treatments-xml/data/0D/3E/D1/0D3ED10470EAFD6BEAB88E42C0226E1B.xml
2024-06-21 12:29:19 +02:00

235 lines
27 KiB
XML

<document ID-CLB-Dataset="297122" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="27821ddc-082b-40c1-825b-c4098cb9da9e" ID-ISBN="0-8018-8221-4" checkinTime="1670905101524" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="0D3ED10470EAFD6BEAB88E42C0226E1B" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Calomyscidae.926-930.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Calomyscus Thomas 1905" docType="treatment" docVersion="3" id="DC7AB3713185F1063B4DEB53CCADC200" lastPageNumber="927" masterDocId="DC7AB3713185F1063B4DEB53CCADC200" masterDocTitle="Order Rodentia - Family Calomyscidae" masterLastPageNumber="930" masterPageNumber="926" pageNumber="927" updateTime="1716775671765" updateUser="ExternalLinkService">
<mods:mods id="2A7D938051AF54C4140E0B7FCECF151E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="30CA73C2469BDDEB0DF551FAAD02EFE4">
<mods:title id="10A084D72D9036E9C0F10133E1D1A937">Order Rodentia - Family Calomyscidae</mods:title>
</mods:titleInfo>
<mods:name id="C7CCE6FA68372FD4A4FEA0F36AF5C0BE" type="personal">
<mods:role id="5D0A29179CF981A12F00DC39049037AF">
<mods:roleTerm id="D716CE20B564EA877985F9010ED56DE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3DB8429CB625BB4E0B525C856E5063B3">Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name id="1C3356978F106B6AFFC71051B30323BE" type="personal">
<mods:role id="BFC6F25596F5D6168C4DC29E7B8FA7DF">
<mods:roleTerm id="581F6666643A16F33D7DCAB99B2AE4EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32F26A02187298BD4DA8507B0217C91D">Reeder, DeeAnn</mods:namePart>
</mods:name>
<mods:typeOfResource id="871E94C90269CD2825D9D40DC9DAE0F1">text</mods:typeOfResource>
<mods:relatedItem id="A706E350AF6A310371A2F9840F68074F" type="host">
<mods:originInfo id="FD09A01B21ABB4192D2EAA0D76FCFD03">
<mods:dateIssued id="E844DCDEE71F5887BC6E52B34E6E04B7">2005</mods:dateIssued>
<mods:publisher id="6B0A8233392EE16A696146FB4413A853">The Johns Hopkins University Press</mods:publisher>
<mods:place id="9FBADE222304B52C9ECF13DD91959D27">
<mods:placeTerm id="13D58418E970ABD2D3F5DDD84B55546E">Baltimore</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo id="7C93433794995E0064ECBB223E2925F3">
<mods:title id="B348F21F88F6ADAE82DF5940669D2AF8">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
</mods:titleInfo>
<mods:part id="8B56D18A924EADAFFE532378960BCFA7">
<mods:extent id="DE8C516DA0A9C18FED058DEC4B23D9A2" unit="page">
<mods:start id="201E166DBC87CB6D14381F5F1CC02284">926</mods:start>
<mods:end id="E1A437DC64B881CA8F6F51D8571B8D67">930</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F8C1D08F9DAA937462E168BE94CDCEBB">book chapter</mods:classification>
<mods:identifier id="5F6B2743771614148834E90F2E16760B" type="ISBN">0-8018-8221-4</mods:identifier>
<mods:identifier id="7BECE558B248C7F68B4776DA526B4972" type="DOI">10.5281/zenodo.7316535</mods:identifier>
</mods:mods>
<treatment id="0D3ED10470EAFD6BEAB88E42C0226E1B" ID-GBIF-Taxon="231547257" LSID="urn:lsid:plazi:treatment:0D3ED10470EAFD6BEAB88E42C0226E1B" httpUri="http://treatment.plazi.org/id/0D3ED10470EAFD6BEAB88E42C0226E1B" lastPageNumber="927" pageNumber="927" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13000055">
