127 lines
19 KiB
XML
127 lines
19 KiB
XML
<document id="0892C4B34100CB0E8FD74CDA16842648" ID-DOI="10.3390/insects15050318" ID-Zenodo-Dep="11120833" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1714997800831" checkinUser="tatiana" docAuthor="Brazidec, Manuel, Lohrmann, Volker & Perrichot, Vincent" docDate="2024" docId="195487B5937D463DFE3BF928D59DFD0D" docLanguage="en" docName="Insects.15.1-12.pdf" docOrigin="Insects (Clifton, N. J.) 15 (5)" docSource="http://dx.doi.org/10.3390/insects15050318" docTitle="Hukawngepyris Brazidec & Lohrmann & Perrichot 2024, gen. nov." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="E56DFFCD937F463EFF87FFDFD045FFA2" masterDocTitle="The First Cretaceous Epyrine Wasp (Hymenoptera: Bethylidae): A New Genus and Species from Early Cenomanian Kachin Amber" masterLastPageNumber="200" masterPageNumber="191" pageNumber="3" updateTime="1714998355944" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D4DC81C27E0FE4D8992FC38EBB9402AD">The First Cretaceous Epyrine Wasp (Hymenoptera: Bethylidae): A New Genus and Species from Early Cenomanian Kachin Amber</mods:title>
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<mods:name id="369A0B756B04B4FBD01D67E14010F138" type="personal">
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<mods:roleTerm id="F7958A9EB5B8D068BC1D3D1441660A79">Author</mods:roleTerm>
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<mods:namePart id="73C04ED4160DAF5339EF39350478FC98">Brazidec, Manuel</mods:namePart>
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<mods:affiliation id="098A45F45D2B98D755CA13F1B9B09B0C">CNRS, Géosciences Rennes, University Rennes, UMR 6118, 35000 Rennes, France & Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="C64CDA5CF8EAB9BA8C04EF1AE2151B66" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="818DF05580CB217BA762A94DE523EBF3">Author</mods:roleTerm>
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<mods:namePart id="75E0A6E5B55FF177B377D26777490807">Lohrmann, Volker</mods:namePart>
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<mods:affiliation id="AEDF2A9323A861A2B9F3211BAA38D89D">Übersee-Museum Bremen, Bahnhofsplatz 13, 28195 Bremen, Germany</mods:affiliation>
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<mods:nameIdentifier id="F5F48C95257FCCDDF47B2706AB03438E" type="email">v.lohrmann@uebersee-museum.de</mods:nameIdentifier>
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<mods:roleTerm id="D35A65CDEB7BD258B7F42D8526D6DEE1">Author</mods:roleTerm>
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<mods:namePart id="3076548D46C311B36DA5E0E0980A0F98">Perrichot, Vincent</mods:namePart>
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<mods:nameIdentifier id="41A7A386F17894381A0D691B6D12E57E" type="LSID">vincent.perrichot@univ-rennes.fr</mods:nameIdentifier>
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<mods:nameIdentifier id="5162BFC268FBC0BD748C082A52D59F1C" type="ORCID">0000-0002-7973-0430</mods:nameIdentifier>
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<mods:affiliation id="C8F39C961F58C224CC7828A3CC5CCEF6">CNRS, Géosciences Rennes, University Rennes, UMR 6118, 35000 Rennes, France</mods:affiliation>
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<mods:typeOfResource id="71223FAFC8D818F530AF81DE736841F6">text</mods:typeOfResource>
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<mods:title id="3AEAA7FF5C07619F509408FC28ED51D2">Insects</mods:title>
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<mods:part id="7C385AFA776718425A4EA29A38FF093B">
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<mods:date id="FE57D5EE37A90830131494B2F22A2599">2024</mods:date>
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<mods:detail id="5E80B344C1C213D10DBAA8E856E47FF8" type="series">
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<mods:title id="E91751631710E3857BCE701813A9B0F8">Clifton, N. J.</mods:title>
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<mods:detail id="76E2A824540654DB186115621B89C990" type="pubDate">
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<mods:number id="3142CE38F9E5FB2AE0FD89F49009DE62">2024-04-30</mods:number>
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<mods:number id="4503927B1266781578DD99B170F9E5FE">15</mods:number>
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<mods:detail id="4D858DF0152C6943DBEDC9A8C824DA79" type="issue">
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<mods:number id="45A8EAA0D2A01D62C4DEDDE77251BEE9">5</mods:number>
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<mods:location id="DB1FB7BD0DB8C863F901F93ED6D426A4">
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<mods:url id="A9F6735688360CC516DB4E15D82C1DA0">http://dx.doi.org/10.3390/insects15050318</mods:url>
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<mods:classification id="864FCBE950C761049293C865EC256462">journal article</mods:classification>
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<mods:identifier id="7DB1D08E4BA19EB5D61581EE88273E66" type="DOI">10.3390/insects15050318</mods:identifier>
