[151,1436,1191,1243]" pageId="7" pageNumber="490">
<emphasis id="3C900A054D6FFFFFFF1AFB7934333FD0" bold="true" box="[151,345,1191,1213]" pageId="7" pageNumber="490">FIGURES 2224.</emphasis>
<taxonomicName id="C9E4AD944D6FFFFFFEE9FB7937213FD0" box="[356,587,1191,1213]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6FFFFFFEE9FB7937213FD0" box="[356,587,1191,1213]" italics="true" pageId="7" pageNumber="490">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6FFFFFFDDBFB7937C63FD0" box="[598,684,1191,1213]" pageId="7" pageNumber="490" rank="species">
<emphasis id="3C900A054D6FFFFFFDDBFB7937C63FD0" bold="true" box="[598,684,1191,1213]" pageId="7" pageNumber="490">sp. nov.</emphasis>
</taxonomicNameLabel>
Female paratype (QCAZI 3441). Spermathecae.
<emphasis id="3C900A054D6FFFFFFB32FB79306F3FD0" bold="true" box="[1215,1285,1191,1213]" pageId="7" pageNumber="490">2223.</emphasis>
Posterior and anterior aspects (apically trilobated apex of the posterior genital lip indicated by arrow).
<emphasis id="3C900A054D6FFFFFFB9EFB1B31593FB6" bold="true" box="[1043,1075,1221,1243]" pageId="7" pageNumber="490">24.</emphasis>
Frontal aspect. Scale bars: 200 µm.
</paragraph>
</caption>
<paragraph id="0E5BD6174D6FFFFFFF4AFAD630273EB9" blockId="7.[151,1437,1288,1961]" pageId="7" pageNumber="490">
<emphasis id="3C900A054D6FFFFFFF4AFAD634A33E4D" box="[199,457,1288,1312]" italics="true" pageId="7" pageNumber="490">Copulatory apparatus:</emphasis>
<figureCitation id="96DFCA924D6FFFFFFE5CFAD637C43E4C" box="[465,686,1288,1313]" captionStart="FIGURES 11 17" captionStartId="5.[151,264,1809,1831]" captionTargetBox="[151,1436,337,1748]" captionTargetId="figure@5.[151,1436,337,1788]" captionTargetPageId="5" captionText="FIGURES 11 17. Cryptocellus chiruisla sp. nov. Male holotype (QCAZI 3440). 11 12. Left leg III, prolateral and retrolateral aspects. 13 14. Left leg I, dorsal and prolateral aspects (metatarsal lateral compression indicated by arrowheads; metatarsal “ elbow-like ” expansion indicated by arrow). 15 17. Copulatory apparatus in prolateral, frontal (note the apparently-striated longitudinal opening of the tarsal process) and retrolateral aspects. Annotations: ac, accessory piece; M-tf, metatarsal tuft; tP, tarsal process; T-vn, tibial ventral “ notch ”. Scale bars: 300 µm (Figs. 11, 12); 500 µm (Figs. 13, 14); 200 µm (Figs. 15 17)." httpUri="https://zenodo.org/record/828735/files/figure.png" pageId="7" pageNumber="490">Figs. 1112, 1517</figureCitation>
. Tarsal process U-shaped; apex directed dorsally, moderately long and gently curved. Tarsal process with a longitudinal opening on ventral subdistal area; pro- and retrolateral edges of the opening are hyaline, and the internal surface of the opening is apparently striated; retrolateral edge more developed than the prolateral; both edges smooth in lateral aspect, without spiniform projections. Accessory piece short (i.e., compared to the tarsal process), predominantly straight, narrowing apically and single-tipped (nonbifid); accessory piece lying along a shallow longitudinal groove on retrolateral aspect of the tarsal process.
</paragraph>
<paragraph id="0E5BD6174D6FFFFFFF4AFA3F37693E95" blockId="7.[151,1437,1288,1961]" box="[199,515,1504,1529]" pageId="7" pageNumber="490">
<emphasis id="3C900A054D6FFFFFFF4AFA3F34183E95" box="[199,370,1505,1528]" italics="true" pageId="7" pageNumber="490">Measurements:</emphasis>
See
<tableCitation id="4366E3AC4D6FFFFFFE24FA3E34963E95" box="[425,508,1504,1529]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Measurements of Cryptocellus chiruisla sp. nov. and Cryptocellus guaviarensis sp. nov. Measurements in millimeters; L = length; W = width; H = height. 1 It excludes pygidium and cucullus lenght. 2 Greatest width. 3 Greatest height. 4 Sum of individual segment lengths, excluding coxa. * Measured at level between coxae II and III, where widest. ** Measured at level of tergite XII, where widest." pageId="7" pageNumber="490">Table 1</tableCitation>
.
