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<mods:title id="75B875CDE60CC34593E2DA6A244321CF">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<paragraph id="EA1E3692C37BC725E12E764AFC64FDBD" blockId="85.[264,990,603,630]" box="[264,990,603,630]" pageId="85" pageNumber="86">
<heading id="B15681FEC37BC725E12E764AFC64FDBD" bold="true" box="[264,990,603,630]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
<emphasis id="D8D5EA80C37BC725E12E764AFC64FDBD" bold="true" box="[264,990,603,630]" pageId="85" pageNumber="86">
<key id="9E3061A3C37BC725E12E764AFC64FDBD" box="[264,990,603,630]" pageId="85" pageNumber="86">
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25.
<taxonomicName id="2DA14D11C37BC725E112764DFD6AFDBD" ID-CoL="6YBD2" authority="Gusarov" authorityName="Gusarov" box="[308,720,603,630]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="geostiboides" status="sp. nov.">
<emphasis id="D8D5EA80C37BC725E112764DFDE4FDBE" bold="true" box="[308,606,604,630]" italics="true" pageId="85" pageNumber="86">Seeversiella geostiboides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37BC725E2FB764AFCA5FDBD" box="[733,799,603,629]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C37BC725E309764AFC6DFDBD" box="[815,983,603,629]" captionStart-0="FIGURES 344 ­ 347" captionStart-1="FIGURES 348 ­ 355" captionStartId-0="85.[264,382,1733,1757]" captionStartId-1="86.[264,383,1647,1671]" captionTargetBox-0="[305,1286,853,1699]" captionTargetBox-1="[290,1290,620,1609]" captionTargetId-0="figure@85.[292,1295,844,1706]" captionTargetId-1="figure@86.[284,1303,609,1620]" captionTargetPageId-0="85" captionTargetPageId-1="86" captionText-0="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri-0="https://zenodo.org/record/156480/files/figure.png" httpUri-1="https://zenodo.org/record/156481/files/figure.png" pageId="85" pageNumber="86">Figs. 344­355</figureCitation>
)
</keyLead>
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</key>
</emphasis>
</heading>
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<subSubSection id="A2BB6519C37BC725E12E76B7FB6AFCD9" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="EA1E3692C37BC725E12E76B7FBDAFD21" blockId="85.[264,1323,678,785]" pageId="85" pageNumber="86">
<emphasis id="D8D5EA80C37BC725E12E76B7FE04FD08" bold="true" box="[264,446,678,704]" pageId="85" pageNumber="86">
<typeStatus id="351A8830C37BC725E12E76B7FEFFFD08" box="[264,325,678,704]" pageId="85" pageNumber="86">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C37BC725E1ED76B6FD81FD09" box="[459,571,679,705]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C37BC725E24E76B7FC1FFD08" bold="true" box="[616,933,678,704]" pageId="85" pageNumber="86">
<collectingCountry id="92B67602C37BC725E24E76B7FC9BFD08" box="[616,801,678,704]" name="Costa Rica" pageId="85" pageNumber="86">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
2.0 km E Villa Mills,
<geoCoordinate id="8F955055C37BC725E48976B6FA91FD09" box="[1199,1323,679,705]" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C37BC725E12E76DEFE22FD21" box="[264,408,719,745]" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="15" value="-83.69722">83°41'50&quot;W</geoCoordinate>
,
<quantity id="2D599B77C37BC725E18276DEFDBAFD21" box="[420,512,719,745]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="85" pageNumber="86" unit="m" value="2750.0">2750 m</quantity>
, oak forest litter (R.Anderson),
<date id="9E1F1052C37BC725E35376DEFC56FD21" box="[885,1004,719,745]" pageId="85" pageNumber="86" value="1998-02-15">15.ii.1998</date>
(
<collectionCode id="8CB0AE57C37BC725E3DA76DEFBEEFD21" box="[1020,1108,719,745]" pageId="85" pageNumber="86">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C37BC725E11E76E6FB6AFCD9" blockId="85.[264,1323,678,785]" box="[312,1232,758,785]" pageId="85" pageNumber="86">
<typeStatus id="351A8830C37BC725E11E76E6FE14FCD9" box="[312,430,759,785]" pageId="85" pageNumber="86" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C37BC725E1E676E7FD44FCD8" bold="true" box="[448,766,758,784]" pageId="85" pageNumber="86">
<collectingCountry id="92B67602C37BC725E1E676E7FDC0FCD8" box="[448,634,758,784]" name="Costa Rica" pageId="85" pageNumber="86">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
, same data as the