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<mods:identifier id="4F8AF912BB7F8510408BA13A58447F3B" type="Zenodo-Dep">11120833</mods:identifier>
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<treatment id="195487B5937D463DFE3BF928D59DFD0D" ID-DOI="http://doi.org/10.5281/zenodo.11120811" ID-Zenodo-Dep="11120811" LSID="urn:lsid:plazi:treatment:195487B5937D463DFE3BF928D59DFD0D" httpUri="http://treatment.plazi.org/id/195487B5937D463DFE3BF928D59DFD0D" lastPageId="3" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="D9E76528937D463CFE3BF928D36AF8AC" box="[444,815,1783,1807]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="914236A3937D463CFE3BF928D36AF8AC" blockId="2.[443,1497,1641,2075]" box="[444,815,1783,1807]" pageId="2" pageNumber="3">
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<emphasis id="A389EAB1937D463CFE3BF928D249F8AD" bold="true" box="[444,524,1783,1807]" pageId="2" pageNumber="3">Genus</emphasis>
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<taxonomicName id="56FD4D20937D463CFD95F928D2F8F8AD" authority="Brazidec & Lohrmann & Perrichot, 2024" authorityName="Brazidec & Lohrmann & Perrichot" authorityYear="2024" box="[530,701,1783,1807]" class="Insecta" family="Bethylidae" genus="Hukawngepyris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="A389EAB1937D463CFD95F928D2F8F8AD" box="[530,701,1783,1807]" italics="true" pageId="2" pageNumber="3">Hukawngepyris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8BA57CA937D463CFD43F928D36AF8AC" box="[708,815,1783,1807]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9E76528937D463CFE3BF8C7D59CF892" box="[444,1497,1816,1840]" pageId="2" pageNumber="3" type="description">
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<paragraph id="914236A3937D463CFE3BF8C7D59CF892" blockId="2.[443,1497,1641,2075]" box="[444,1497,1816,1840]" pageId="2" pageNumber="3">https://zoobank.org/5E418042-63B6-4688-91C9-4E2D6D9D0465 (accessed on 25 April 2024).</paragraph>
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</subSubSection>
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<subSubSection id="D9E76528937D463CFE3BF8E5D27FF837" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="914236A3937D463CFE3BF8E5D27FF837" blockId="2.[443,1497,1641,2075]" pageId="2" pageNumber="3">
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<emphasis id="A389EAB1937D463CFE3BF8E5D206F8F0" bold="true" box="[444,579,1850,1874]" pageId="2" pageNumber="3">Etymology.</emphasis>
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The name is a combination of Hukawng, as a reference to the Hukawng Valley where Kachin amber is widely found, and
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<taxonomicName id="56FD4D20937D463CFC37F883D3B2F8D1" authorityName="Westwood" authorityYear="1832" box="[944,1015,1884,1907]" class="Insecta" family="Zygaenidae" genus="Epyris" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="A389EAB1937D463CFC37F883D3B2F8D1" box="[944,1015,1884,1907]" italics="true" pageId="2" pageNumber="3">Epyris</emphasis>
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</taxonomicName>
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, the type genus of the
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<taxonomicName id="56FD4D20937D463CFA85F884D535F8D1" authorityName="Kieffer" authorityYear="1914" box="[1282,1392,1883,1907]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Epyrinae">Epyrinae</taxonomicName>
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. Gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="D9E76528937D463CFE3CF841D436F815" box="[443,1139,1950,1975]" pageId="2" pageNumber="3" type="type_taxon">
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<paragraph id="914236A3937D463CFE3CF841D436F815" blockId="2.[443,1497,1641,2075]" box="[443,1139,1950,1975]" pageId="2" pageNumber="3">
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<emphasis id="A389EAB1937D463CFE3CF841D21EF814" bold="true" box="[443,603,1950,1974]" pageId="2" pageNumber="3">Type species.</emphasis>
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<taxonomicName id="56FD4D20937D463CFDE4F841D327F814" authority="Brazidec & Lohrmann & Perrichot, 2024" authorityName="Brazidec & Lohrmann & Perrichot" authorityYear="2024" box="[611,866,1950,1974]" class="Insecta" family="Bethylidae" genus="Hukawngepyris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
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<emphasis id="A389EAB1937D463CFDE4F841D327F814" box="[611,866,1950,1974]" italics="true" pageId="2" pageNumber="3">Hukawngepyris setosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8BA57CA937D463CFCEFF840D380F814" box="[872,965,1951,1974]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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, by monotypy.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D9E76528937D463DFE3BF860D59DFD0D" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
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||
