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<document id="1AF6177D962A40A6B43945B3481D26FF" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C36DC735E12E75C8FED2FA2E" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella lativentris Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="70" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="68" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="39BD41E616270CD8686AE066F8DF0A40">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C36DC733E12E75C8FC7FFE3B" box="[264,965,473,499]" pageId="67" pageNumber="68" type="key">
<paragraph id="EA1E3692C36DC733E12E75C8FC7FFE3B" blockId="67.[264,965,473,499]" box="[264,965,473,499]" pageId="67" pageNumber="68">
<heading id="B15681FEC36DC733E12E75C8FC7FFE3B" bold="true" box="[264,965,473,499]" fontSize="11" level="1" pageId="67" pageNumber="68" reason="1">
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<keyStep id="51552E37C36DC733E12E75C8FC7FFE3B" box="[264,965,473,499]" pageId="67" pageNumber="68">
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<emphasis id="D8D5EA80C36DC733E12E75C8FC7FFE3B" bold="true" box="[264,965,473,499]" pageId="67" pageNumber="68">
19.
<taxonomicName id="2DA14D11C36DC733E11275C8FD0DFE3B" ID-CoL="6YBD8" authority="Gusarov" authorityName="Gusarov" box="[308,695,473,499]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="D8D5EA80C36DC733E11275C8FDFEFE3B" bold="true" box="[308,580,473,499]" italics="true" pageId="67" pageNumber="68">Seeversiella lativentris</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36DC733E2E575C8FCBCFE3B" box="[707,774,473,499]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C36DC733E33375C8FC07FE3B" box="[789,957,473,499]" captionStart-0="FIGURES 265 ­ 268" captionStart-1="FIGURES 269 ­ 277" captionStartId-0="67.[264,382,1514,1538]" captionStartId-1="68.[264,383,1662,1686]" captionTargetBox-0="[285,1297,756,1480]" captionTargetBox-1="[286,1302,460,1628]" captionTargetId-0="figure@67.[276,1311,746,1488]" captionTargetId-1="figure@68.[277,1309,449,1635]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionText-0="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri-0="https://zenodo.org/record/156466/files/figure.png" httpUri-1="https://zenodo.org/record/156467/files/figure.png" pageId="67" pageNumber="68">Figs. 265­277</figureCitation>
)
</emphasis>
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<subSubSection id="A2BB6519C36DC733E12E760FFECFFD79" pageId="67" pageNumber="68" type="materials_examined">
<paragraph id="EA1E3692C36DC733E12E760FFC7FFDA9" blockId="67.[264,1323,542,689]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E12E760FFE04FDF0" bold="true" box="[264,446,542,568]" pageId="67" pageNumber="68">
<typeStatus id="351A8830C36DC733E12E760FFEF9FDF0" box="[264,323,542,568]" pageId="67" pageNumber="68">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C36DC733E1F2760EFDFEFDF1" box="[468,580,543,569]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C36DC733E250760FFC78FDF0" bold="true" box="[630,962,542,568]" pageId="67" pageNumber="68">
<collectingCountry id="92B67602C36DC733E250760FFC8BFDF0" box="[630,817,542,568]" name="Costa Rica" pageId="67" pageNumber="68">COSTA RICA</collectingCountry>
: San Jose:
</emphasis>
<quantity id="2D599B77C36DC733E3F0760EFBA9FDF1" box="[982,1043,543,569]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="km" value="2.0">2 km</quantity>
S Villa Mills,
<quantity id="2D599B77C36DC733E4EE760EFA9CFDF1" box="[1224,1318,543,569]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="67" pageNumber="68" unit="m" value="3000.0">3000 m</quantity>
, ridgetop oak forest litter (R.Anderson),
<date id="9E1F1052C36DC733E2FC7656FCEBFDA9" box="[730,849,583,609]" pageId="67" pageNumber="68" value="1996-02-10">10.ii.1996</date>
(
<collectionCode id="8CB0AE57C36DC733E3477656FC03FDA9" box="[865,953,583,609]" pageId="67" pageNumber="68">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C36DC733E11E767EFECFFD79" blockId="67.[264,1323,542,689]" pageId="67" pageNumber="68">
<typeStatus id="351A8830C36DC733E11E767EFE15FD41" box="[312,431,623,649]" pageId="67" pageNumber="68" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C36DC733E1EF767FFCA4FD40" bold="true" box="[457,798,622,648]" pageId="67" pageNumber="68">
