treatments-xml/data/62/08/87/62088784C360C722E12E727AFED2FE96.xml
2024-06-21 12:38:20 +02:00

231 lines
36 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="C2D8BF2C698280E2DAA0AD3F0202E970" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C360C722E12E727AFED2FE96" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella sulcicollis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="83" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="79" updateTime="1698203169024" updateUser="ExternalLinkService">
<mods:mods id="B40C9462CEADE213AF03DFDA58D87196" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1F4D32B4CB57190A1537D8E2A3A2131F">
<mods:title id="21FE1559D26C65FED59C6C55E033A700">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="DBE3CC44B9683ABE81D43F3DBE372F0C" type="personal">
<mods:role id="635B1B26AE31A18EBA0FBB6398197B28">
<mods:roleTerm id="8095F4C05CAE40BA7159377BC7A7DABA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB0CC3DA300AA96D17D7482BD7129115">Gusarov, Vladimir I.</mods:namePart>
</mods:name>
<mods:typeOfResource id="95F58B736EBF12D0B7308AB4D60F19DF">text</mods:typeOfResource>
<mods:relatedItem id="116B47F8EBE51C729AA6DE228A1828C0" type="host">
<mods:titleInfo id="A778A39C2B67B573118DEC33482CB5CC">
<mods:title id="855B80122203D0053819177A057C5676">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B296104F2D471B2C77D0604C93F39908">
<mods:date id="B0B23D1D7AEE96E19DFC96990E7A16E2">2003</mods:date>
<mods:detail id="DFBFD78BDA5478EFFBF53AB030647807" type="volume">
<mods:number id="5CF9AC97DDD6833E0228C38EB27DB0B1">142</mods:number>
</mods:detail>
<mods:extent id="A225E59E455D9E572C1C9D47CF845800" unit="page">
<mods:start id="A57F4164E66E455EE0F5168EC12114E5">1</mods:start>
<mods:end id="9B5899C8A97712C72121A08C5D9CC4F5">102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C3BCDA76BF34878077393870FDE12EC3">journal article</mods:classification>
<mods:identifier id="DFB50525E465F868A7478E727A9DC7D3" type="CLB-Dataset">51371</mods:identifier>
<mods:identifier id="0311203A6532683104E682375E7A546C" type="DOI">10.5281/zenodo.156420</mods:identifier>
<mods:identifier id="78B0F4D994F0F6162E02D8C762C3EDBB" type="GBIF-Dataset">76c096e3-d27c-4abc-a504-a62ae8c7ba92</mods:identifier>
<mods:identifier id="58D451C29A2F7408252EA491EB054919" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="41E6207FC234A9296BC3B41D7B64F16E" type="Zenodo-Dep">156420</mods:identifier>
</mods:mods>
<treatment id="62088784C360C722E12E727AFED2FE96" ID-DOI="http://doi.org/10.5281/zenodo.6274687" ID-Zenodo-Dep="6274687" LSID="urn:lsid:plazi:treatment:62088784C360C722E12E727AFED2FE96" httpUri="http://treatment.plazi.org/id/62088784C360C722E12E727AFED2FE96" lastPageId="82" lastPageNumber="83" pageId="78" pageNumber="79">
<subSubSection id="A2BB6519C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79" type="key">
<paragraph id="EA1E3692C360C73EE12E727AFC7FF94D" blockId="78.[264,965,1643,1670]" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<heading id="B15681FEC360C73EE12E727AFC7FF94D" bold="true" box="[264,965,1643,1670]" fontSize="11" level="1" pageId="78" pageNumber="79" reason="1">
<emphasis id="D8D5EA80C360C73EE12E727AFC7FF94D" bold="true" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<key id="9E3061A3C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<keyStep id="51552E37C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<keyLead id="515095A7C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
23.
