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<document id="E7C2FF21B073CA9AB673BFCE0E1F8B12" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C33BC768E12E75F2FA96FC46" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella texana Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="22" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="E07DBB083B01C9ADB70C079D63B84E12">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="76612D3394E1ABA0046ABDC0AC39B7F4">Gusarov, Vladimir I.</mods:namePart>
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<mods:typeOfResource id="897B83A4C7B897FDC32EE3D701FBF05A">text</mods:typeOfResource>
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<mods:date id="C28E784CBB91466984B223453E8AC894">2003</mods:date>
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<treatment id="62088784C33BC768E12E75F2FA96FC46" ID-DOI="http://doi.org/10.5281/zenodo.6274638" ID-Zenodo-Dep="6274638" LSID="urn:lsid:plazi:treatment:62088784C33BC768E12E75F2FA96FC46" httpUri="http://treatment.plazi.org/id/62088784C33BC768E12E75F2FA96FC46" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="A2BB6519C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22" type="key">
<paragraph id="EA1E3692C33BC765E12E75F2FCCBFE35" blockId="21.[264,881,483,510]" box="[264,881,483,510]" pageId="21" pageNumber="22">
<heading id="B15681FEC33BC765E12E75F2FCCBFE35" bold="true" box="[264,881,483,510]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<key id="9E3061A3C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22">
<keyStep id="51552E37C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E75F2FCCBFE35" bold="true" box="[264,881,483,510]" pageId="21" pageNumber="22">
<keyLead id="515095A7C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22">
2.
<taxonomicName id="2DA14D11C33BC765E10075F5FD3AFE35" ID-CoL="6YBBD" authority="Gusarov" authorityName="Gusarov" box="[294,640,483,510]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C33BC765E10075F5FDB7FE36" bold="true" box="[294,525,484,510]" italics="true" pageId="21" pageNumber="22">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC33BC765E2AB75F2FD6AFE35" box="[653,720,483,509]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C33BC765E2F975F2FCD0FE35" box="[735,874,483,509]" captionStart-0="FIGURES 37 ­ 40" captionStart-1="FIGURES 41 ­ 44" captionStart-2="FIGURES 45 ­ 53" captionStartId-0="21.[264,383,1773,1797]" captionStartId-1="22.[264,383,1298,1322]" captionStartId-2="23.[264,383,1314,1338]" captionTargetBox-0="[274,1312,896,1738]" captionTargetBox-1="[293,1296,576,1260]" captionTargetBox-2="[313,1274,323,1277]" captionTargetId-0="figure@21.[264,1323,884,1747]" captionTargetId-1="figure@22.[281,1306,569,1271]" captionTargetId-2="figure@23.[306,1280,313,1287]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 male tergum 8; 38 male sternum 8; 39 female tergum 8; 40 female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-2="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156428/files/figure.png" httpUri-1="https://zenodo.org/record/156429/files/figure.png" httpUri-2="https://zenodo.org/record/156430/files/figure.png" pageId="21" pageNumber="22">Figs. 37­53</figureCitation>
)
</keyLead>
</emphasis>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C33BC765E12E763FFD6EFCD9" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="EA1E3692C33BC765E12E763FFC83FD51" blockId="21.[264,1325,558,785]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E763FFE7BFD80" bold="true" box="[264,449,558,584]" pageId="21" pageNumber="22">
<typeStatus id="351A8830C33BC765E12E763FFEFFFD80" box="[264,325,558,584]" pageId="21" pageNumber="22">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C33BC765E1F3763EFDFFFD81" box="[469,581,559,585]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C33BC765E25E763FFC75FD80" bold="true" box="[632,975,558,584]" pageId="21" pageNumber="22">
<collectingCountry id="92B67602C33BC765E25E763FFCDCFD80" box="[632,870,558,584]" name="United States of America" pageId="21" pageNumber="22">UNITED STATES</collectingCountry>
: Texas:
</emphasis>
Culberson Co.:
<collectingCountry id="92B67602C33BC765E48A763EFA91FD81" box="[1196,1323,559,585]" name="Guadeloupe" pageId="21" pageNumber="22">Guadalupe</collectingCountry>
Mts. N. P., The Bowl, jct. Tejas Tr. and Juniper Tr.,
<quantity id="2D599B77C33BC765E3457646FC05FDB9" box="[867,959,599,625]" metricMagnitude="3" metricUnit="m" metricValue="2.407" pageId="21" pageNumber="22" unit="m" value="2407.0">2407 m</quantity>
, berlese maple, oak, pine, ironwood litter (R.Anderson),
<date id="9E1F1052C33BC765E219766EFD7FFD51" box="[575,709,639,665]" pageId="21" pageNumber="22" value="1989-07-31">31.vii.1989</date>
(