</paragraph>
<paragraph id="0E5BD6174D6FFFFFFF4AF9DA375D3D08" blockId="7.[151,1437,1288,1961]" pageId="7" pageNumber="490">
<emphasis id="3C900A054D6FFFFFFF4AF9DA344F3D70" bold="true" box="[199,293,1540,1565]" pageId="7" pageNumber="490">Female.</emphasis>
<figureCitation id="96DFCA924D6FFFFFFEA1F9DA34DE3D70" box="[300,436,1540,1565]" captionStart-0="FIGURES 18 21" captionStart-1="FIGURES 22 24" captionStartId-0="6.[151,264,1452,1474]" captionStartId-1="7.[151,264,1191,1213]" captionTargetBox-0="[157,1421,193,1422]" captionTargetBox-1="[158,1426,243,1090]" captionTargetId-0="figure@6.[157,1421,193,1422]" captionTargetId-1="figure@7.[151,1436,221,1126]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 18 21. Cryptocellus chiruisla sp. nov. Female paratype (QCAZI 3441). 18. Carapace, dorsal aspect. 19. Prosoma, ventral aspect (sternal region). 20. Opisthosoma, dorsal aspect. 21. Opisthosoma, pygidium. Scale bars: 500 µm." captionText-1="FIGURES 22 24. Cryptocellus chiruisla sp. nov. Female paratype (QCAZI 3441). Spermathecae. 22 23. Posterior and anterior aspects (apically trilobated apex of the posterior genital lip indicated by arrow). 24. Frontal aspect. Scale bars: 200 µm." httpUri-0="https://zenodo.org/record/828737/files/figure.png" httpUri-1="https://zenodo.org/record/828739/files/figure.png" pageId="7" pageNumber="490">Figs. 1824</figureCitation>
. Similar to the male in several aspects, but with sex-related differences. Spermathecae as in
<figureCitation id="96DFCA924D6FFFFFFF3BF9F6342A3D2D" box="[182,320,1576,1601]" captionStart="FIGURES 22 24" captionStartId="7.[151,264,1191,1213]" captionTargetBox="[158,1426,243,1090]" captionTargetId="figure@7.[151,1436,221,1126]" captionTargetPageId="7" captionText="FIGURES 22 24. Cryptocellus chiruisla sp. nov. Female paratype (QCAZI 3441). Spermathecae. 22 23. Posterior and anterior aspects (apically trilobated apex of the posterior genital lip indicated by arrow). 24. Frontal aspect. Scale bars: 200 µm." httpUri="https://zenodo.org/record/828739/files/figure.png" pageId="7" pageNumber="490">Figs. 2224</figureCitation>
, wider than long, not touching each other; posterior genital lip with apex sub-triangular, apically trilobated. Leg length formula: 2341.
