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<document ID-DOI="http://dx.doi.org/10.3897/jhr.53.9890" ID-GBIF-Dataset="e6b7983e-dc70-447a-bb89-c93b74d6f249" ID-GBIF-Taxon="154473352" ID-Pensoft-Pub="1314-2607-53-77" ID-Pensoft-UUID="CB6EB5480F1F2F28FFF91521801EFF8C" ID-ZBK="989B7FE41F5048A784148D0AF4F4C064" ID-Zenodo-Dep="575132" ID-ZooBank="989B7FE41F5048A784148D0AF4F4C064" LSID="urn:lsid:zoobank.org:act:9C8A4DD6-E635-4B7A-A8EB-02646151D521" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2607-53-77" ModsDocOrigin="Journal of Hymenoptera Research " ModsDocTitle="New Dryocosmus Giraud species associated with Cyclobalanopsis and non-Quercus host plants from the Eastern Palaearctic (Hymenoptera, Cynipidae, Cynipini)" checkinTime="1553125490764" checkinUser="pensoft" docAuthor="Tang, Chang-Ti, Miko, Istvan, Nicholls, James A., Schweger, Szabina, Yang, Man-Miao, Stone, Graham N., Sinclair, Frazer, Bozso, Miklos, Melika, George &amp; Penzes, Zsolt" docDate="2016" docId="5E28449C08EA28F1B58C01E3B18A4C14" docLanguage="en" docName="JourHymenoptRes 53: 77-162" docOrigin="Journal of Hymenoptera Research 53" docPubDate="2016-12-19" docSource="http://dx.doi.org/10.3897/jhr.53.9890" docTitle="Dryocosmus moriius Tang &amp; Melika 2016, sp. n." docType="treatment" docUuid="9C8A4DD6-E635-4B7A-A8EB-02646151D521" docUuidSource="ZooBank" docVersion="4" id="CB6EB5480F1F2F28FFF91521801EFF8C" lastPageId="58" lastPageNumber="135" masterDocId="CB6EB5480F1F2F28FFF91521801EFF8C" masterDocTitle="New Dryocosmus Giraud species associated with Cyclobalanopsis and non- Quercus host plants from the Eastern Palaearctic (Hymenoptera, Cynipidae, Cynipini)" masterLastPageNumber="162" masterPageNumber="77" pageId="53" pageNumber="130" updateTime="1643491172415" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New Dryocosmus Giraud species associated with Cyclobalanopsis and non- Quercus host plants from the Eastern Palaearctic (Hymenoptera, Cynipidae, Cynipini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tang, Chang-Ti</mods:namePart>
<mods:affiliation>Department of Entomology, National Chung Hsing University, 250 Kuo Kuang Rd., Taichung 402, Taiwan</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Miko, Istvan</mods:namePart>
<mods:affiliation>Pennsylvania State University, Department of Entomology, Frost Entomological Museum, PA, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nicholls, James A.</mods:namePart>
<mods:affiliation>Institute of Evolutionary Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH 9 3 JT, Scotland, U. K.</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schweger, Szabina</mods:namePart>
<mods:affiliation>University of Szeged, Department of Ecology, Koezep fasor 52 Szeged, Hungary</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yang, Man-Miao</mods:namePart>
<mods:affiliation>Department of Entomology, National Chung Hsing University, 250 Kuo Kuang Rd., Taichung 402, Taiwan</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Stone, Graham N.</mods:namePart>
<mods:affiliation>Institute of Evolutionary Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH 9 3 JT, Scotland, U. K.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sinclair, Frazer</mods:namePart>
<mods:affiliation>Institute of Evolutionary Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH 9 3 JT, Scotland, U. K.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bozso, Miklos</mods:namePart>
<mods:affiliation>Plant Health and Molecular Biology Laboratory, National Food Chain Safety Office, Directorate of Plant Protection, Soil Conservation and Agri-environment, Budaoersi str. 141 - 145, Budapest 1118, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Melika, George</mods:namePart>
<mods:affiliation>Plant Health and Molecular Biology Laboratory, National Food Chain Safety Office, Directorate of Plant Protection, Soil Conservation and Agri-environment, Budaoersi str. 141 - 145, Budapest 1118, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Penzes, Zsolt</mods:namePart>
<mods:affiliation>University of Szeged, Department of Ecology, Koezep fasor 52 Szeged, Hungary</mods:affiliation>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:date>2016</mods:date>
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<mods:number>2016-12-19</mods:number>
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<mods:start>77</mods:start>
<mods:end>162</mods:end>
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<mods:url>http://dx.doi.org/10.3897/jhr.53.9890</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.53.9890</mods:identifier>
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<treatment ID-GBIF-Taxon="154473352" LSID="urn:lsid:zoobank.org:act:9C8A4DD6-E635-4B7A-A8EB-02646151D521" httpUri="http://treatment.plazi.org/id/5E28449C08EA28F1B58C01E3B18A4C14" lastPageId="58" lastPageNumber="135" pageId="53" pageNumber="130">