<collectionCode id="8CB0AE57C33BC765E2F2766EFC96FD51" box="[724,812,639,665]" pageId="21" pageNumber="22">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C33BC765E11E76B6FD6EFCD9" blockId="21.[264,1325,558,785]" pageId="21" pageNumber="22">
<typeStatus id="351A8830C33BC765E11E76B6FE15FD09" box="[312,431,679,705]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C33BC765E1E476B7FCA9FD08" bold="true" box="[450,787,678,704]" pageId="21" pageNumber="22">
<collectingCountry id="92B67602C33BC765E1E476B7FD17FD08" box="[450,685,678,704]" name="United States of America" pageId="21" pageNumber="22">UNITED STATES</collectingCountry>
: Texas:
</emphasis>
Culberson Co.:
<specimenCount id="FCA7FD1BC33BC765E3C076B6FB33FD09" box="[998,1161,679,705]" pageId="21" pageNumber="22" type="generic">14 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C33BC765E11176DEFE1BFD21" box="[311,417,719,745]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC33BC765E19676DEFDFCFD21" box="[432,582,719,745]" pageId="21" pageNumber="22" type="generic">4 specimens</specimenCount>
, ditto but mixed hardwood litter;
<specimenCount id="FCA7FD1BC33BC765E3C276DEFBC0FD21" box="[996,1146,719,745]" pageId="21" pageNumber="22" type="generic">8 specimens</specimenCount>
, ditto but oak, conifer litter,
<date id="9E1F1052C33BC765E18176E6FD9FFCD9" box="[423,549,759,785]" pageId="21" pageNumber="22" value="1988-09-13">13.ix.1988</date>
(all
<collectionCode id="8CB0AE57C33BC765E25676E6FD7DFCD9" box="[624,711,759,785]" pageId="21" pageNumber="22">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC33BC765E12E72FCFC32F882" httpUri="https://zenodo.org/record/156428/files/figure.png" pageId="21" pageNumber="22" targetBox="[274,1312,896,1738]" targetPageId="21">
<paragraph id="EA1E3692C33BC765E12E72FCFC32F882" blockId="21.[264,1323,1773,1866]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E72FCFE7FF8CD" bold="true" box="[264,453,1773,1797]" pageId="21" pageNumber="22">FIGURES 37­40</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C33BC765E2D072FFFB90F8CD" authority="Gusarov" authorityName="Gusarov" box="[758,1066,1774,1797]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C33BC765E2D072FFFC7DF8CD" box="[758,967,1774,1797]" italics="true" pageId="21" pageNumber="22">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C33BC765E41E72FCFBCCF8CD" bold="true" box="[1080,1142,1773,1797]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="C3E657FBC33BC765E41E72FCFBCCF8CD" box="[1080,1142,1773,1797]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 37 male tergum 8; 38 male sternum 8; 39 female tergum 8; 40 female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C338C766E11E750AFC9CF969" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="EA1E3692C338C766E11E750AFD8AFE36" blockId="22.[264,1323,283,510]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C338C766E1E8750DFD0FFEFD" box="[462,693,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E1E8750DFD0FFEFD" box="[462,693,284,309]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C338C766E4F5750DFEFDFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C338C766E4F5750DFEFDFE95" italics="true" pageId="22" pageNumber="23">Seeversiella</emphasis>
</taxonomicName>
by having brown or brownish yellow body; small eyes (temples 2.1­2.7 times as long as eyes); glossy pronotum with weak microsculpture; wings reduced, 2 times as long as elytra; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C338C766E4E77585FE92FE1E" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 41­ 52</figureCitation>
), especially the hook­shaped distal sclerites of internal sac (
<figureCitation id="729A2A17C338C766E3C175ADFBCCFE1E" box="[999,1142,444,470]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 48, 51</figureCitation>
); and the shape of spermatheca (
<figureCitation id="729A2A17C338C766E1EC75F5FD98FE36" box="[458,546,484,510]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
).
</paragraph>
<caption id="BEDE661AC338C766E12E7103FDCFFA59" httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="22" pageNumber="23" targetBox="[293,1296,576,1260]" targetPageId="22">
<paragraph id="EA1E3692C338C766E12E7103FDCFFA59" blockId="22.[264,1323,1298,1425]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E12E7103FE7DFAE2" bold="true" box="[264,455,1298,1322]" pageId="22" pageNumber="23">FIGURES 41­44</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C338C766E2527103FC10FAE1" authority="Gusarov" authorityName="Gusarov" box="[628,938,1298,1321]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C338C766E2527103FCFFFAE1" box="[628,837,1298,1321]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C338C766E39E7103FC4CFAE2" bold="true" box="[952,1014,1298,1322]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="C3E657FBC338C766E39E7103FC4CFAE2" box="[952,1014,1298,1322]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44).