</paragraph>
<paragraph id="0E5BD6174D6FFFFFFF4AF9AF37693DE5" blockId="7.[151,1437,1288,1961]" box="[199,515,1648,1673]" pageId="7" pageNumber="490">
<emphasis id="3C900A054D6FFFFFFF4AF9AF34183DE5" box="[199,370,1649,1672]" italics="true" pageId="7" pageNumber="490">Measurements:</emphasis>
See
<tableCitation id="4366E3AC4D6FFFFFFE24F9AE34963DE5" box="[425,508,1648,1673]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Measurements of Cryptocellus chiruisla sp. nov. and Cryptocellus guaviarensis sp. nov. Measurements in millimeters; L = length; W = width; H = height. 1 It excludes pygidium and cucullus lenght. 2 Greatest width. 3 Greatest height. 4 Sum of individual segment lengths, excluding coxa. * Measured at level between coxae II and III, where widest. ** Measured at level of tergite XII, where widest." pageId="7" pageNumber="490">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46FE859C4D6FFFFFFF4AF94A34C53CC4" pageId="7" pageNumber="490" type="distribution">
<paragraph id="0E5BD6174D6FFFFFFF4AF94A36B63DC0" blockId="7.[151,1437,1288,1961]" box="[199,988,1684,1709]" pageId="7" pageNumber="490">
<emphasis id="3C900A054D6FFFFFFF4AF94A340A3DC0" bold="true" box="[199,352,1684,1709]" pageId="7" pageNumber="490">Distribution.</emphasis>
Known only from the
<typeStatus id="D15F68B54D6FFFFFFDEDF94B37FA3DC0" box="[608,656,1685,1709]" pageId="7" pageNumber="490">type</typeStatus>
locality in
<collectingCountry id="76F396874D6FFFFFFC82F94A36063DC0" box="[783,876,1684,1709]" name="Ecuador" pageId="7" pageNumber="490">Ecuador</collectingCountry>
(
<figureCitation id="96DFCA924D6FFFFFFCF1F94A36A53DC0" box="[892,975,1684,1709]" captionStart="FIGURE 41" captionStartId="14.[151,250,863,885]" captionTargetBox="[151,1436,193,842]" captionTargetId="figure@14.[151,1436,193,842]" captionTargetPageId="14" captionText="FIGURE 41. Map of Northwestern South America, plotting the known distribution of Cryptocellus chiruisla sp. nov. and Cryptocellus guaviarensis sp. nov." httpUri="https://zenodo.org/record/828747/files/figure.png" pageId="7" pageNumber="490">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="0E5BD6174D6FFFFFFF4AF96634C53CC4" blockId="7.[151,1437,1288,1961]" pageId="7" pageNumber="490">
<materialsCitation id="BE8CDC4A4D6FFFFFFF4AF96634C53CC4" ID-GBIF-Occurrence="1572325257" country="Ecuador" county="The" latitude="-1.4088888" location="Paz" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" pageId="7" pageNumber="490" specimenCount="2" stateProvince="Rio Negro" typeStatus="holotype">
<emphasis id="3C900A054D6FFFFFFF4AF966347B3DBC" bold="true" box="[199,273,1720,1745]" pageId="7" pageNumber="490">Notes.</emphasis>
Two other ricinuleid species have been described from
<collectingCountry id="76F396874D6FFFFFFC0CF966368A3DBC" box="[897,992,1720,1745]" name="Ecuador" pageId="7" pageNumber="490">Ecuador</collectingCountry>
,
<taxonomicName id="C9E4AD944D6FFFFFFC66F96634B93D98" authority="Botero-Trujillo &amp; Valdez-Mondragon, 2016" authorityName="Botero-Trujillo &amp; Valdez-Mondragon" authorityYear="2016" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="chimaera">
<emphasis id="3C900A054D6FFFFFFC66F96631813DBD" box="[1003,1259,1720,1744]" italics="true" pageId="7" pageNumber="490">Cryptocellus chimaera</emphasis>
<bibRefCitation id="6A75ABE64D6FFFFFFB7FF96634B93D98" author="Botero-Trujillo" pageId="7" pageNumber="490" refString="Botero-Trujillo, R. &amp; Valdez-Mondragon, A. (2016) A remarkable new species of the magnus species-group of Cryptocellus (Arachnida, Ricinulei) from Ecuador, with observations on the taxonomy of the New World genera. Zootaxa, 4107 (3), 321 - 337." type="journal article" year="2016">Botero-Trujillo &amp; Valdez-Mondragón, 2016</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="C9E4AD944D6FFFFFFD86F90236D93D98" authority="Cooreman, 1976" authorityName="Cooreman" authorityYear="1976" box="[523,947,1756,1781]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="3C900A054D6FFFFFFD86F90237843D99" box="[523,750,1756,1780]" italics="true" pageId="7" pageNumber="490">Cryptocellus leleupi</emphasis>
<bibRefCitation id="6A75ABE64D6FFFFFFD78F90336D93D98" author="Cooreman" box="[757,947,1756,1781]" pageId="7" pageNumber="490" refString="Cooreman, J. (1976) Description d'une stase nymphale d'un Ricinulei de l'Ecuador, Cryptocellus leleupi n. sp. In: Leleup, N. (Ed.), Mission Zoologique Belge aux iles Galapagos et un Ecuador (N. et J. Leleup, 1964 - 1965). Uol. 3. Koninklijk Museum voor Midden-Afrika, Bruxelles, pp. 25 - 52." type="journal article" year="1976">Cooreman, 1976</bibRefCitation>
</taxonomicName>
.