<subSubSection id="F815E3FC5F716B4FBDA4DCF51F360CE0" pageNumber="927" type="nomenclature">
<paragraph id="03CE2885270A94ABD73CAFE05A586B9F" pageNumber="927">
<taxonomicName id="F485B4B88BA13EB039BE18C48745D2CB" ID-CoL="62KQC" authority="Thomas 1905" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="genus">
<emphasis id="34A8150CBE24EEFE9C87087F803F680F" italics="true" pageNumber="927">Calomyscus</emphasis>
Thomas 1905
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="4FF0B28C4F9D909B5E15259802007592" pageNumber="927" type="reference_group">
<paragraph id="F1141A1202904E1C18B3277DF6DE4857" pageNumber="927">
<treatmentCitation id="F3B856C05F9992111BE9DBFAB8B4E62F" author="Thomas" authority="Thomas 1905" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" issue="24" journal="Abstr. Proc. Zool. Soc. Lond." kingdom="Animalia" order="Rodentia" page="23" pageNumber="927" phylum="Chordata" rank="genus" title="Calomyscus" volume="1905" year="1905">
<taxonomicName id="ADAA945807071DFB3EF9ED4EB5B86961" ID-CoL="62KQC" authority="Thomas 1905" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="genus">
<emphasis id="DE903941745A39BA1A5DFA9F6F743CE0" italics="true" pageNumber="927">Calomyscus</emphasis>
Thomas 1905
</taxonomicName>
,
<bibCitation id="60CA62CFBD469483F58E0D4C98F4C511" author="Thomas" issue="24" journal="Abstr. Proc. Zool. Soc. Lond." pageNumber="927" pagination="23" title="Calomyscus" volume="1905" year="1905">Abstr. Proc. Zool. Soc. Lond., 1905 (24): 23</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2F7A8DFD7C448E61815141D417094CBC" pageNumber="927" type="type_taxon">
<paragraph id="A8309700D1A84BFF82F3F77666F7CE77" pageNumber="927">
<emphasis id="850E62FAB18644084466A3266A0B5FC8" inLineHeading="true" pageNumber="927">Type Species:</emphasis>
<taxonomicName id="44B032BA397686183CF4B484455CB3B9" authority="Thomas 1905" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="bailwardi">
<emphasis id="6CB0F31B44463A4FC6F23CEB70F0A031" italics="true" pageNumber="927">Calomyscus bailwardi</emphasis>
Thomas 1905
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="83F693DB3BBAB698B9B43C5D48749AAA" pageNumber="927" type="child_taxa">
<paragraph id="48066BD22EB7A8FB966983FA6A3D7CD0" pageNumber="927">
<emphasis id="F76F3F3F5B886F68E8D57F2FAA101F70" inLineHeading="true" pageNumber="927">Species and subspecies:</emphasis>
8 species:
</paragraph>
<paragraph id="C1B6CACB740EA77DE458909A67E58610" pageNumber="927">
Species
<taxonomicName id="13E7CD80EB320CBC8C11080A871EBAFF" authority="Thomas 1905" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="bailwardi">
<emphasis id="2A6AB8874B5DFCA1A8E0C3A532BF801B" italics="true" pageNumber="927">Calomyscus bailwardi</emphasis>
Thomas 1905
</taxonomicName>
</paragraph>
<paragraph id="477B6E37C71B49E2946BA09DE5B87CD2" pageNumber="927">
Species
<taxonomicName id="0CC1611E182CF49A86159EECD55A7F0F" authority="Thomas 1920" authorityName="Thomas" authorityYear="1920" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="baluchi">
