484 lines
53 KiB
XML
484 lines
53 KiB
XML
<document ID-CLB-Dataset="5518" ID-DOI="http://dx.doi.org/10.3897/phytokeys.179.65990" ID-GBIF-Dataset="ebc27064-6919-4f59-87e1-f773500962f1" ID-PMC="PMC8289813" ID-Pensoft-Pub="1314-2003-179-99" ID-Pensoft-UUID="F9007B81A3025C47AF6B5D7A9EF49B0E" ID-PubMed="34316268" ModsDocID="1314-2003-179-99" checkinTime="1626109678468" checkinUser="pensoft" docAuthor="Clark, John L. & Jost, Lou" docDate="2021" docId="D3BAC67DCB745A49AA88BB9A52AE8F15" docLanguage="en" docName="PhytoKeys 179: 99-110" docOrigin="PhytoKeys 179" docPubDate="2021-07-12" docSource="http://dx.doi.org/10.3897/phytokeys.179.65990" docTitle="Kohleria anisophylla Wiehler 1978" docType="treatment" docVersion="6" id="F9007B81A3025C47AF6B5D7A9EF49B0E" lastPageNumber="99" masterDocId="F9007B81A3025C47AF6B5D7A9EF49B0E" masterDocTitle="New circumscriptions add two northern Andean species to Kohleria (Gesneriaceae)" masterLastPageNumber="110" masterPageNumber="99" pageNumber="99" updateTime="1732939505993" updateUser="ExternalLinkService">
|
|
<mods:mods id="40960E0DDDF9672E27AD1E16891E7199" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="417AD55072F63907AB2473F867B29FF5">
|
|
<mods:title id="8153F2B08C50F5D8706AAE9F9F058E83">New circumscriptions add two northern Andean species to Kohleria (Gesneriaceae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="63FD2D24E13EBA81FED5003CF8617FB9" type="personal">
|
|
<mods:role id="D7FA801DB617C99F8FA18EC174D9F384">
|
|
<mods:roleTerm id="27A19DDD3AF3874DD99F0945AA850A89">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="108DBCCC81817C3B8ECC6849FE4F3AB8">Clark, John L.</mods:namePart>
|
|
<mods:nameIdentifier id="F560F424661EF3235670DF3D7E0DC758" type="ORCID">https://orcid.org/0000-0002-1414-6380</mods:nameIdentifier>
|
|
<mods:affiliation id="B520F25E718C53FAF9BBAFE61BDFC319">Science Department, The Lawrenceville School, Lawrenceville, NJ 08648, USA</mods:affiliation>
|
|
<mods:nameIdentifier id="12B5E62A49E2B85E8BFB42947105FEC0" type="email">jclark@lawrenceville.org</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="9E5C08FFC293552592E8A032FBB92B78" type="personal">
|
|
<mods:role id="C2105873FC9C3D1996A02DA600AFA3E4">
|
|
<mods:roleTerm id="E933B1E7513CB1250C425223D649C37C">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="BB0BB24BF2D0AAA04158843864B16FCE">Jost, Lou</mods:namePart>
|
|
<mods:affiliation id="06CD71F748A78C6DC96E13E2134D7BA7">Lou Jost, Fundacion EcoMinga, Banos, Tungurahua, Ecuador</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource id="4F4F72DC0BEFF017F28DE9E03156D853">text</mods:typeOfResource>
|
|
<mods:relatedItem id="DBA4DD8B150A542BF07A5F0524801D5B" type="host">
|
|
<mods:titleInfo id="D12877E599F4096DC1E640554DE8CB83">
|
|
<mods:title id="22D5965D57891CB74E010F28A2DCD3EA">PhytoKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="10D5A71EC0C9238FDC098E976C135891">
|
|
<mods:date id="8AADDC9AD872C1CED1CDB9F27B6D4FA1">2021</mods:date>
|
|
<mods:detail id="0586EB33CA8E76E3994F52086D67AF45" type="pubDate">
|
|
<mods:number id="3F5AC415B4C8C18AA8E0497938BA0BDE">2021-07-12</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="F712B56F4DACF102EC727A6063EB1EF4" type="volume">
|
|
<mods:number id="D8E15E9F44210B1F39FB3A09B5BD8127">179</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="BD60750441BE06B90E2391871AC0FDF9" unit="page">
|
|
<mods:start id="E13C6C8373D63A6089931C53C7784C46">99</mods:start>
|
|
<mods:end id="EA7FB4FB604C474F8AA2D05753DBA2B1">110</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="67B46427948FF906CA3D3D00AA149368">
|
|
<mods:url id="F47ED1C26E5E8A0C163F214027D4D6FD">http://dx.doi.org/10.3897/phytokeys.179.65990</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="5665E1CED71B7EB81E9092A13E9C3240">journal article</mods:classification>
|
|
<mods:identifier id="B3E677674BE1CF08CDD551C834C96F49" type="DOI">http://dx.doi.org/10.3897/phytokeys.179.65990</mods:identifier>
|
|
<mods:identifier id="7F05D081F59D68DDD175530C66497630" type="Pensoft-Pub">1314-2003-179-99</mods:identifier>
|
|