<typeStatus id="351A8830C37BC725E3D076E6FBE7FCD9" box="[1014,1117,759,785]" pageId="85" pageNumber="86" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C37BC725E44A76E6FB7EFCD9" box="[1132,1220,759,785]" pageId="85" pageNumber="86">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC37BC725E12E72D4FB91F8E9" httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="85" pageNumber="86" targetBox="[305,1286,853,1699]" targetPageId="85">
<paragraph id="EA1E3692C37BC725E12E72D4FB91F8E9" blockId="85.[264,1323,1733,1825]" pageId="85" pageNumber="86">
<emphasis id="D8D5EA80C37BC725E12E72D4FE64F915" bold="true" box="[264,478,1733,1757]" pageId="85" pageNumber="86">FIGURES 344­347</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C37BC725E32272D4FBC9F914" authority="Gusarov" authorityName="Gusarov" box="[772,1139,1733,1756]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="geostiboides" status="sp. nov.">
<emphasis id="D8D5EA80C37BC725E32272D4FBABF914" box="[772,1041,1733,1756]" italics="true" pageId="85" pageNumber="86">Seeversiella geostiboides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37BC725E45972D4FB00F915" box="[1151,1210,1733,1757]" pageId="85" pageNumber="86" rank="species">
<emphasis id="D8D5EA80C37BC725E45972D4FB00F915" bold="true" box="[1151,1210,1733,1757]" pageId="85" pageNumber="86">sp. n.</emphasis>
</taxonomicNameLabel>
(344­345, holotype; 346­347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C378C727E11E750AFB4DFE1E" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="diagnosis">
<paragraph id="EA1E3692C378C726E11E750AFA91FE36" blockId="86.[264,1324,283,510]" pageId="86" pageNumber="87">
<emphasis id="D8D5EA80C378C726E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="86" pageNumber="87">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C378C726E1EE750DFD48FEFD" box="[456,754,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C378C726E1EE750DFD48FEFD" box="[456,754,284,309]" italics="true" pageId="86" pageNumber="87">Seeversiella geostiboides</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C378C726E4DE750DFED7FE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C378C726E4DE750DFED7FE95" italics="true" pageId="86" pageNumber="87">Seeversiella</emphasis>
</taxonomicName>
by having uniformly brown body; small eyes (temples 3.5­4.0 times as long as eyes); pronotal pubescence in lateral portions of the disc directed posteriorly; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking clear medial emargination at the posterior margin of male tergum 8 (
<figureCitation id="729A2A17C378C726E34E75ADFC6AFE1E" box="[872,976,444,470]" captionStart="FIGURES 344 ­ 347" captionStartId="85.[264,382,1733,1757]" captionTargetBox="[305,1286,853,1699]" captionTargetId="figure@85.[292,1295,844,1706]" captionTargetPageId="85" captionText="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="86" pageNumber="87">Fig. 344</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C378C726E30875F5FC69FE36" box="[814,979,484,510]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="86" pageNumber="87">Figs. 348­354</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C378C726E49175F5FAA4FE36" box="[1207,1310,484,510]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="86" pageNumber="87">Fig. 355</figureCitation>
).
</paragraph>
<caption id="BEDE661AC378C726E12E727EFC5AF8FB" httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="86" pageNumber="87" targetBox="[290,1290,620,1609]" targetPageId="86">
<paragraph id="EA1E3692C378C726E12E727EFC5AF8FB" blockId="86.[264,1324,1647,1843]" pageId="86" pageNumber="87">
<emphasis id="D8D5EA80C378C726E12E727EFE5BF94F" bold="true" box="[264,481,1647,1671]" pageId="86" pageNumber="87">FIGURES 348­355</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C378C726E2A1727EFC46F94E" authority="Gusarov" authorityName="Gusarov" box="[647,1020,1647,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="geostiboides" status="sp. nov.">
<emphasis id="D8D5EA80C378C726E2A1727EFC2DF94E" box="[647,919,1647,1670]" italics="true" pageId="86" pageNumber="87">Seeversiella geostiboides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC378C726E42C727EFBF3F94F" box="[1034,1097,1647,1671]" pageId="86" pageNumber="87" rank="species">
<emphasis id="D8D5EA80C378C726E42C727EFBF3F94F" bold="true" box="[1034,1097,1647,1671]" pageId="86" pageNumber="87">sp. n.</emphasis>
</taxonomicNameLabel>
(348­354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353­354), 0.1 mm (349, 351­352, 355).