<emphasis id="B7A51BEF64A32042BB37080C5BEDB958" italics="true" pageNumber="927">Calomyscus baluchi</emphasis>
Thomas 1920
</taxonomicName>
</paragraph>
<paragraph id="D956AAE74BDFB1194980BB75EDC86485" pageNumber="927">
Species
<taxonomicName id="8639B11129B2A2130BFE312F39530D1F" authority="Goodwin 1938" authorityName="Goodwin" authorityYear="1938" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="elburzensis">
<emphasis id="AB9E7FA222B78CA0582A31D7EB78EC63" italics="true" pageNumber="927">Calomyscus elburzensis</emphasis>
<bibRefCitation id="CB1C6884F98E567143010EFA1826AAF4" author="Goodwin, G. G." refId="ref125418" refString="Goodwin, G. G. 1938. Five new rodents from the eastern Elburz Mountains and a new race of hare from Teheran. American Museum Novitates, 1950: 1 - 5." year="1938">Goodwin 1938</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="AA2C6403D8B97426564C2568ABD7CEE3" pageNumber="927">
Species
<taxonomicName id="20785E0587381081183548630E5C955C" authority="Schlitter and Setzer 1973" authorityName="Schlitter and Setzer" authorityYear="1973" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="grandis">
<emphasis id="52795E8A0C67711AC6DC99DEA3C58865" italics="true" pageNumber="927">Calomyscus grandis</emphasis>
<bibRefCitation id="5BC2D0459C9C0ADFA6A7A82653BAE83A" author="Schlitter, D. A. &amp; H. W. Setzer" refId="ref319843" refString="Schlitter, D. A., and H. W. Setzer. 1973. New rodents (Mammalia: Cricetidae, Muridae) from Iran and Pakistan. Proceedings of the Biological Society of Washington, 86: 163 - 174." year="1973">Schlitter and Setzer 1973</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="6A2286F0666D6F40F882CC66D1D98CFD" pageNumber="927">
Species
<taxonomicName id="5D0E8BD00AEBB5355F9173B517698224" authority="Thomas 1920" authorityName="Thomas" authorityYear="1920" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="hotsoni">
<emphasis id="61F3DEA1A0D05319A6B277E098591F62" italics="true" pageNumber="927">Calomyscus hotsoni</emphasis>
Thomas 1920
</taxonomicName>
</paragraph>
<paragraph id="945D0F4E524FFB4810120D4C61ACCE0F" pageNumber="927">
Species
<taxonomicName id="63582C6F18E5224B1FEFDD5592108CBD" authority="Kashkarov 1925" authorityName="Kashkarov" authorityYear="1925" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="mystax">
<emphasis id="0A6F4DD79AD50A9BB8235876CF8F39A1" italics="true" pageNumber="927">Calomyscus mystax</emphasis>
Kashkarov 1925
</taxonomicName>
</paragraph>
<paragraph id="610A229A2B28A07565351C51E7252516" pageNumber="927">
Species
<taxonomicName id="08DDC077522D08D79F396E31703EAC87" authority="Peshev 1991" authorityName="Peshev" authorityYear="1991" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="tsolovi">
<emphasis id="A18B3C97001FB310B9D7A7723FBC862E" italics="true" pageNumber="927">Calomyscus tsolovi</emphasis>
<bibRefCitation id="262262D60D7A08A1BA834FDAE775499A" author="Peshev, D." refId="ref281321" refString="Peshev, D. 1991. On the systematic position of the mouse-like hamster Calomyscus bailwardi Thomas, 1905 (Cricetidae, Rodentia) from the Near East and Middle Asia. Mammalia, 55: 107 - 112." year="1991">Peshev 1991</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="99033E51E6732EBC88EA2A56657BC98A" pageNumber="927">