<collectingCounty id="E73AAE9B4D6FFFFFFC32F90236813D98" box="[959,1003,1756,1781]" pageId="7" pageNumber="490">The</collectingCounty>
former species belongs to the
<taxonomicName id="C9E4AD944D6FFFFFFAC8F90330F63D99" box="[1349,1436,1757,1780]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="3C900A054D6FFFFFFAC8F90330F63D99" box="[1349,1436,1757,1780]" italics="true" pageId="7" pageNumber="490">magnus</emphasis>
</taxonomicName>
species-groups, whereas the latter is a
<emphasis id="3C900A054D6FFFFFFDC9F8DF37833C75" box="[580,745,1792,1816]" italics="true" pageId="7" pageNumber="490">nomen dubium</emphasis>
of uncertain systematic position (
<bibRefCitation id="6A75ABE64D6FFFFFFBEEF8DE30253C75" author="Platnick" box="[1123,1359,1792,1817]" pageId="7" pageNumber="490" refString="Platnick, N. I. &amp; Paz, N. (1979) On the Cryptocellus magnus group (Arachnida, Ricinulei). American Museum Novitates, 2677, 1 - 9." type="journal article" year="1979">
<collectingMunicipality id="EE3F4C6D4D6FFFFFFBEEF8DE31AA3C74" box="[1123,1216,1792,1817]" pageId="7" pageNumber="490">Platnick</collectingMunicipality>
&amp;
<location id="0B3B80CC4D6FFFFFFB69F8DF30673C75" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFB69F8DF30673C75" box="[1252,1293,1793,1816]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="Paz" pageId="7" pageNumber="490" stateProvince="Rio Negro">Paz</location>
1979
</bibRefCitation>
).
<location id="0B3B80CC4D6FFFFFFAEFF8DF30F63C75" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFAEFF8DF30F63C75" box="[1378,1436,1793,1816]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="Even" pageId="7" pageNumber="490" stateProvince="Rio Negro">Even</location>
though the adult morphology of
<taxonomicName id="C9E4AD944D6FFFFFFD85F8FB371C3C51" box="[520,630,1828,1852]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="3C900A054D6FFFFFFD85F8FB371C3C51" box="[520,630,1828,1852]" italics="true" pageId="7" pageNumber="490">C. leleupi</emphasis>
</taxonomicName>
is unknown (it was described from a protonymph), the
<typeStatus id="D15F68B54D6FFFFFFB7DF8FB304A3C50" box="[1264,1312,1829,1853]" pageId="7" pageNumber="490">type</typeStatus>
locality of this species (
<location id="0B3B80CC4D6FFFFFFEAAF89634153C0C" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFEAAF89634153C0C" box="[295,383,1864,1889]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="Oriente" pageId="7" pageNumber="490" stateProvince="Rio Negro">Oriente</location>
,
<collectingRegion id="CC2018F54D6FFFFFFE01F896376F3C0C" box="[396,517,1864,1889]" country="Argentina" name="Rio Negro" pageId="7" pageNumber="490">Rio Negro</collectingRegion>
, approx.