<collectingCountry id="92B67602C36DC733E1EF767FFD3DFD40" box="[457,647,622,648]" name="Costa Rica" pageId="67" pageNumber="68">COSTA RICA</collectingCountry>
: San Jose:
</emphasis>
<specimenCount id="FCA7FD1BC36DC733E310767EFC66FD41" box="[822,988,623,649]" pageId="67" pageNumber="68" type="generic">78 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C36DC733E4E2767EFA91FD41" box="[1220,1323,623,649]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C36DC733E1367686FED2FD79" box="[272,360,663,689]" pageId="67" pageNumber="68">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC36DC733E12E71FBFD63F98F" httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="67" pageNumber="68" targetBox="[285,1297,756,1480]" targetPageId="67">
<paragraph id="EA1E3692C36DC733E12E71FBFD63F98F" blockId="67.[264,1323,1514,1607]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E12E71FBFE5AF9CA" bold="true" box="[264,480,1514,1538]" pageId="67" pageNumber="68">FIGURES 265­268</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C36DC733E32971FAFBD2F9C9" authority="Gusarov" authorityName="Gusarov" box="[783,1128,1514,1538]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="D8D5EA80C36DC733E32971FAFBBEF9CA" box="[783,1028,1515,1538]" italics="true" pageId="67" pageNumber="68">Seeversiella lativentris</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C36DC733E45371FBFB09F9CA" bold="true" box="[1141,1203,1514,1538]" pageId="67" pageNumber="68">
<taxonomicNameLabel id="C3E657FBC36DC733E45371FBFB09F9CA" box="[1141,1203,1514,1538]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C36DC735E11E729CFC43FE1D" lastPageId="69" lastPageNumber="70" pageId="67" pageNumber="68" type="diagnosis">
<paragraph id="EA1E3692C36DC733E11E729CFDA5F8A7" blockId="67.[264,1323,1677,1903]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E11E729CFE02F96F" bold="true" box="[312,440,1677,1703]" pageId="67" pageNumber="68">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C36DC733E1E1729CFD69F96E" box="[455,723,1677,1702]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36DC733E1E1729CFD69F96E" box="[455,723,1677,1702]" italics="true" pageId="67" pageNumber="68">Seeversiella lativentris</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C36DC733E4F5729CFEFDF906" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C36DC733E4F5729CFEFDF906" italics="true" pageId="67" pageNumber="68">Seeversiella</emphasis>
</taxonomicName>
by having brown body with lighter pronotum and elytra; temples 1.5­1.8 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking clear medial emargination at posterior margin of male tergum 7 (
<figureCitation id="729A2A17C36DC733E47B7314FB79F8D7" box="[1117,1219,1797,1823]" captionStart="FIGURES 265 ­ 268" captionStartId="67.[264,382,1514,1538]" captionTargetBox="[285,1297,756,1480]" captionTargetId="figure@67.[276,1311,746,1488]" captionTargetPageId="67" captionText="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="67" pageNumber="68">Fig. 265</figureCitation>
); tergum 8 with three pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C36DC733E463733CFB54F88F" box="[1093,1262,1837,1863]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="67" pageNumber="68">Figs. 269­276</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C36DC733E18D7344FDA8F8A7" box="[427,530,1877,1903]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="67" pageNumber="68">Fig. 277</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36AC734E11E750DFBDCFE96" blockId="68.[264,1323,284,350]" pageId="68" pageNumber="69">
<taxonomicName id="2DA14D11C36AC734E11E750DFDF9FEFD" box="[312,579,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36AC734E11E750DFDF9FEFD" box="[312,579,284,309]" italics="true" pageId="68" pageNumber="69">Seeversiella lativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36AC734E2FA750DFCC7FEFD" box="[732,893,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36AC734E2FA750DFCC7FEFD" box="[732,893,284,309]" italics="true" pageId="68" pageNumber="69">S. curtipennis</emphasis>
</taxonomicName>
in having stronger basal constriction of the apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C36AC734E3667555FC5DFE96" box="[832,999,324,350]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="68" pageNumber="69">Figs. 269­270</figureCitation>
; 251­252).