</paragraph>
</caption>
<paragraph id="EA1E3692C379C727E11E750DFB4DFE1E" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E750DFDD8FEFD" box="[312,610,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E750DFDD8FEFD" box="[312,610,284,309]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C379C727E38C750DFBD6FEFD" box="[938,1132,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C379C727E38C750DFBD6FEFD" box="[938,1132,284,309]" italics="true" pageId="87" pageNumber="88">S. impressicollis</emphasis>
</taxonomicName>
by not asperate pronotal punctation; in males pronotum without broad medial impression, lateral portions of posterior margin of tergum 3 extending as short and blunt projections (as in
<figureCitation id="729A2A17C379C727E4E2757DFAA6FE4E" box="[1220,1308,364,390]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="87" pageNumber="88">Fig. 19</figureCitation>
), and tergum 7 with medial carina; the apex of median lobe in parameral view more rounded (
<figureCitation id="729A2A17C379C727E13775ADFE02FE1E" box="[273,440,444,470]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 348­349</figureCitation>
; 311­312); and spermatheca with larger distal portion (
<figureCitation id="729A2A17C379C727E41A75ADFB17FE1E" box="[1084,1197,444,470]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 355</figureCitation>
; 320).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C379C728E11E75F5FDC7FE4E" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
<paragraph id="EA1E3692C379C727E11E75F5FDAAFDBE" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E75F5FDDCFE35" box="[312,614,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E75F5FDDCFE35" box="[312,614,484,509]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C379C727E3E275F5FBE5FE35" box="[964,1119,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C379C727E3E275F5FBE5FE35" box="[964,1119,484,509]" italics="true" pageId="87" pageNumber="88">S. sulcicollis</emphasis>
</taxonomicName>
by not asperate pronotal punctation; in males pronotum without medial furrow and posterior margin of tergum 8 without denticles; the apex of median lobe without divergent carinae (in parameral view) (
<figureCitation id="729A2A17C379C727E17D764DFDB9FDBE" box="[347,515,604,630]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 348­349</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C379C727E11E7695FC14FCDE" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E7695FDDBFD55" box="[312,609,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E7695FDDBFD55" box="[312,609,644,669]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C379C727E3267695FC6BFD55" box="[768,977,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C379C727E3267695FC6BFD55" box="[768,977,644,669]" italics="true" pageId="87" pageNumber="88">S. microphthalma</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C379C727E42B7695FB08FD55" box="[1037,1202,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C379C727E42B7695FB08FD55" box="[1037,1202,644,669]" italics="true" pageId="87" pageNumber="88">S. paramoana</emphasis>
</taxonomicName>
in having pubescence in the lateral portions of pronotum directed posteriorly; in narrower apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C379C727E28676C5FCF0FD26" box="[672,842,724,750]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 348­349</figureCitation>
; 335­336; 377­378); and different shape of the distal portion of spermatheca (
<figureCitation id="729A2A17C379C727E29E76EDFC93FCDE" box="[696,809,764,790]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 355</figureCitation>
; 341; 381).
</paragraph>
<paragraph id="EA1E3692C379C727E11E7735FB7DFC45" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E7735FDE5FCF5" box="[312,607,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E7735FDE5FCF5" box="[312,607,804,829]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C379C727E2DE7735FCDCFCF5" box="[760,870,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C379C727E2DE7735FCDCFCF5" box="[760,870,804,829]" italics="true" pageId="87" pageNumber="88">S. flavida</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C379C727E3B97735FBB0FCF5" box="[927,1034,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C379C727E3B97735FBB0FCF5" box="[927,1034,804,829]" italics="true" pageId="87" pageNumber="88">S. adusta</emphasis>
</taxonomicName>
in having pubescence in the lateral portions of pronotum directed posteriorly; tergum 8 with four pairs of macrosetae; and different shape of the distal portion of spermatheca (
<figureCitation id="729A2A17C379C727E3F77765FBF8FC46" box="[977,1090,884,910]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 355</figureCitation>
; 358; 371).
</paragraph>
<paragraph id="EA1E3692C379C727E11E778AFC05FC7E" blockId="87.[264,1324,284,1870]" box="[312,959,923,950]" pageId="87" pageNumber="88">
<emphasis id="D8D5EA80C379C727E11E778AFE6AFC7D" bold="true" box="[312,464,923,949]" pageId="87" pageNumber="88">Description.</emphasis>
Length
<quantity id="2D599B77C379C727E21F778DFD2CFC7E" box="[569,662,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="87" pageNumber="88" unit="mm" value="2.6">2.6 mm</quantity>
. Body uniformly brown.