<geoCoordinate id="6BD0B0D04D6FFFFFFDE0F896379C3C0D" box="[621,758,1864,1889]" direction="south" orientation="latitude" pageId="7" pageNumber="490" precision="15" value="-1.4088888">01°2432″ S</geoCoordinate>
<geoCoordinate id="6BD0B0D04D6FFFFFFD70F89636FB3C0D" box="[765,913,1864,1889]" direction="west" orientation="longitude" pageId="7" pageNumber="490" precision="15" value="-78.19111">78°1128″ W</geoCoordinate>
) is located in the
<location id="0B3B80CC4D6FFFFFFBD6F89731C53C0D" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFBD6F89731C53C0D" box="[1115,1199,1865,1888]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="Eastern" pageId="7" pageNumber="490" stateProvince="Rio Negro">Eastern</location>
slopes of the
<location id="0B3B80CC4D6FFFFFFAC3F89630F23C0C" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFAC3F89630F23C0C" box="[1358,1432,1864,1889]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="Andes" pageId="7" pageNumber="490" stateProvince="Rio Negro">Andes</location>
, some
<quantity id="C91C7BF24D6FFFFFFF54F8B334443CE8" box="[217,302,1900,1925]" metricMagnitude="5" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="490" unit="km" value="270.0">270 km</quantity>
<location id="0B3B80CC4D6FFFFFFEB9F8B234C73CE9" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFEB9F8B234C73CE9" box="[308,429,1900,1925]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="South East" pageId="7" pageNumber="490" stateProvince="Rio Negro">South East</location>
of that of
<taxonomicName id="C9E4AD944D6FFFFFFDACF8B337CF3CE9" box="[545,677,1900,1924]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D6FFFFFFDACF8B337CF3CE9" box="[545,677,1900,1924]" italics="true" pageId="7" pageNumber="490">C. chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D6FFFFFFD21F8B2366F3CE8" box="[684,773,1900,1925]" pageId="7" pageNumber="490" rank="species">
<emphasis id="3C900A054D6FFFFFFD21F8B2366F3CE8" bold="true" box="[684,773,1900,1925]" pageId="7" pageNumber="490">sp. nov.</emphasis>
</taxonomicNameLabel>
<location id="0B3B80CC4D6FFFFFFC81F8B236523CE8" LSID="urn:lsid:plazi:treatment:864D67014D6AFFF1FF1AFF49310F38C8:0B3B80CC4D6FFFFFFC81F8B236523CE8" box="[780,824,1900,1925]" country="Ecuador" county="The" latitude="-1.4088888" longLatPrecision="21" longitude="-78.19111" municipality="Platnick" name="The" pageId="7" pageNumber="490" stateProvince="Rio Negro">The</location>
probability that the
<typeStatus id="D15F68B54D6FFFFFFB97F8B231ED3CE8" box="[1050,1159,1900,1925]" pageId="7" pageNumber="490" type="holotype">holotypes</typeStatus>
of both species could be conspecific is negligible.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="5A9B869F4D60FFF0FF1AFF4637C13A79" pageId="8" pageNumber="491">
<paragraph id="0E5BD6174D60FFF0FF1AFF4637C13A79" blockId="8.[151,1436,151,276]" pageId="8" pageNumber="491">
<emphasis id="3C900A054D60FFF0FF1AFF4634663BC2" bold="true" box="[151,268,152,175]" pageId="8" pageNumber="491">TABLE 1.</emphasis>
Measurements of
<taxonomicName id="C9E4AD944D60FFF0FE55FF4937A23BC2" box="[472,712,151,175]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="491" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D60FFF0FE55FF4937A23BC2" box="[472,712,151,175]" italics="true" pageId="8" pageNumber="491">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D60FFF0FD5CFF4636403BC2" box="[721,810,152,175]" pageId="8" pageNumber="491" rank="species">
<emphasis id="3C900A054D60FFF0FD5CFF4636403BC2" bold="true" box="[721,810,152,175]" pageId="8" pageNumber="491">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="C9E4AD944D60FFF0FCECFF4931133BC2" box="[865,1145,151,175]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="491" phylum="Arthropoda" rank="species" species="guaviarensis" status="sp. nov.">
<emphasis id="3C900A054D60FFF0FCECFF4931133BC2" box="[865,1145,151,175]" italics="true" pageId="8" pageNumber="491">Cryptocellus guaviarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D60FFF0FB0FFF4631B03BC2" box="[1154,1242,152,175]" pageId="8" pageNumber="491" rank="species">
<emphasis id="3C900A054D60FFF0FB0FFF4631B03BC2" bold="true" box="[1154,1242,152,175]" pageId="8" pageNumber="491">sp. nov.</emphasis>
</taxonomicNameLabel>
Measurements in millimeters;
<emphasis id="3C900A054D60FFF0FE92FF6434443BBC" box="[287,302,186,209]" italics="true" pageId="8" pageNumber="491">L</emphasis>
= length;
<emphasis id="3C900A054D60FFF0FE13FF6434DE3BBC" box="[414,436,186,209]" italics="true" pageId="8" pageNumber="491">W</emphasis>
= width;
<emphasis id="3C900A054D60FFF0FD93FF64375B3BBC" box="[542,561,186,209]" italics="true" pageId="8" pageNumber="491">H</emphasis>
= height. 1 It excludes pygidium and cucullus lenght. 2 Greatest width. 3 Greatest height. 4 Sum of individual segment lengths, excluding coxa. * Measured at level between coxae II and III, where widest. ** Measured at level of tergite XII, where widest.