<subSubSection pageId="53" pageNumber="130" type="nomenclature">
<paragraph pageId="53" pageNumber="130">
<taxonomicName LSID="http://zoobank.org/9C8A4DD6-E635-4B7A-A8EB-02646151D521" authority="Tang &amp; Melika" authorityName="Tang &amp; Melika" authorityYear="2016" class="Insecta" family="Cynipidae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus moriius" order="Hymenoptera" pageId="53" pageNumber="130" phylum="Arthropoda" rank="species" species="moriius" status="sp. n.">Dryocosmus moriius Tang &amp; Melika</taxonomicName>
<taxonomicNameLabel pageId="53" pageNumber="130">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figures 122129" captionStartId="F23" captionText="Figures 122 - 129. Dryocosmus moriius sp. n. 122 - 124 head, female: 122 frontal view 123 dorsal view 124 posterior view 125 head, frontal view, male 126 - 127 antenna: 126 female 127 male 128 - 129 female: 128 mesosoma, dorsal view 129 fore wing, part, female (rad = radial cell)." figureDoi="10.3897/jhr.53.9890.figures122-129" httpUri="https://binary.pensoft.net/fig/115562" pageId="53" pageNumber="130">Figures 122-129</figureCitation>
<figureCitation captionStart="Figures 130135" captionStartId="F24" captionText="Figures 130 - 135. Dryocosmus moriius sp. n. 130 mesosoma, female, lateral view 131 metascutellum and propodeum, female, posterodorsal view 132 - 133 metasoma, lateral view: 132 female 133 male 134 - 135 galls (photos by C. - T. Tang) (vsh = ventral spine of hypopygium)." figureDoi="10.3897/jhr.53.9890.figures130-135" httpUri="https://binary.pensoft.net/fig/115563" pageId="53" pageNumber="130">, 130-135</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="54" lastPageNumber="131" pageId="53" pageNumber="130" type="type material">
<paragraph pageId="53" pageNumber="130">Type material.</paragraph>
<paragraph lastPageId="54" lastPageNumber="131" pageId="53" pageNumber="130">
HOLOTYPE female: TAIWAN: Nantou Co., Mt. Liying, Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="53" pageNumber="130" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="53" pageNumber="130">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT474),
<geoCoordinate degrees="24" direction="north" minutes="04" orientation="latitude" precision="1" seconds="49.68" value="24.080467">24°04'49.68&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="14.03" value="121.17056">121°10'14.03&quot;E</geoCoordinate>
, 2217m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 26.IV.2011, leg. Chang-Ti Tang. PARATYPES: 31 females and 8 males: 3 female paratypes with the same labels as holotype; 2 male and 2 female paratypes: TAIWAN: Nantou Co., Mt. Liying, Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="53" pageNumber="130" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="53" pageNumber="130">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT474),
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,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="14.03" value="121.17056">121°10'14.03&quot;E</geoCoordinate>
, 2217m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 27.IV.2011, leg. Chang-Ti Tang; 3 female paratypes: TAIWAN: Nantou Co., Mt. Liying, Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="53" pageNumber="130" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="53" pageNumber="130">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT474),
<geoCoordinate degrees="24" direction="north" minutes="04" orientation="latitude" precision="1" seconds="49.68" value="24.080467">24°04'49.68&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="14.03" value="121.17056">121°10'14.03&quot;E</geoCoordinate>
, 2217m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 28.IV.2011, leg. Chang-Ti Tang; 1 female paratype: TAIWAN: Nantou Co., Mt. Liying, Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="53" pageNumber="130" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="53" pageNumber="130">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT474),
<geoCoordinate degrees="24" direction="north" minutes="04" orientation="latitude" precision="1" seconds="49.68" value="24.080467">24°04'49.68&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="14.03" value="121.17056">121°10'14.03&quot;E</geoCoordinate>
, 2217m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 2.V.2011, leg. Chang-Ti Tang; 1 female paratype: TAIWAN: Nantou Co., Mt. Liying, Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="53" pageNumber="130" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="53" pageNumber="130">Quercus morii</emphasis>
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, 25.IV.2011 (TWT474),
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,