</paragraph>
</caption>
<paragraph id="EA1E3692C338C766E11E71F9FC9CF969" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<taxonomicName id="2DA14D11C338C766E11E71F9FDA4F9C9" box="[312,542,1512,1537]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E11E71F9FDA4F9C9" box="[312,542,1512,1537]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C338C766E35E71F9FBAFF9C9" box="[888,1045,1512,1537]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C338C766E35E71F9FBAFF9C9" box="[888,1045,1512,1537]" italics="true" pageId="22" pageNumber="23">S. globicollis</emphasis>
</taxonomicName>
in having lighter body colour; more distinct microsculpture of head and pronotum; elytra shorter than pronotum; reduced wings, 2 times as long as elytra; apex of median lobe bent paramerally (in lateral view) (
<figureCitation id="729A2A17C338C766E17B724EFE53F9B1" box="[349,489,1631,1657]" captionStart="FIGURES 41 ­ 44" captionStartId="22.[264,383,1298,1322]" captionTargetBox="[293,1296,576,1260]" captionTargetId="figure@22.[281,1306,569,1271]" captionTargetPageId="22" captionText="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="22" pageNumber="23">Figs. 43­44</figureCitation>
; 26­27), shorter copulatory piece of internal sac (
<figureCitation id="729A2A17C338C766E41B724EFB42F9B1" box="[1085,1272,1631,1657]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 46, 49­50</figureCitation>
; 29, 34) and spermatheca with umbilicus (
<figureCitation id="729A2A17C338C766E2E77296FCA3F969" box="[705,793,1671,1697]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C338C768E11E72A1FDD5FCF6" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="EA1E3692C338C766E11E72A1FBF5F8D1" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<taxonomicName id="2DA14D11C338C766E11E72A1FDA5F901" box="[312,543,1712,1737]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E11E72A1FDA5F901" box="[312,543,1712,1737]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
differs from the other similar species of
<taxonomicName id="2DA14D11C338C766E43872A1FB16F901" box="[1054,1196,1712,1737]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C338C766E43872A1FB16F901" box="[1054,1196,1712,1737]" italics="true" pageId="22" pageNumber="23">Seeversiella</emphasis>
</taxonomicName>
with long elytra and weak pronotal microsculpture in having strongly sclerotized hook­shaped distal sclerites of internal sac and J­shaped spermatheca with large umbilicus.
</paragraph>
<paragraph id="EA1E3692C338C766E11E7336FB32F8A1" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E11E7336FE6AF889" bold="true" box="[312,464,1831,1857]" pageId="22" pageNumber="23">Description.</emphasis>
Length
<quantity id="2D599B77C338C766E2677336FD75F889" box="[577,719,1831,1857]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="22" pageNumber="23" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Body from brownish yellow to brown, head and abdominal segments 6­7 often darker, legs and mouthparts brownish yellow.
</paragraph>
<caption id="BEDE661AC339C767E12E7133FDC5FA2E" httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="23" pageNumber="24" targetBox="[313,1274,323,1277]" targetPageId="23">
<paragraph id="EA1E3692C339C767E12E7133FDC5FA2E" blockId="23.[264,1323,1314,1545]" pageId="23" pageNumber="24">
<emphasis id="D8D5EA80C339C767E12E7133FE7DFAF2" bold="true" box="[264,455,1314,1338]" pageId="23" pageNumber="24">FIGURES 45­53</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C339C767E2497133FC1DFAF1" authority="Gusarov" authorityName="Gusarov" box="[623,935,1314,1337]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C339C767E2497133FCFBFAF1" box="[623,833,1314,1337]" italics="true" pageId="23" pageNumber="24">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C339C767E3907133FC4FFAF2" bold="true" box="[950,1013,1314,1338]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="C3E657FBC339C767E3907133FC4FFAF2" box="[950,1013,1314,1338]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48­49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48­49, 53), 0.1 mm (46­47, 50­52).
