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<document id="89CB7BA19A3E98609EEADE790C56B32A" ID-DOI="10.5281/zenodo.213360" ID-GBIF-Dataset="b1a33963-aa99-478f-8b76-50c13ada801c" ID-ISSN="1175-5326" ID-Zenodo-Dep="213360" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460549395632" checkinUser="plazi" docAuthor="Sathiandran, Nithya &amp; Sabu, Thomas K." docDate="2012" docId="0D07755309360F58FF3702DAFEA5AE68" docLanguage="en" docName="zt03526p058.pdf" docOrigin="Zootaxa 3526" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus militaris" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="F13E0D2B09370F59FFA0036DFD7CAA5A" masterDocTitle="New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India" masterLastPageNumber="58" masterPageNumber="53" pageNumber="54" updateTime="1698318061882" updateUser="plazi">
<mods:mods id="FF32378D81F54AD0C17112088A982FFC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CE4873BDC80965C85598D1F4F9C73913">
<mods:title id="07BC683DCF46AE32A5028D3EE6EB8A90">New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India</mods:title>
</mods:titleInfo>
<mods:name id="D1D11FDFDB7C2AE2CE863BE3D0A88C00" type="personal">
<mods:role id="B0D9B0364C3DA56F56DB9710843910AA">
<mods:roleTerm id="621759318505383AD224FD8D865C1522">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19541E9A7BB02158E25229E3E5BD49D9">Sathiandran, Nithya</mods:namePart>
</mods:name>
<mods:name id="E2C7A621596BA4338BA95BD23BA7B0BF" type="personal">
<mods:role id="C42524A635855E130DC85B23AF069224">
<mods:roleTerm id="F8FEEDFCB4D713DE0504C5FF8502936A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FB0CA4D4396A9C69504713629C0C83D">Sabu, Thomas K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3D988CD0B2D5A7B5C2DE3D2EA3162D60">text</mods:typeOfResource>
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<mods:titleInfo id="534760906041CD3BADB8851D2078DD07">
<mods:title id="31687BDD6C13305C8227E23DA35DE252">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FD86A8798C2FF67DB517879089938194">
<mods:date id="A749FB8CAE4730F011F4F0D38C08C365">2012</mods:date>
<mods:detail id="155ECFC5DCC2787CD9C5933D33C23ED1" type="volume">
<mods:number id="AE14ADF9B73D28ED2116038AD52D7842">3526</mods:number>
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<mods:start id="5376B3C317BBA90966F681BF1B5D04C0">53</mods:start>
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<mods:classification id="FAE38F0AE354732367B073EDDBBD3ACC">journal article</mods:classification>
<mods:identifier id="5770656D5A3A947EB44319D087CE88A6" type="DOI">10.5281/zenodo.213360</mods:identifier>
<mods:identifier id="7324C66BF2761CA9257611BD45B5C411" type="GBIF-Dataset">b1a33963-aa99-478f-8b76-50c13ada801c</mods:identifier>
<mods:identifier id="D05A0A796326C45766ED52382F688271" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C4357BC106EB545F3B933C22225D2142" type="Zenodo-Dep">213360</mods:identifier>
</mods:mods>
<treatment id="0D07755309360F58FF3702DAFEA5AE68" ID-DOI="http://doi.org/10.5281/zenodo.6175208" ID-GBIF-Taxon="119529156" ID-Zenodo-Dep="6175208" LSID="urn:lsid:plazi:treatment:0D07755309360F58FF3702DAFEA5AE68" httpUri="http://treatment.plazi.org/id/0D07755309360F58FF3702DAFEA5AE68" lastPageNumber="54" pageId="1" pageNumber="54">
<subSubSection id="CDB497CE09360F58FF3702DAFE89A924" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="8511C44509360F58FF3702DAFF2AAB8B" blockId="1.[151,989,439,499]" box="[151,598,439,465]" pageId="1" pageNumber="54">
<heading id="DE59732909360F58FF3702DAFF2AAB8B" bold="true" box="[151,598,439,465]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
<emphasis id="B7DA185709360F58FF3702DAFF2AAB8B" bold="true" box="[151,598,439,465]" pageId="1" pageNumber="54">
<taxonomicName id="42AEBFC609360F58FF3702DAFCDBAB8B" ID-CoL="7XNJS" box="[151,423,439,465]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="B7DA185709360F58FF3702DAFCDBAB8B" bold="true" box="[151,423,439,465]" italics="true" pageId="1" pageNumber="54">Onthophagus militaris</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="8511C44509360F58FF3702B7FEA1ABA8" blockId="1.[151,989,439,499]" box="[151,989,474,499]" pageId="1" pageNumber="54">