<pageBreakToken pageId="54" pageNumber="131" start="start">2217</pageBreakToken>
m, ovipositing female on young leaves of host plant, leg. Chang-Ti Tang; 2 male and 1 female paratypes: TAIWAN: Nantou Co., near Cuifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT498),
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,
<geoCoordinate degrees="121" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.47" value="121.19013">121°11'24.47&quot;E</geoCoordinate>
, 2285m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 1.V.2011, leg. Chang-Ti Tang; 1 male and 3 female paratypes: TAIWAN: Nantou Co., near Cuifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT498),
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,
<geoCoordinate degrees="121" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.47" value="121.19013">121°11'24.47&quot;E</geoCoordinate>
, 2285m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 2.V.2011, leg. Chang-Ti Tang; 1 male and 1 female paratypes: TAIWAN: Nantou Co., near Cuifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT498),
<geoCoordinate degrees="24" direction="north" minutes="06" orientation="latitude" precision="1" seconds="06.63" value="24.101843">24°06'06.63&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.47" value="121.19013">121°11'24.47&quot;E</geoCoordinate>
, 2285m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 3.V.2011, leg. Chang-Ti Tang; 3 female paratypes: TAIWAN: Nantou Co., near Cuifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT498),
<geoCoordinate degrees="24" direction="north" minutes="06" orientation="latitude" precision="1" seconds="06.63" value="24.101843">24°06'06.63&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.47" value="121.19013">121°11'24.47&quot;E</geoCoordinate>
, 2285m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 6.V.2011, leg. Chang-Ti Tang; 2 female paratypes: TAIWAN: Nantou Co., near Cuifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT498),
<geoCoordinate degrees="24" direction="north" minutes="06" orientation="latitude" precision="1" seconds="06.63" value="24.101843">24°06'06.63&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.47" value="121.19013">121°11'24.47&quot;E</geoCoordinate>
, 2285m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 9.V.2011, leg. Chang-Ti Tang; 1 female paratype: TAIWAN: Nantou Co., near Cuifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT498),
<geoCoordinate degrees="24" direction="north" minutes="06" orientation="latitude" precision="1" seconds="06.63" value="24.101843">24°06'06.63&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.47" value="121.19013">121°11'24.47&quot;E</geoCoordinate>
, 2285m, ex small green oval bud gall cover by bud-scales (TWTb13), em. 13.V.2011, leg. Chang-Ti Tang; 1 male and 3 female paratypes: TAIWAN: Nantou Co., near Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT502),
<geoCoordinate degrees="24" direction="north" minutes="05" orientation="latitude" precision="1" seconds="19.33" value="24.088703">24°05'19.33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="17.98" value="121.17166">121°10'17.98&quot;E</geoCoordinate>
, 2120m, ex small oval grenn bud gall cover by bud-scales (TWTb13), em. 27.IV.2011, leg. Chang-Ti Tang; 1 male and 4 female paratypes: TAIWAN: Nantou Co., near Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT502),
<geoCoordinate degrees="24" direction="north" minutes="05" orientation="latitude" precision="1" seconds="19.33" value="24.088703">24°05'19.33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="17.98" value="121.17166">121°10'17.98&quot;E</geoCoordinate>
, 2120m, ex small oval grenn bud gall cover by bud-scales (TWTb13), em. 28.IV.2011, leg. Chang-Ti Tang; 2 female paratypes: TAIWAN: Nantou Co., near Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT502),
<geoCoordinate degrees="24" direction="north" minutes="05" orientation="latitude" precision="1" seconds="19.33" value="24.088703">24°05'19.33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="17.98" value="121.17166">121°10'17.98&quot;E</geoCoordinate>
, 2120m, ex small oval grenn bud gall cover by bud-scales (TWTb13), em. 29.IV.2011, leg. Chang-Ti Tang; 1 female paratype: TAIWAN: Nantou Co., near Meifeng, Renai Township, ex
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
, 25.IV.2011 (TWT502),
<geoCoordinate degrees="24" direction="north" minutes="05" orientation="latitude" precision="1" seconds="19.33" value="24.088703">24°05'19.33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="10" orientation="longitude" precision="1" seconds="17.98" value="121.17166">121°10'17.98&quot;E</geoCoordinate>
, 2120m, ex small oval grenn bud gall cover by bud-scales (TWTb13), em. 3.V.2011, leg. Chang-Ti Tang.