<mods:identifier id="CBFD596FE5DD2C37C839F400894FEE14" type="Pensoft-UUID">F9007B81A3025C47AF6B5D7A9EF49B0E</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="D3BAC67DCB745A49AA88BB9A52AE8F15" ID-GBIF-Taxon="183501758" LSID="urn:lsid:plazi:treatment:D3BAC67DCB745A49AA88BB9A52AE8F15" httpUri="http://treatment.plazi.org/id/D3BAC67DCB745A49AA88BB9A52AE8F15" lastPageNumber="99" pageId="0" pageNumber="99" scope_family="Gesneriaceae">
|
|
<subSubSection id="F8D3B84866A8766FE8C9AE652F11A432" pageId="0" pageNumber="99" type="nomenclature">
|
|
<paragraph id="835CC28D940742E9E63AABBADC86BF62" pageId="0" pageNumber="99">
|
|
<taxonomicName id="0492C7A3B5078204DC0CAA8CAD249A3D" ID-CoL="3RC4N" LSID="D3BAC67D-CB74-5A49-AA88-BB9A52AE8F15" authority="(Fritsch) Wiehler" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">Kohleria anisophylla (Fritsch) Wiehler</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2654A9637B46C59231F7A1FE63E76577" pageId="0" pageNumber="99" type="reference_group">
|
|
<paragraph id="433E003363902E1D0F9FAE4170B7F156" pageId="0" pageNumber="99">
|
|
<taxonomicName id="83339F7900FCCA2503C8B23C1C0A6429" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">Kohleria anisophylla</taxonomicName>
|
|
(Fritsch) Wiehler.
|
|
</paragraph>
|
|
<paragraph id="F314C51C66CCC905A6647F01595299EA" pageId="0" pageNumber="99">
|
|
<taxonomicName id="C782A979E2BE4CCF3C63BD5261890D6D" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">Kohleria anisophylla</taxonomicName>
|
|
(Fritsch) Wiehler, Selbyana 5: 62. 1978. Type: Based on
|
|
<taxonomicName id="FEC298B26A12C11CE4567FBD0BB8F7A1" authorityName="Fritsch" authorityYear="1913" class="Magnoliopsida" family="Gesneriaceae" genus="Diastema" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diastema anisophyllum" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophyllum">Diastema anisophyllum</taxonomicName>
|
|
Fritsch.
|
|
</paragraph>
|
|
<paragraph id="EF8649B6E87FBE8005D668D46D5AE265" pageId="0" pageNumber="99">
|
|
<taxonomicName id="5D349B5CEC0A95129BEFAA4B28F3CB60" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa subsp. var. var. anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="villosa" subSpecies="var." variety="anisophylla">Kohleria villosa var. anisophylla</taxonomicName>
|
|
(Fritsch) Kvist & Skog, Smithsonian Contr. Bot. 79: 70. 1992. Type: Based on
|
|
<taxonomicName id="AF48C0C6E1BB4E138A822A62265C8B43" authorityName="Fritsch" authorityYear="1913" class="Magnoliopsida" family="Gesneriaceae" genus="Diastema" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diastema anisophyllum" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophyllum">Diastema anisophyllum</taxonomicName>
|
|
Fritsch. Basionym.
|
|
</paragraph>
|
|
<paragraph id="17880FBE07F012AA4F6C457257458236" pageId="0" pageNumber="99">
|
|
<taxonomicName id="19513F7D5D87BB0D040E6B9E0BB658CC" authorityName="Fritsch" authorityYear="1913" class="Magnoliopsida" family="Gesneriaceae" genus="Diastema" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diastema anisophyllum" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophyllum">Diastema anisophyllum</taxonomicName>
|
|
Fritsch, Bot. Jahrb. Syst. 50: 408. 1913 (
|
|
<normalizedToken id="496ABC87EE75F9FEB67389BD6E30C4DB" originalValue="“1914”">"1914"</normalizedToken>
|
|
). Type: Colombia. [
|
|
<normalizedToken id="04683900310F3325943EAE926CB6CC9A" originalValue="Nariño">Narino</normalizedToken>
|
|
] Piedra Ancha, West of Andes of Tuquerres,
|
|
<emphasis id="ACA79C2ACA5B5BE9090E22985BEDEC91" italics="true" pageId="0" pageNumber="99">F.C. Lehmann 5843</emphasis>
|
|
(B, holotype not extant, lectotype K000509983, designated by
|
|
<bibRefCitation id="C0A024E5EDA4BA9D1B66FDFEC680EA7F" author="Wiehler, H" journalOrPublisher="Selbyana" pageId="0" pageNumber="99" pagination="61 - 93" refId="B10" refString="Wiehler, H, 1978. Miscellaneous transfers and new species of neotropical Gesneriaceae. Selbyana 5 (1): 61 - 93" title="Miscellaneous transfers and new species of neotropical Gesneriaceae." volume="5" year="1978">Wiehler (1978</bibRefCitation>
|
|
: 62), isolectotype K000509984).