</paragraph>
<caption id="BEDE661AC36AC734E12E726FFCA5F88A" httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="68" pageNumber="69" targetBox="[286,1302,460,1628]" targetPageId="68">
<paragraph id="EA1E3692C36AC734E12E726FFCA5F88A" blockId="68.[264,1323,1662,1895]" pageId="68" pageNumber="69">
<emphasis id="D8D5EA80C36AC734E12E726FFE58F95E" bold="true" box="[264,482,1662,1686]" pageId="68" pageNumber="69">FIGURES 269­277</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C36AC734E2AC726FFC52F95D" authority="Gusarov" authorityName="Gusarov" box="[650,1000,1662,1685]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="D8D5EA80C36AC734E2AC726FFC3BF95D" box="[650,897,1662,1685]" italics="true" pageId="68" pageNumber="69">Seeversiella lativentris</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36AC734E3D3726FFB8FF95E" box="[1013,1077,1662,1686]" pageId="68" pageNumber="69" rank="species">
<emphasis id="D8D5EA80C36AC734E3D3726FFB8FF95E" bold="true" box="[1013,1077,1662,1686]" pageId="68" pageNumber="69">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274­276), 0.1 mm (270, 272­273, 277).
</paragraph>
</caption>
<paragraph id="EA1E3692C36AC734E12E7341FE46F8AF" blockId="68.[264,1323,1662,1895]" box="[264,508,1872,1895]" pageId="68" pageNumber="69">ML medial lamellae.</paragraph>
<paragraph id="EA1E3692C36BC735E11E750DFC43FE1D" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
<taxonomicName id="2DA14D11C36BC735E11E750DFDFEFEFD" box="[312,580,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36BC735E11E750DFDFEFEFD" box="[312,580,284,309]" italics="true" pageId="69" pageNumber="70">Seeversiella lativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36BC735E2C7750DFC3AFEFD" box="[737,896,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C36BC735E2C7750DFC3AFEFD" box="[737,896,284,309]" italics="true" pageId="69" pageNumber="70">S. luridicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C36BC735E39C750DFBD6FEFD" box="[954,1132,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C36BC735E39C750DFBD6FEFD" box="[954,1132,284,309]" italics="true" pageId="69" pageNumber="70">S. micralymma</emphasis>
</taxonomicName>
in having matte pronotum with strong isodiametric microsculpture; tergum 8 with three pairs of macrosetae; medial lamellae of internal sac long (
<figureCitation id="729A2A17C36BC735E2CB757DFCEEFE4E" box="[749,852,364,390]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 274</figureCitation>
), the apices of lamellae protruding from retracted sac (
<figureCitation id="729A2A17C36BC735E18B7585FDAEFE66" box="[429,532,404,430]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 269</figureCitation>
); different shape of the median lobe (
<figureCitation id="729A2A17C36BC735E3E97585FBCDFE66" box="[975,1143,404,430]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Figs. 269­270</figureCitation>
; 282­283; 295­ 296) and L­shaped spermatheca with large umbilicus (
<figureCitation id="729A2A17C36BC735E3A375ADFC53FE1E" box="[901,1001,444,470]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 277</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36BC735E11E75F2FDC4FAA6" pageId="69" pageNumber="70" type="description">
<paragraph id="EA1E3692C36BC735E11E75F2FAB0FDEE" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E11E75F2FE6AFE35" bold="true" box="[312,464,483,509]" pageId="69" pageNumber="70">Description.</emphasis>
Length 2.0­
<quantity id="2D599B77C36BC735E24A75F5FD71FE36" box="[620,715,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="69" pageNumber="70" unit="mm" value="2.6">2.6 mm</quantity>
. Head and abdomen brown to dark brown; pronotum reddish brown; elytra reddish brown or yellowish brown; antennae and legs brown.