</paragraph>
<paragraph pageId="54" pageNumber="131">The female holotype, 9 female and 2 male paratypes are deposited in NMNS, 9 female and 3 male paratypes in PHMB, 3 female and 1 male paratypes in USNM, 10 female and 2 male paratypes in NCHU.</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="131" type="etymology">
<paragraph pageId="54" pageNumber="131">Etymology.</paragraph>
<paragraph pageId="54" pageNumber="131">
Named after the host plant,
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="54" pageNumber="131">Quercus morii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="55" lastPageNumber="132" pageId="54" pageNumber="131" type="diagnosis">
<paragraph pageId="54" pageNumber="131">Diagnosis.</paragraph>
<paragraph pageId="54" pageNumber="131">
<taxonomicName authorityName="Tang &amp; Melika" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus moriius" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="moriius">
<emphasis italics="true" pageId="54" pageNumber="131">Dryocosmus moriius</emphasis>
</taxonomicName>
is most similar to
<taxonomicName authorityName="Ide, Wachi &amp; Abe" authorityYear="2013" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus sakureiensis" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="sakureiensis">
<emphasis italics="true" pageId="54" pageNumber="131">Dryocosmus sakureiensis</emphasis>
</taxonomicName>
, both sharing the presence of a basal lobe on the metatarsal claw.
</paragraph>
<paragraph pageId="54" pageNumber="131">
<taxonomicName authorityName="Ide, Wachi &amp; Abe" authorityYear="2013" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus sakureiensis" order="Hymenoptera" pageId="54" pageNumber="131" phylum="Arthropoda" rank="species" species="sakureiensis">
<emphasis italics="true" pageId="54" pageNumber="131">Dryocosmus sakureiensis</emphasis>
</taxonomicName>
: Female antenna with 12 flagellomeres; eye height / malar distance = 2.62; intertorular distance / eye torulus distance = 0.67; radial cell length / radial cell width = 3.43; 2nd metasomal tergite length dorsal view / length of metasoma dorsal view&gt; 0.50; prominent part of ventral spine of hypopygium length ventral view / prominent part of ventral spine of hypopygium width ventral view = 2.20.
</paragraph>
<paragraph pageId="55" pageNumber="132">
<taxonomicName authorityName="Tang &amp; Melika" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus moriius" order="Hymenoptera" pageId="55" pageNumber="132" phylum="Arthropoda" rank="species" species="moriius">
<emphasis italics="true" pageId="55" pageNumber="132">
<pageBreakToken pageId="55" pageNumber="132" start="start">Dryocosmus</pageBreakToken>
moriius
</emphasis>
</taxonomicName>
: Female antenna with 13 flagellomeres, eye height / malar distance = 2.0; intertorular distance / eye torulus distance = 0.36; radial cell length / radial cell width = 4.5; 2nd metasomal tergite length dorsal view / length of metasoma dorsal view = 0.33; prominent part of ventral spine of hypopygium length ventral view / prominent part of ventral spine of hypopygium width ventral view = 1.3.