<taxonomicName id="2DA14D11C360C73EE112727DFD0DF94D" ID-CoL="6YBBC" authority="Gusarov" authorityName="Gusarov" box="[308,695,1643,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C360C73EE112727DFDFEF94E" bold="true" box="[308,580,1644,1670]" italics="true" pageId="78" pageNumber="79">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC360C73EE2E5727AFCBCF94D" box="[707,774,1643,1669]" pageId="78" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C360C73EE333727AFC06F94D" box="[789,956,1643,1669]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="78" pageNumber="79">Figs. 321­330</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C720E12E72A7FC86F8D7" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="EA1E3692C360C73EE12E72A7FECFF8E8" blockId="78.[264,1323,1718,1904]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE12E72A7FE04F918" bold="true" box="[264,446,1718,1744]" pageId="78" pageNumber="79">
<typeStatus id="351A8830C360C73EE12E72A7FEFFF918" box="[264,325,1718,1744]" pageId="78" pageNumber="79">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C360C73EE1EA72A7FD86F918" box="[460,572,1718,1744]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C360C73EE24C72A7FBA6F918" bold="true" box="[618,1052,1718,1744]" pageId="78" pageNumber="79">
<collectingCountry id="92B67602C360C73EE24C72A7FC99F918" box="[618,803,1718,1744]" name="Costa Rica" pageId="78" pageNumber="79">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89, Int. Amer. Hwy.,
<geoCoordinate id="8F955055C360C73EE1D972CFFDC1F930" box="[511,635,1758,1784]" direction="north" orientation="latitude" pageId="78" pageNumber="79" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C360C73EE2A472CFFCA8F930" box="[642,786,1758,1784]" direction="west" orientation="longitude" pageId="78" pageNumber="79" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C360C73EE33872CFFCC0F930" box="[798,890,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="78" pageNumber="79" unit="m" value="3300.0">3300 m</quantity>
, forest litter (R.Anderson),
<date id="9E1F1052C360C73EE49A72CFFA90F930" box="[1212,1322,1758,1784]" pageId="78" pageNumber="79" value="1997-06-08">8.vi.1997</date>
(
<collectionCode id="8CB0AE57C360C73EE1367317FED2F8E8" box="[272,360,1798,1824]" pageId="78" pageNumber="79">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C360C73FE11E733FFD83FE1E" blockId="78.[264,1323,1718,1904]" lastBlockId="79.[264,1324,284,470]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<typeStatus id="351A8830C360C73EE11E733FFE14F880" box="[312,430,1838,1864]" pageId="78" pageNumber="79" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C360C73EE1E6733FFD44F880" bold="true" box="[448,766,1838,1864]" pageId="78" pageNumber="79">
<collectingCountry id="92B67602C360C73EE1E6733FFDC0F880" box="[448,634,1838,1864]" name="Costa Rica" pageId="78" pageNumber="79">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
, same data as the
<typeStatus id="351A8830C360C73EE3D0733FFBE7F880" box="[1014,1117,1838,1864]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C360C73EE44A733FFB7EF880" box="[1132,1220,1838,1864]" pageId="78" pageNumber="79">KSEM</collectionCode>
).
<emphasis id="D8D5EA80C360C73EE11E7347FE02F8B8" bold="true" box="[312,440,1878,1904]" pageId="78" pageNumber="79">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C360C73EE1E17346FD69F8B8" box="[455,723,1879,1904]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C360C73EE1E17346FD69F8B8" box="[455,723,1879,1904]" italics="true" pageId="78" pageNumber="79">Seeversiella sulcicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C360C73FE4F57346FEFDFEFD" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" lastPageId="79" lastPageNumber="80" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C360C73FE4F57346FEFDFEFD" italics="true" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">Seeversiella</emphasis>
</taxonomicName>
by having reddish brown body; temples 1.8­2.3 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by having small medial emargination delimited by two short and blunt projections at the posterior margin of the male tergum 8 (
<figureCitation id="729A2A17C361C73FE1AE7585FE49FE66" box="[392,499,404,430]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="79" pageNumber="80">Fig. 330</figureCitation>
); tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C361C73FE1A575ADFD96FE1E" box="[387,556,444,470]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="79" pageNumber="80">Figs. 321­328</figureCitation>
).
</paragraph>
<caption id="BEDE661AC361C73FE12E729EFB76F923" httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="79" pageNumber="80" targetBox="[283,1308,537,1644]" targetPageId="79">
<paragraph id="EA1E3692C361C73FE12E729EFB76F923" blockId="79.[264,1322,1679,1771]" pageId="79" pageNumber="80">
<emphasis id="D8D5EA80C361C73FE12E729EFE67F96F" bold="true" box="[264,477,1679,1703]" pageId="79" pageNumber="80">FIGURES 321­324</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C361C73FE25B729EFC68F96E" authority="Gusarov" authorityName="Gusarov" box="[637,978,1679,1702]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C361C73FE25B729EFCCAF96E" box="[637,880,1679,1702]" italics="true" pageId="79" pageNumber="80">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C361C73FE3FA729EFBADF96F" bold="true" box="[988,1047,1679,1703]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="C3E657FBC361C73FE3FA729EFBADF96F" box="[988,1047,1679,1703]" pageId="79" pageNumber="80" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324).