Species
<taxonomicName id="9DB0408C1EDAEF298A1F845208178728" authority="Vorontsov and Kartavseva 1979" authorityName="Vorontsov and Kartavseva" authorityYear="1979" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" pageNumber="927" phylum="Chordata" rank="species" species="urartensis">
<emphasis id="0114803DB2E916769B392FD15C59904B" italics="true" pageNumber="927">Calomyscus urartensis</emphasis>
Vorontsov and Kartavseva 1979
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="B88C7EA2A5F3439E4FC4E008E96A18D5" pageNumber="927" type="discussion">
<paragraph id="7A820BA2158912C9A83C9F1B097FED46" pageNumber="927">
<emphasis id="7562C30998010604C9E541AB260DA2FB" inLineHeading="true" pageNumber="927">Discussion:</emphasis>
<paragraph id="56F586EB97D89A6FB3393EF7F7A4C765" pageNumber="927">
Although several taxa were originally described as species,
<taxonomicName id="094F1EC09514BF2FE26A35583286E355" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="175C1F87B80DC52246582E784C076F6C" italics="true" pageNumber="927">Calomyscus</emphasis>
</taxonomicName>
was long considered monotypic (
<bibRefCitation id="C125225CB340732A5C479E419EFAD267" author="Ellerman, J. R." refId="ref92857" refString="Ellerman, J. R. 1941. The families and genera of living rodents. Vol. II. Family Muridae. British Museum (Natural History), London, 690 pp." year="1941">Ellerman, 1941</bibRefCitation>
,
<bibRefCitation id="5656EE3EF9DCC9D44A4DA36D2FF93B5B" author="Ellerman, J. R." refId="ref93118" refString="Ellerman, J. R. 1961. Rodentia. Volume 3, in The fauna of India including Pakistan, Burma and Ceylon. Mammalia. Second ed. Manager of Publications, Zoological Survey of India, Calcutta, vol. 3 (in 2 parts), 1: 1 - 482; 2: 483 - 884." year="1961">1961</bibRefCitation>
;
<bibRefCitation id="8C66650761DC3505DE5CAECF2CBFBF9E" author="Ellerman, J. R. &amp; T. C. S. Morrison-Scott" refId="ref93180" refString="Ellerman, J. R., and T. C. S. Morrison-Scott. 1951. Checklist of Palaearctic and Indian mammals 1758 to 1946. Trustees of the British Museum (Natural History), London, 810 pp." year="1951">Ellerman and Morrison-Scott, 1951</bibRefCitation>
;
<bibRefCitation id="F1DDC8FAD0840A1306387D044821775B" author="Peshev, D." refId="ref281321" refString="Peshev, D. 1991. On the systematic position of the mouse-like hamster Calomyscus bailwardi Thomas, 1905 (Cricetidae, Rodentia) from the Near East and Middle Asia. Mammalia, 55: 107 - 112." year="1991">Peshev, 1991</bibRefCitation>
). Character variation in the genus was concisely summarized by
<bibRefCitation id="FC463A47A9D7ABC84172235E2C6E8B6D" author="Corbet, G. B." refId="ref67548" refString="Corbet, G. B. 1978 c. The mammals of the Palaearctic region: A taxonomic review. British Museum (Natural History), London, 314 pp." year="1978">
Corbet (1978
<emphasis id="2E4A1A5ED4BF38CCBB97F5DB06A2BCF4" italics="true" pageNumber="927">c</emphasis>
:89)
</bibRefCitation>
: &quot;There is considerable variation in colour, size and proportions but this seems to form a mosaic pattern and it seems unlikely that any major, discrete regional groups can be recognized.&quot; However, Vorontsov et al. (1979) comprehensively revised the genus and treated most former subspecies of