</paragraph>
</subSubSection>
<subSubSection lastPageId="58" lastPageNumber="135" pageId="56" pageNumber="133" type="description">
<paragraph pageId="56" pageNumber="133">
<pageBreakToken pageId="56" pageNumber="133" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="56" pageNumber="133">Sexual female: Head color: Head black, except for light brown clypeus; mandibles, maxillary and labial palps yellowish; scape, pedicel, F1-F4 light brown, subsequent flagellomeres progressively darker. Mesosoma and metasoma color: mesosoma black, except for lighter tegula; metasoma dark brown; legs yellowish.</paragraph>
<paragraph lastPageId="57" lastPageNumber="134" pageId="56" pageNumber="133">
Head sculpture: coriaceous. Head shape anterior view: rounded. Head width / head length: 2.10. Head width / head height: 1.20. Head width / maximum mesosoma width: &lt;1. Gena sculpture: coriaceous. Gena reflectivity: NOT CODED. Gena length / eye width: 0.94. Gena shape: broadened posterior to eye, visible in frontal view. Malar striae count: present. Malar striae dorsal limit: torulus-eye line on lower face and lower eye margin on malar area. Malar area: alutaceous. Malar sulcus: absent. Eye height / malar distance: 2.0. Inner margins of eyes: NOT CODED. Median ocellus shape: NOT CODED. Lateral ocellus shape: NOT CODED. POL / OOL: 1.20. OOL / diameter of lateral ocellus: 2.45. OOL / LOL: 1.97. Diameter of lateral ocellus / diameter of median ocellus: 1. Transfacial distance / eye height: 1.30. Diameter of torulus / intertorular distance: 1.72. Intertorular distance / eye torulus distance: 0.4. Eye-torulus distance / diameter of torulus: 1.25. Lower face sculpture: NOT CODED. Lower face pilosity color: NOT CODED. Lower face pilosity density: NOT CODED. Clypeus convexity: NOT CODED. Clypeus sculpture: NOT CODED. Clypeus ventral margin shape: straight. Clypeus reflectivity: matt. Clypeus shape anterior view: rectangular. Clypeus pilosity: NOT CODED. Clypeus pilosity density: NOT CODED. Clypeus pilosity color: NOT CODED. Anterior tentorial pit: small, indistinct. Epistomal sulcus: indistinct. Clypeo-pleurostomal line: indistinct. Frons sculpture: alutaceous. Frons pilosity density: rare. Frons reflectivity: NOT CODED. Frons pilosity color: white. Impression around central ocellus: present. Interocellar area sculpture: alutaceous. Interocellar area pilosity density: rare. Interocellar area reflectivity: glossy. Interocellar area pilosity color: white. Vertex sculpture: alutaceous. Vertex pilosity density: rare. Vertex reflectivity: glossy. Vertex pilosity color: white. Occiput sculpture: alutaceous. Occiput pilosity density: rare. Occiput reflectivity: glossy. Occiput pilosity color: white. Postocciput sculpture: smooth. Postocciput reflectivity: glossy. Postocciput pilosity: present. Postocciput pilosity density: rare. Postocciput pilosity color: white. Median impression of postocciput dorsal to occipital foramen: present. Posterior tentorial pit: distinct, ovate, deep. Impression abjacent ventrally to posterior tentorial pit: absent. Postgena sculpture: alutaceous. Postgena reflectivity: glossy. Postgena pilosity color: white. Postgena pilosity density: NOT CODED. Postgena pilosity count: absent medially, present laterally. Postgenal bridge / height of occipital foramen: 1. Postgenal bridge / length of oral foramen: &lt;1. Postgenal bridge sculpture: NOT CODED. Number of flagellomeres (female): 13. Antenna length / body length: &lt;1. Pedicel length / pedicel width: longer than wide. Pedicel length / length of broadened part of scape: 1.08. Combined length of scape and pedicel / first flagellomere length: NOT CODED. F1 length / F2 length: 1.1. F1 length / pedicel length: 2.1. F1 length / F3 length: NOT CODED. Flagellomeres relative length: F2-F8 progressively shorter. F4-F7 relative length: NOT CODED. F3 length/F2 length: NOT CODED.