</paragraph>
<paragraph id="EA1E3692C379C727E11E77D5FD49FB9E" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and poorly visible punctation, distance between punctures equals 2 times their diameter. Temples 3.5­4.0 times as long as eyes. Antennal article 2 as long as 3, article 4 slightly transverse, 5­10 transverse (ratio 1.6­2.0).</paragraph>
<paragraph id="EA1E3692C379C727E11E7075FEE8FA76" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C379C727E40F7075FB69FBB6" box="[1065,1235,1124,1150]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="87" pageNumber="88" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47­0.50 mm</quantity>
, length
<quantity id="2D599B77C379C727E12E709DFE0AFB6E" box="[264,432,1164,1190]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.6" metricValueMin="4.3" pageId="87" pageNumber="88" unit="mm" value="0.445" valueMax="0.46" valueMin="0.43">0.43­0.46 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine punctation, distance between punctures equals 1­2 times their diameter. Pronotal pubescence in lateral portions of the disc directed posteriorly. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with poorly visible microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C379C727E11E71DDFE71F966" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
Abdominal terga glossy, terga 3­5 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 2­3 times their diameter; tergum 7 with microsculpture consisting of isodiametric or transverse meshes and with fine and sparse punctation, distance between punctures equals 3­7 times. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C379C727E1377285FE05F966" box="[273,447,1684,1710]" captionStart="FIGURES 344 ­ 347" captionStartId="85.[264,382,1733,1757]" captionTargetBox="[305,1286,853,1699]" captionTargetId="figure@85.[292,1295,844,1706]" captionTargetPageId="85" captionText="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="87" pageNumber="88">Figs. 344, 346</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C379C727E11E72ADFB01F886" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
In the only known male the posterior angles of tergum 3 not projecting as spines, lateral portions of posterior margin slightly extending as short and blunt projections (as in
<figureCitation id="729A2A17C379C727E12E731DFEDAF8EE" box="[264,352,1804,1830]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="87" pageNumber="88">Fig. 19</figureCitation>
); tergum
<quantity id="2D599B77C379C727E1F2731DFDB9F8EE" box="[468,515,1804,1830]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="87" pageNumber="88" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina 2/3 as long as the tergum; posterior margin of tergum 8 without clear medial emargination (
<figureCitation id="729A2A17C379C727E4617325FB15F886" box="[1095,1199,1844,1870]" captionStart="FIGURES 344 ­ 347" captionStartId="85.[264,382,1733,1757]" captionTargetBox="[305,1286,853,1699]" captionTargetId="figure@85.[292,1295,844,1706]" captionTargetPageId="85" captionText="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="87" pageNumber="88">Fig. 344</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C376C728E11E750DFD48FE96" blockId="88.[264,1323,284,510]" pageId="88" pageNumber="89">
Aedeagus as in
<figureCitation id="729A2A17C376C728E1C8750DFD2EFEFE" box="[494,660,284,310]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="88" pageNumber="89">Figs. 348­354</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C376C728E2587555FD5FFE96" box="[638,741,324,350]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="88" pageNumber="89">Fig. 352</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C376C728E11E757DFDC7FE4E" blockId="88.[264,1323,284,510]" box="[312,637,364,390]" pageId="88" pageNumber="89">
Spermatheca as in
<figureCitation id="729A2A17C376C728E232757DFDC3FE4E" box="[532,633,364,390]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="88" pageNumber="89">Fig. 355</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C376C728E11E7582FED2FE36" pageId="88" pageNumber="89" type="distribution">
<paragraph id="EA1E3692C376C728E11E7582FBCFFE66" blockId="88.[264,1323,284,510]" box="[312,1141,403,430]" pageId="88" pageNumber="89">
<emphasis id="D8D5EA80C376C728E11E7582FE62FE65" bold="true" box="[312,472,403,429]" pageId="88" pageNumber="89">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C376C728E3547585FC48FE66" box="[882,1010,404,430]" name="Costa Rica" pageId="88" pageNumber="89">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C376C728E4247585FBD3FE66" box="[1026,1129,404,430]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="88" pageNumber="89">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C376C728E11E75AAFED2FE36" blockId="88.[264,1323,284,510]" pageId="88" pageNumber="89">
<emphasis id="D8D5EA80C376C728E11E75AAFDBFFE1D" bold="true" box="[312,517,443,469]" pageId="88" pageNumber="89">Natural History.</emphasis>
<taxonomicName id="2DA14D11C376C728E23375ADFC84FE1D" box="[533,830,444,469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C376C728E23375ADFC84FE1D" box="[533,830,444,469]" italics="true" pageId="88" pageNumber="89">Seeversiella geostiboides</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C376C728E12E75F5FEDEFE36" box="[264,356,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="88" pageNumber="89" unit="m" value="2750.0">2750 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>