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7625FB0DFD56" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">Head surface matte, with fine and dense isodiametric microsculpture, and fine and poorly visible punctation. Temples 1.5­1.8 times as long as eyes. Antennal article 2 as long as 3, article 4 transverse (ratio 1.2), 5­10 strongly transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C36BC735E11E76BDFD2EFC7E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C36BC735E40D76BDFB6CFD0E" box="[1067,1238,684,710]" metricMagnitude="-4" metricUnit="m" metricValue="4.3999999999999995" metricValueMax="4.699999999999999" metricValueMin="4.1" pageId="69" pageNumber="70" unit="mm" value="0.43999999999999995" valueMax="0.47" valueMin="0.41">0.41­0.47 mm</quantity>
, length
<quantity id="2D599B77C36BC735E12E76C5FE09FD26" box="[264,435,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.3" metricValueMin="3.4" pageId="69" pageNumber="70" unit="mm" value="0.385" valueMax="0.43" valueMin="0.34">0.34­0.43 mm</quantity>
, width to length ratio 1.1, surface matte, with fine and dense isodiametric microsculpture, and fine and poorly visible punctation. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.3), 1.7 times wider than long, matte, with fine and weak isodiametric microsculpture, and fine punctation, distance between punctures equals ½­1 time their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C36BC735E11E77D5FC16FB9E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­4 times their diameter on terga 3­5 and 3­5 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C36BC735E2D4702DFC1AFB9E" box="[754,928,1084,1110]" captionStart="FIGURES 265 ­ 268" captionStartId="67.[264,382,1514,1538]" captionTargetBox="[285,1297,756,1480]" captionTargetId="figure@67.[276,1311,746,1488]" captionTargetPageId="67" captionText="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="69" pageNumber="70">Figs. 265, 267</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7075FEFFFB3E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
In large males posterior angles of tergum 3 projecting as spines up to half as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to ½ as long as tergum length; posterior margin of tergum 8 without clear emargination (
<figureCitation id="729A2A17C36BC735E4DD70A5FE82FB3E" captionStart="FIGURES 265 ­ 268" captionStartId="67.[264,382,1514,1538]" captionTargetBox="[285,1297,756,1480]" captionTargetId="figure@67.[276,1311,746,1488]" captionTargetPageId="67" captionText="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="69" pageNumber="70">Fig. 265</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7115FD48FA8E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
Aedeagus as in
<figureCitation id="729A2A17C36BC735E1C87115FD2FFAD6" box="[494,661,1284,1310]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Figs. 269­276</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C36BC735E258713DFD5FFA8E" box="[638,741,1324,1350]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 273</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7145FDC4FAA6" blockId="69.[264,1324,284,1510]" box="[312,638,1364,1390]" pageId="69" pageNumber="70">
Spermatheca as in
<figureCitation id="729A2A17C36BC735E2327145FDC0FAA6" box="[532,634,1364,1390]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 277</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36BC735E11E716AFED2FA2E" pageId="69" pageNumber="70" type="distribution">
<paragraph id="EA1E3692C36BC735E11E716AFBCFFA5E" blockId="69.[264,1324,284,1510]" box="[312,1141,1403,1430]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E11E716AFE62FA5D" bold="true" box="[312,472,1403,1429]" pageId="69" pageNumber="70">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C36BC735E354716DFC48FA5E" box="[882,1010,1404,1430]" name="Costa Rica" pageId="69" pageNumber="70">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C36BC735E424716DFBD3FA5E" box="[1026,1129,1404,1430]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="69" pageNumber="70">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E71B2FED2FA2E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E11E71B2FDBDFA75" bold="true" box="[312,519,1443,1469]" pageId="69" pageNumber="70">Natural History.</emphasis>
<taxonomicName id="2DA14D11C36BC735E23A71B5FC91FA75" box="[540,811,1444,1469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36BC735E23A71B5FC91FA75" box="[540,811,1444,1469]" italics="true" pageId="69" pageNumber="70">Seeversiella lativentris</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C36BC735E12E71DDFEDEFA2E" box="[264,356,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="69" pageNumber="70" unit="m" value="3000.0">3000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>