<pageBreakToken pageId="57" pageNumber="134" start="start">F</pageBreakToken>
3 length / F4 length: NOT CODED. F8 length / F7 length: NOT CODED. F9 length /F8 length: 1.30. F10 length / F9 length:&gt;1. F9-F11 relative length: NOT CODED. F10 length / F11 length: 1.00. F12 length / F11 length: 0.9. F13 length / F12 length: NOT CODED. Placoid sensilla present on: F3-F13.
</paragraph>
<paragraph lastPageId="58" lastPageNumber="135" pageId="57" pageNumber="134">
Mesosoma length lateral view / mesosoma height lateral view: 1.14. Pronotum sculpture: alutaceous with some rugae laterally. Pronotal dorsal row of setae count: present. Transverse pronotal sulcus depth: deep. Transverse pronotal sulcus sculpture: foevolate. Mesoscutum sculpture: smooth. Mesoscutum reflectivity: glossy. Adnotaular setae: present. Mesoscutum length / transscutal line: 1.35. Notaulus limits: well-impressed, posterior end adjacent to posterior margin of mesoscutum, anterior end adjacent to anterior margin of mesoscutum. Notaulus sculpture: smooth. Notaulus posterior region width / anterior region width: NOT CODED. Mesoscutal suprahumeral sulcus anterior end vs notaulus anterior end: NOT CODED. Mesoscutal suprahumeral sulcus sculpture: NOT CODED. Median mesoscutal line: present. Median mesoscutal line shape: triangular. Parapsidal line: absent. Parapsidal line distinctness: NOT CODED. Anteroadmedian line: absent. Antero-admedian line length / mesoscutum median length: NOT CODED. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae): rugose medially, foveolate laterally and posteriorly. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae) shape: trapezoid. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae) pilosity color: white. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae) pilosity density: rare. Mesoscutellar axillar complex posterior margin vs metanotum: overhanging. Mesoscutellar-axillar complex length / mesoscutellar disc width:&gt;1. Disc of mesoscutellum sculpture: rugose. Disc of mesoscutellum maximum width: in posterior 1/4th. Scutellar fovea sculpture: smooth without longitudinal rugae. Scutellar fovea shape: transversely ovate. Scutellar fovea reflectivity: glossy. Foveal septum: narrow. Anterior pits on foveal septum: NOT CODED. Scutellar fovea maximum diameter / scutellar fovea minimum diameter: 2.25. Scutellar fovea minimum diameter / foveal septum width: NOT CODED. Postacetabular suclus count: NOT CODED. Mesopleuron sculpture: coriaceous, striae marking transepisternal line absent. Mesopleuron reflectivity: matt. Mesopleuron pilosity: dense setae present just ventral to transepisternal line. Speculum sculpture: smooth. Speculum reflectivity: glossy. Mesopleural triangle sculpture: rugose. Mesopleural triangle reflectivity: NOT CODED. Mesopleural triangle pilosity: present. Mesopleural triangle pilosity color: NOT CODED. Mesopleural trinagle pilosity density: rare. Dorsoaxillar area sculpture: smooth. Dorsoaxillar area reflectivity: glossy. Dorsoaxillar area pilosity color: white. Dorsoaxillar area pilosity density: rare. Lateroaxillar area sculpture: smooth. Lateroaxillar area reflectivity: glossy. Lateroaxillar area pilosity density: rare. Lateroaxillar area pilosity color: white. Subaxillular bar sculpture: smooth. Subaxillular bar reflectivity: glossy. Posterior height of subaxillular bar / height of metanotal trough: 1. Metapleural sulcus anterior end: reaches mesometapleural suture slightly
<pageBreakToken pageId="58" pageNumber="135" start="start">ventrally</pageBreakToken>
of its mid-height. Metascutellum sculpture: coriaceous. Metanotal trough sculpture: smooth. Metanotal trough reflectivity: glossy. Metanotal trough pilosity: NOT CODED. Metanotal trough pilosity density: NOT CODED. Ventral impressed area of metanotum sculpture: smooth without striae. Metascutellum height / ventral impressed area of metanotum height: 1. Central propodeal area sculpture: rugose. Central propodeal area reflectivity: NOT CODED. Lateral propodeal carina shape: broad, high, slightly curved laterally in mid-height. Lateral propodeal area sculpture: rugose. Lateral propodeal area pilosity: present. Lateral propodeal area pilosity color: NOT CODED. Lateral propodeal area pilosity density: rare. Nucha sculpture: with delicate longitudinal rugae dorsally and laterally. Radial cell length / radial cell width: 4.5. Rs+M vs basalis: reaches basalis in lower half of its height. Rs+M color: dark brown. Areolet: large, triangular, conspicuous. Marginal cilia: long. Rs distal end vs wing margin: adjacent to wing margin. R1 distal end vs wing margin: adjacent to wing margin. Basal lobe on metatarsal claw: present. Basal lobe on tarsal claw shape: blunt.