<taxonomicName id="BDFED16AF7A1E75B39FFD44BF42AF1C9" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bailwardi">
<emphasis id="A92590A4F5E1114789C9681DED685180" italics="true" pageNumber="927">C. bailwardi</emphasis>
</taxonomicName>
as separate species. The morphological and geographic integrity of some has been subsequently tested with additional chromosomal data (
<bibRefCitation id="286A4D091065F2D72823B810DD88A70D" author="Graphodatsky, A. S. &amp; O. V. Sablina &amp; M. N. Meyer &amp; V. G. Malikov &amp; E. A. Isakova &amp; V. A. Trifonov &amp; A. V. Polyakov &amp; T. P. Lushnikova &amp; N. V. Vorobieva &amp; N. A. Serdyukova &amp; P. L. Perelman &amp; P. M. Borodin &amp; P. Benda &amp; D. Frynta &amp; L. Leikepova &amp; P. Munclinger &amp; J. Pialek &amp; J. Sadlova &amp; J. Zima" refId="ref127509" refString="Graphodatsky, A. S., O. V. Sablina, M. N. Meyer, V. G. Malikov, E. A. Isakova, V. A. Trifonov, A. V. Polyakov, T. P. Lushnikova, N. V. Vorobieva, N. A. Serdyukova, P. L. Perelman, P. M. Borodin, P. Benda, D. Frynta, L. Leikepova, P. Munclinger, J. Pialek, J. Sadlova, and J. Zima. 2000. Comparative cytogenetics of hamsters of the genus Calomyscus. Cytogenetics and Cell Genetics, 88: 296 - 304." year="2000">Graphodatsky et al., 2000</bibRefCitation>
;
<bibRefCitation id="086DDC7C810AB6AA028F04F5DC450D98" author="Malikov, V. G. &amp; M. N. Meyer &amp; A. S. Grafodatsky &amp; A. V. Polyakov &amp; O. V. Sablina &amp; A. Sh. Vaziri &amp; F. Nazari &amp; J. Zima" refId="ref223074" refString="Malikov, V. G., M. N. Meyer, A. S. Grafodatsky, A. V. Polyakov, O. V. Sablina, A. Sh. Vaziri, F. Nazari, and J. Zima. 1999. On a taxonomic position of some karyomorphs belonging to genus Calomyscus (Rodentia, Cricetidae). Proceedings of the Zoological Institute RAS, 281: 27 - 32." year="1999">Malikov et al., 1999</bibRefCitation>
;
<bibRefCitation id="14E20008E303E44B45D78A4107BAA3BD" author="Meyer, M. N. &amp; V. G. Malikov" refId="ref238852" refString="Meyer, M. N., and V. G. Malikov. 1995. [On the distribution, taxonomic status and biology of mouselike hamsters of the genus Calomyscus (Rodentia, Cricetidae).] Zoologiskii Zhurnal, 74 (7): 96 - 100 (in Russian with English summary)." year="1995">Meyer and Malikov, 1995</bibRefCitation>
, 2000), mitochondrial cytochrome
<emphasis id="18D3986FA4352F9A1634A5E2525DE5EC" italics="true" pageNumber="927">b</emphasis>
sequences (
<bibRefCitation id="E619321E0EF2CE695E35B5AD87150CB9" author="Morshed, S. &amp; J. L. Patton" refId="ref247929" refString="Morshed, S., and J. L. Patton. 2002. New records of mammals from Iran with systematic comments on hedgehogs (Erinaceidae) and mouse-like hamsters (Calomyscus, Muridae). Zoology in the Middle East, 26: 49 - 58." year="2002">Morshed and Patton, 2002</bibRefCitation>
), comparative postnatal growth and development (
<bibRefCitation id="13520BFDC1BA06A10B6F79D9DD20B575" author="Meyer, M. N. &amp; V. G. Malikov" refId="ref238910" refString="Meyer, M. N., and V. G. Malikov. 1996. [Peculiarities of biology and postnatal otogenesis in Calomyscus (Cricetidae, Calomyscus).] Zoologiskii Zhurnal, 75 (12): 1852 - 1862 (in Russian with English summary)." year="1996">Meyer and Malikov, 1996</bibRefCitation>