|
|
</paragraph>
|
|
<paragraph id="B3ABE63DC37BF59060E615F5D3B5B428" pageId="0" pageNumber="99">
|
|
<taxonomicName id="2C379EFED3B456042CBE32687F4CA80F" class="Magnoliopsida" family="Gesneriaceae" genus="Nematanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nematanthus erianthus" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="erianthus">Nematanthus erianthus</taxonomicName>
|
|
Bentham, Pl. Hartw: 231. 1846. Type: Ecuador. Pichincha: Quito towards Nanegal,
|
|
<emphasis id="46A1BA386578B1BA9495F94A81E8804E" italics="true" pageId="0" pageNumber="99">Hartweg s.n.</emphasis>
|
|
(holotype K000509985).
|
|
</paragraph>
|
|
<paragraph id="96DA44B8FF06E61917A4D74BB9A4EF7F" pageId="0" pageNumber="99">
|
|
<taxonomicName id="DEC019C02D9D125B42576CD834C82A94" class="Magnoliopsida" family="Gesneriaceae" genus="Columnea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Columnea eriantha" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="eriantha">Columnea eriantha</taxonomicName>
|
|
(Bentham) Hanstein, Linnaea 34: 391. 1865. Type: Based on
|
|
<taxonomicName id="E649B0E43C9CB739F89E118E012C0BA1" class="Magnoliopsida" family="Gesneriaceae" genus="Nematanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nematanthus erianthus" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="erianthus">Nematanthus erianthus</taxonomicName>
|
|
Fritsch.
|
|
</paragraph>
|
|
<paragraph id="19965269E59099D70BAFEDC683FAE2EE" pageId="0" pageNumber="99">
|
|
<taxonomicName id="19AE8B59DDAAC6049ECA746E3E875E2D" authorityName="Fritsch" authorityYear="1913" class="Magnoliopsida" family="Gesneriaceae" genus="Diastema" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diastema anisophyllum subsp. var. var. quitense" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="anisophyllum" subSpecies="var." variety="quitense">Diastema anisophyllum Fritsch var. quitense</taxonomicName>
|
|
Fritsch. Bot. Jahrb. Syst. 50(4): 408. 1913 (
|
|
<normalizedToken id="3E5F364C1DD4BA2D9A6F9820B79FE448" originalValue="“1914”">"1914"</normalizedToken>
|
|
). Type: Ecuador. [Pichincha] Quito,
|
|
<emphasis id="35C305538AA0F9C6BA124D454471B4AE" italics="true" pageId="0" pageNumber="99">W. Jameson s.n.</emphasis>
|
|
(holotype W).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A83CC5270310E8B7CF0A10195AB0A0FF" pageId="0" pageNumber="99" type="remarks">
|
|
<paragraph id="CE807DA500E690FDFAC093DAF3D1ADFB" pageId="0" pageNumber="99">Remarks.</paragraph>
|
|
<paragraph id="91318754BF38FFE15ED327C2EFB25DE1" pageId="0" pageNumber="99">
|
|
<taxonomicName id="E08A6F783C20EEF43DDDFDD91261B956" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="33CBF9EDCB939D71A4E4318128607A1C" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation id="DFF9B06DB4885CC479A70135E22A2D1F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Kohleria anisophylla (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A, B Clark et al. 10981 C Clark 10948 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure3" httpUri="https://binary.pensoft.net/fig/564819" pageId="0" pageNumber="99">3</figureCitation>
|
|
) was previously recognized by
|
|
<bibRefCitation id="F6F961B9EBC47311C3B3FDC6409376D6" DOI="https://doi.org/10.5479/si.0081024X.79" author="Kvist, L" journalOrPublisher="Smithsonian Contributions to Botany" pageId="0" pageNumber="99" pagination="1 - 83" refId="B4" refString="Kvist, L, Skog, LE, 1992. Revision of Kohleria (Gesneriaceae). Smithsonian Contributions to Botany 79 (79): 1 - 83, DOI: https://doi.org/10.5479/si.0081024X.79" title="Revision of Kohleria (Gesneriaceae)." url="https://doi.org/10.5479/si.0081024X.79" volume="79" year="1992">Kvist and Skog (1992)</bibRefCitation>
|
|
as a variety of
|
|
<taxonomicName id="A003EB52D36BBA6A34A775684A1FC4CF" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="9E9D5F3F7A64C2DC30588C01E2FAC516" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation id="8ABEB868DBFC4216EE81BBBCC03184AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Kohleria villosa (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D erect herbaceous habit with isophyllous leaves (A Clark et al. 14295 B Clark 7331 C Clark et al. 7400 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure4" httpUri="https://binary.pensoft.net/fig/564820" pageId="0" pageNumber="99">4</figureCitation>
|
|
). The strongly anisophyllous leaves and dorsiventral shoots (Fig.