</paragraph>
</caption>
<paragraph id="EA1E3692C339C767E12E71E3FDCDF9C1" blockId="23.[264,1323,1314,1545]" box="[264,631,1522,1545]" pageId="23" pageNumber="24">DS distal sclerite of internal sac.</paragraph>
<paragraph id="EA1E3692C339C767E11E7274FBD5F93F" blockId="23.[264,1324,1637,1903]" pageId="23" pageNumber="24">
Head surface glossy, with weak isodiametric microsculpture, with fine, weak and poorly visible (at
<date id="9E1F1052C339C767E1F0729CFDBEF96F" box="[470,516,1677,1703]" pageId="23" pageNumber="24" value="1970-10">70x</date>
) punctation, distance between punctures equals 2­3 times their diameter. Temples 2.1­2.7 times as long as eyes. Antennal article 2 longer than article 3, article 4 slightly transverse, 5­10 transverse or strongly transverse (ratio 1.5­2.0).
</paragraph>
<paragraph id="EA1E3692C339C768E11E7314FE14FE66" blockId="23.[264,1324,1637,1903]" lastBlockId="24.[264,1324,284,910]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C339C767E40D7314FB6CF8D7" box="[1067,1238,1797,1823]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.9" metricValueMin="4.4" pageId="23" pageNumber="24" unit="mm" value="0.46499999999999997" valueMax="0.49" valueMin="0.44">0.44­0.49 mm</quantity>
, length
<quantity id="2D599B77C339C767E12E733CFE0FF88F" box="[264,437,1837,1863]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="23" pageNumber="24" unit="mm" value="0.435" valueMax="0.46" valueMin="0.41">0.41­0.46 mm</quantity>
, width to length ratio 1.1, surface glossy, with weak isodiametric microsculpture; punctation as on head. Elytra wider and shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.1), 1.4 times wider than long, glossy, with weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings reduced, 2 times as long as elytra.
</paragraph>
<paragraph id="EA1E3692C336C768E11E75ADFCA7FD86" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (37, 39).</paragraph>
<paragraph id="EA1E3692C336C768E11E764DFA92FD0E" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
In males posterior angles of tergum 3 projecting as spines up to half as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to half as long as tergum length. Posterior margin of male tergum 8 without emargination (
<figureCitation id="729A2A17C336C768E4E576BDFAA0FD0E" box="[1219,1306,684,710]" captionStart="FIGURES 37 ­ 40" captionStartId="21.[264,383,1773,1797]" captionTargetBox="[274,1312,896,1738]" captionTargetId="figure@21.[264,1323,884,1747]" captionTargetPageId="21" captionText="FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 male tergum 8; 38 male sternum 8; 39 female tergum 8; 40 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156428/files/figure.png" pageId="24" pageNumber="25">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E76C5FAA7FCDE" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
Aedeagus as in
<figureCitation id="729A2A17C336C768E1C976C5FDC3FD26" box="[495,633,724,750]" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Figs. 41­52</figureCitation>
. Distal sclerites of internal sac hook­shaped (
<figureCitation id="729A2A17C336C768E4B676C5FAA5FD26" box="[1168,1311,724,750]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Figs. 48, 51</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C336C768E49E76EDFAAAFCDE" box="[1208,1296,764,790]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7735FDD5FCF6" blockId="24.[264,1324,284,910]" box="[312,623,804,830]" pageId="24" pageNumber="25">
Spermatheca as in
<figureCitation id="729A2A17C336C768E2337735FDD6FCF6" box="[533,620,804,830]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Fig. 53</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E11E775AFA96FC46" pageId="24" pageNumber="25" type="distribution">
<paragraph id="EA1E3692C336C768E11E775AFB31FCAE" blockId="24.[264,1324,284,910]" box="[312,1163,843,870]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E775AFE62FCAD" bold="true" box="[312,472,843,869]" pageId="24" pageNumber="25">Distribution.</emphasis>
Known from the
<collectingCountry id="92B67602C336C768E296775DFC95FCAE" box="[688,815,844,870]" name="Guadeloupe" pageId="24" pageNumber="25">Guadalupe</collectingCountry>
Mountains, Texas (
<figureCitation id="729A2A17C336C768E430775DFBC4FCAE" box="[1046,1150,844,870]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="24" pageNumber="25">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7762FA96FC46" blockId="24.[264,1324,284,910]" box="[312,1324,883,910]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E7762FDBFFC45" bold="true" box="[312,517,883,909]" pageId="24" pageNumber="25">Natural History.</emphasis>
<taxonomicName id="2DA14D11C336C768E2357765FD4CFC45" box="[531,758,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C336C768E2357765FD4CFC45" box="[531,758,884,909]" italics="true" pageId="24" pageNumber="25">Seeversiella texana</emphasis>
</taxonomicName>
was collected at altitude of
<quantity id="2D599B77C336C768E41B7765FB2CFC46" box="[1085,1174,884,910]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="25" unit="m" value="2400.0">2400 m</quantity>
in leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>