(as defined by
<bibRefCitation id="E13FB9B409360F58FE9C02B6FCBBABA8" author="Arrow" box="[316,455,474,499]" pageId="1" pageNumber="54" refString="Arrow, G. J. (1931) The Fauna of British India including Ceylon and Burma, Coleoptera: Lamellicornia (Coprinae), Taylor and Francis, London. 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
,
<bibRefCitation id="E13FB9B409360F58FE7302B7FF02ABA8" author="Balthasar" box="[467,638,474,498]" pageId="1" pageNumber="54" refString="Balthasar, V. (1963) Monographic der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region (Coleoptera: Lamellicornia). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague. Volume I, 391 pp, Vo l u m e I I, 627 pp." type="book" year="1963">Balthasar 1963</bibRefCitation>
, Schoolmeesters &amp;
<bibRefCitation id="E13FB9B409360F58FCFD02B7FEA9ABA8" author="Sabu" box="[861,981,474,498]" pageId="1" pageNumber="54" refString="Schoolmeester, P. &amp; Sabu, K. T. (2006) A new Onthophagus s pecies from Kerala, India (Coleoptera: Scarabaeidae: Scarabaeinae). Phegea, 34, 73 - 75." type="journal article" year="2006">Sabu 2006</bibRefCitation>
)
</paragraph>
<paragraph id="8511C44509360F58FF37014FFE33A804" blockId="1.[151,1436,546,1075]" pageId="1" pageNumber="54">
Medium size (
<quantity id="425669A009360F58FE9E014FFCDFA860" box="[318,419,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="54" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
), uniformly greenish blue or head and pronotum dark coppery or metallic green. Elytra black with yellow spots or yellow with black spots or patches.
</paragraph>
<paragraph id="8511C44509360F58FF670106FCD1A948" blockId="1.[151,1436,546,1075]" pageId="1" pageNumber="54">
Upper surface shiny and body clothed with setae. Clypeus deeply or shallowly emarginate anteriorly or truncate with anterior margin reflexed. Head with ocular lobes gently rounded. Pronotum coarsely and closely punctate, granulate, or punctate behind granules. Elytra finely striate, bearing scattered punctures or granules in double rows. Pygidium distinctly punctate, clothed with setae. Metasternum smooth in the middle with lateral punctures (
<bibRefCitation id="E13FB9B409360F58FEB20197FCDCA948" author="Arrow" box="[274,416,762,786]" pageId="1" pageNumber="54" refString="Arrow, G. J. (1931) The Fauna of British India including Ceylon and Burma, Coleoptera: Lamellicornia (Coprinae), Taylor and Francis, London. 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
).
</paragraph>
<paragraph id="8511C44509360F58FF670073FE89A924" blockId="1.[151,1436,546,1075]" pageId="1" pageNumber="54">
<emphasis id="B7DA185709360F58FF670073FF6CA96D" bold="true" box="[199,528,798,823]" pageId="1" pageNumber="54">Geographical distribution:</emphasis>
<collectingCountry id="FDB984D509360F58FD810073FF26A96C" box="[545,602,798,822]" name="India" pageId="1" pageNumber="54">India</collectingCountry>
(Jharkand: Chota Nagpur; Karnataka: Bangalore; Kerala: Nilambur, Ranipuram, Shendurney, Silent valley, Thekkady, Wayanad; Orissa; Tamilnadu: Anamalai hills, Nilgiri hills, Yercaud);
<collectingCountry id="FDB984D509360F58FEAB000BFC01A924" box="[267,381,870,894]" name="Sri Lanka" pageId="1" pageNumber="54">Sri Lanka</collectingCountry>
: Kandy (
<bibRefCitation id="E13FB9B409360F58FE40000BFF11A924" author="Arrow" box="[480,621,870,894]" pageId="1" pageNumber="54" refString="Arrow, G. J. (1931) The Fauna of British India including Ceylon and Burma, Coleoptera: Lamellicornia (Coprinae), Taylor and Francis, London. 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
,
<bibRefCitation id="E13FB9B409360F58FDD8000BFE58A924" author="Balthasar" box="[632,804,870,894]" pageId="1" pageNumber="54" refString="Balthasar, V. (1963) Monographic der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region (Coleoptera: Lamellicornia). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague. Volume I, 391 pp, Vo l u m e I I, 627 pp." type="book" year="1963">Balthasar 1963</bibRefCitation>