</paragraph>
</caption>
<caption id="BEDE661AC37EC720E12E7229FE10F934" httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="80" pageNumber="81" targetBox="[280,1306,336,1561]" targetPageId="80">
<paragraph id="EA1E3692C37EC720E12E7229FE10F934" blockId="80.[264,1323,1592,1823]" pageId="80" pageNumber="81">
<emphasis id="D8D5EA80C37EC720E12E7229FE67F998" bold="true" box="[264,477,1592,1616]" pageId="80" pageNumber="81">FIGURES 325­330</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C37EC720E3B77229FB5FF987" authority="Gusarov" authorityName="Gusarov" box="[913,1253,1592,1615]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C37EC720E3B77229FB3EF987" box="[913,1156,1592,1615]" italics="true" pageId="80" pageNumber="81">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37EC720E4C97229FA91F998" box="[1263,1323,1592,1616]" pageId="80" pageNumber="81" rank="species">
<emphasis id="D8D5EA80C37EC720E4C97229FA91F998" bold="true" box="[1263,1323,1592,1616]" pageId="80" pageNumber="81">sp. n.</emphasis>
</taxonomicNameLabel>
(325, 327­330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326­328), 0.4 mm (329­330).
</paragraph>
</caption>
<paragraph id="EA1E3692C37EC720E12E7319FC86F8D7" blockId="80.[264,1323,1592,1823]" box="[264,828,1800,1823]" pageId="80" pageNumber="81">SLD sclerite of lateral diverticulum of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37FC721E11E750DFDB1F8EE" pageId="81" pageNumber="82" type="description">
<paragraph id="EA1E3692C37FC721E11E750DFE76FDEE" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
<taxonomicName id="2DA14D11C37FC721E11E750DFDFCFEFD" box="[312,582,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37FC721E11E750DFDFCFEFD" box="[312,582,284,309]" italics="true" pageId="81" pageNumber="82">Seeversiella sulcicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37FC721E2C1750DFC12FEFD" box="[743,936,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C37FC721E2C1750DFC12FEFD" box="[743,936,284,309]" italics="true" pageId="81" pageNumber="82">S. impressicollis</emphasis>
</taxonomicName>
in having larger body; in males pronotum with medial furrow, lateral portions of posterior margin of tergum 3 extending as short and obtuse projections (as in
<figureCitation id="729A2A17C37FC721E298757DFCA9FE4E" box="[702,787,364,390]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="81" pageNumber="82">Fig. 19</figureCitation>
), tergum 7 with short medial carina; posterior margin of tergum 8 with two medial denticles (
<figureCitation id="729A2A17C37FC721E3647585FC11FE66" box="[834,939,404,430]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
); apex of median lobe with two divergent carinae (in parameral view) (
<figureCitation id="729A2A17C37FC721E2C375ADFC28FE1E" box="[741,914,444,470]" captionStart="FIGURES 321 ­ 324" captionStartId="79.[264,382,1679,1703]" captionTargetBox="[283,1308,537,1644]" captionTargetId="figure@79.[272,1316,529,1652]" captionTargetPageId="79" captionText="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="81" pageNumber="82">Figs. 321­322</figureCitation>
); copulatory piece of internal sac longer (
<figureCitation id="729A2A17C37FC721E14175F5FDA6FE36" box="[359,540,484,510]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 326, 328</figureCitation>
; 317); sclerites of lateral diverticula of internal sac more robust (
<figureCitation id="729A2A17C37FC721E137761DFE38FDEE" box="[273,386,524,550]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 326</figureCitation>
; 318).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7625FEFFFD26" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
<taxonomicName id="2DA14D11C37FC721E11E7625FDFEFD85" box="[312,580,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37FC721E11E7625FDFEFD85" box="[312,580,564,589]" italics="true" pageId="81" pageNumber="82">Seeversiella sulcicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37FC721E2C77625FC08FD85" box="[737,946,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37FC721E2C77625FC08FD85" box="[737,946,564,589]" italics="true" pageId="81" pageNumber="82">S. microphthalma</emphasis>
</taxonomicName>
in having more asperate pronotal punctation; in males pronotum with medial furrow, posterior angles of tergum 3 not projecting as spines, posterior margin of tergum 8 with two medial denticles (
<figureCitation id="729A2A17C37FC721E4947695FAA6FD56" box="[1202,1308,644,670]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
); and the apex of median lobe with two divergent carinae (in parameral view) (
<figureCitation id="729A2A17C37FC721E48976BDFE82FD26" captionStart="FIGURES 321 ­ 324" captionStartId="79.[264,382,1679,1703]" captionTargetBox="[283,1308,537,1644]" captionTargetId="figure@79.[272,1316,529,1652]" captionTargetPageId="79" captionText="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="81" pageNumber="82">Figs. 321­ 322</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E76EAFC1AFCDE" blockId="81.[264,1324,284,1870]" box="[312,928,763,790]" pageId="81" pageNumber="82">
<emphasis id="D8D5EA80C37FC721E11E76EAFE6AFCDD" bold="true" box="[312,464,763,789]" pageId="81" pageNumber="82">Description.</emphasis>
Length 3.0 mm. Body reddish brown.
