treatments-xml/data/82/A9/16/82A916CBF94853B8B3104C9EA3739D59.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.96.47792" ID-GBIF-Dataset="46dee71c-5ee5-4a2b-9ca9-c6ec9c73f07c" ID-GBIF-Taxon="182236566" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="A8EDB50ED5705717A80AD87E91F4AE96" ID-ZooBank="C5560B3FB5A0462380EBFE2C483D4774" ModsDocID="1860-0743-96-1-1" checkinTime="1621423348088" checkinUser="pensoft" docAuthor="Tripathy, Basudev, Sajan, Sheikh &amp; Cowie, Robert H." docDate="2019" docId="82A916CBF94853B8B3104C9EA3739D59" docLanguage="en" docName="ZoosystEvol 96(1): 1-23" docOrigin="Zoosystematics and Evolution 96 (1)" docPubDate="2019-01-14" docSource="http://dx.doi.org/10.3897/zse.96.47792" docTitle="Ampullaria conica subsp. var. var. expansa Nevill 1877" docType="treatment" docVersion="3" id="A8EDB50ED5705717A80AD87E91F4AE96" lastPageNumber="1" masterDocId="A8EDB50ED5705717A80AD87E91F4AE96" masterDocTitle="Illustrated catalogue of types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the National Zoological Collection of the Zoological Survey of India, with lectotype designations" masterLastPageNumber="23" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1643454254309" updateUser="ExternalLinkService">
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<mods:title>Illustrated catalogue of types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the National Zoological Collection of the Zoological Survey of India, with lectotype designations</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
<mods:affiliation>Malacology Division, Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sajan, Sheikh</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
<mods:affiliation>Malacology Division, Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
<mods:nameIdentifier type="email">sksajan.sajan@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cowie, Robert H.</mods:namePart>
<mods:affiliation>Pacific Biosciences Research Center, University of Hawaii, 3050 Maile Way, Gilmore 408, Honolulu, Hawaii 96822, USA</mods:affiliation>
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<mods:date>2019</mods:date>
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<treatment ID-GBIF-Taxon="182236566" LSID="urn:lsid:plazi:treatment:82A916CBF94853B8B3104C9EA3739D59" httpUri="http://treatment.plazi.org/id/82A916CBF94853B8B3104C9EA3739D59" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="82A916CB-F948-53B8-B310-4C9EA3739D59" authority="Nevill, 1877" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ampullaria conica subsp. var. var. expansa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="conica" subSpecies="var." variety="expansa">Ampullaria conica var. expansa Nevill, 1877</taxonomicName>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Ampullaria conica var. expansa Nevill, 1877; NZSI M. 2426." figureDoi="10.3897/zse.96.47792.figure4" httpUri="https://binary.pensoft.net/fig/371731" pageId="0" pageNumber="1">Figs 4</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Collection labels. A. Pachylabra angelica Annandale, 1920; NZSI M. 11649 / 2. B. Ampullaria erronea Nevill, 1877; NZSI M. 2404. C. Ampullaria conica var. expansa Nevill, 1877; NZSI M. 2426." figureDoi="10.3897/zse.96.47792.figure12" httpUri="https://binary.pensoft.net/fig/371739" pageId="0" pageNumber="1">, 12C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ampullaria conica subsp. var. var. expansa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="conica" subSpecies="var." variety="expansa">Ampullaria conica var. expansa</taxonomicName>
Nevill, 1877. 
<emphasis italics="true" pageId="0" pageNumber="1">Cat. Moll. Ind. Mus.</emphasis>
Fasc. E: 9.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="current taxonomic status">
<paragraph pageId="0" pageNumber="1">Current taxonomic status.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Mousson" baseAuthorityYear="1848" class="Gastropoda" family="Ampullariidae" genus="Pila" kingdom="Animalia" lsidName="Pila scutata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scutata">
<emphasis italics="true" pageId="0" pageNumber="1">Pila scutata</emphasis>
</taxonomicName>
(Mousson, 1848), probable synonym, pending further taxonomic research.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
<normalizedToken originalValue="“Pegu”">&quot;Pegu&quot;</normalizedToken>
[= Bago, Bago Region, Myanmar] (Nevill, 1877: 9); W. Theobald, Esq., leg. (see Remarks).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype (original designation): NZSI M.2426 (registered 8 January 1894). Paratypes: specimens listed by 
<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill (1877</bibRefCitation>
: 9, 1885: 5), not found in the ZSI collections, perhaps misplaced when the collections were moved from the Indian Museum to ZSI (8-10 spms); the shell figured by Hanley and Theobald (1874 in 1870-1876: pl. 114, fig. 5), location unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="shell dimensions">
<paragraph pageId="0" pageNumber="1">Shell dimensions.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype: SH 56.6 mm, SW 49.7 mm, AH 40.6 mm, AW 33.8 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill (1877</bibRefCitation>
: 9) listed seven specimens from 