</paragraph>
<paragraph pageId="58" pageNumber="135">Metasoma length / head+mesosoma length: &lt;1. Metasoma lateral height / metasoma lateral length: &lt;1. 2nd metasomal tergite length dorsal view / length of metasoma dorsal view: 1/3. Second metasomal tergite pilosity: NOT CODED. Second metasomal tergite sculpture: smooth. Second metasomal tergite reflectivity: glossy. Second metasomal tergite pilosity density: NOT CODED. Metasomal tergites 3-6 sculpture: smooth, micropunctate. Metasomal tergites 3-6 reflectivity: glossy. Metasomal tergites 3-6 pilosity: absent. Prominent part of ventral spine of hypopygium length ventral view / Prominent part of ventral spine of hypopygium width ventral view: 1.30. Hypopygial setae apical end: extending beyond posterior end of ventral spine of hypopygium.</paragraph>
<paragraph pageId="58" pageNumber="135">Body length: 2.60-2.80 mm (n=5).</paragraph>
<paragraph pageId="58" pageNumber="135">Male: Similar to female. Color: Head and mesosoma black, metasoma dark brown, antenna light brown to dark brown. Eye size vs female eye size: eye larger in male. Anterior tentorial pit size: large, distinct. Diameter of torulus / intertorular distance: NOT CODED. Diameter of lateral ocellus vs diameter of female lateral ocellus: NOT CODED. Flagellomeres count: 13. Antenna length / body length:&gt;1. F1 shape: F1 curved and broadened apically. F1 length / F2 length: 0.94. Flagellomeres relative length: F2&gt;F3&gt;F4&gt;F5&gt;F6&gt;F9&gt;F10&gt;F11&gt;F12. F13 length / F12 length: 1.2. Placoid sensilla present on: F2-F13. Body length: 2.6-2.8 mm (n=4).</paragraph>
<paragraph pageId="58" pageNumber="135">
Gall (Figs
<figureCitation captionStart="Figures 130135" captionStartId="F24" captionText="Figures 130 - 135. Dryocosmus moriius sp. n. 130 mesosoma, female, lateral view 131 metascutellum and propodeum, female, posterodorsal view 132 - 133 metasoma, lateral view: 132 female 133 male 134 - 135 galls (photos by C. - T. Tang) (vsh = ventral spine of hypopygium)." figureDoi="10.3897/jhr.53.9890.figures130-135" httpUri="https://binary.pensoft.net/fig/115563" pageId="58" pageNumber="135">134-135</figureCitation>
): The gall closely resembles the sexual galls of
<taxonomicName authorityName="Ide, Wachi &amp; Abe" authorityYear="2013" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus sakureiensis" order="Hymenoptera" pageId="58" pageNumber="135" phylum="Arthropoda" rank="species" species="sakureiensis">
<emphasis italics="true" pageId="58" pageNumber="135">Dryocosmus sakureiensis</emphasis>
</taxonomicName>
. The gall is unilocular, ovate, 2.8-3.4 mm long and 1.8-2.1 mm wide (n=5). Galls are at the base of young shoots, usually by bud scales. Galls are green and yellowish white in the central area.