</paragraph>
</caption>
<subSubSection id="46FE859C4D60FFF1FD2EFEEE310F38C8" lastPageId="9" lastPageNumber="492" pageId="8" pageNumber="491" type="description">
<paragraph id="0E5BD6174D60FFF0FD2EFEEE30F23A2B" blockId="8.[159,1432,304,2015]" box="[675,1432,304,326]" pageId="8" pageNumber="491">
<taxonomicName id="C9E4AD944D60FFF0FD2EFEEE36EF3A2B" box="[675,901,304,326]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="491" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">
<emphasis id="3C900A054D60FFF0FD2EFEEE36EF3A2B" box="[675,901,304,326]" italics="true" pageId="8" pageNumber="491">Cryptocellus chiruisla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D60FFF0FC06FEEE36B13A2B" box="[907,987,304,326]" pageId="8" pageNumber="491" rank="species">
<emphasis id="3C900A054D60FFF0FC06FEEE36B13A2B" bold="true" box="[907,987,304,326]" pageId="8" pageNumber="491">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="C9E4AD944D60FFF0FBB4FEEE302B3A2B" box="[1081,1345,304,326]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="491" phylum="Arthropoda" rank="species" species="guaviarensis" status="sp. nov.">
<emphasis id="3C900A054D60FFF0FBB4FEEE302B3A2B" box="[1081,1345,304,326]" italics="true" pageId="8" pageNumber="491">Cryptocellus guaviarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D60FFF0FACAFEEE30F23A2B" box="[1351,1432,304,326]" pageId="8" pageNumber="491" rank="species">
<emphasis id="3C900A054D60FFF0FACAFEEE30F23A2B" bold="true" box="[1351,1432,304,326]" pageId="8" pageNumber="491">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="0E5BD6174D60FFF0FD2EFE87310F3AF5" blockId="8.[159,1432,304,2015]" pageId="8" pageNumber="491">Male Female Male Total body L:1 4.50 4.47 4.50</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FE70310F3AA9" blockId="8.[159,1432,304,2015]" box="[159,1125,430,452]" pageId="8" pageNumber="491">Cucullus: L: 0.87 0.87 0.83</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FE05310F3A80" blockId="8.[159,1432,304,2015]" box="[457,1125,471,496]" pageId="8" pageNumber="491">W:2 1.17 1.13 1.13</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FDDD310F3974" blockId="8.[159,1432,304,2015]" box="[159,1125,515,537]" pageId="8" pageNumber="491">Carapace: L: 1.83 1.77 1.70</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FDEE3112392E" blockId="8.[159,1432,304,2015]" box="[457,1144,556,581]" pageId="8" pageNumber="491">W:2 1.83 * 1.83 * 1.70 *</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FD87310F3902" blockId="8.[159,1432,304,2015]" box="[159,1125,600,625]" pageId="8" pageNumber="491">Opisthosoma: Total L:1 2.67 2.70 2.80</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FD56310F39AB" blockId="8.[159,1432,304,2015]" pageId="8" pageNumber="491">W:2 2.50 ** 2.57 ** 2.30 ** Median plate XI: L: 0.97 0.97 0.93</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FD03310F399D" blockId="8.[159,1432,304,2015]" box="[457,1125,729,754]" pageId="8" pageNumber="491">W:2 1.23 1.57 1.27</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FCDB310F3876" blockId="8.[159,1432,304,2015]" box="[159,1125,773,795]" pageId="8" pageNumber="491">Median plate XII: L: 0.70 0.70 0.77</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FCEE310F3828" blockId="8.[159,1432,304,2015]" box="[457,1125,815,837]" pageId="8" pageNumber="491">W: 0.93 1.20 1.00</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FC86310F3803" blockId="8.[159,1432,304,2015]" box="[159,1125,856,878]" pageId="8" pageNumber="491">Median plate XIII: L: 0.80 0.87 0.90</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FC5D310F38F5" blockId="8.