), and multivariate analyses of cranial and dental measurements (
<bibRefCitation id="558E74DEEA521D92B87060FEA49F48A6" author="Lebedev, V. S. &amp; I. Ya. Pavlinov &amp; M. N. Meyer &amp; V. G. Malikov" refId="ref208246" refString="Lebedev, V. S., I. Ya. Pavlinov, M. N. Meyer, and V. G. Malikov. 1998. [Craniometric analysis of mouse-like hamsters of the genus Calomyscus (Cricetidae).] Zoologschiskii Zhurnal, 77 (6): 721 - 731 (in Russian with English abstract)." year="1998">Lebedev et al., 1998</bibRefCitation>
). While much research has focused on the geographical distribution of different chromosomal morphologies and their taxonomic significance (summarized in
<bibRefCitation id="B421C61A43EF1C7CA345CD305B129037" author="Graphodatsky, A. S. &amp; O. V. Sablina &amp; M. N. Meyer &amp; V. G. Malikov &amp; E. A. Isakova &amp; V. A. Trifonov &amp; A. V. Polyakov &amp; T. P. Lushnikova &amp; N. V. Vorobieva &amp; N. A. Serdyukova &amp; P. L. Perelman &amp; P. M. Borodin &amp; P. Benda &amp; D. Frynta &amp; L. Leikepova &amp; P. Munclinger &amp; J. Pialek &amp; J. Sadlova &amp; J. Zima" refId="ref127509" refString="Graphodatsky, A. S., O. V. Sablina, M. N. Meyer, V. G. Malikov, E. A. Isakova, V. A. Trifonov, A. V. Polyakov, T. P. Lushnikova, N. V. Vorobieva, N. A. Serdyukova, P. L. Perelman, P. M. Borodin, P. Benda, D. Frynta, L. Leikepova, P. Munclinger, J. Pialek, J. Sadlova, and J. Zima. 2000. Comparative cytogenetics of hamsters of the genus Calomyscus. Cytogenetics and Cell Genetics, 88: 296 - 304." year="2000">Graphodatsky et al., 2000</bibRefCitation>
), the multivariate analysis by
<bibRefCitation id="715031473831684D5EE357BC0D9B39B7" author="Lebedev, V. S. &amp; I. Ya. Pavlinov &amp; M. N. Meyer &amp; V. G. Malikov" refId="ref208246" refString="Lebedev, V. S., I. Ya. Pavlinov, M. N. Meyer, and V. G. Malikov. 1998. [Craniometric analysis of mouse-like hamsters of the genus Calomyscus (Cricetidae).] Zoologschiskii Zhurnal, 77 (6): 721 - 731 (in Russian with English abstract)." year="1998">Lebedev et al. (1998)</bibRefCitation>
demonstrated distinct morphological clusters that correspond to the karyotypic differences. More integrative studies of this kind are required to better understand species diversity and geographic distributions.
</paragraph>
<paragraph id="9650675A32DEA62E0A77C63CB04868AC" pageNumber="927">
All species occupy well drained, barren, rocky habitats in foothills and mountains (
<bibRefCitation id="258BA8701723037134692870B20A4C82" author="Malikov, V. G. &amp; M. N. Meyer &amp; A. S. Grafodatsky &amp; A. V. Polyakov &amp; O. V. Sablina &amp; A. Sh. Vaziri &amp; F. Nazari &amp; J. Zima" refId="ref223074" refString="Malikov, V. G., M. N. Meyer, A. S. Grafodatsky, A. V. Polyakov, O. V. Sablina, A. Sh. Vaziri, F. Nazari, and J. Zima. 1999. On a taxonomic position of some karyomorphs belonging to genus Calomyscus (Rodentia, Cricetidae). Proceedings of the Zoological Institute RAS, 281: 27 - 32." year="1999">Malikov et al., 1999</bibRefCitation>
;