<paragraph id="914236A3937D463DFE3BF860D59DFD0D" blockId="2.[443,1497,1641,2075]" lastBlockId="3.[443,1497,261,921]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<emphasis id="A389EAB1937D463CFE3BF860D278F875" bold="true" box="[444,573,1983,2007]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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Females of the new genus stand out of all known bethylid genera by the following combination of characters: Vertex, gena, pronotum, mesopleuron, and mesoscutellum foveolate; head and anterior mesosoma with scattered long erect setae (
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<figureCitation id="09C62A26937D463CFB78F7DCD52EF7B9" box="[1279,1387,2051,2075]" captionStart="Figure 2" captionStartId="3.[444,514,1899,1921]" captionTargetBox="[130,1456,950,1864]" captionTargetId="figure-282@3.[130,1456,950,1864]" captionTargetPageId="3" captionText="Figure 2. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, habitus in left lateroventral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11120837" httpUri="https://zenodo.org/record/11120837/files/figure.png" pageId="2" pageNumber="3">Figures 2</figureCitation>
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and
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<figureCitation id="09C62A26937D463CFA22F7DCD58FF7B9" box="[1445,1482,2051,2075]" captionStart="Figure 3" captionStartId="4.[444,514,1820,1842]" captionTargetBox="[108,1476,413,1780]" captionTargetId="figure-128@4.[108,1476,413,1781]" captionTargetPageId="4" captionText="Figure 3. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) head in left lateral view; (B) head and partial mesosoma in dorsal view (black arrow: notaulus; white arrow: parapsidal signum); (C) mesosoma in left lateroventral view (black arrow: propleural epicoxal sulcus; white arrow: protibial spur). Scale bars: A = 0.25 mm, B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120839" httpUri="https://zenodo.org/record/11120839/files/figure.png" pageId="2" pageNumber="3">3A</figureCitation>
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); compound eyes without setation, 0.6 × HL; malar space convergent, longer than VOL; median clypeal lobe triangular, projecting; mandible elongate, quadridentate; antenna filiform (
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<figureCitation id="09C62A26937C463DFDA6FE97D2D7FEC2" box="[545,658,328,352]" captionStart="Figure 3" captionStartId="4.[444,514,1820,1842]" captionTargetBox="[108,1476,413,1780]" captionTargetId="figure-128@4.[108,1476,413,1781]" captionTargetPageId="4" captionText="Figure 3. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) head in left lateral view; (B) head and partial mesosoma in dorsal view (black arrow: notaulus; white arrow: parapsidal signum); (C) mesosoma in left lateroventral view (black arrow: propleural epicoxal sulcus; white arrow: protibial spur). Scale bars: A = 0.25 mm, B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120839" httpUri="https://zenodo.org/record/11120839/files/figure.png" pageId="3" pageNumber="4">Figure 3B</figureCitation>
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); scape and pedicel bearing long thick setae; pedicel shorter than each of the flagellomeres; anterior ocellus crossing supra-ocular line; distance between anterior and posterior ocelli shorter than width of ocellar triangle; occipital carina complete. Pronotum longer than wide, with anterior angle rounded; without longitudinal sulcus (
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<figureCitation id="09C62A26937C463DFAD4FE72D58CFE67" box="[1363,1481,429,453]" captionStart="Figure 3" captionStartId="4.[444,514,1820,1842]" captionTargetBox="[108,1476,413,1780]" captionTargetId="figure-128@4.[108,1476,413,1781]" captionTargetPageId="4" captionText="Figure 3. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) head in left lateral view; (B) head and partial mesosoma in dorsal view (black arrow: notaulus; white arrow: parapsidal signum); (C) mesosoma in left lateroventral view (black arrow: propleural epicoxal sulcus; white arrow: protibial spur). Scale bars: A = 0.25 mm, B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120839" httpUri="https://zenodo.org/record/11120839/files/figure.png" pageId="3" pageNumber="4">Figure 3B</figureCitation>
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); notaulus and parapsidal signum present; sulcus of the mesoscuto-mesoscutellar suture present; mesoscutellum not contacting metapectal–propodeal complex (
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<figureCitation id="09C62A26937C463DFA8DFE2FD5C5FDAA" box="[1290,1408,496,520]" captionStart="Figure 3" captionStartId="4.[444,514,1820,1842]" captionTargetBox="[108,1476,413,1780]" captionTargetId="figure-128@4.[108,1476,413,1781]" captionTargetPageId="4" captionText="Figure 3. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) head in left lateral view; (B) head and partial mesosoma in dorsal view (black arrow: notaulus; white arrow: parapsidal signum); (C) mesosoma in left lateroventral view (black arrow: propleural epicoxal sulcus; white arrow: protibial spur). Scale bars: A = 0.25 mm, B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120839" httpUri="https://zenodo.org/record/11120839/files/figure.png" pageId="3" pageNumber="4">Figure 3C</figureCitation>