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7735FDB1FC7E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Temples 1.8­2.3 times as long as eyes. Antennal article 2 as long as 3, articles 4­6 subquadrate, 7­ 10 slightly transverse.</paragraph>
<paragraph id="EA1E3692C37FC721E11E77D5FCD3FB3E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C37FC721E40D77D5FB6CFC16" box="[1067,1238,964,990]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="81" pageNumber="82" unit="mm" value="0.665" valueMax="0.67" valueMin="0.66">0.66­0.67 mm</quantity>
, length
<quantity id="2D599B77C37FC721E12E77FDFE0BFBCE" box="[264,433,1004,1030]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.0" metricValueMin="5.8999999999999995" pageId="81" pageNumber="82" unit="mm" value="0.595" valueMax="0.6" valueMin="0.59">0.59­0.60 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­1.5 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7115FDD5FA2E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Abdominal terga glossy, terga 3­5 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 1­4 times their diameter; tergum 7 with microsculpture consisting of isodiametric or slightly transverse meshes and with fine and sparse punctation, distance between punctures equals 3­6 times their diameter. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C37FC721E1DD71DDFDD9FA2E" box="[507,611,1484,1510]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E71E5FD43F966" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
In males pronotum with medial furrow; lateral portions of posterior margin of tergum 3 extending as short and blunt projections up to 1/8 as long as tergum (measured medially) (as in
<figureCitation id="729A2A17C37FC721E1777255FE1DF996" box="[337,423,1604,1630]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="81" pageNumber="82">Fig. 19</figureCitation>
); tergum
<quantity id="2D599B77C37FC721E23F7255FDFDF996" box="[537,583,1604,1630]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="81" pageNumber="82" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina, up to 1/4 as long as the tergum; posterior margin of tergum 8 with small medial emargination delimited by two short and blunt projections (
<figureCitation id="729A2A17C37FC721E2A37285FD57F966" box="[645,749,1684,1710]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E72ADFD48F936" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Aedeagus as in
<figureCitation id="729A2A17C37FC721E1C872ADFD2EF91E" box="[494,660,1724,1750]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 321­328</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C37FC721E25872F5FD5FF936" box="[638,741,1764,1790]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 325</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E731DFDB1F8EE" blockId="81.[264,1324,284,1870]" box="[312,523,1804,1830]" pageId="81" pageNumber="82">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37FC722E11E7322FED2FE96" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="distribution">
<paragraph id="EA1E3692C37FC721E11E7322FBCDF886" blockId="81.[264,1324,284,1870]" box="[312,1143,1843,1870]" pageId="81" pageNumber="82">
<emphasis id="D8D5EA80C37FC721E11E7322FE62F885" bold="true" box="[312,472,1843,1869]" pageId="81" pageNumber="82">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C37FC721E3547325FC49F886" box="[882,1011,1844,1870]" name="Costa Rica" pageId="81" pageNumber="82">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C37FC721E4257325FBD0F886" box="[1027,1130,1844,1870]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="81" pageNumber="82">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37CC722E11E750AFED2FE96" blockId="82.[264,1323,283,350]" pageId="82" pageNumber="83">
<emphasis id="D8D5EA80C37CC722E11E750AFDB2FEFD" bold="true" box="[312,520,283,309]" pageId="82" pageNumber="83">Natural History.</emphasis>
<taxonomicName id="2DA14D11C37CC722E23A750DFC91FEFD" box="[540,811,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37CC722E23A750DFC91FEFD" box="[540,811,284,309]" italics="true" pageId="82" pageNumber="83">Seeversiella sulcicollis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C37CC722E12E7555FEDEFE96" box="[264,356,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="82" pageNumber="83" unit="m" value="3300.0">3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>