</paragraph>
<caption ID-Zenodo-Dep="984497" doi="10.3897/jhr.53.9890.figures122-129" httpUri="https://binary.pensoft.net/fig/115562" pageId="58" pageNumber="135" start="Figures 122129" startId="F23">
<paragraph pageId="58" pageNumber="135">
<emphasis bold="true" pageId="58" pageNumber="135">Figures 122-129.</emphasis>
<taxonomicName authorityName="Tang &amp; Melika" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus moriius" order="Hymenoptera" pageId="58" pageNumber="135" phylum="Arthropoda" rank="species" species="moriius">
<emphasis italics="true" pageId="58" pageNumber="135">Dryocosmus moriius</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="58" pageNumber="135">122-124</emphasis>
head, female:
<emphasis bold="true" pageId="58" pageNumber="135">122</emphasis>
frontal view
<emphasis bold="true" pageId="58" pageNumber="135">123</emphasis>
dorsal view
<emphasis bold="true" pageId="58" pageNumber="135">124</emphasis>
posterior view
<emphasis bold="true" pageId="58" pageNumber="135">125</emphasis>
head, frontal view, male
<emphasis bold="true" pageId="58" pageNumber="135">126-127</emphasis>
antenna:
<emphasis bold="true" pageId="58" pageNumber="135">126</emphasis>
female
<emphasis bold="true" pageId="58" pageNumber="135">127</emphasis>
male
<emphasis bold="true" pageId="58" pageNumber="135">128-129</emphasis>
female:
<emphasis bold="true" pageId="58" pageNumber="135">128</emphasis>
mesosoma, dorsal view
<emphasis bold="true" pageId="58" pageNumber="135">129</emphasis>
fore wing, part, female (rad=radial cell).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="984499" doi="10.3897/jhr.53.9890.figures130-135" httpUri="https://binary.pensoft.net/fig/115563" pageId="58" pageNumber="135" start="Figures 130135" startId="F24">
<paragraph pageId="58" pageNumber="135">
<emphasis bold="true" pageId="58" pageNumber="135">Figures 130-135.</emphasis>
<taxonomicName authorityName="Tang &amp; Melika" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus moriius" order="Hymenoptera" pageId="58" pageNumber="135" phylum="Arthropoda" rank="species" species="moriius">
<emphasis italics="true" pageId="58" pageNumber="135">Dryocosmus moriius</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="58" pageNumber="135">130</emphasis>
mesosoma, female, lateral view
<emphasis bold="true" pageId="58" pageNumber="135">131</emphasis>
metascutellum and propodeum, female, posterodorsal view
<emphasis bold="true" pageId="58" pageNumber="135">132-133</emphasis>
metasoma, lateral view:
<emphasis bold="true" pageId="58" pageNumber="135">132</emphasis>
female
<emphasis bold="true" pageId="58" pageNumber="135">133</emphasis>
male
<emphasis bold="true" pageId="58" pageNumber="135">134-135</emphasis>
galls (photos by C.-T. Tang) (vsh=ventral spine of hypopygium).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="58" pageNumber="135" type="biology_ecology">
<paragraph pageId="58" pageNumber="135">Biology.</paragraph>
<paragraph pageId="58" pageNumber="135">
Only the sexual generation is known. Galls are induced on young shoots on
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="58" pageNumber="135" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="58" pageNumber="135">Quercus morii</emphasis>
</taxonomicName>
. Galls were collected in April and adults emerged from galls under laboratory conditions immediately after field collection.
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="135" type="distribution">
<paragraph pageId="58" pageNumber="135">Distribution.</paragraph>
<paragraph pageId="58" pageNumber="135">
Taiwan: Renai Township, Nantou County.
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Hexapoda" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quercus morii" order="Hymenoptera" pageId="58" pageNumber="135" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="58" pageNumber="135">Quercus morii</emphasis>
</taxonomicName>
is endemic to Taiwan, suggesting that
<taxonomicName class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus morii" order="Hymenoptera" pageId="58" pageNumber="135" phylum="Arthropoda" rank="species" species="morii">
<emphasis italics="true" pageId="58" pageNumber="135">Dryocosmus morii</emphasis>
</taxonomicName>
is also endemic to Taiwan.
</paragraph>
</subSubSection>
</treatment>
</document>