<bibRefCitation id="F81DA3C8CD43D0C92DE0ACCF745661D8" author="Morshed, S. &amp; J. L. Patton" refId="ref247929" refString="Morshed, S., and J. L. Patton. 2002. New records of mammals from Iran with systematic comments on hedgehogs (Erinaceidae) and mouse-like hamsters (Calomyscus, Muridae). Zoology in the Middle East, 26: 49 - 58." year="2002">Morshed and Patton, 2002</bibRefCitation>
), as is depicted in the descriptions and photographs of collection localities for
<collectionCode id="7C38012383346D011309DE0EE7192831" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" type="">AMNH</collectionCode>
and
<collectionCode id="9694FF81E2B23AA624D6B7B8C654C181" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
specimens from
<collectingCountry id="0BB0633B7F50C764C647CD0534130045" name="Iran">Iran</collectingCountry>
and
<collectingCountry id="3F3BCEED1FDAE3248C9551384CF1B4C1" name="Afghanistan">Afghanistan</collectingCountry>
(
<bibRefCitation id="2C5A639961E98F9420904E8446FAD834" author="Goodwin, G. G." refId="ref125418" refString="Goodwin, G. G. 1938. Five new rodents from the eastern Elburz Mountains and a new race of hare from Teheran. American Museum Novitates, 1950: 1 - 5." year="1938">Goodwin, 1938</bibRefCitation>
;
<bibRefCitation id="4AA9B83BB2AEC536E17CA79893044C75" author="Hassinger, J." refId="ref142365" refString="Hassinger, J. 1968. Introduction to the mammal survey of the 1965 Street Expedition to Afghanistan. Fieldiana: Zoology, 55: 1 - 81." year="1968">Hassinger, 1968</bibRefCitation>
;
<bibRefCitation id="04EBED5812C1AF21EA923A38FACD9CBF" author="Lay, D. M." refId="ref207358" refString="Lay, D. M. 1967. A study of the mammals of Iran resulting from the Street expedition of 1962 - 63. Fieldiana: Zoology, 54: 1 - 282." year="1967">Lay, 1967</bibRefCitation>
). Populations appear to be patchily distributed and some are geographically isolated, &quot;which promotes the effect of random genetic drift and may be an important factor of rapid karyotype evolution&quot; (
<bibRefCitation id="962530DAAF39A4551BABE111C0A89A30" author="Graphodatsky, A. S. &amp; O. V. Sablina &amp; M. N. Meyer &amp; V. G. Malikov &amp; E. A. Isakova &amp; V. A. Trifonov &amp; A. V. Polyakov &amp; T. P. Lushnikova &amp; N. V. Vorobieva &amp; N. A. Serdyukova &amp; P. L. Perelman &amp; P. M. Borodin &amp; P. Benda &amp; D. Frynta &amp; L. Leikepova &amp; P. Munclinger &amp; J. Pialek &amp; J. Sadlova &amp; J. Zima" refId="ref127509" refString="Graphodatsky, A. S., O. V. Sablina, M. N. Meyer, V. G. Malikov, E. A. Isakova, V. A. Trifonov, A. V. Polyakov, T. P. Lushnikova, N. V. Vorobieva, N. A. Serdyukova, P. L. Perelman, P. M. Borodin, P. Benda, D. Frynta, L. Leikepova, P. Munclinger, J. Pialek, J. Sadlova, and J. Zima. 2000. Comparative cytogenetics of hamsters of the genus Calomyscus. Cytogenetics and Cell Genetics, 88: 296 - 304." year="2000">Graphodatsky et al., 2000:303</bibRefCitation>
). Natural hybridization between populations with very different karyotypes seems usual in
<taxonomicName id="2CF4EE767B0A5B1AD6ADD4F742EF28AD" authorityName="Thomas" authorityYear="1905" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="2D52987EBA44121E02B92E7CDF4F1524" italics="true" pageNumber="927">Calomyscus</emphasis>
</taxonomicName>
(see account of