<normalizedToken originalValue="“Pegu”">&quot;Pegu&quot;</normalizedToken>
collected by Theobald, and one from 
<normalizedToken originalValue="“Tenasserim?”">&quot;Tenasserim?&quot;</normalizedToken>
and one from 
<normalizedToken originalValue="“Mandalay?”">&quot;Mandalay?&quot;</normalizedToken>
each with &quot;No history&quot;. He also noted that the shell illustrated by Hanley and Theobald (1874 in 1870-1876: pl. 114, fig. 5) was referable to this taxon; it is therefore also to be considered a paratype. Subsequently, 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.23978" author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" refId="B22" refString="Nevill, G, 1885. Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [=Kolkata], x + 306 pp. https://doi.org/10.5962/bhl.title.23978" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [= Kolkata], x + 306 pp." url="https://doi.org/10.5962/bhl.title.23978" year="1885">Nevill (1885</bibRefCitation>
: 5) listed the seven Pegu specimens from Theobald (with &quot;type var.&quot;) but three (not one) from &quot;Tenasserim (?)&quot; and one from &quot;Mandalay (?)&quot;, all four from the collection of Stoliczka. Thus it is not clear whether there were eight or ten paratypes, as either 
<normalizedToken originalValue="“one”">&quot;one&quot;</normalizedToken>
or 
<normalizedToken originalValue="“three”">&quot;three&quot;</normalizedToken>
from 
<normalizedToken originalValue="“Tenasserim?”">&quot;Tenasserim?&quot;</normalizedToken>
could be in error. 
<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill (1877</bibRefCitation>
: 10) gave measurements for the holotype and the shell height, at least, matches that of the present specimen (NZSI M.2426); shell width was given as 48 mm, while the present measurement is 45.6 mm, probably within the margin of error. We therefore consider it to be the holotype. None of the paratypes could be found in NZSI and the location of the specimen illustrated by Hanley and Theobald is unknown. The old specimen label associated with the holotype (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Collection labels. A. Pachylabra angelica Annandale, 1920; NZSI M. 11649 / 2. B. Ampullaria erronea Nevill, 1877; NZSI M. 2404. C. Ampullaria conica var. expansa Nevill, 1877; NZSI M. 2426." figureDoi="10.3897/zse.96.47792.figure12" httpUri="https://binary.pensoft.net/fig/371739" pageId="0" pageNumber="1">12C</figureCitation>
) is almost unreadable. However, a newer label (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Collection labels. A. Pachylabra angelica Annandale, 1920; NZSI M. 11649 / 2. B. Ampullaria erronea Nevill, 1877; NZSI M. 2404. C. Ampullaria conica var. expansa Nevill, 1877; NZSI M. 2426." figureDoi="10.3897/zse.96.47792.figure12" httpUri="https://binary.pensoft.net/fig/371739" pageId="0" pageNumber="1">12C</figureCitation>
) indicates Dr F. Stoliczka as the collector, although both this label and the register indicate 
<normalizedToken originalValue="“Pegu”">&quot;Pegu&quot;</normalizedToken>
as the locality, with the collector indicated in the register as unknown, i.e. with a question mark (
<normalizedToken originalValue="“?”">&quot;?&quot;</normalizedToken>
). Nevertheless, given that 
<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill (1877</bibRefCitation>
: 9, 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.23978" author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" refId="B22" refString="Nevill, G, 1885. Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [=Kolkata], x + 306 pp. https://doi.org/10.5962/bhl.title.23978" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [= Kolkata], x + 306 pp." url="https://doi.org/10.5962/bhl.title.23978" year="1885">1885</bibRefCitation>
: 5) was consistent in noting Theobald as the collector of the Pegu specimens, we consider that Theobald was indeed the collector of the holotype and that the label noting Stoliczka as the collector reflects an inadvertent error, perhaps associated with Stoliczka having collected the 
<normalizedToken originalValue="“Tenasserim”">&quot;Tenasserim&quot;</normalizedToken>
and 
<normalizedToken originalValue="“Mandalay”">&quot;Mandalay&quot;</normalizedToken>
specimens. 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3940.1.1" author="Cowie, RH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 92" refId="B8" refString="Cowie, RH, 2015. The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda. Zootaxa 3940 (1): 1 - 92, DOI: https://doi.org/10.11646/zootaxa.3940.1.1" title="The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda." url="https://doi.org/10.11646/zootaxa.3940.1.1" volume="3940" year="2015">Cowie (2015</bibRefCitation>
: 36) treated 
<taxonomicName authorityName="K.Miller" authorityYear="1879" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria expansa" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansa">
<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria expansa</emphasis>
</taxonomicName>
Nevill as a synonym of either 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria ampullacea" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ampullacea">
<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria ampullacea</emphasis>
</taxonomicName>
Linnaeus, following 