,
<bibRefCitation id="E13FB9B409360F58FC8F000BFE95A924" author="Sabu" box="[815,1001,870,895]" pageId="1" pageNumber="54" refString="Sabu, T. K., Nithya, S. &amp; Vinod K. V. (2011) Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa, 2830, 29 - 38." type="journal article" year="2011">
Sabu
<emphasis id="B7DA185709360F58FCCE0005FEDAA924" box="[878,934,870,895]" italics="true" pageId="1" pageNumber="54">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CDB497CE09360F58FF6700E6FEA5AE68" pageId="1" pageNumber="54" type="discussion">
<paragraph id="8511C44509360F58FF6700E6FEA5AE68" blockId="1.[151,1436,546,1075]" pageId="1" pageNumber="54">
<emphasis id="B7DA185709360F58FF6700E6FC41A9F9" bold="true" box="[199,317,907,931]" pageId="1" pageNumber="54">Remarks:</emphasis>
All species in this group are brightly colored, setose, and are rare in collections. They are predominantly high altitude species as seven of the nine species are found in the montane rain forest ecoregion of the moist south Western Ghats in south
<collectingCountry id="FDB984D509360F58FDF300BFFFF1A9B0" box="[595,653,978,1002]" name="India" pageId="1" pageNumber="54">India</collectingCountry>
and
<collectingCountry id="FDB984D509360F58FD6300BFFE48A9B0" box="[707,820,978,1002]" name="Sri Lanka" pageId="1" pageNumber="54">Sri Lanka</collectingCountry>
; four species (
<taxonomicName id="42AEBFC609360F58FC7600BEF9CDA9B0" authority="Gill" authorityName="Gill" box="[982,1201,978,1003]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="lemniscatus">
<emphasis id="B7DA185709360F58FC7600BEF900A9B1" box="[982,1148,979,1003]" italics="true" pageId="1" pageNumber="54">O. lemniscatus</emphasis>
Gill
</taxonomicName>
,
<taxonomicName id="42AEBFC609360F58FB1C00BEF8E9A9B1" authority="Arrow" authorityName="Arrow" box="[1212,1429,979,1003]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="madoqua">
<emphasis id="B7DA185709360F58FB1C00BEF83BA9B1" box="[1212,1351,979,1003]" italics="true" pageId="1" pageNumber="54">O. madoqua</emphasis>
Arrow
</taxonomicName>
,
<taxonomicName id="42AEBFC609360F58FF37009AFF1BAE54" authority="Schoolmeesters &amp; Sabu" authorityName="Schoolmeesters &amp; Sabu" box="[151,615,1014,1038]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="devagiriensis">
<emphasis id="B7DA185709360F58FF37009AFC33AE54" box="[151,335,1014,1038]" italics="true" pageId="1" pageNumber="54">O. devagiriensis</emphasis>
Schoolmeesters &amp; Sabu
</taxonomicName>
, and
<taxonomicName id="42AEBFC609360F58FD03009AFED9AE54" authority="Nithya &amp; Sabu" authorityName="Nithya &amp; Sabu" box="[675,933,1014,1038]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="tnai">
<emphasis id="B7DA185709360F58FD03009AFF8DAE54" box="[675,753,1015,1038]" italics="true" pageId="1" pageNumber="54">O. tnai</emphasis>
Nithya &amp; Sabu
</taxonomicName>
) are endemics to the Western Ghats and one species (
<taxonomicName id="42AEBFC609360F58FF580776FF79AE68" authority="Boucomont" authorityName="Boucomont" box="[248,517,1050,1075]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="B7DA185709360F58FF580776FC05AE69" box="[248,377,1051,1075]" italics="true" pageId="1" pageNumber="54">O. militaris</emphasis>
Boucomont
</taxonomicName>
) endemic to
<collectingCountry id="FDB984D509360F58FD360777FE7FAE68" box="[662,771,1050,1074]" name="Sri Lanka" pageId="1" pageNumber="54">Sri Lanka</collectingCountry>
(
<bibRefCitation id="E13FB9B409360F58FCB20777FEB2AE68" author="Sabu" box="[786,974,1050,1075]" pageId="1" pageNumber="54" refString="Sabu, T. K., Nithya, S. &amp; Vinod K. V. (2011) Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa, 2830, 29 - 38." type="journal article" year="2011">
Sabu
<emphasis id="B7DA185709360F58FCF30771FEF7AE69" box="[851,907,1051,1075]" italics="true" pageId="1" pageNumber="54">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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