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); metapostnotal median carina complete, straight, extending on propodeal declivity (
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<figureCitation id="09C62A26937C463DFAD4FDCED589FD8B" box="[1363,1484,529,553]" captionStart="Figure 4" captionStartId="5.[444,514,1715,1737]" captionTargetBox="[109,1477,260,1674]" captionTargetId="figure-150@5.[109,1477,260,1679]" captionTargetPageId="5" captionText="Figure 4. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) mesosoma in dorsolateral view; (B) metasoma in left posterolateral view; (C) drawing of fore and hind wings (dashed lines indicate flexion lines; pt = pterostigma). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120841" httpUri="https://zenodo.org/record/11120841/files/figure.png" pageId="3" pageNumber="4">Figure 4A</figureCitation>
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). Forewing with C, Sc + R, M + Cu, A, Rs&M, cu-a, and 2r-rs&Rs tubular; 2r-rs&Rs tubular and curved, distally reaching anterior wing margin. Hind wing with three straight proximal and three hook-like distal hamuli (
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<figureCitation id="09C62A26937C463DFC10FDA9D442FD2C" box="[919,1031,630,654]" captionStart="Figure 2" captionStartId="3.[444,514,1899,1921]" captionTargetBox="[130,1456,950,1864]" captionTargetId="figure-282@3.[130,1456,950,1864]" captionTargetPageId="3" captionText="Figure 2. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, habitus in left lateroventral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11120837" httpUri="https://zenodo.org/record/11120837/files/figure.png" pageId="3" pageNumber="4">Figures 2</figureCitation>
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and
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<figureCitation id="09C62A26937C463DFBC5FDA9D423FD2C" box="[1090,1126,630,654]" captionStart="Figure 4" captionStartId="5.[444,514,1715,1737]" captionTargetBox="[109,1477,260,1674]" captionTargetId="figure-150@5.[109,1477,260,1679]" captionTargetPageId="5" captionText="Figure 4. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) mesosoma in dorsolateral view; (B) metasoma in left posterolateral view; (C) drawing of fore and hind wings (dashed lines indicate flexion lines; pt = pterostigma). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120841" httpUri="https://zenodo.org/record/11120841/files/figure.png" pageId="3" pageNumber="4">4C</figureCitation>
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). Protibial spur long and stout (
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<figureCitation id="09C62A26937C463DFE44FD48D27DFD0D" box="[451,568,663,687]" captionStart="Figure 3" captionStartId="4.[444,514,1820,1842]" captionTargetBox="[108,1476,413,1780]" captionTargetId="figure-128@4.[108,1476,413,1781]" captionTargetPageId="4" captionText="Figure 3. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) head in left lateral view; (B) head and partial mesosoma in dorsal view (black arrow: notaulus; white arrow: parapsidal signum); (C) mesosoma in left lateroventral view (black arrow: propleural epicoxal sulcus; white arrow: protibial spur). Scale bars: A = 0.25 mm, B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120839" httpUri="https://zenodo.org/record/11120839/files/figure.png" pageId="3" pageNumber="4">Figure 3C</figureCitation>
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); tarsal claw bifid (
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<figureCitation id="09C62A26937C463DFC93FD48D3CDFD0D" box="[788,904,663,687]" captionStart="Figure 4" captionStartId="5.[444,514,1715,1737]" captionTargetBox="[109,1477,260,1674]" captionTargetId="figure-150@5.[109,1477,260,1679]" captionTargetPageId="5" captionText="Figure 4. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) mesosoma in dorsolateral view; (B) metasoma in left posterolateral view; (C) drawing of fore and hind wings (dashed lines indicate flexion lines; pt = pterostigma). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120841" httpUri="https://zenodo.org/record/11120841/files/figure.png" pageId="3" pageNumber="4">Figure 4B</figureCitation>
|
||
). Metasomal tergum 2 entirely smooth (
|
||
<figureCitation id="09C62A26937C463DFADFFD48D58EFD0D" box="[1368,1483,663,687]" captionStart="Figure 4" captionStartId="5.[444,514,1715,1737]" captionTargetBox="[109,1477,260,1674]" captionTargetId="figure-150@5.[109,1477,260,1679]" captionTargetPageId="5" captionText="Figure 4. Hukawngepyris setosus gen. et sp. nov., female holotype, MB.I.8638, (A) mesosoma in dorsolateral view; (B) metasoma in left posterolateral view; (C) drawing of fore and hind wings (dashed lines indicate flexion lines; pt = pterostigma). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11120841" httpUri="https://zenodo.org/record/11120841/files/figure.png" pageId="3" pageNumber="4">Figure 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |