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<mods:title id="DF3F2CEC2FC09B3A5208F45B442E414C">Baronniesia delioti gen. n. sp. n., a new subterranean Leptodirini from the French Pyrenees (Coleoptera: Leiodidae: Cholevinae)</mods:title>
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<mods:namePart id="7BDE95BEB6EEA71540EEC6922FBDC3E9">Fresneda, Javier</mods:namePart>
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<mods:namePart id="A1889F27B7AE69A90A74931022288112">Bourdeau, Charles</mods:namePart>
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<mods:namePart id="5E0A627078786AE0382B095C26C8C9F7">Faille, Arnaud</mods:namePart>
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<mods:date id="A936CC70C822319362B29052547C3F78">2009</mods:date>
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<taxonomicName id="42B04D5AB872FFBCFF654C9DFD591DF3" ID-CoL="8X74W" authority="Fresneda, Bourdeau &amp; Faille" authorityName="Fresneda, Bourdeau &amp; Faille" box="[151,669,693,719]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB872FFBCFF654C9DFEEF1DF3" bold="true" box="[151,299,693,719]" italics="true" pageId="1" pageNumber="2">Baronniesia</emphasis>
Fresneda, Bourdeau &amp; Faille
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gen. n.
</emphasis>
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<emphasis id="B7C4EACBB872FFBCFF654D2AFEE91C26" bold="true" box="[151,301,770,794]" pageId="1" pageNumber="2">
<typeStatus id="5A0B887BB872FFBCFF654D2AFF0B1C26" box="[151,207,770,794]" pageId="1" pageNumber="2">Type</typeStatus>
species:
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<taxonomicName id="42B04D5AB872FFBCFEC14D2AFCFE1C25" authority="Fresneda, Bourdeau &amp; Faille" authorityName="Fresneda, Bourdeau &amp; Faille" box="[307,826,770,793]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="delioti" status="sp. nov.">
<emphasis id="B7C4EACBB872FFBCFEC14D2AFE3B1C25" box="[307,511,770,793]" italics="true" pageId="1" pageNumber="2">Baronniesia delioti</emphasis>
Fresneda, Bourdeau &amp; Faille
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<paragraph id="850F36D9B872FFBCFF654D60FE3419E0" blockId="1.[151,1437,840,2027]" pageId="1" pageNumber="2">
<emphasis id="B7C4EACBB872FFBCFF654D60FEEB1C5E" bold="true" box="[151,303,840,866]" pageId="1" pageNumber="2">Description.</emphasis>
The 
<typeStatus id="5A0B887BB872FFBCFE984D61FE591C5F" box="[362,413,841,867]" pageId="1" pageNumber="2">type</typeStatus>
species of this new genus is characterized by its large size, 4.0–5.0 mm long, being one of the largest species of Leptodirini known from the Pyrenean chain. Sexual dimorphism is very accentuated: male with dilated pentamerous protarsi, antennomeres 5 to 9 exceptionally wide, forming a sort of blade (
<figureCitation id="1D8B2A5CB872FFBCFA9E4DBEFF601CEB" captionStart="FIGURES 1 – 2" captionStartId="2.[151,269,1590,1614]" captionTargetBox="[168,1413,341,1573]" captionTargetId="figure@2.[159,1427,330,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Baronniesia delioti gen. n. sp. n. 1 ― male; 2 ― female." httpUri="https://zenodo.org/record/185599/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
), filiform in females (
<figureCitation id="1D8B2A5CB872FFBCFE5E4D95FE311CEB" box="[428,501,957,983]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,269,1590,1614]" captionTargetBox="[168,1413,341,1573]" captionTargetId="figure@2.[159,1427,330,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Baronniesia delioti gen. n. sp. n. 1 ― male; 2 ― female." httpUri="https://zenodo.org/record/185599/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
). Elytra strongly punctured; punctures wide and rough, forming poorly defined transverse striae, especially in the front area. Mesoventral carina low, forming an obtuse angle, and not extended on metaventrite. Male genital segment reduced, forming a slightly sclerotized ring with two very long lateral appendices. Width of aedeagus (
<figureCitation id="1D8B2A5CB872FFBCFD6E4A19FD251B77" box="[668,737,1073,1099]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,269,1501,1525]" captionTargetBox="[375,1213,477,1469]" captionTargetId="figure@3.[365,1221,465,1477]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Aedeagus of Baronniesia delioti gen. n. sp. n. 3 — dorsal view; 4 — lateral view." httpUri="https://zenodo.org/record/185600/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
) decreasing from base to apex in dorsal view, apex rounded with central part truncated; lateral margins sinuate; aedeagus slightly bent in lateral view (
<figureCitation id="1D8B2A5CB872FFBCFB4E4A70FAC01B4E" box="[1212,1284,1112,1138]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,269,1501,1525]" captionTargetBox="[375,1213,477,1469]" captionTargetId="figure@3.[365,1221,465,1477]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Aedeagus of Baronniesia delioti gen. n. sp. n. 3 — dorsal view; 4 — lateral view." httpUri="https://zenodo.org/record/185600/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
), forming an obtuse angle; terminal tip strongly curved; basal part proportionally large. Lateral styles of aedeagus very thin through their entire length; apex simple, with only three spines (
<figureCitation id="1D8B2A5CB872FFBCFC634A8DFC1E1B83" box="[913,986,1189,1215]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
). Inner sac of aedeagus with the three sections typical of the 
<taxonomicName id="42B04D5AB872FFBCFE404AE4FDFA1BD9" box="[434,574,1228,1253]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB872FFBCFE404AE4FDFA1BD9" box="[434,574,1228,1253]" italics="true" pageId="1" pageNumber="2">Speonomus</emphasis>
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phyletic lineage (
<figureCitation id="1D8B2A5CB872FFBCFCED4AE4FCAF1BDA" box="[799,875,1228,1254]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
) (
<bibRefCitation id="E1214B28B872FFBCFC774AE4FBEA1BDA" author="Jeannel" box="[901,1070,1228,1254]" pageId="1" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel 1910</bibRefCitation>
, 
<bibRefCitation id="E1214B28B872FFBCFBC94AE4FB161BDA" author="Dupre" box="[1083,1234,1228,1254]" pageId="1" pageNumber="14" refString="Dupre, E. (1992) Analyse comparee du sac interne des Bathysciinae (Col., Catopidae): interet taxonomique, considerations sur l'evolution fonctionnelle du sac. Memoires de Biospeologie, 19, 169 - 186." type="journal article" year="1992">Dupré 1992</bibRefCitation>
, 
<bibRefCitation id="E1214B28B872FFBCFB2D4AE4FA5C1BDA" author="Fresneda" box="[1247,1432,1228,1254]" pageId="1" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda 1998</bibRefCitation>
, 
<bibRefCitation id="E1214B28B872FFBCFF654ADAFE0E1A30" author="Fresneda" box="[151,458,1266,1292]" pageId="1" pageNumber="14" refString="Fresneda, J. &amp; Salgado, J. M. (2000) Revision de los generos de Leptodirinae de la Seccion Speonomus del sur de los Pirineos. II: generos Aranzadiella Espanol, 1972, Euryspeonomus Jeannel, 1919, Kobiella Espanol y Belles, 1980 y Speocharidius Jeannel, 1919 (Coleoptera, Cholevidae). Memoires de Biospeologie, 27, 41 - 52." type="journal article" year="2000">Fresneda &amp; Salgado 2000</bibRefCitation>
); reinforcement bands (BRA) only in apical area, very long and reaching the base of the section, which is round and more sclerotized, inserted on two additional large feather-like structures located at the external apical part of the FDM (
<figureCitation id="1D8B2A5CB872FFBCFD394B68FCD11A66" box="[715,789,1344,1370]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
: BDAM). Median area with dorsal (
<figureCitation id="1D8B2A5CB872FFBCFB354B68FAD61A66" box="[1223,1298,1344,1370]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
: FDM) and ventral feather-like structures (
<figureCitation id="1D8B2A5CB872FFBCFE0D4B4EFD8C1ABC" box="[511,584,1382,1408]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
: FVM), two additional slightly curved feather-like structures forming an inverted “V” located above FDM and FVM (
<figureCitation id="1D8B2A5CB872FFBCFD414BA5FD391A9B" box="[691,765,1421,1447]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
: IVP), between BDAM. Apical part of PY in basal area (
<figureCitation id="1D8B2A5CB872FFBCFF6D4B9CFF2C1AF2" box="[159,232,1460,1486]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
: FAPY), above connection nodule (NL), very thin and elongate; feather-like structures associated with PY with strongly sclerotized and large basal nodule; ventral basal complex very robust and sclerotized. Eighth female urosternite with short and robust apophyse and round apex (
<figureCitation id="1D8B2A5CB872FFBCFC4F4829FBD21927" box="[957,1046,1537,1563]" captionStart="FIGURES 15 – 16" captionStartId="8.[151,275,1121,1145]" captionTargetBox="[151,1435,194,1097]" captionTargetId="figure@8.[151,1436,194,1097]" captionTargetPageId="8" captionText="FIGURES 15 – 16. Baronniesia delioti gen. n. sp. n. 15 — urite VIII of female; 16 — female genital segment with spermathecal complex." httpUri="https://zenodo.org/record/185603/files/figure.png" pageId="1" pageNumber="2">Fig. 15</figureCitation>
). Spermatheca of “ 
<typeStatus id="5A0B887BB872FFBCFB0A4829FAEA1927" box="[1272,1326,1537,1563]" pageId="1" pageNumber="2">type</typeStatus>
1” 
<emphasis id="B7C4EACBB872FFBCFAA84829FA581926" box="[1370,1436,1537,1562]" italics="true" pageId="1" pageNumber="2">sensu</emphasis>
<bibRefCitation id="E1214B28B872FFBCFF654800FE8F197E" author="Perreau" box="[151,331,1576,1602]" pageId="1" pageNumber="15" refString="Perreau, M. (1989) De la phylogenie des Cholevidae et des familles apparentees (Coleoptera, Cholevidae). Archives des Sciences Geneve, 39 (3), 579 - 590." type="journal article" year="1989">Perreau (1989)</bibRefCitation>
(
<figureCitation id="1D8B2A5CB872FFBCFEA94800FE70197E" box="[347,436,1576,1602]" captionStart="FIGURES 15 – 16" captionStartId="8.[151,275,1121,1145]" captionTargetBox="[151,1435,194,1097]" captionTargetId="figure@8.[151,1436,194,1097]" captionTargetPageId="8" captionText="FIGURES 15 – 16. Baronniesia delioti gen. n. sp. n. 15 — urite VIII of female; 16 — female genital segment with spermathecal complex." httpUri="https://zenodo.org/record/185603/files/figure.png" pageId="1" pageNumber="2">Fig. 16</figureCitation>
), thin and elongated, slightly curved and with two well-differentiated lobes; apical lobe spherical and smaller than basal lobe, with spermathecal gland inserted at the same place as spermiduct. Spermiduct 10–12 times longer than spermatheca, which gets larger close to the insertion with spermatheca and crosses a sclerotized disc at the contact with bursa copulatrix. Main part of spermiduct close to spermatheca strongly curved.
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B872FFBFFF3448C0FB141E1A" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="etymology">
<paragraph id="850F36D9B872FFBCFF3448C0FC771816" blockId="1.[151,1437,840,2027]" pageId="1" pageNumber="2">
<emphasis id="B7C4EACBB872FFBCFF3448C0FE97183E" bold="true" box="[198,339,1768,1794]" pageId="1" pageNumber="2">Etymology.</emphasis>
The name of the new genus refers to the mountainous massif of Baronnies, located north of the central Pyrenees, between the Aure valley and the Adour valley.
</paragraph>
<paragraph id="850F36D9B872FFBFFF34491EFB141E1A" blockId="1.[151,1437,840,2027]" lastBlockId="2.[151,1437,152,294]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="B7C4EACBB872FFBCFF34491EFE61186C" bold="true" box="[198,421,1846,1872]" pageId="1" pageNumber="2">Taxonomic notes.</emphasis>
<taxonomicName id="42B04D5AB872FFBCFE40491FFD84186C" box="[434,576,1847,1872]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB872FFBCFE40491FFD84186C" box="[434,576,1847,1872]" italics="true" pageId="1" pageNumber="2">Baronniesia</emphasis>
</taxonomicName>
belongs to the Infraflagellates of the 
<taxonomicName id="42B04D5AB872FFBCFBEE491FFB66186C" box="[1052,1186,1847,1872]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB872FFBCFBEE491FFB66186C" box="[1052,1186,1847,1872]" italics="true" pageId="1" pageNumber="2">Speonomus</emphasis>
</taxonomicName>
“phyletic series” of 
<bibRefCitation id="E1214B28B872FFBCFF654975FE8C184B" author="Jeannel" box="[151,328,1885,1911]" pageId="1" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel (1910)</bibRefCitation>
. However, the new genus is morphologically very isolated from the rest of the Pyrenean fauna. At first glance, it looks like a large 
<taxonomicName id="42B04D5AB872FFBCFDB249ACFBC518A2" authority="Jeannel, 1919" authorityName="Jeannel" authorityYear="1919" box="[576,1025,1924,1950]" class="Insecta" family="Leiodidae" genus="Euryspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="breuili">
<emphasis id="B7C4EACBB872FFBCFDB249ACFC9018A1" box="[576,852,1924,1949]" italics="true" pageId="1" pageNumber="2">Euryspeonomus breuili</emphasis>
<bibRefCitation id="E1214B28B872FFBCFCAE49ACFBC518A2" author="Jeannel" box="[860,1025,1924,1950]" pageId="1" pageNumber="15" refString="Jeannel, R. (1919) Bathysciinae nouveaux des Pyrenees espagnoles. Boletin de la real Sociedad espanola de Historia natural, 19, 129 - 137." type="journal article" year="1919">Jeannel, 1919</bibRefCitation>
</taxonomicName>
(Sierra de Aralar, Navarra, 
<collectingCountry id="FDA77649B872FFBCFABD49ACFA5018A2" box="[1359,1428,1924,1950]" name="Spain" pageId="1" pageNumber="2">Spain</collectingCountry>
) but an accurate examination shows that some genital structures are very different, and similar to 
<taxonomicName id="42B04D5AB872FFBCFAE34983FF2918D7" authority="Abeille" authorityName="Abeille" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB872FFBCFAE34983FA5818F8" box="[1297,1436,1963,1988]" italics="true" pageId="1" pageNumber="2">Antrocharis</emphasis>
Abeille
</taxonomicName>
de Perrin, 1878 (Ariège, 
<collectingCountry id="FDA77649B872FFBCFDE749F9FDA318D7" box="[533,615,2001,2027]" name="France" pageId="1" pageNumber="2">France</collectingCountry>
) and related taxa. The lateral styles of the aedeagus of 
<taxonomicName id="42B04D5AB872FFBCFB1149F9FAB518D6" box="[1251,1393,2001,2026]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB872FFBCFB1149F9FAB518D6" box="[1251,1393,2001,2026]" italics="true" pageId="1" pageNumber="2">Baronniesia</emphasis>
</taxonomicName>
are very thin through their entire length, with only three spines at the apex. The inner sac of the aedeagus has two additional feather-like structures forming an inverted “V” above the FDM and FVM, and the apical part of PY is very thin and elongate. All these characters allow to link the new genus to the Pyrenean genera of Leptodirini of the 
<taxonomicName id="42B04D5AB871FFBFFE9F4F24FE3C1E19" box="[365,504,268,293]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB871FFBFFE9F4F24FE3C1E19" box="[365,504,268,293]" italics="true" pageId="2" pageNumber="3">Antrocharis</emphasis>
</taxonomicName>
group (
<emphasis id="B7C4EACBB871FFBFFDA64F24FD511E19" box="[596,661,268,293]" italics="true" pageId="2" pageNumber="3">sensu</emphasis>
<bibRefCitation id="E1214B28B871FFBFFD6F4F24FC4B1E1A" author="Fresneda" box="[669,911,268,294]" pageId="2" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB871FFBFFCFC4F24FC851E19" box="[782,833,268,293]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. 2007
</bibRefCitation>
), which is redefined below.
</paragraph>
</subSubSection>
<caption id="D1CF6651B871FFBFFF65481EFBCD1971" box="[151,1033,1590,1614]" httpUri="https://zenodo.org/record/185599/files/figure.png" pageId="2" pageNumber="3" targetBox="[168,1413,341,1573]" targetPageId="2">
<paragraph id="850F36D9B871FFBFFF65481EFBCD1971" blockId="2.[151,1033,1590,1614]" box="[151,1033,1590,1614]" pageId="2" pageNumber="3">
<emphasis id="B7C4EACBB871FFBFFF65481EFE861972" bold="true" box="[151,322,1590,1614]" pageId="2" pageNumber="3">FIGURES 1–2.</emphasis>
Habitus of 
<taxonomicName id="42B04D5AB871FFBFFE32481EFD481971" box="[448,652,1590,1613]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="delioti" status="gen. nov.">
<emphasis id="B7C4EACBB871FFBFFE32481EFD481971" box="[448,652,1590,1613]" italics="true" pageId="2" pageNumber="3">Baronniesia delioti</emphasis>
</taxonomicName>
<emphasis id="B7C4EACBB871FFBFFD60481EFCE41972" bold="true" box="[658,800,1590,1614]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="ACF757B0B871FFBFFD60481EFD191972" box="[658,733,1590,1614]" pageId="2" pageNumber="3" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
1―male; 2―female.
</paragraph>
</caption>
<subSubSection id="CDAA6552B871FFBFFF344857FEEE18BD" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="850F36D9B871FFBFFF344857FA9819FC" blockId="2.[151,1437,1663,2037]" pageId="2" pageNumber="3">
The 
<taxonomicName id="42B04D5AB871FFBFFF0948A8FE4219A5" ID-CoL="8X73K" box="[251,390,1664,1689]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB871FFBFFF0948A8FE4219A5" box="[251,390,1664,1689]" italics="true" pageId="2" pageNumber="3">Antrocharis</emphasis>
</taxonomicName>
group was characterized by the following characters (the terminology of the structures of the inner sac of the aedeagus follows 
<bibRefCitation id="E1214B28B871FFBFFDBD488EFD2F19FC" author="Jeannel" box="[591,747,1702,1728]" pageId="2" pageNumber="15" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63 (1), 1 - 436." type="journal article" year="1924">Jeannel 1924</bibRefCitation>
, 
<bibRefCitation id="E1214B28B871FFBFFD05488EFC4119FC" author="Belles" box="[759,901,1702,1728]" pageId="2" pageNumber="13" refString="Belles, X. (1984) Estudi de l'armadura genital en les poblacions de Speonomus (Parvospeonomus) delarouzeei s. l. i S. (P.) vilarrubiasi Zariquiey (Col., Catopidae, Bathysciinae) que conviuen a la Cova del Far (Susqueda, La Selva). Exploracions, 8, 7 - 13." type="journal article" year="1984">Bellés 1984</bibRefCitation>
, 
<bibRefCitation id="E1214B28B871FFBFFC63488EFBDA19FC" author="Dupre" box="[913,1054,1702,1728]" pageId="2" pageNumber="14" refString="Dupre, E. (1989) Bathysciinae (Col. Catopidae) des Pyrenees Occidentales: I. Etude du sac interne et morphologie comparee des Speonomus de la region de St-Jean-Pied-de-Port, redefinition du sous-genre Phacomorphus et creation du sous-genre Phacomorphoides, considerations phylogeniques et biogeographiques locales, considerations generales sur la systematique des Bathysciinae. Ikartzaleak, 13, 55 - 99." type="journal article" year="1989">Dupré 1989</bibRefCitation>
, 
<bibRefCitation id="E1214B28B871FFBFFBD9488EFBA219FC" author="Dupre" box="[1067,1126,1702,1728]" pageId="2" pageNumber="14" refString="Dupre, E. (1992) Analyse comparee du sac interne des Bathysciinae (Col., Catopidae): interet taxonomique, considerations sur l'evolution fonctionnelle du sac. Memoires de Biospeologie, 19, 169 - 186." type="journal article" year="1992">1992</bibRefCitation>
and 
<bibRefCitation id="E1214B28B871FFBFFB6C488EFA8B19FC" author="Fresneda" box="[1182,1359,1702,1728]" pageId="2" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda 1998</bibRefCitation>
):
</paragraph>
<paragraph id="850F36D9B871FFBFFF3448E5FEEE18BD" blockId="2.[151,1437,1663,2037]" pageId="2" pageNumber="3">
1) The median lobe of the aedeagus in lateral view is curved forming right or obtuse angle, with a strong depression or dorsal sinuosity in the apical third. Until now this character was considered a synapomorphy of the group. The median lobe of aedeagus of 
<taxonomicName id="42B04D5AB871FFBFFD664932FCE6180F" ID-CoL="8X74W" box="[660,802,1818,1843]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB871FFBFFD664932FCE6180F" box="[660,802,1818,1843]" italics="true" pageId="2" pageNumber="3">Baronniesia</emphasis>
</taxonomicName>
in lateral view is curved forming an obtuse angle, but without any depression or sinuosity, decreasing regularly from base to apex, and strongly curved in the central part (
<figureCitation id="1D8B2A5CB871FFBFFF26494FFED918BD" box="[212,285,1895,1921]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,269,1501,1525]" captionTargetBox="[375,1213,477,1469]" captionTargetId="figure@3.[365,1221,465,1477]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Aedeagus of Baronniesia delioti gen. n. sp. n. 3 — dorsal view; 4 — lateral view." httpUri="https://zenodo.org/record/185600/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B871FFBEFF3449A6FE421EA6" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="850F36D9B871FFBFFF3449A6FA8C18C9" blockId="2.[151,1437,1663,2037]" pageId="2" pageNumber="3">
2) Lateral styles of the aedeagus are shorter than the median lobe, very thin through their entire length and each carrying three long spines at the apex (one shorter), without any penicillum, cavity or membranous lamina (
<figureCitation id="1D8B2A5CB871FFBFFF0449F3FEAD18C9" box="[246,361,2011,2037]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="2" pageNumber="3">Figs. 8–9</figureCitation>
). The lateral stylus of 
<taxonomicName id="42B04D5AB871FFBFFD9C49F4FD3818C9" box="[622,764,2012,2037]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB871FFBFFD9C49F4FD3818C9" box="[622,764,2012,2037]" italics="true" pageId="2" pageNumber="3">Baronniesia</emphasis>
</taxonomicName>
conforms to this morphological structure (
<figureCitation id="1D8B2A5CB871FFBFFB0049F3FAFE18C9" box="[1266,1338,2011,2037]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="850F36D9B870FFBEFF344EB0FBA51FC3" blockId="3.[151,1436,152,410]" pageId="3" pageNumber="4">
3) Presence of two additional strongly sclerotized feather-like structures forming an inverted “V”, above the FDM and FVM (
<figureCitation id="1D8B2A5CB870FFBEFE794E96FE3A1FE4" box="[395,510,190,216]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="3" pageNumber="4">Figs. 5–6</figureCitation>
: IVP). These two feather-like structures are present in 
<taxonomicName id="42B04D5AB870FFBEFB724E97FACA1FE4" box="[1152,1294,191,216]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFB724E97FACA1FE4" box="[1152,1294,191,216]" italics="true" pageId="3" pageNumber="4">Baronniesia</emphasis>
</taxonomicName>
, which also has two additional structures located laterally of the inverted “V” (
<figureCitation id="1D8B2A5CB870FFBEFC514ECDFC281FC3" box="[931,1004,229,255]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
: BDAM).
</paragraph>
<paragraph id="850F36D9B870FFBEFF344F24FE421EA6" blockId="3.[151,1436,152,410]" pageId="3" pageNumber="4">
4) The apical part of PY (FAPY), above the connection nodule (NL), is very thin and elongate. 
<taxonomicName id="42B04D5AB870FFBEFF654F1AFEED1E77" box="[151,297,306,331]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFF654F1AFEED1E77" box="[151,297,306,331]" italics="true" pageId="3" pageNumber="4">Baronniesia</emphasis>
</taxonomicName>
shares this character with all genera of the 
<taxonomicName id="42B04D5AB870FFBEFCCE4F1AFC0F1E77" box="[828,971,306,331]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFCCE4F1AFC0F1E77" box="[828,971,306,331]" italics="true" pageId="3" pageNumber="4">Antrocharis</emphasis>
</taxonomicName>
group (
<figureCitation id="1D8B2A5CB870FFBEFBD94F1AFBB21E70" box="[1067,1142,306,332]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
: FAPY). In all the other genera of the Pyrenean Leptodirini (not members of the 
<taxonomicName id="42B04D5AB870FFBEFCC84F71FC031E4E" box="[826,967,345,370]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFCC84F71FC031E4E" box="[826,967,345,370]" italics="true" pageId="3" pageNumber="4">Antrocharis</emphasis>
</taxonomicName>
group) the apical part of PY is shorter, broader and conical.
</paragraph>
</subSubSection>
<caption id="D1CF6651B870FFBEFF654BF5FB5D1AC8" box="[151,1177,1501,1525]" httpUri="https://zenodo.org/record/185600/files/figure.png" pageId="3" pageNumber="4" targetBox="[375,1213,477,1469]" targetPageId="3">
<paragraph id="850F36D9B870FFBEFF654BF5FB5D1AC8" blockId="3.[151,1177,1501,1525]" box="[151,1177,1501,1525]" pageId="3" pageNumber="4">
<emphasis id="B7C4EACBB870FFBEFF654BF5FE871AC9" bold="true" box="[151,323,1501,1525]" pageId="3" pageNumber="4">FIGURES 3–4.</emphasis>
Aedeagus of 
<taxonomicName id="42B04D5AB870FFBEFE244BF6FD661AC9" box="[470,674,1502,1525]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="delioti" status="gen. nov.">
<emphasis id="B7C4EACBB870FFBEFE244BF6FD661AC9" box="[470,674,1502,1525]" italics="true" pageId="3" pageNumber="4">Baronniesia delioti</emphasis>
</taxonomicName>
<emphasis id="B7C4EACBB870FFBEFD5B4BF5FCF21AC9" bold="true" box="[681,822,1501,1525]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="ACF757B0B870FFBEFD5B4BF5FD371AC9" box="[681,755,1501,1525]" pageId="3" pageNumber="4" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
3—dorsal view; 4—lateral view.
</paragraph>
</caption>
<subSubSection id="CDAA6552B870FFB9FF34480FFC1C18CA" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="850F36D9B870FFBEFF34480FFCAD1873" blockId="3.[151,1437,1575,2026]" pageId="3" pageNumber="4">
The 
<taxonomicName id="42B04D5AB870FFBEFEF5480FFE56197C" ID-CoL="8X73K" box="[263,402,1575,1600]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFEF5480FFE56197C" box="[263,402,1575,1600]" italics="true" pageId="3" pageNumber="4">Antrocharis</emphasis>
</taxonomicName>
group includes four genera: 
<taxonomicName id="42B04D5AB870FFBEFCD0480FFC69197C" ID-CoL="8X73K" box="[802,941,1575,1600]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFCD0480FFC69197C" box="[802,941,1575,1600]" italics="true" pageId="3" pageNumber="4">Antrocharis</emphasis>
</taxonomicName>
(two species), 
<taxonomicName id="42B04D5AB870FFBEFB76480FFEB1195B" ID-CoL="8X7FT" authority="Giachino &amp; Gueorguiev, 1989" authorityName="Giachino &amp; Gueorguiev" authorityYear="1989" class="Insecta" family="Leiodidae" genus="Gesciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFB76480FFB37197C" box="[1156,1267,1575,1600]" italics="true" pageId="3" pageNumber="4">Gesciella</emphasis>
<bibRefCitation id="E1214B28B870FFBEFAF4480FFEB1195B" author="Giachino" pageId="3" pageNumber="15" refString="Giachino, P. M. &amp; Gueorguiev, V. B. (1989) Un nouveau genre de Bathysciinae des Pyrenees francaises (Coleoptera, Bathysciinae). Revue suisse de Zoologie, 96 (2), 403 - 410." type="journal article" year="1989">Giachino &amp; Guéorguiev, 1989</bibRefCitation>
</taxonomicName>
(monospecific genus), 
<taxonomicName id="42B04D5AB870FFBEFD6E4866FBD3195B" ID-CoL="8X7NS" authority="Foures, 1954" authorityName="Foures" authorityYear="1954" box="[668,1047,1613,1639]" class="Insecta" family="Leiodidae" genus="Paratroglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFD6E4866FCAF195B" box="[668,875,1614,1639]" italics="true" pageId="3" pageNumber="4">Paratroglophyes</emphasis>
<bibRefCitation id="E1214B28B870FFBEFC864865FBD3195B" author="Foures" box="[884,1047,1613,1639]" pageId="3" pageNumber="14" refString="Foures, H. (1954) Diagnose preliminaire de nouveaux Coleopteres cavernicoles de la region du mont Valier (Ariege). Bulletin de la Societe linneenne de Lyon, 23, 3 - 5." type="journal article" year="1954">Fourès, 1954</bibRefCitation>
</taxonomicName>
(two species) and 
<taxonomicName id="42B04D5AB870FFBEFAF64866FF2A19B2" ID-CoL="8X7XP" authority="Abeille" authorityName="Abeille" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFAF64866FA58195B" box="[1284,1436,1614,1639]" italics="true" pageId="3" pageNumber="4">Troglophyes</emphasis>
Abeille
</taxonomicName>
de Perrin, 
<quantity id="42489B3CB870FFBEFE83485CFE0F19B1" box="[369,459,1652,1678]" metricMagnitude="1" metricUnit="m" metricValue="4.81076" pageId="3" pageNumber="4" unit="in" value="1894.0">
1894 
<emphasis id="B7C4EACBB870FFBEFE46485CFE0F19B1" box="[436,459,1652,1677]" italics="true" pageId="3" pageNumber="4">in</emphasis>
</quantity>
pars sensu 
<bibRefCitation id="E1214B28B870FFBEFDAB485CFC8A19B2" author="Fresneda" box="[601,846,1652,1678]" pageId="3" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB870FFBEFD39485CFCC319B1" box="[715,775,1652,1677]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
2007
</bibRefCitation>
: 
<taxonomicName id="42B04D5AB870FFBEFCA8485CFBB619B2" ID-CoL="5994W" authority="Coiffait, 1953" authorityName="Coiffait" authorityYear="1953" box="[858,1138,1652,1678]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B7C4EACBB870FFBEFCA8485CFC0019B1" box="[858,964,1652,1677]" italics="true" pageId="3" pageNumber="4">T. aubryi</emphasis>
<bibRefCitation id="E1214B28B870FFBEFC3E485CFBB619B2" author="Coiffait" box="[972,1138,1652,1678]" pageId="3" pageNumber="14" refString="Coiffait, H. (1953) Quatre nouveaux coleopteres cavernicoles du massif du Montvallier (Ariege). Notes Biospeologiques, 8, 27 - 32." type="journal article" year="1953">Coiffait, 1953</bibRefCitation>
</taxonomicName>
with the subspecies 
<emphasis id="B7C4EACBB870FFBEFA9A485CFEEC1988" italics="true" pageId="3" pageNumber="4">T. a. vallierensis</emphasis>
<bibRefCitation id="E1214B28B870FFBEFEC348B3FE251989" author="Coiffait" box="[305,481,1691,1717]" pageId="3" pageNumber="14" refString="Coiffait, H. (1953) Quatre nouveaux coleopteres cavernicoles du massif du Montvallier (Ariege). Notes Biospeologiques, 8, 27 - 32." type="journal article" year="1953">Coiffait, 1953</bibRefCitation>
(
<bibRefCitation id="E1214B28B870FFBEFE0148B3FD391989" author="Fresneda" box="[499,765,1691,1717]" pageId="3" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB870FFBEFD9D48B3FD741988" box="[623,688,1691,1716]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
2007
</bibRefCitation>
). 
<bibRefCitation id="E1214B28B870FFBEFCE148B3FC161989" author="Coiffait" box="[787,978,1691,1717]" pageId="3" pageNumber="14" refString="Coiffait, H. (1953) Quatre nouveaux coleopteres cavernicoles du massif du Montvallier (Ariege). Notes Biospeologiques, 8, 27 - 32." type="journal article" year="1953">Coiffait (1953)</bibRefCitation>
indicates that the two subspecies of 
<taxonomicName id="42B04D5AB870FFBEFF6548EAFEBD19E7" ID-CoL="5994W" box="[151,377,1730,1755]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B7C4EACBB870FFBEFF6548EAFEBD19E7" box="[151,377,1730,1755]" italics="true" pageId="3" pageNumber="4">Troglophyes aubryi</emphasis>
</taxonomicName>
have the apex of the styles each with three spines and a penicillum composed of 7–8 setae. We did not see a penicillum in the numerous specimens from the Anglade cave (
<emphasis id="B7C4EACBB870FFBEFB9B48C1FAC1183E" box="[1129,1285,1769,1794]" italics="true" pageId="3" pageNumber="4">locus typicus</emphasis>
of 
<taxonomicName id="42B04D5AB870FFBEFADD48C1FF221814" ID-CoL="8YFVB" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="aubryi" subSpecies="aubryi">
<emphasis id="B7C4EACBB870FFBEFADD48C1FF221814" italics="true" pageId="3" pageNumber="4">T. aubryi aubryi</emphasis>
</taxonomicName>
) and various MSS (“ 
<emphasis id="B7C4EACBB870FFBEFE194927FC8F1814" box="[491,843,1807,1832]" italics="true" pageId="3" pageNumber="4">Milieu souterrain superficiel</emphasis>
” 
<bibRefCitation id="E1214B28B870FFBEFC924927FB991815" author="Juberthie" box="[864,1117,1807,1833]" pageId="3" pageNumber="15" refString="Juberthie, C., Delay, B. &amp; Bouillon, M. (1980) Extension du milieu souterrain en zone non calcaire: Description d'un nouveau milieu et de son peuplement par les coleopteres troglobies. Memoires de Biospeologie, 7, 19 - 52." type="journal article" year="1980">
Juberthie 
<emphasis id="B7C4EACBB870FFBEFC2A4927FBCB1814" box="[984,1039,1807,1832]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. 1980
</bibRefCitation>
) in the area; all examined specimens have only three spines without penicillum (
<figureCitation id="1D8B2A5CB870FFBEFCE1491DFC9E1873" box="[787,858,1845,1871]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="850F36D9B870FFBEFF344974FD7518D6" blockId="3.[151,1437,1575,2026]" pageId="3" pageNumber="4">
<taxonomicName id="42B04D5AB870FFBEFF344974FD6A184A" ID-CoL="3BHT8" authority="Coiffait, 1950" authorityName="Coiffait" authorityYear="1950" box="[198,686,1884,1910]" class="Insecta" family="Leiodidae" genus="Eskualdunella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="delespierrei">
<emphasis id="B7C4EACBB870FFBEFF344974FDC41849" box="[198,512,1884,1909]" italics="true" pageId="3" pageNumber="4">Eskualdunella delespierrei</emphasis>
<bibRefCitation id="E1214B28B870FFBEFDFA4974FD6A184A" author="Coiffait" box="[520,686,1884,1910]" pageId="3" pageNumber="14" refString="Coiffait, H. (1950 b) Trois Bathysciites nouveaux de Larrau (Basses-Pyrenees). Notes Biospeologiques, 5, 65 - 70." type="journal article" year="1950" yearSuffix="b">Coiffait, 1950</bibRefCitation>
</taxonomicName>
b, an enigmatic species from the French Basque country, known by a single specimen, seems to have a similar structure of the stylus apex, as the members of the 
<taxonomicName id="42B04D5AB870FFBEFAE349ABFA5818A0" ID-CoL="8X73K" box="[1297,1436,1923,1948]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFAE349ABFA5818A0" box="[1297,1436,1923,1948]" italics="true" pageId="3" pageNumber="4">Antrocharis</emphasis>
</taxonomicName>
group: three spines without penicillum (
<bibRefCitation id="E1214B28B870FFBEFD9B4982FCDE18F8" author="Coiffait" box="[617,794,1962,1988]" pageId="3" pageNumber="14" refString="Coiffait, H. (1950 b) Trois Bathysciites nouveaux de Larrau (Basses-Pyrenees). Notes Biospeologiques, 5, 65 - 70." type="journal article" year="1950" yearSuffix="b">Coiffait 1950b</bibRefCitation>
: 
<figureCitation id="1D8B2A5CB870FFBEFCD74982FCBE18F8" box="[805,890,1962,1988]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
; 1955). But in 
<taxonomicName id="42B04D5AB870FFBEFBDA4982FB1418FF" ID-CoL="8XL99" box="[1064,1232,1962,1987]" class="Insecta" family="Leiodidae" genus="Eskualdunella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB870FFBEFBDA4982FB1418FF" box="[1064,1232,1962,1987]" italics="true" pageId="3" pageNumber="4">Eskualdunella</emphasis>
</taxonomicName>
one of the spines is longer and more robust than the two others.
</paragraph>
<caption id="D1CF6651B877FFB9FF654BA9FCCF193C" httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="4" pageNumber="5" targetBox="[441,1145,181,1397]" targetPageId="4">
<paragraph id="850F36D9B877FFB9FF654BA9FCCF193C" blockId="4.[151,1437,1409,1536]" pageId="4" pageNumber="5">
<emphasis id="B7C4EACBB877FFB9FF654BA9FE891AA5" bold="true" box="[151,333,1409,1433]" pageId="4" pageNumber="5">FIGURES 5–6.</emphasis>
Inner sac of aedeagus of Leptodirini. 5— 
<taxonomicName id="42B04D5AB877FFB9FCC04BA9FBCB1AA4" ID-CoL="68DT6" box="[818,1039,1409,1432]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="delioti" status="gen. nov.">
<emphasis id="B7C4EACBB877FFB9FCC04BA9FBCB1AA4" box="[818,1039,1409,1432]" italics="true" pageId="4" pageNumber="5">Baronniesia delioti</emphasis>
</taxonomicName>
<emphasis id="B7C4EACBB877FFB9FBE54BA9FB761AA5" bold="true" box="[1047,1202,1409,1433]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="ACF757B0B877FFB9FBE54BA9FBAC1AA5" box="[1047,1128,1409,1433]" pageId="4" pageNumber="5" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
and 6— 
<taxonomicName id="42B04D5AB877FFB9FAE74BA9FE751A87" ID-CoL="5DJF2" authority="Abeille" authorityName="Abeille" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="querilhaci" subSpecies="dispar">
<emphasis id="B7C4EACBB877FFB9FAE74BA9FE9C1A87" italics="true" pageId="4" pageNumber="5">Antrocharis querilhaci dispar</emphasis>
Abeille
</taxonomicName>
de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “V” feather-like structures; PY, “ 
<emphasis id="B7C4EACBB877FFB9FD784BC1FD3C193C" box="[650,760,1513,1536]" italics="true" pageId="4" pageNumber="5">piece en Y</emphasis>
”.
</paragraph>
</caption>
<paragraph id="850F36D9B877FFB9FF34481AFC1C18CA" blockId="4.[151,1437,1586,2038]" pageId="4" pageNumber="5">
In the genus 
<taxonomicName id="42B04D5AB877FFB9FE96481AFD591970" ID-CoL="8X754" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" box="[356,669,1586,1612]" class="Insecta" family="Leiodidae" genus="Bathysciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FE96481AFE2A1977" box="[356,494,1586,1611]" italics="true" pageId="4" pageNumber="5">Bathysciola</emphasis>
<bibRefCitation id="E1214B28B877FFB9FE05481AFD591970" author="Jeannel" box="[503,669,1586,1612]" pageId="4" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
, the species of 
<emphasis id="B7C4EACBB877FFB9FCAB481AFC5E1977" box="[857,922,1586,1611]" italics="true" pageId="4" pageNumber="5">ovata</emphasis>
group (
<bibRefCitation id="E1214B28B877FFB9FC0B481AFB581970" author="Perreau" box="[1017,1180,1586,1612]" pageId="4" pageNumber="15" refString="Perreau, M. (2000) Catalogue des Coleopteres Leiodidae Cholevinae et Platypsyllinae. Memoires de la Societe entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau 2000</bibRefCitation>
, 
<bibRefCitation id="E1214B28B877FFB9FB58481AFF13194F" author="Fresneda" pageId="4" pageNumber="14" refString="Fresneda, J. &amp; Salgado, J. M. (2006) The genus Bathysciola Jeannel, 1910 in the Iberian Peninsula and Pyrenees. Taxonomic revision of the sections IV, VI and VII (Jeannel, 1924) (Coleoptera, Cholevidae, Leptodirinae). Graellsia, 62 (1), 25 - 74." type="journal article" year="2006">Fresneda &amp; Salgado 2006</bibRefCitation>
) have the apex of the stylus with only three spines, but the mesoventral keel is extended over the metaventrite, and transverse striae on the elytra are missing. Those characters separate the 
<emphasis id="B7C4EACBB877FFB9FB3E48A8FAC919A5" box="[1228,1293,1664,1689]" italics="true" pageId="4" pageNumber="5">ovata</emphasis>
group from the 
<taxonomicName id="42B04D5AB877FFB9FF36488FFE9619FC" ID-CoL="8X73K" box="[196,338,1703,1728]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FF36488FFE9619FC" box="[196,338,1703,1728]" italics="true" pageId="4" pageNumber="5">Antrocharis</emphasis>
</taxonomicName>
group and 
<emphasis id="B7C4EACBB877FFB9FE29488FFDB019FC" box="[475,628,1703,1728]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="42B04D5AB877FFB9FE29488FFDB419FC" ID-CoL="8X74W" box="[475,624,1703,1728]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Baronniesia</taxonomicName>
.
</emphasis>
The species of 
<emphasis id="B7C4EACBB877FFB9FCC0488FFC4819FC" box="[818,908,1703,1728]" italics="true" pageId="4" pageNumber="5">madoni</emphasis>
and 
<emphasis id="B7C4EACBB877FFB9FC35488FFBFB19FC" box="[967,1087,1703,1728]" italics="true" pageId="4" pageNumber="5">zariquieyi</emphasis>
groups are closely related to 
<emphasis id="B7C4EACBB877FFB9FF6548E5FF1C19DA" box="[151,216,1741,1766]" italics="true" pageId="4" pageNumber="5">ovata</emphasis>
group (
<bibRefCitation id="E1214B28B877FFB9FEC148E5FDED19DB" author="Fresneda" box="[307,553,1741,1767]" pageId="4" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB877FFB9FE5548E5FE1F19DA" box="[423,475,1741,1766]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2007
</bibRefCitation>
), and have a variable number of spines in the lateral stylus of the aedeagus (between 5 and 14). In the phylogenetic reconstruction by 
<bibRefCitation id="E1214B28B877FFB9FCB848DCFB891832" box="[842,1101,1780,1806]" pageId="4" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article">
Fresneda 
<emphasis id="B7C4EACBB877FFB9FC4E48DCFC331831" box="[956,1015,1780,1805]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(2007)
</bibRefCitation>
these groups of 
<taxonomicName id="42B04D5AB877FFB9FAE048DCFA581831" ID-CoL="8X754" box="[1298,1436,1780,1805]" class="Insecta" family="Leiodidae" genus="Bathysciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FAE048DCFA581831" box="[1298,1436,1780,1805]" italics="true" pageId="4" pageNumber="5">Bathysciola</emphasis>
</taxonomicName>
are not closely related to the 
<taxonomicName id="42B04D5AB877FFB9FE0A4933FD431808" ID-CoL="8X73K" box="[504,647,1819,1844]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FE0A4933FD431808" box="[504,647,1819,1844]" italics="true" pageId="4" pageNumber="5">Antrocharis</emphasis>
</taxonomicName>
group.The other Pyrenean 
<taxonomicName id="42B04D5AB877FFB9FC204933FBA41808" ID-CoL="8X754" box="[978,1120,1819,1844]" class="Insecta" family="Leiodidae" genus="Bathysciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FC204933FBA41808" box="[978,1120,1819,1844]" italics="true" pageId="4" pageNumber="5">Bathysciola</emphasis>
</taxonomicName>
(
<emphasis id="B7C4EACBB877FFB9FB804933FB281808" box="[1138,1260,1819,1844]" italics="true" pageId="4" pageNumber="5">lapidicola</emphasis>
, 
<emphasis id="B7C4EACBB877FFB9FB0E4933FA501808" box="[1276,1428,1819,1844]" italics="true" pageId="4" pageNumber="5">meridionalis</emphasis>
, 
<emphasis id="B7C4EACBB877FFB9FF654969FECC1866" box="[151,264,1857,1882]" italics="true" pageId="4" pageNumber="5">larcennei</emphasis>
and 
<emphasis id="B7C4EACBB877FFB9FEB14969FE741866" box="[323,432,1857,1882]" italics="true" pageId="4" pageNumber="5">schiodtei</emphasis>
groups) have four spines in the apex of the lateral stylus of the aedeagus (the only exception is one species of the 
<emphasis id="B7C4EACBB877FFB9FDE54940FD7518BD" box="[535,689,1896,1921]" italics="true" pageId="4" pageNumber="5">meridionalis</emphasis>
group, 
<taxonomicName id="42B04D5AB877FFB9FCE24940FAE618BE" ID-CoL="KY5V" authority="Fresneda &amp; Salgado, 2006" authorityName="Fresneda &amp; Salgado" authorityYear="2006" box="[784,1314,1896,1922]" class="Insecta" family="Leiodidae" genus="Bathysciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="finismillennii">
<emphasis id="B7C4EACBB877FFB9FCE24940FC1018BD" box="[784,980,1896,1921]" italics="true" pageId="4" pageNumber="5">B. finismillennii</emphasis>
<bibRefCitation id="E1214B28B877FFB9FC2E4940FAE618BE" author="Fresneda" box="[988,1314,1896,1922]" pageId="4" pageNumber="14" refString="Fresneda, J. &amp; Salgado, J. M. (2006) The genus Bathysciola Jeannel, 1910 in the Iberian Peninsula and Pyrenees. Taxonomic revision of the sections IV, VI and VII (Jeannel, 1924) (Coleoptera, Cholevidae, Leptodirinae). Graellsia, 62 (1), 25 - 74." type="journal article" year="2006">Fresneda &amp; Salgado, 2006</bibRefCitation>
</taxonomicName>
with nine spines), and belong to a clade of Leptodirini restricted to the Spanish Basque country (
<emphasis id="B7C4EACBB877FFB9FB4849A7FB251894" box="[1210,1249,1935,1960]" italics="true" pageId="4" pageNumber="5">see</emphasis>
<bibRefCitation id="E1214B28B877FFB9FB1B49A6FF1318F3" author="Fresneda" pageId="4" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB877FFB9FAAC49A7FA501894" box="[1374,1428,1935,1960]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2007
</bibRefCitation>
): 
<taxonomicName id="42B04D5AB877FFB9FF18499DFDAC18F3" ID-CoL="8X7H9" authority="Belles &amp; Deliot, 1983" authorityName="Belles &amp; Deliot" authorityYear="1983" box="[234,616,1973,1999]" class="Insecta" family="Leiodidae" genus="Josettekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FF18499DFEA418F2" box="[234,352,1973,1998]" italics="true" pageId="4" pageNumber="5">Josettekia</emphasis>
<bibRefCitation id="E1214B28B877FFB9FE94499DFDAC18F3" author="Belles" box="[358,616,1973,1999]" pageId="4" pageNumber="13" refString="Belles, X. &amp; Deliot, P. (1983) Nouveaux Bathysciinae (Coleopteres Catopidae) des Pyrenees francaises et espagnoles. Memoires de Biospeologie, 10, 237 - 243." type="journal article" year="1983">Bellés &amp; Déliot, 1983</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB877FFB9FD86499DFC0118F3" ID-CoL="8X7VQ" authority="Jeannel, 1919" authorityName="Jeannel" authorityYear="1919" box="[628,965,1973,1999]" class="Insecta" family="Leiodidae" genus="Speocharidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FD86499DFCD818F2" box="[628,796,1973,1998]" italics="true" pageId="4" pageNumber="5">Speocharidius</emphasis>
<bibRefCitation id="E1214B28B877FFB9FCD0499DFC0118F3" author="Jeannel" box="[802,965,1973,1999]" pageId="4" pageNumber="15" refString="Jeannel, R. (1919) Bathysciinae nouveaux des Pyrenees espagnoles. Boletin de la real Sociedad espanola de Historia natural, 19, 129 - 137." type="journal article" year="1919">Jeannel, 1919</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB877FFB9FC23499DFAF018F3" ID-CoL="8X7DW" authority="Jeannel, 1919" authorityName="Jeannel" authorityYear="1919" box="[977,1332,1973,1999]" class="Insecta" family="Leiodidae" genus="Euryspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FC23499DFB4818F2" box="[977,1164,1973,1998]" italics="true" pageId="4" pageNumber="5">Euryspeonomus</emphasis>
<bibRefCitation id="E1214B28B877FFB9FB61499DFAF018F3" author="Jeannel" box="[1171,1332,1973,1999]" pageId="4" pageNumber="15" refString="Jeannel, R. (1919) Bathysciinae nouveaux des Pyrenees espagnoles. Boletin de la real Sociedad espanola de Historia natural, 19, 129 - 137." type="journal article" year="1919">Jeannel, 1919</bibRefCitation>
</taxonomicName>
(all with four spines) and 
<taxonomicName id="42B04D5AB877FFB9FEA949F4FD6618CA" ID-CoL="8X73X" authority="Espanol, 1972" authorityName="Espanol" authorityYear="1972" box="[347,674,2012,2038]" class="Insecta" family="Leiodidae" genus="Aranzadiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB877FFB9FEA949F4FE3618C9" box="[347,498,2012,2037]" italics="true" pageId="4" pageNumber="5">Aranzadiella</emphasis>
<bibRefCitation id="E1214B28B877FFB9FE0849F4FD6618CA" author="Espanol" box="[506,674,2012,2038]" pageId="4" pageNumber="14" refString="Espanol, F. (1972) Nuevos datos sobre los Bathysciinae de Guipuzcoa (Col. Catopidae). Eos, 47, 59 - 66." type="journal article" year="1972">Español, 1972</bibRefCitation>
</taxonomicName>
(with about thirty spines).
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B876FFBAFF344EB0FD3F1C7F" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="850F36D9B876FFB8FF344EB0FDEE1FC3" blockId="5.[151,1436,152,255]" pageId="5" pageNumber="6">In all other Pyrenean Leptodirini the number of spines in the apex of the lateral stylus of the aedeagus varies from two to five, but they all have a penicillum, which could be inserted in a depression or not, or be inserted on a membranous lamina.</paragraph>
<caption id="D1CF6651B876FFB8FF6548D1FB4A1845" httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="5" pageNumber="6" targetBox="[164,1428,301,1753]" targetPageId="5">
<paragraph id="850F36D9B876FFB8FF6548D1FB4A1845" blockId="5.[151,1437,1785,1913]" pageId="5" pageNumber="6">
<emphasis id="B7C4EACBB876FFB8FF6548D1FEA6182D" bold="true" box="[151,354,1785,1809]" pageId="5" pageNumber="6">FIGURES 7–14.</emphasis>
Apices of lateral styles of aedeagus of Leptodirini. 7— 
<taxonomicName id="42B04D5AB876FFB8FBF148D2FB2D182D" box="[1027,1257,1786,1809]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="delioti" status="gen. nov.">
<emphasis id="B7C4EACBB876FFB8FBF148D2FB2D182D" box="[1027,1257,1786,1809]" italics="true" pageId="5" pageNumber="6">Baronniesia delioti</emphasis>
</taxonomicName>
<emphasis id="B7C4EACBB876FFB8FB0048D1FA51182D" bold="true" box="[1266,1429,1785,1809]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="ACF757B0B876FFB8FB0048D1FA83182D" box="[1266,1351,1785,1809]" pageId="5" pageNumber="6" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
; 8— 
<taxonomicName id="42B04D5AB876FFB8FF324934FDFB180F" authority="Coiffait" authorityName="Coiffait" box="[192,575,1820,1843]" class="Insecta" family="Leiodidae" genus="Paratroglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="B7C4EACBB876FFB8FF324934FE18180F" box="[192,476,1820,1843]" italics="true" pageId="5" pageNumber="6">Paratroglophyes jeanneli</emphasis>
Coiffait
</taxonomicName>
; 9— 
<taxonomicName id="42B04D5AB876FFB8FD864934FBC6180F" authority="Coiffait" authorityName="Coiffait" box="[628,1026,1820,1843]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="aubryi" subSpecies="aubryi">
<emphasis id="B7C4EACBB876FFB8FD864934FC5B180F" box="[628,927,1820,1843]" italics="true" pageId="5" pageNumber="6">Troglophyes aubryi aubryi</emphasis>
Coiffait
</taxonomicName>
; 10— 
<taxonomicName id="42B04D5AB876FFB8FBB74934FABE180F" authority="Abeille" authorityName="Abeille" box="[1093,1402,1820,1843]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gavoyi">
<emphasis id="B7C4EACBB876FFB8FBB74934FAE4180F" box="[1093,1312,1820,1843]" italics="true" pageId="5" pageNumber="6">Troglophyes gavoyi</emphasis>
Abeille
</taxonomicName>
de Perrin; 11— 
<taxonomicName id="42B04D5AB876FFB8FEE94917FD0C186A" authority="Fairmaire" authorityName="Fairmaire" box="[283,712,1855,1878]" class="Insecta" family="Leiodidae" genus="Parvospeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="delarouzeei">
<emphasis id="B7C4EACBB876FFB8FEE94917FD92186A" box="[283,598,1855,1878]" italics="true" pageId="5" pageNumber="6">Parvospeonomus delarouzeei</emphasis>
Fairmaire
</taxonomicName>
; 12— 
<taxonomicName id="42B04D5AB876FFB8FCFB4917FA5C186A" authority="Coiffait" authorityName="Coiffait" box="[777,1432,1855,1878]" class="Insecta" family="Leiodidae" genus="Phacomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="sioberi" subGenus="Phacomorphoides" subSpecies="sioberi">
<emphasis id="B7C4EACBB876FFB8FCFB4917FC6F186A" box="[777,939,1855,1878]" italics="true" pageId="5" pageNumber="6">Phacomorphus</emphasis>
(
<emphasis id="B7C4EACBB876FFB8FC494917FBB9186A" box="[955,1149,1855,1878]" italics="true" pageId="5" pageNumber="6">Phacomorphoides</emphasis>
) 
<emphasis id="B7C4EACBB876FFB8FB7F4917FAED186A" box="[1165,1321,1855,1878]" italics="true" pageId="5" pageNumber="6">sioberi sioberi</emphasis>
(Coiffait)
</taxonomicName>
; 13— 
<taxonomicName id="42B04D5AB876FFB8FF3E494AFD451845" authority="Abeille" authorityName="Abeille" box="[204,641,1890,1913]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="curvipes" subSpecies="subcurvipes">
<emphasis id="B7C4EACBB876FFB8FF3E494AFDE81845" box="[204,556,1890,1913]" italics="true" pageId="5" pageNumber="6">Speonomus curvipes subcurvipes</emphasis>
Abeille
</taxonomicName>
de Perrin; 14— 
<taxonomicName id="42B04D5AB876FFB8FCDE494AFB4A1845" authority="Coiffait." authorityName="Coiffait." box="[812,1166,1890,1913]" class="Insecta" family="Leiodidae" genus="Paraspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B7C4EACBB876FFB8FCDE494AFBE91845" box="[812,1069,1890,1913]" italics="true" pageId="5" pageNumber="6">Paraspeonomus vandeli</emphasis>
Coiffait.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="850F36D9B875FFBBFF344EB0FA811F8E" blockId="6.[151,1437,152,1995]" box="[198,1349,152,178]" pageId="6" pageNumber="7">
There are two main 
<typeStatus id="5A0B887BB875FFBBFE404EB0FE341F8E" box="[434,496,152,178]" pageId="6" pageNumber="7">types</typeStatus>
of lateral stylus of the aedeagus among the rest of Pyrenean Leptodirini:
</paragraph>
<paragraph id="850F36D9B875FFBBFF344E96FBE71D94" blockId="6.[151,1437,152,1995]" pageId="6" pageNumber="7">
1) The 
<taxonomicName id="42B04D5AB875FFBBFED34E97FE751FE4" box="[289,433,191,216]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFED34E97FE751FE4" box="[289,433,191,216]" italics="true" pageId="6" pageNumber="7">Troglophyes</emphasis>
</taxonomicName>
<typeStatus id="5A0B887BB875FFBBFE494E96FE351FE4" box="[443,497,190,216]" pageId="6" pageNumber="7">type</typeStatus>
. The apex of the lateral stylus of the aedeagus with two or (more frequently) three long spines, with a dense penicillum formed by numerous short setae. This penicillum may be inserted in a cavity or directly on the integument of the apex of the stylus (
<figureCitation id="1D8B2A5CB875FFBBFC544F24FBF31E1A" box="[934,1079,268,294]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="6" pageNumber="7">Figs. 10–12</figureCitation>
). In the 
<emphasis id="B7C4EACBB875FFBBFB6A4F24FAEB1E19" box="[1176,1327,268,293]" italics="true" pageId="6" pageNumber="7">speluncarum</emphasis>
group of 
<taxonomicName id="42B04D5AB875FFBBFF654F1AFE021E70" authority="Jeannel, 1908" authorityName="Jeannel" authorityYear="1908" box="[151,454,306,332]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFF654F1AFED91E77" box="[151,285,306,331]" italics="true" pageId="6" pageNumber="7">Speonomus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFED64F1AFE021E70" author="Jeannel" box="[292,454,306,332]" pageId="6" pageNumber="15" refString="Jeannel, R. (1908) Etude sur le genre Speonomus Jeann. (Silphides cavernicoles pyreneens) et sur sa distribution geographique. L'Abeille, 31, 57 - 102." type="journal article" year="1908">Jeannel, 1908</bibRefCitation>
</taxonomicName>
the third spine is often concealed by the penicillum. The following groups have the lateral stylus of the 
<taxonomicName id="42B04D5AB875FFBBFE7F4F71FDE71E4E" box="[397,547,345,370]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFE7F4F71FDE71E4E" box="[397,547,345,370]" italics="true" pageId="6" pageNumber="7">Troglophyes</emphasis>
</taxonomicName>
<typeStatus id="5A0B887BB875FFBBFDDE4F71FDA21E4F" box="[556,614,345,371]" pageId="6" pageNumber="7">type</typeStatus>
: 
<taxonomicName id="42B04D5AB875FFBBFD804F71FC7B1E4F" authority="Jeannel, 1906" authorityName="Jeannel" authorityYear="1906" box="[626,959,345,371]" class="Insecta" family="Leiodidae" genus="Bathysciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFD804F71FCCD1E4E" box="[626,777,345,370]" italics="true" pageId="6" pageNumber="7">Bathysciella</emphasis>
<bibRefCitation id="E1214B28B875FFBBFCE34F71FC7B1E4F" author="Jeannel" box="[785,959,345,371]" pageId="6" pageNumber="15" refString="Jeannel, R. (1906) Contribution la faune cavernicole des Basses-Pyrenees (Col.). Bulletin de la Societe entomologique de France, 11, 22 - 24." type="journal article" year="1906">Jeannel, 1906</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="42B04D5AB875FFBBFC3E4F71FA5C1E4F" authority="Fresneda &amp; Hernando, 1994" authorityName="Fresneda &amp; Hernando" authorityYear="1994" box="[972,1432,345,371]" class="Insecta" family="Leiodidae" genus="Bellesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFC3E4F71FBEB1E4E" box="[972,1071,345,370]" italics="true" pageId="6" pageNumber="7">Bellesia</emphasis>
Fresneda &amp; Hernando, 1994
</taxonomicName>
; 
<taxonomicName id="42B04D5AB875FFBBFF654FA8FE391EA6" authority="Fresneda, 1998" authorityName="Fresneda" authorityYear="1998" box="[151,509,384,410]" class="Insecta" family="Leiodidae" genus="Ceretophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFF654FA8FEF51EA5" box="[151,305,384,409]" italics="true" pageId="6" pageNumber="7">Ceretophyes</emphasis>
<bibRefCitation id="E1214B28B875FFBBFEC84FA8FE391EA6" author="Fresneda" box="[314,509,384,410]" pageId="6" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B7C4EACBB875FFBBFDF84FA8FD6E1EA5" box="[522,682,384,409]" italics="true" pageId="6" pageNumber="7">speluncarum</emphasis>
(
<bibRefCitation id="E1214B28B875FFBBFD4E4FA8FCAF1EA6" author="Jeannel" box="[700,875,384,410]" pageId="6" pageNumber="15" refString="Jeannel, R. (1947) Coleopteres cavernicoles nouveaux avec une etude de la phylogenie des Speonomus. Notes Biospeologiques, 1 (1), 83 - 95." type="journal article" year="1947">Jeannel, 1947</bibRefCitation>
) and 
<taxonomicName id="42B04D5AB875FFBBFC404FA8FB161EA6" authority="Perreau, 2000" authorityName="Perreau" authorityYear="2000" box="[946,1234,384,410]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ehlersi">
<emphasis id="B7C4EACBB875FFBBFC404FA8FBC31EA5" box="[946,1031,384,409]" italics="true" pageId="6" pageNumber="7">ehlersi</emphasis>
(
<bibRefCitation id="E1214B28B875FFBBFBEA4FA8FB0E1EA6" author="Perreau" box="[1048,1226,384,410]" pageId="6" pageNumber="15" refString="Perreau, M. (2000) Catalogue des Coleopteres Leiodidae Cholevinae et Platypsyllinae. Memoires de la Societe entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau, 2000</bibRefCitation>
)
</taxonomicName>
groups, and the subgenus 
<taxonomicName id="42B04D5AB875FFBBFEF94F8FFDA81EFC" authority="Jeannel, 1924" authorityName="Jeannel" authorityYear="1924" box="[267,620,422,448]" class="Insecta" family="Leiodidae" genus="Machaeroscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFEF94F8FFE001EFC" box="[267,452,423,448]" italics="true" pageId="6" pageNumber="7">Machaeroscelis</emphasis>
<bibRefCitation id="E1214B28B875FFBBFE394F8EFDA81EFC" author="Jeannel" box="[459,620,422,448]" pageId="6" pageNumber="15" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63 (1), 1 - 436." type="journal article" year="1924">Jeannel, 1924</bibRefCitation>
</taxonomicName>
of 
<taxonomicName id="42B04D5AB875FFBBFD604F8FFCD31EFC" box="[658,791,423,448]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFD604F8FFCD31EFC" box="[658,791,423,448]" italics="true" pageId="6" pageNumber="7">Speonomus</emphasis>
</taxonomicName>
; 
<taxonomicName id="42B04D5AB875FFBBFCD44F8FFB461EFC" authority="Jeannel, 1908" authorityName="Jeannel" authorityYear="1908" box="[806,1154,422,448]" class="Insecta" family="Leiodidae" genus="Phacomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFCD44F8FFC1C1EFC" box="[806,984,423,448]" italics="true" pageId="6" pageNumber="7">Phacomorphus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC2D4F8EFB461EFC" author="Jeannel" box="[991,1154,422,448]" pageId="6" pageNumber="15" refString="Jeannel, R. (1908) Etude sur le genre Speonomus Jeann. (Silphides cavernicoles pyreneens) et sur sa distribution geographique. L'Abeille, 31, 57 - 102." type="journal article" year="1908">Jeannel, 1908</bibRefCitation>
</taxonomicName>
, including the subgenus 
<taxonomicName id="42B04D5AB875FFBBFF654FE5FDDC1EDB" authority="Dupre, 1989" authorityName="Dupre" authorityYear="1989" box="[151,536,461,487]" class="Insecta" family="Leiodidae" genus="Phacomorphoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFF654FE5FEB21EDA" box="[151,374,461,486]" italics="true" pageId="6" pageNumber="7">Phacomorphoides</emphasis>
<bibRefCitation id="E1214B28B875FFBBFE8C4FE5FDDC1EDB" author="Dupre" box="[382,536,461,487]" pageId="6" pageNumber="14" refString="Dupre, E. (1989) Bathysciinae (Col. Catopidae) des Pyrenees Occidentales: I. Etude du sac interne et morphologie comparee des Speonomus de la region de St-Jean-Pied-de-Port, redefinition du sous-genre Phacomorphus et creation du sous-genre Phacomorphoides, considerations phylogeniques et biogeographiques locales, considerations generales sur la systematique des Bathysciinae. Ikartzaleak, 13, 55 - 99." type="journal article" year="1989">Dupré, 1989</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1D8B2A5CB875FFBBFDD84FE5FD431EDB" box="[554,647,461,487]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="6" pageNumber="7">Fig. 12</figureCitation>
); 
<taxonomicName id="42B04D5AB875FFBBFD6D4FE5FB4B1EDB" authority="Belles &amp; Escola, 1977" authorityName="Belles &amp; Escola" authorityYear="1977" box="[671,1167,461,487]" class="Insecta" family="Leiodidae" genus="Parvospeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFD6D4FE5FCB61EDA" box="[671,882,461,486]" italics="true" pageId="6" pageNumber="7">Parvospeonomus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC884FE5FB4B1EDB" author="Belles" box="[890,1167,461,487]" pageId="6" pageNumber="13" refString="Belles, X. &amp; Escola, O. (1977) Nuevos datos sobre los Speonomus del grupo delarouzeei (Fairmaire, 1860): Parvospeonomus subgen. nov. (Col. Bathysciinae). Speleon, 23, 33 - 37." type="journal article" year="1977">Bellés &amp; Escolà, 1977</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1D8B2A5CB875FFBBFB534FE5FB381EDB" box="[1185,1276,461,487]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
); 
<taxonomicName id="42B04D5AB875FFBBFAE74FE5FEFC1D32" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" class="Insecta" family="Leiodidae" genus="Perriniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFAE74FE5FA581EDA" box="[1301,1436,461,486]" italics="true" pageId="6" pageNumber="7">Perriniella</emphasis>
<bibRefCitation id="E1214B28B875FFBBFF654FDCFEFC1D32" author="Jeannel" box="[151,312,500,526]" pageId="6" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFE824FDCFD9A1D32" authority="Abeille" authorityName="Abeille" box="[368,606,500,526]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFE824FDCFE3A1D31" box="[368,510,500,525]" italics="true" pageId="6" pageNumber="7">Troglophyes</emphasis>
Abeille
</taxonomicName>
de Perrin, 1894 (
<figureCitation id="1D8B2A5CB875FFBBFCDA4FDCFCBB1D32" box="[808,895,500,526]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
). 
<taxonomicName id="42B04D5AB875FFBBFC674FDCFBE11D31" box="[917,1061,500,525]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFC674FDCFBE11D31" box="[917,1061,500,525]" italics="true" pageId="6" pageNumber="7">Troglophyes</emphasis>
</taxonomicName>
is a para- or polyphyletic genus (
<bibRefCitation id="E1214B28B875FFBBFF6D4C32FE671D08" author="Fresneda" box="[159,419,538,564]" pageId="6" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB875FFBBFEEB4C32FE941D0F" box="[281,336,538,563]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2007
</bibRefCitation>
). All the easternmost species of 
<taxonomicName id="42B04D5AB875FFBBFCC44C32FC091D0F" box="[822,973,538,563]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFCC44C32FC091D0F" box="[822,973,538,563]" italics="true" pageId="6" pageNumber="7">Troglophyes</emphasis>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFC2F4C32FB3A1D08" authority="Jeannel, 1906" authorityName="Jeannel" authorityYear="1906" box="[989,1278,538,564]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bedeli">
<emphasis id="B7C4EACBB875FFBBFC2F4C32FB8C1D0F" box="[989,1096,538,563]" italics="true" pageId="6" pageNumber="7">T. bedeli</emphasis>
<bibRefCitation id="E1214B28B875FFBBFBA34C32FB3A1D08" author="Jeannel" box="[1105,1278,538,564]" pageId="6" pageNumber="15" refString="Jeannel, R. (1906) Contribution la faune cavernicole des Basses-Pyrenees (Col.). Bulletin de la Societe entomologique de France, 11, 22 - 24." type="journal article" year="1906">Jeannel, 1906</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B7C4EACBB875FFBBFAF94C32FA591D0F" box="[1291,1437,538,563]" italics="true" pageId="6" pageNumber="7">
T. g. 
<taxonomicName id="42B04D5AB875FFBBFAB94C32FA591D0F" authority="Abeille" authorityName="Abeille" box="[1355,1437,538,563]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gavoyi">gavoyi</taxonomicName>
</emphasis>
Abeille de Perrin, 1894, 
<emphasis id="B7C4EACBB875FFBBFE304C69FDA71D66" box="[450,611,577,602]" italics="true" pageId="6" pageNumber="7">T. g. alluaudi</emphasis>
<bibRefCitation id="E1214B28B875FFBBFD9E4C69FCD11D67" author="Jeannel" box="[620,789,577,603]" pageId="6" pageNumber="15" refString="Jeannel, R. (1911) Biospeologica XIX. Revision des Bathysciinae (Coleopteres, Silphides). Morphologie, distribution geographique, Systematique. Archives de Zoologie experimentale et generale, (5) 7, 1 - 641." type="journal article" year="1911">Jeannel, 1911</bibRefCitation>
, 
<taxonomicName id="42B04D5AB875FFBBFCD04C69FB9F1D67" authority="Chobaut, 1903" authorityName="Chobaut" authorityYear="1903" box="[802,1115,577,603]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ludovici">
<emphasis id="B7C4EACBB875FFBBFCD04C69FC651D66" box="[802,929,577,602]" italics="true" pageId="6" pageNumber="7">T. ludovici</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC5B4C69FB9F1D67" author="Chobaut" box="[937,1115,577,603]" pageId="6" pageNumber="14" refString="Chobaut, A. (1903) Description de deux Coleopteres cavernicoles nouveaux du Midi de la France. Bulletin de la Societe entomologique de France, 8, 263 - 265." type="journal article" year="1903">Chobaut, 1903</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFB654C69FF101DBE" authority="Reitter, 1908" authorityName="Reitter" authorityYear="1908" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="oblongulus">
<emphasis id="B7C4EACBB875FFBBFB654C69FAFF1D66" box="[1175,1339,577,602]" italics="true" pageId="6" pageNumber="7">T. oblongulus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFAB64C69FF101DBE" author="Reitter" pageId="6" pageNumber="15" refString="Reitter, E. (1908) Dichotomische Ubersicht der Blinden Silphiden-Gattungen. Wiener entomologische Zeitung, 27 (2 - 3), 103 - 118." type="journal article" year="1908">Reitter, 1908</bibRefCitation>
</taxonomicName>
, are closely related to 
<taxonomicName id="42B04D5AB875FFBBFE284C40FCE21DBE" authority="Fresneda, 1998" authorityName="Fresneda" authorityYear="1998" box="[474,806,616,642]" class="Insecta" family="Leiodidae" genus="Ceretophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFE284C40FDA81DBD" box="[474,620,616,641]" italics="true" pageId="6" pageNumber="7">Ceretophyes</emphasis>
<bibRefCitation id="E1214B28B875FFBBFD804C40FCE21DBE" author="Fresneda" box="[626,806,616,642]" pageId="6" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFCAC4C40FB4D1DBE" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" box="[862,1161,616,642]" class="Insecta" family="Leiodidae" genus="Perriniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFCAC4C40FC1B1DBD" box="[862,991,616,641]" italics="true" pageId="6" pageNumber="7">Perriniella</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC144C40FB4D1DBE" author="Jeannel" box="[998,1161,616,642]" pageId="6" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
, unlike the 
<taxonomicName id="42B04D5AB875FFBBFAE34C40FA581DBD" box="[1297,1436,616,641]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFAE34C40FA581DBD" box="[1297,1436,616,641]" italics="true" pageId="6" pageNumber="7">Antrocharis</emphasis>
</taxonomicName>
group of genera and 
<taxonomicName id="42B04D5AB875FFBBFE7B4CA7FDD41D94" box="[393,528,655,680]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="aubryi" subSpecies="aubryi">
<emphasis id="B7C4EACBB875FFBBFE7B4CA7FDD41D94" box="[393,528,655,680]" italics="true" pageId="6" pageNumber="7">T. a. aubryi</emphasis>
</taxonomicName>
and 
<emphasis id="B7C4EACBB875FFBBFDBB4CA7FCCD1D94" box="[585,777,655,680]" italics="true" pageId="6" pageNumber="7">T. a. vallierensis</emphasis>
of the central Pyrenees.
</paragraph>
<paragraph id="850F36D9B875FFBBFF344C9DFBD71BA3" blockId="6.[151,1437,152,1995]" pageId="6" pageNumber="7">
2) The 
<taxonomicName id="42B04D5AB875FFBBFEDA4C9DFE721DF2" box="[296,438,693,718]" class="Insecta" family="Leiodidae" genus="Trocharanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFEDA4C9DFE721DF2" box="[296,438,693,718]" italics="true" pageId="6" pageNumber="7">Trocharanis</emphasis>
</taxonomicName>
<typeStatus id="5A0B887BB875FFBBFE374C9DFE3F1DF3" box="[453,507,693,719]" pageId="6" pageNumber="7">type</typeStatus>
. The apex of the lateral stylus of the aedeagus is with three long spines (sometimes five). One can be longer than the other two (
<figureCitation id="1D8B2A5CB875FFBBFCDE4CF4FC471DCA" box="[812,899,732,758]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="6" pageNumber="7">Fig. 13</figureCitation>
: 
<taxonomicName id="42B04D5AB875FFBBFC7D4CF4FAD61DC9" box="[911,1298,732,757]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="curvipes" subSpecies="subcurvipes">
<emphasis id="B7C4EACBB875FFBBFC7D4CF4FAD61DC9" box="[911,1298,732,757]" italics="true" pageId="6" pageNumber="7">Speonomus curvipes subcurvipes</emphasis>
</taxonomicName>
(Abeille de Perrin, 1878)), or two longer than the third one, all three of different size (
<figureCitation id="1D8B2A5CB875FFBBFBE44D2AFBB41C20" box="[1046,1136,770,796]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
: 
<taxonomicName id="42B04D5AB875FFBBFB8E4D2AFEF91C7F" authority="Coiffait, 1952" authorityName="Coiffait" authorityYear="1952" class="Insecta" family="Leiodidae" genus="Paraspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B7C4EACBB875FFBBFB8E4D2AFA581C27" box="[1148,1436,770,795]" italics="true" pageId="6" pageNumber="7">Paraspeonomus vandeli</emphasis>
<bibRefCitation id="E1214B28B875FFBBFF654D01FEF91C7F" author="Coiffait" box="[151,317,809,835]" pageId="6" pageNumber="14" refString="Coiffait, H. (1952) Note sur les Bathysciites. Description d'un genre nouveau et de formes nouvelles. Notes Biospeologiques, 7, 41 - 44." type="journal article" year="1952">Coiffait, 1952</bibRefCitation>
</taxonomicName>
) or all subequal. A dense penicillum formed by numerous long setae inserted on the side or on a more or less developed membranous lamina is always present; in some species the long setae of the penicillum look like a pedunculus or thin extension of the membranous lamina (e.g. in 
<taxonomicName id="42B04D5AB875FFBBFB494D5FFF1F1C8B" authority="Foures, 1954" authorityName="Foures" authorityYear="1954" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carrerei">
<emphasis id="B7C4EACBB875FFBBFB494D5FFAFE1CAC" box="[1211,1338,887,912]" italics="true" pageId="6" pageNumber="7">S. carrerei</emphasis>
<bibRefCitation id="E1214B28B875FFBBFAB14D5EFF1F1C8B" author="Foures" pageId="6" pageNumber="14" refString="Foures, H. (1954) Diagnose preliminaire de nouveaux Coleopteres cavernicoles de la region du mont Valier (Ariege). Bulletin de la Societe linneenne de Lyon, 23, 3 - 5." type="journal article" year="1954">Fourès, 1954</bibRefCitation>
</taxonomicName>
). This group is formed by the genus 
<taxonomicName id="42B04D5AB875FFBBFD5E4DB5FBF41C8B" authority="Coiffait, 1952" authorityName="Coiffait" authorityYear="1952" box="[684,1072,925,951]" class="Insecta" family="Leiodidae" genus="Paraspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFD5E4DB5FCB01C8A" box="[684,884,925,950]" italics="true" pageId="6" pageNumber="7">Paraspeonomus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC8F4DB5FBF41C8B" author="Coiffait" box="[893,1072,925,951]" pageId="6" pageNumber="14" refString="Coiffait, H. (1952) Note sur les Bathysciites. Description d'un genre nouveau et de formes nouvelles. Notes Biospeologiques, 7, 41 - 44." type="journal article" year="1952">Coiffait, 1952</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFBCC4DB5FB0F1C8A" box="[1086,1227,925,950]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFBCC4DB5FB0F1C8A" box="[1086,1227,925,950]" italics="true" pageId="6" pageNumber="7">Speonomus</emphasis>
</taxonomicName>
of 
<emphasis id="B7C4EACBB875FFBBFB054DB5FAA31C8A" box="[1271,1383,925,950]" italics="true" pageId="6" pageNumber="7">pyreneus</emphasis>
and 
<emphasis id="B7C4EACBB875FFBBFF654DECFEDF1CE1" box="[151,283,964,989]" italics="true" pageId="6" pageNumber="7">zophosinus</emphasis>
groups (
<bibRefCitation id="E1214B28B875FFBBFE764DECFDE31CE2" author="Jeannel" box="[388,551,964,990]" pageId="6" pageNumber="15" refString="Jeannel, R. (1947) Coleopteres cavernicoles nouveaux avec une etude de la phylogenie des Speonomus. Notes Biospeologiques, 1 (1), 83 - 95." type="journal article" year="1947">Jeannel 1947</bibRefCitation>
), subgenus 
<taxonomicName id="42B04D5AB875FFBBFD464DECFBE21CE2" authority="Coiffait, 1959" authorityName="Coiffait" authorityYear="1959" box="[692,1062,964,990]" class="Insecta" family="Leiodidae" genus="Metaspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFD464DECFCB11CE1" box="[692,885,964,989]" italics="true" pageId="6" pageNumber="7">Metaspeonomus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC8C4DECFBE21CE2" author="Coiffait" box="[894,1062,964,990]" pageId="6" pageNumber="14" refString="Coiffait, H. (1959) Note sur les Bathysciitae de la region pyreneenne et de Catalogne. Annales de Speleologie, 14 (1 - 2), 159 - 179." type="journal article" year="1959">Coiffait, 1959</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFB934DECFA5C1CE2" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" box="[1121,1432,964,990]" class="Insecta" family="Leiodidae" genus="Trocharanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFB934DECFB361CE1" box="[1121,1266,964,989]" italics="true" pageId="6" pageNumber="7">Trocharanis</emphasis>
<bibRefCitation id="E1214B28B875FFBBFB0B4DECFA5C1CE2" author="Reitter" box="[1273,1432,964,990]" pageId="6" pageNumber="15" refString="Reitter, E. (1885) Bestimmungs-Tabellen der europaischen Coleopteren, 12 Necrophaga. Verhandlungen des naturforschenden Vereines in Brunn, [1884], 23, 3 - 122." type="journal article" year="1885">Reitter, 1885</bibRefCitation>
</taxonomicName>
. All the genera of the southern side of central Pyrenees and coastal ridges of Catalonia (
<collectingCountry id="FDA77649B875FFBBFB514DC2FB2D1B38" box="[1187,1257,1002,1028]" name="Spain" pageId="6" pageNumber="7">Spain</collectingCountry>
) belong to this group: 
<taxonomicName id="42B04D5AB875FFBBFF1C4A39FDF61B17" authority="Fresneda, 2000" authorityName="Fresneda" authorityYear="2000" box="[238,562,1041,1067]" class="Insecta" family="Leiodidae" genus="Lagariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFF1C4A39FEA91B16" box="[238,365,1041,1066]" italics="true" pageId="6" pageNumber="7">Lagariella</emphasis>
Fresneda, 2000
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFDCD4A39FBB31B17" authority="Fresneda, Hernando &amp; Lagar, 1994" authorityName="Fresneda, Hernando &amp; Lagar" authorityYear="1994" box="[575,1143,1041,1067]" class="Insecta" family="Leiodidae" genus="Naspunius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFDCD4A39FD7A1B16" box="[575,702,1041,1066]" italics="true" pageId="6" pageNumber="7">Naspunius</emphasis>
<bibRefCitation id="E1214B28B875FFBBFD354A39FBB31B17" author="Fresneda" box="[711,1143,1041,1067]" pageId="6" pageNumber="14" refString="Fresneda, J., Hernando, C. &amp; Lagar, A. (1994) Contribucio al coneixement de Speonomus eseranus Lagar, 1974 (Coleoptera, Cholevidae, Leptodirinae) i descripcio del subgenere Naspunius nov. Bolleti de la Societat d'Historia natural de les Balears, 37, 109 - 116." type="journal article" year="1994">Fresneda, Hernando &amp; Lagar, 1994</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFB764A39FF131B6E" authority="Fresneda, 1998" authorityName="Fresneda" authorityYear="1998" class="Insecta" family="Leiodidae" genus="Pallaresiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFB764A39FADB1B16" box="[1156,1311,1041,1066]" italics="true" pageId="6" pageNumber="7">Pallaresiella</emphasis>
<bibRefCitation id="E1214B28B875FFBBFADA4A39FF131B6E" author="Fresneda" pageId="6" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFF164A10FDEC1B6E" authority="Fresneda, 1998" authorityName="Fresneda" authorityYear="1998" box="[228,552,1080,1106]" class="Insecta" family="Leiodidae" genus="Salgadoia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFF164A10FEA41B6D" box="[228,352,1080,1105]" italics="true" pageId="6" pageNumber="7">Salgadoia</emphasis>
<bibRefCitation id="E1214B28B875FFBBFE9B4A10FDEC1B6E" author="Fresneda" box="[361,552,1080,1106]" pageId="6" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFDC44A10FC421B6E" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" box="[566,902,1080,1106]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFDC44A10FD151B6D" box="[566,721,1080,1105]" italics="true" pageId="6" pageNumber="7">Speonomites</emphasis>
<bibRefCitation id="E1214B28B875FFBBFD2B4A10FC421B6E" author="Jeannel" box="[729,902,1080,1106]" pageId="6" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFC664A10FB291B6E" authority="Fresneda, 1998" authorityName="Fresneda" authorityYear="1998" box="[916,1261,1080,1106]" class="Insecta" family="Leiodidae" genus="Stygiophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFC664A10FBE21B6D" box="[916,1062,1080,1105]" italics="true" pageId="6" pageNumber="7">Stygiophyes</emphasis>
<bibRefCitation id="E1214B28B875FFBBFBDC4A10FB291B6E" author="Fresneda" box="[1070,1261,1080,1106]" pageId="6" pageNumber="14" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFB094A10FEFC1B44" authority="Jeannel, 1924" authorityName="Jeannel" authorityYear="1924" class="Insecta" family="Leiodidae" genus="Trapezodirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFB094A10FA591B6D" box="[1275,1437,1080,1105]" italics="true" pageId="6" pageNumber="7">Trapezodirus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFF654A76FEFC1B44" author="Jeannel" box="[151,312,1118,1144]" pageId="6" pageNumber="15" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63 (1), 1 - 436." type="journal article" year="1924">Jeannel, 1924</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFE9D4A77FD051B44" authority="Reitter, 1908" authorityName="Reitter" authorityYear="1908" box="[367,705,1118,1144]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFE9D4A77FDE71B44" box="[367,547,1119,1144]" italics="true" pageId="6" pageNumber="7">Troglocharinus</emphasis>
<bibRefCitation id="E1214B28B875FFBBFDD84A76FD051B44" author="Reitter" box="[554,705,1118,1144]" pageId="6" pageNumber="15" refString="Reitter, E. (1908) Dichotomische Ubersicht der Blinden Silphiden-Gattungen. Wiener entomologische Zeitung, 27 (2 - 3), 103 - 118." type="journal article" year="1908">Reitter, 1908</bibRefCitation>
</taxonomicName>
(except the three taxa of the subgenus 
<taxonomicName id="42B04D5AB875FFBBFB774A77FF161BA3" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" class="Insecta" family="Leiodidae" genus="Antrocharidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFB774A77FAF21B44" box="[1157,1334,1119,1144]" italics="true" pageId="6" pageNumber="7">Antrocharidius</emphasis>
<bibRefCitation id="E1214B28B875FFBBFACE4A76FF161BA3" author="Jeannel" pageId="6" pageNumber="15" refString="Jeannel, R. (1910) Biospeologica XIV. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale, 45 (1), 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
with a secondary loss of penicillum but with the membranous lamina).
</paragraph>
<paragraph id="850F36D9B875FFBBFF344A84FDF81AE8" blockId="6.[151,1437,152,1995]" pageId="6" pageNumber="7">
Dilated antennae in male of 
<taxonomicName id="42B04D5AB875FFBBFDE44A84FD601BF9" box="[534,676,1196,1221]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFDE44A84FD601BF9" box="[534,676,1196,1221]" italics="true" pageId="6" pageNumber="7">Baronniesia</emphasis>
</taxonomicName>
is a character shared with 
<taxonomicName id="42B04D5AB875FFBBFC104A84FF161BD0" authority="Belles &amp; Deliot, 1983" authorityName="Belles &amp; Deliot" authorityYear="1983" class="Insecta" family="Leiodidae" genus="Josettekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="angelinae">
<emphasis id="B7C4EACBB875FFBBFC104A84FB171BF9" box="[994,1235,1196,1221]" italics="true" pageId="6" pageNumber="7">Josettekia angelinae</emphasis>
<bibRefCitation id="E1214B28B875FFBBFB294A84FF161BD0" author="Belles" pageId="6" pageNumber="13" refString="Belles, X. &amp; Deliot, P. (1983) Nouveaux Bathysciinae (Coleopteres Catopidae) des Pyrenees francaises et espagnoles. Memoires de Biospeologie, 10, 237 - 243." type="journal article" year="1983">Bellés &amp; Déliot, 1983</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFEFE4AFBFD131BD0" authority="Coiffait, 1952" authorityName="Coiffait" authorityYear="1952" box="[268,727,1234,1260]" class="Insecta" family="Leiodidae" genus="Paraspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B7C4EACBB875FFBBFEFE4AFBFDE21BD0" box="[268,550,1235,1260]" italics="true" pageId="6" pageNumber="7">Paraspeonomus vandeli</emphasis>
<bibRefCitation id="E1214B28B875FFBBFDDD4AFAFD131BD0" author="Coiffait" box="[559,727,1234,1260]" pageId="6" pageNumber="14" refString="Coiffait, H. (1952) Note sur les Bathysciites. Description d'un genre nouveau et de formes nouvelles. Notes Biospeologiques, 7, 41 - 44." type="journal article" year="1952">Coiffait, 1952</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="42B04D5AB875FFBBFD114AFBFC101BD0" box="[739,980,1235,1260]" class="Insecta" family="Leiodidae" genus="Josettekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="angelinae">
<emphasis id="B7C4EACBB875FFBBFD114AFBFC101BD0" box="[739,980,1235,1260]" italics="true" pageId="6" pageNumber="7">Josettekia angelinae</emphasis>
</taxonomicName>
has the 9th (especially) and 10th male antennomeres dilated. However, in 
<taxonomicName id="42B04D5AB875FFBBFDB94AD1FD011A2E" box="[587,709,1273,1298]" class="Insecta" family="Leiodidae" genus="Josettekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFDB94AD1FD011A2E" box="[587,709,1273,1298]" italics="true" pageId="6" pageNumber="7">Josettekia</emphasis>
</taxonomicName>
this is not a generic character, as the second species of the genus, 
<taxonomicName id="42B04D5AB875FFBBFF1F4B08FDA61A06" authority="Bolivar, 1921" authorityName="Bolivar" authorityYear="1921" box="[237,610,1312,1338]" class="Insecta" family="Leiodidae" genus="Josettekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mendizabali">
<emphasis id="B7C4EACBB875FFBBFF1F4B08FE5A1A05" box="[237,414,1312,1337]" italics="true" pageId="6" pageNumber="7">J. mendizabali</emphasis>
(
<bibRefCitation id="E1214B28B875FFBBFE5C4B08FD9D1A06" author="Bolivar" box="[430,601,1312,1338]" pageId="6" pageNumber="14" refString="Bolivar, C. (1921) Silphidae. In: Bolivar, C. &amp; Jeannel, R. Coleopteros cavernicolas nuevos de las provincias vascas. Boletin de la Real Sociedad espanola de Historia natural, tomo del Centenario, 50, 509 - 539." type="journal article" year="1921">Bolívar, 1921</bibRefCitation>
)
</taxonomicName>
, does not show any dilatation of the male antennae. Both species of 
<taxonomicName id="42B04D5AB875FFBBFF654B6FFEC81A5C" box="[151,268,1351,1376]" class="Insecta" family="Leiodidae" genus="Josettekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFF654B6FFEC81A5C" box="[151,268,1351,1376]" italics="true" pageId="6" pageNumber="7">Josettekia</emphasis>
</taxonomicName>
have the apex of the lateral stylus with four short spines and lack the two inverted “V” feather-like structures in the inner sac of aedeagus. In 
<taxonomicName id="42B04D5AB875FFBBFD764B45FD331ABA" box="[644,759,1389,1414]" class="Insecta" family="Leiodidae" genus="Paraspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B7C4EACBB875FFBBFD764B45FD331ABA" box="[644,759,1389,1414]" italics="true" pageId="6" pageNumber="7">P. vandeli</emphasis>
</taxonomicName>
only the fourth antennal article of male is dilated, but the lateral stylus of the aedeagus has a membranous lamina and a penicillum, and the inner sac lacks the two inverted “V” feather-like structures.