[159,1432,304,2015]" box="[457,1125,898,920]" pageId="8" pageNumber="491">W: 0.93 1.00 0.90</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FC75310F38AC" blockId="8.[159,1432,304,2015]" box="[159,1125,939,961]" pageId="8" pageNumber="491">Suture line coxae II: L: 0.10 0.12 0.18</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FC0A310F3887" blockId="8.[159,1432,304,2015]" box="[159,1125,980,1002]" pageId="8" pageNumber="491">Suture line coxae III: L: 0.18 0.15 0.23</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FC23310F3F7E" blockId="8.[159,1432,304,2015]" box="[159,1125,1021,1043]" pageId="8" pageNumber="491">Pedipalp: Femur L: 0.95 0.87 0.75</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FBF4310F3F50" blockId="8.[159,1432,304,2015]" box="[457,1125,1063,1087]" pageId="8" pageNumber="491">Femur H:3 0.37 0.40 0.37</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FB8D310F3F04" blockId="8.[159,1432,304,2015]" box="[457,1125,1107,1129]" pageId="8" pageNumber="491">Tibia L: 1.18 1.33 1.18</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FBA0310F3FFF" blockId="8.[159,1432,304,2015]" box="[457,1125,1148,1172]" pageId="8" pageNumber="491">Tibia H:3 0.17 0.18 0.17</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FB77310F3FD3" blockId="8.[159,1432,304,2015]" box="[159,1125,1192,1217]" pageId="8" pageNumber="491">Leg I: Total L:4 4.57 3.93 4.31</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FB0A310F3F87" blockId="8.[159,1432,304,2015]" box="[457,1125,1236,1258]" pageId="8" pageNumber="491">Trochanter L: 0.33 0.23 0.27</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FB20310F3E7E" blockId="8.[159,1432,304,2015]" box="[457,1125,1277,1299]" pageId="8" pageNumber="491">Femur L: 0.90 0.80 0.87</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FAF4310F3E51" blockId="8.[159,1432,304,2015]" box="[457,1125,1318,1343]" pageId="8" pageNumber="491">Femur W:2 0.53 0.38 0.52</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FA8C310F3E05" blockId="8.[159,1432,304,2015]" box="[457,1125,1362,1384]" pageId="8" pageNumber="491">Patella L: 0.77 0.70 0.80</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FAA2310F3EFF" blockId="8.[159,1432,304,2015]" box="[457,1125,1404,1426]" pageId="8" pageNumber="491">Tibia L: 1.07 0.73 0.87</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FA78310F3ED6" blockId="8.[159,1432,304,2015]" box="[457,1125,1445,1467]" pageId="8" pageNumber="491">Metatarsus L: 1.00 1.00 0.97</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44FA11310F3E89" blockId="8.[159,1432,304,2015]" box="[457,1125,1486,1508]" pageId="8" pageNumber="491">Tarsus L: 0.50 0.47 0.53</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12FA27310F3D63" blockId="8.[159,1432,304,2015]" box="[159,1125,1527,1552]" pageId="8" pageNumber="491">Leg II: Total L:4 6.84 6.53 7.24</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F9FA310F3D57" blockId="8.[159,1432,304,2015]" box="[457,1125,1572,1594]" pageId="8" pageNumber="491">Trochanter L: 0.40 0.33 0.37</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F990310F3D0E" blockId="8.[159,1432,304,2015]" box="[457,1125,1613,1635]" pageId="8" pageNumber="491">Femur L: 1.37 1.33 1.50</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F9A4310F3DE1" blockId="8.[159,1432,304,2015]" box="[457,1125,1654,1679]" pageId="8" pageNumber="491">Femur W:2 0.42 0.45 0.42</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F97B310F3DD5" blockId="8.[159,1432,304,2015]" box="[457,1125,1698,1722]" pageId="8" pageNumber="491">Femur H:3 0.48 0.50 0.48</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F910310F3D89" blockId="8.