<taxonomicName id="15D459248318C7227BADC6A1CE0DD5A4" authorityName="Goodwin" authorityYear="1938" class="Mammalia" family="Calomyscidae" genus="Calomyscus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="elburzensis">
<emphasis id="D49A9AECB310A3EA62EB86D09C89EFF3" italics="true" pageNumber="927">C. elburzensis</emphasis>
</taxonomicName>
) and no unequivocal evidence suggests that chromosomal change is responsible for speciation in the populations studied (
<bibRefCitation id="C56DB128DBD90333EE640492890157AC" author="Graphodatsky, A. S. &amp; O. V. Sablina &amp; M. N. Meyer &amp; V. G. Malikov &amp; E. A. Isakova &amp; V. A. Trifonov &amp; A. V. Polyakov &amp; T. P. Lushnikova &amp; N. V. Vorobieva &amp; N. A. Serdyukova &amp; P. L. Perelman &amp; P. M. Borodin &amp; P. Benda &amp; D. Frynta &amp; L. Leikepova &amp; P. Munclinger &amp; J. Pialek &amp; J. Sadlova &amp; J. Zima" refId="ref127509" refString="Graphodatsky, A. S., O. V. Sablina, M. N. Meyer, V. G. Malikov, E. A. Isakova, V. A. Trifonov, A. V. Polyakov, T. P. Lushnikova, N. V. Vorobieva, N. A. Serdyukova, P. L. Perelman, P. M. Borodin, P. Benda, D. Frynta, L. Leikepova, P. Munclinger, J. Pialek, J. Sadlova, and J. Zima. 2000. Comparative cytogenetics of hamsters of the genus Calomyscus. Cytogenetics and Cell Genetics, 88: 296 - 304." year="2000">Graphodatsky et al., 2000</bibRefCitation>
). Modern geographic range of the genus (SW
<collectingCountry id="344743E8738F1687BDEFAECECCCC367F" name="Syria">Syria</collectingCountry>
, S Caucusus,
<collectingCountry id="AC614720029D86C2F5C64210F62F0E64" name="Iran">Iran</collectingCountry>
, S
<collectingCountry id="26A9AF4F9D2DB4097F4A2E450AB2FBBA" name="Turkmenistan">Turkmenistan</collectingCountry>
,
<collectingCountry id="23CCA08CCCBB3C4AC335DAC1F0CA04E7" name="Afghanistan">Afghanistan</collectingCountry>
, and W
<collectingCountry id="4737BD7D3C86577C534E829DCF7E20E6" name="Pakistan">Pakistan</collectingCountry>
) is a remnant of a broader distribution that in the late Miocene extended as far west as
<collectingCountry id="BD9E09F17C596A402ADD233E979CA7F2" name="Spain">Spain</collectingCountry>
(see
<bibRefCitation id="E47FB9ACB5DCE59C6861209ADDACB57A" author="Agusti, J." refId="ref2926" refString="Agusti, J. 1989. The Miocene rodent succession in Eastern Spain: A zoogeographical appraisal. Pp. 375 - 404, in European Neogene Mammal Chronology (E. H. Lindsay, V. Fahlbusch, and P. Mein, eds.). Plenum Press, New York, 658 pp." year="1989">Agusti, 1989</bibRefCitation>
;
<bibRefCitation id="1A7047BFDCAE5A96539D4D5B0E716275" author="Wessels, W." refId="ref387208" refString="Wessels, W. 1998. Gerbillidae from the Miocene and Pliocene of Europe. Mitteilungen- - Bayerische Staatssammlung fur Palaontolgie und Historische Geologie, 38: 187 - 207." year="1998">Wessels, 1998</bibRefCitation>
,
<bibRefCitation id="0B74ED655AB7D1F0C8B64D7FC9100CF3" author="Wessels, W." refId="ref387239" refString="Wessels, W. 1999. Family Gerbillidae. Pp. 395 - 400, in The Miocene land mammals of Europe (G. E. Rossner and K. Heissig, eds.). Dr. Friedrich Pfeil, Munchen, 515 pp." year="1999">1999</bibRefCitation>
)
</paragraph>
.
</paragraph>
</subSubSection>
</treatment>
</document>