</paragraph>
<paragraph id="850F36D9B875FFBBFF344BC8FBFC180C" blockId="6.[151,1437,152,1995]" pageId="6" pageNumber="7">
<emphasis id="B7C4EACBB875FFBBFF344BC8FE181AC6" bold="true" box="[198,476,1504,1530]" pageId="6" pageNumber="7">Phylogenetic analyses.</emphasis>
<taxonomicName id="42B04D5AB875FFBBFE164BC9FDB61AC6" box="[484,626,1505,1530]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B7C4EACBB875FFBBFE164BC9FDB61AC6" box="[484,626,1505,1530]" italics="true" pageId="6" pageNumber="7">Baronniesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACF757B0B875FFBBFD8B4BC9FD0C1AC7" box="[633,712,1505,1531]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
and the genus 
<taxonomicName id="42B04D5AB875FFBBFC854BC9FC221AC6" box="[887,998,1505,1530]" class="Insecta" family="Leiodidae" genus="Gesciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFC854BC9FC221AC6" box="[887,998,1505,1530]" italics="true" pageId="6" pageNumber="7">Gesciella</emphasis>
</taxonomicName>
(omitted from the previous analyses) were added to the data matrix of 
<bibRefCitation id="E1214B28B875FFBBFDE84820FCCD191E" author="Fresneda" box="[538,777,1544,1570]" pageId="6" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB875FFBBFD7E4820FD7B191D" box="[652,703,1544,1569]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2007
</bibRefCitation>
(Appendix 1: this paper) with the aim of including both genera in a formal phylogenetic analysis. See 
<bibRefCitation id="E1214B28B875FFBBFD4E4806FC771974" author="Fresneda" box="[700,947,1582,1608]" pageId="6" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB875FFBBFCDD4807FCA71974" box="[815,867,1583,1608]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2007
</bibRefCitation>
: 335–344 for the list of the 34 characters and their states. For a list of studied species see Appendix 
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<bibRefCitation id="E1214B28B875FFBBFC57487DFB601953" author="Fresneda" box="[933,1188,1621,1647]" pageId="6" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB875FFBBFBEE487DFB9E1952" box="[1052,1114,1621,1646]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2007
</bibRefCitation>
. For composition of groups of terminal taxa sharing identical character states and lumped together for the purposes of analysis see Appendix 
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this paper. Despite the fact that three new characters were added to the matrix of 
<bibRefCitation id="E1214B28B875FFBBFB03488AFF2319DF" pageId="6" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article">
Fresneda 
<emphasis id="B7C4EACBB875FFBBFA90488BFA5F1980" box="[1378,1435,1699,1724]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(2007)
</bibRefCitation>
(see below), this could not help to resolve relationships between 
<taxonomicName id="42B04D5AB875FFBBFBF548E1FB5E19DE" box="[1031,1178,1737,1762]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFBF548E1FB5E19DE" box="[1031,1178,1737,1762]" italics="true" pageId="6" pageNumber="7">Baronniesia</emphasis>
</taxonomicName>
and the most closely related taxa (the 
<taxonomicName id="42B04D5AB875FFBBFE9E48D8FDC41835" box="[364,512,1776,1801]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFE9E48D8FDC41835" box="[364,512,1776,1801]" italics="true" pageId="6" pageNumber="7">Antrocharis</emphasis>
</taxonomicName>
group). In the new character matrix (Appendix 1) 
<taxonomicName id="42B04D5AB875FFBBFB7748D8FAD91835" box="[1157,1309,1776,1801]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFB7748D8FAD91835" box="[1157,1309,1776,1801]" italics="true" pageId="6" pageNumber="7">Baronniesia</emphasis>
</taxonomicName>
shares all character states with 
<taxonomicName id="42B04D5AB875FFBBFE7F493FFDDC180C" box="[397,536,1815,1840]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFE7F493FFDDC180C" box="[397,536,1815,1840]" italics="true" pageId="6" pageNumber="7">Antrocharis</emphasis>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFDD5493FFD51180C" box="[551,661,1815,1840]" class="Insecta" family="Leiodidae" genus="Gesciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFDD5493FFD51180C" box="[551,661,1815,1840]" italics="true" pageId="6" pageNumber="7">Gesciella</emphasis>
</taxonomicName>
, 
<taxonomicName id="42B04D5AB875FFBBFD56493FFCAC180C" box="[676,872,1815,1840]" class="Insecta" family="Leiodidae" genus="Paratroglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFD56493FFCAC180C" box="[676,872,1815,1840]" italics="true" pageId="6" pageNumber="7">Paratroglophyes</emphasis>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB875FFBBFC53493FFBF5180C" box="[929,1073,1815,1840]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB875FFBBFC53493FFBF5180C" box="[929,1073,1815,1840]" italics="true" pageId="6" pageNumber="7">Troglophyes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="850F36D9B875FFBAFF344915FD311FE4" blockId="6.[151,1437,152,1995]" lastBlockId="7.[151,1437,152,835]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Character 35. 
<emphasis id="B7C4EACBB875FFBBFE984915FD5F186A" box="[362,667,1853,1878]" italics="true" pageId="6" pageNumber="7">Lateral styles of aedeagus</emphasis>
: very thin along their entire length (0) and always shorter than the median lobe (
<figureCitation id="1D8B2A5CB875FFBBFEC8494CFE6A1842" box="[314,430,1892,1918]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,269,1501,1525]" captionTargetBox="[375,1213,477,1469]" captionTargetId="figure@3.[365,1221,465,1477]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Aedeagus of Baronniesia delioti gen. n. sp. n. 3 — dorsal view; 4 — lateral view." httpUri="https://zenodo.org/record/185600/files/figure.png" pageId="6" pageNumber="7">Figs. 3–4</figureCitation>
); or strong along their entire length (1), with the apex more or less thickened and of length ranging from shorter to longer than the median lobe. Coded as missing data (?) for the species with character states 26 (1), 27 (1), 28 (0), 29 (0), 30 (1) and 31 (1), 
<emphasis id="B7C4EACBB875FFBBFC8F4999FC7A18F6" box="[893,958,1969,1994]" italics="true" pageId="6" pageNumber="7">sensu</emphasis>
<bibRefCitation id="E1214B28B875FFBBFC374999FB0F18F7" box="[965,1227,1969,1995]" pageId="6" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article">
Fresneda 
<emphasis id="B7C4EACBB875FFBBFBC44999FBAD18F6" box="[1078,1129,1969,1994]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. (2007)
</bibRefCitation>
. These species have the apex of the lateral stylus of the aedeagus with two to five spines; when they have three, there is always a penicillum, cavity or membranous lamina.
</paragraph>
<paragraph id="850F36D9B874FFBAFF344ECDFC3A1E4F" blockId="7.[151,1437,152,835]" pageId="7" pageNumber="8">
Character 36. 
<emphasis id="B7C4EACBB874FFBAFE824ECDFB9E1FC2" box="[368,1114,229,254]" italics="true" pageId="7" pageNumber="8">Apical part of the median region of the inner sac of aedeagus</emphasis>
: with two very sclerotized pieces forming an inverted “V” located above FDM and FVM (
<figureCitation id="1D8B2A5CB874FFBAFC8F4F24FC011E1A" box="[893,965,268,294]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
: IVP) (1); without inverted “V”, or with other structures (0). In the species with the inner sac of the aedeagus lacking copulating armature (
<bibRefCitation id="E1214B28B874FFBAFAE64F1AFF3F1E4F" author="Fresneda" pageId="7" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB874FFBAFA754F1AFF6A1E4E" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2007
</bibRefCitation>
: character 16, state 0) character 36 was coded as missing data (?).
</paragraph>
<paragraph id="850F36D9B874FFBAFF344FA8FB511EDB" blockId="7.[151,1437,152,835]" pageId="7" pageNumber="8">
Character 37. 
<emphasis id="B7C4EACBB874FFBAFE834FA8FCF81EA5" box="[369,828,384,409]" italics="true" pageId="7" pageNumber="8">Basal region of inner sac of aedeagus</emphasis>
: apical piece of PY (FAPY) above the connection nodule (NL) very thin and elongated (
<figureCitation id="1D8B2A5CB874FFBAFD824F8EFD7A1EFC" box="[624,702,422,448]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,276,1409,1433]" captionTargetBox="[441,1145,181,1397]" captionTargetId="figure@4.[441,1146,181,1398]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri="https://zenodo.org/record/185601/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
: FAPY) (1) or short and conical (0). In species without PY (
<bibRefCitation id="E1214B28B874FFBAFF6C4FE5FE571EDB" author="Fresneda" box="[158,403,461,487]" pageId="7" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB874FFBAFEE04FE5FE811EDA" box="[274,325,461,486]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2007
</bibRefCitation>
: character 17, state 0) character 37 was coded as missing data (?).
</paragraph>
<paragraph id="850F36D9B874FFBAFF344FDCFAF31D94" blockId="7.[151,1437,152,835]" pageId="7" pageNumber="8">
The extended data matrix was analysed using the same methods as in 
<bibRefCitation id="E1214B28B874FFBAFBF34FDCFACD1D32" box="[1025,1289,500,526]" pageId="7" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article">
Fresneda 
<emphasis id="B7C4EACBB874FFBAFB814FDCFB631D31" box="[1139,1191,500,525]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. (2007)
</bibRefCitation>
: the shortest trees were heuristically searched in PAUP 4.0b10 (
<bibRefCitation id="E1214B28B874FFBAFD014C32FC681D08" author="Swofford" box="[755,940,538,564]" pageId="7" pageNumber="15" refString="Swofford, D. L. (2002) PAU P *. Phylogenetic Analysis using Parsimony (* and other methods), Version 4.0 b 10. Sinauer Associates, Sunderland, MA." type="book chapter" year="2002">Swofford 2002</bibRefCitation>
) with 10,000 tree-bisection-reconnection (
<collectingCountry id="FDA77649B874FFBAFF6C4C69FF161D67" box="[158,210,577,603]" name="Tuvalu" pageId="7" pageNumber="8">TBR</collectingCountry>
) replicas, swapping on all multiple starting trees, and saving all of them. To obtain a higher resolution, data were reweighted successively according to the rescaled consistency index (
<bibRefCitation id="E1214B28B874FFBAFB884C40FA871DBE" author="Swofford" box="[1146,1347,616,642]" pageId="7" pageNumber="15" refString="Swofford, D. L. (2002) PAU P *. Phylogenetic Analysis using Parsimony (* and other methods), Version 4.0 b 10. Sinauer Associates, Sunderland, MA." type="book chapter" year="2002">Swofford, 2002</bibRefCitation>
), and a heuristic search conducted on the initial set of the shortest trees (see 
<bibRefCitation id="E1214B28B874FFBAFC484CA6FB6D1D94" author="Fresneda" box="[954,1193,654,680]" pageId="7" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article" year="2007">
Fresneda 
<emphasis id="B7C4EACBB874FFBAFBDE4CA7FB9B1D94" box="[1068,1119,655,680]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2007
</bibRefCitation>
for details).
</paragraph>
<paragraph id="850F36D9B874FFBAFF344C9DFD3F1C7F" blockId="7.[151,1437,152,835]" pageId="7" pageNumber="8">
The topology of the strict consensus of the 330 reweighted trees (consistency index 0.67, retention index 0.9) was identical to that of 
<bibRefCitation id="E1214B28B874FFBAFE1A4CF4FD2B1DCA" box="[488,751,732,758]" pageId="7" pageNumber="14" refString="Fresneda, J., Salgado, J. M. &amp; Ribera, I. (2007) Phylogeny of Western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32, 332 - 358." type="journal article">
Fresneda 
<emphasis id="B7C4EACBB874FFBAFDAE4CF4FD541DC9" box="[604,656,732,757]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. (2007)
</bibRefCitation>
(
<figureCitation id="1D8B2A5CB874FFBAFCF24CF4FC931DCA" box="[768,855,732,758]" captionStart="FIGURE 17" captionStartId="9.[151,255,1891,1915]" captionTargetBox="[233,1355,186,1872]" captionTargetId="figure@9.[233,1360,180,1876]" captionTargetPageId="9" captionText="FIGURE 17. Phylogram of the strict consensus of the 330 trees obtained after successive reweighting of the characters (in the tree, the character weights were reset to unity, so the scale of the branch lengths is uniform across the tree). Taxa with identical character states were lumped in a single terminal taxon except the genera of the Antrocharis group (see Appendix 2)." httpUri="https://zenodo.org/record/185604/files/figure.png" pageId="7" pageNumber="8">Fig. 17</figureCitation>
) and, thus, was not affected by addition of three new characters. 
<taxonomicName id="42B04D5AB874FFBAFEAE4D2AFE0A1C27" box="[348,462,770,795]" class="Insecta" family="Leiodidae" genus="Gesciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB874FFBAFEAE4D2AFE0A1C27" box="[348,462,770,795]" italics="true" pageId="7" pageNumber="8">Gesciella</emphasis>
</taxonomicName>
and 
<taxonomicName id="42B04D5AB874FFBAFDF94D2AFD591C27" box="[523,669,770,795]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB874FFBAFDF94D2AFD591C27" box="[523,669,770,795]" italics="true" pageId="7" pageNumber="8">Baronniesia</emphasis>
</taxonomicName>
appear in the same polytomy with the remaining genera of the 
<taxonomicName id="42B04D5AB874FFBAFF654D01FEE61C7E" box="[151,290,809,834]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB874FFBAFF654D01FEE61C7E" box="[151,290,809,834]" italics="true" pageId="7" pageNumber="8">Antrocharis</emphasis>
</taxonomicName>
group, as they share all character states.
</paragraph>
</subSubSection>
</treatment>
</document>