[159,1432,304,2015]" box="[457,1125,1742,1764]" pageId="8" pageNumber="491">Patella L: 1.00 0.97 1.07</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F929310F3C60" blockId="8.[159,1432,304,2015]" box="[457,1125,1783,1805]" pageId="8" pageNumber="491">Tibia L: 1.13 1.17 1.23</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F8FC310F3C5A" blockId="8.[159,1432,304,2015]" box="[457,1125,1825,1847]" pageId="8" pageNumber="491">Metatarsus L: 1.47 1.40 1.57</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F895310F3C0D" blockId="8.[159,1432,304,2015]" box="[457,1125,1866,1888]" pageId="8" pageNumber="491">Tarsus L: 1.47 1.33 1.50</paragraph>
<paragraph id="0E5BD6174D60FFF0FF12F8AA310F3CE4" blockId="8.[159,1432,304,2015]" box="[159,1125,1907,1932]" pageId="8" pageNumber="491">Leg III: Total L:4 6.03 5.23 6.26</paragraph>
<paragraph id="0E5BD6174D60FFF0FE44F841310F3CD8" blockId="8.[159,1432,304,2015]" box="[457,1125,1951,1973]" pageId="8" pageNumber="491">1st trochanter L: 0.30 0.30 0.37</paragraph>
<paragraph id="0E5BD6174D60FFF1FBECF816310F3941" blockId="8.[159,1432,304,2015]" lastBlockId="9.[159,1432,160,933]" lastPageId="9" lastPageNumber="492" pageId="8" pageNumber="491">
<emphasis id="3C900A054D60FFF1FBECF81636EF3BB2" italics="true" lastPageId="9" lastPageNumber="492" pageId="8" pageNumber="491">
......continued on the next page
<taxonomicName id="C9E4AD944D61FFF1FD2EFF1736EF3BB2" box="[675,901,201,223]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="492" phylum="Arthropoda" rank="species" species="chiruisla" status="sp. nov.">Cryptocellus chiruisla</taxonomicName>
</emphasis>
<taxonomicNameLabel id="27A3B77E4D61FFF1FC06FF1736B13BB2" box="[907,987,201,223]" pageId="9" pageNumber="492" rank="species">
<emphasis id="3C900A054D61FFF1FC06FF1736B13BB2" bold="true" box="[907,987,201,223]" pageId="9" pageNumber="492">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="C9E4AD944D61FFF1FBB4FF17302B3BB2" box="[1081,1345,201,223]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="492" phylum="Arthropoda" rank="species" species="guaviarensis" status="sp. nov.">
<emphasis id="3C900A054D61FFF1FBB4FF17302B3BB2" box="[1081,1345,201,223]" italics="true" pageId="9" pageNumber="492">Cryptocellus guaviarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27A3B77E4D61FFF1FACAFF1730F23BB2" box="[1351,1432,201,223]" pageId="9" pageNumber="492" rank="species">
<emphasis id="3C900A054D61FFF1FACAFF1730F23BB2" bold="true" box="[1351,1432,201,223]" pageId="9" pageNumber="492">sp. nov.</emphasis>
</taxonomicNameLabel>
Male Female Male 2nd trochanter L: 0.43 0.33 0.43 Femur L: 1.20 1.33 1.20 Femur W:2 0.50 0.48 0.43 Patella L: 0.87 0.77 0.93 Tibia L: 0.83 0.70 0.90 Metatarsus L: 1.13 0.93 1.13 Tarsus L: 1.27 0.87 1.30
</paragraph>
<caption id="5A9B869F4D61FFF1FF12FF7E34E63BDB" box="[159,396,160,182]" pageId="9" pageNumber="492">
<paragraph id="0E5BD6174D61FFF1FF12FF7E34E63BDB" blockId="9.[159,1432,160,933]" box="[159,396,160,182]" pageId="9" pageNumber="492">
<emphasis id="3C900A054D61FFF1FF12FF7E34663BDB" bold="true" box="[159,268,160,182]" pageId="9" pageNumber="492">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="0E5BD6174D61FFF1FF12FD9E310F38C8" blockId="9.[159,1432,160,933]" pageId="9" pageNumber="492">Leg IV: Total L:4 5.39 5.16 5.70 1st trochanter L: 0.33 0.33 0.40 2nd trochanter L: 0.33 0.43 0.33 Femur L: 1.33 1.10 1.30 Femur W:2 0.77 0.42 0.55 Patella L: 0.87 0.77 0.90 Tibia L: 0.60 0.73 0.77 Metatarsus L: 1.03 0.97 1.00 Tarsus L: 0.90 0.83 1.00</paragraph>
</subSubSection>
</treatment>
</document>