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a066303" author="Sowerby, GB" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="56 - 64" refId="B29" refString="Sowerby, GB, 1910. Notes on the family Ampullariidae. Proceedings of the Malacological Society of London 9 (1): 56 - 64, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a066303" title="Notes on the family Ampullariidae." url="https://doi.org/10.1093/oxfordjournals.mollus.a066303" volume="9" year="1910">Sowerby (1910</bibRefCitation>
: 56), or 
<emphasis italics="true" pageId="0" pageNumber="1">gracilis</emphasis>
Lea, following 
<bibRefCitation author="Prashad, B" journalOrPublisher="Nature in Singapore" pageId="0" pageNumber="1" publicationUrl="http://faunaofindia.nic.in/PDFVolumes/memoirs/008/02/0069-0089.pdf" refId="B26" refString="Prashad, B, 1925. Revision of the Indian Ampullariidae. Memoirs of the Indian Museum 8(2): 69-89, pls. 13-15. http://faunaofindia.nic.in/PDFVolumes/memoirs/008/02/0069-0089.pdf" title="Revision of the Indian Ampullariidae. Memoirs of the Indian Museum 8 (2): 69 - 89, pls. 13 - 15." url="http://faunaofindia.nic.in/PDFVolumes/memoirs/008/02/0069-0089.pdf" year="1925">Prashad (1925</bibRefCitation>
: 81), but refrained from making a definitive decision, pending further research. The holotype of 
<taxonomicName authorityName="K.Miller" authorityYear="1879" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria expansa" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansa">
<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria expansa</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Ampullaria conica var. expansa Nevill, 1877; NZSI M. 2426." figureDoi="10.3897/zse.96.47792.figure4" httpUri="https://binary.pensoft.net/fig/371731" pageId="0" pageNumber="1">4</figureCitation>
) is clearly not 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria ampullacea" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ampullacea">
<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria ampullacea</emphasis>
</taxonomicName>
, based on its shell shape. In fact, it seems more likely that 
<taxonomicName authorityName="K.Miller" authorityYear="1879" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria expansa" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansa">
<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria expansa</emphasis>
</taxonomicName>
is a junior synonym of the widespread 
<taxonomicName baseAuthorityName="Mousson" baseAuthorityYear="1848" class="Gastropoda" family="Ampullariidae" genus="Pila" kingdom="Animalia" lsidName="Pila scutata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scutata">
<emphasis italics="true" pageId="0" pageNumber="1">Pila scutata</emphasis>
</taxonomicName>
(Mousson, 1848), which is known from Myanmar (
<bibRefCitation author="Low, MEY" journalOrPublisher="Nature in Singapore" pageId="0" pageNumber="1" pagination="55 - 60" publicationUrl="https://lkcnhm.nus.edu.sg/app/uploads/2017/06/2013nis055-060-5.pdf" refId="B17" refString="Low, MEY, Tan, SK, Ng, TH, 2013. Pila conica (Wood, 1828), or Pila scutata (Mousson, 1848)? The correct name for the native apple snail of Singapore (Gastropoda: Ampullariidae). Nature in Singapore 6: 55 - 60, https://lkcnhm.nus.edu.sg/app/uploads/2017/06/2013nis055-060-5.pdf" title="Pila conica (Wood, 1828), or Pila scutata (Mousson, 1848)? The correct name for the native apple snail of Singapore (Gastropoda: Ampullariidae)." url="https://lkcnhm.nus.edu.sg/app/uploads/2017/06/2013nis055-060-5.pdf" volume="6" year="2013">Low et al. 2013</bibRefCitation>
: 56; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3940.1.1" author="Cowie, RH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 92" refId="B8" refString="Cowie, RH, 2015. The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda. Zootaxa 3940 (1): 1 - 92, DOI: https://doi.org/10.11646/zootaxa.3940.1.1" title="The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda." url="https://doi.org/10.11646/zootaxa.3940.1.1" volume="3940" year="2015">Cowie 2015</bibRefCitation>
: 47). Nonetheless, we refrain from formally synonymising 
<taxonomicName authorityName="K.Miller" authorityYear="1879" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria expansa" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansa">
<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria expansa</emphasis>
</taxonomicName>
Nevill with 
<taxonomicName baseAuthorityName="Mousson" baseAuthorityYear="1848" class="Gastropoda" family="Ampullariidae" genus="Pila" kingdom="Animalia" lsidName="Pila scutata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scutata">
<emphasis italics="true" pageId="0" pageNumber="1">Pila scutata</emphasis>
</taxonomicName>
Mousson, pending further, taxonomic research.
</paragraph>
<caption doi="10.3897/zse.96.47792.figure4" httpUri="https://binary.pensoft.net/fig/371731" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
Holotype of 
<taxonomicName authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria conica subsp. var. var. expansa" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="conica" subSpecies="var." variety="expansa">Ampullaria conica var. expansa</taxonomicName>
Nevill, 1877; NZSI M.2426.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>