|
|
<figureCitation id="EE7933F6E897E5C991C5C381F5AAE767" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Kohleria anisophylla (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A, B Clark et al. 10981 C Clark 10948 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure3" httpUri="https://binary.pensoft.net/fig/564819" pageId="0" pageNumber="99">3D</figureCitation>
|
|
) are more similar to
|
|
<taxonomicName id="2C665F74B5C4AABAFBA7E9D26AD86A79" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria hypertrichosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="hypertrichosa">
|
|
<emphasis id="4287450BAA0E5867121D72D675E7ED4D" italics="true" pageId="0" pageNumber="99">Kohleria hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation id="63CEB7709C0F806B591B4E309C9A0B54" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Kohleria hypertrichosa J. L. Clark & L. E. Skog A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A Clark et al. 15900 B Clark 6539 C Clark et al. 10310 D Clark et al. 14942). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure5" httpUri="https://binary.pensoft.net/fig/564821" pageId="0" pageNumber="99">5D</figureCitation>
|
|
) than
|
|
<taxonomicName id="EFEBFD01423C71AA8CFFD72F611703F4" lsidName="K. villosa" pageId="0" pageNumber="99" rank="species" species="villosa">
|
|
<emphasis id="A9FC44A16BF8C2D4C95AFC9BB4132F8C" italics="true" pageId="0" pageNumber="99">K. villosa</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation id="DB89FAF853B21DE1641C4DFD01A3FEDF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Kohleria villosa (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D erect herbaceous habit with isophyllous leaves (A Clark et al. 14295 B Clark 7331 C Clark et al. 7400 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure4" httpUri="https://binary.pensoft.net/fig/564820" pageId="0" pageNumber="99">4D</figureCitation>
|
|
). All three species are found on the northwestern Andean slopes of Ecuador. Only
|
|
<taxonomicName id="144B76015DFCD0BA3B8B9C428E03DC40" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="A80A431B7FAFCB732E24C04E48226E27" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
is documented from Colombia (
|
|
<normalizedToken id="606AA0BDC79A06436D0CA6BB1DD3C5A8" originalValue="Nariño">Narino</normalizedToken>
|
|
department).
|
|
<bibRefCitation id="40F08015DF3EF9794E01C4248A477BF4" author="Wiehler, H" journalOrPublisher="Selbyana" pageId="0" pageNumber="99" pagination="61 - 93" refId="B10" refString="Wiehler, H, 1978. Miscellaneous transfers and new species of neotropical Gesneriaceae. Selbyana 5 (1): 61 - 93" title="Miscellaneous transfers and new species of neotropical Gesneriaceae." volume="5" year="1978">Wiehler (1978)</bibRefCitation>
|
|
made the combination
|
|
<taxonomicName id="F981BDEEDCC81CD7642AE4A7BC1E0D9D" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="8C2D39D15AED68BE1AB0DDAF0C7B6B78" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
and recognized it at the rank of species.
|
|
<bibRefCitation id="EF7D60962312574409150D0E190C9B2F" DOI="https://doi.org/10.5479/si.0081024X.79" author="Kvist, L" journalOrPublisher="Smithsonian Contributions to Botany" pageId="0" pageNumber="99" pagination="1 - 83" refId="B4" refString="Kvist, L, Skog, LE, 1992. Revision of Kohleria (Gesneriaceae). Smithsonian Contributions to Botany 79 (79): 1 - 83, DOI: https://doi.org/10.5479/si.0081024X.79" title="Revision of Kohleria (Gesneriaceae)." url="https://doi.org/10.5479/si.0081024X.79" volume="79" year="1992">Kvist and Skog (1992)</bibRefCitation>
|
|
recognized this taxon as
|
|
<taxonomicName id="6AE7DC5C5E1318D5C253168572A489B0" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa subsp. var. var. anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="villosa" subSpecies="var." variety="anisophylla">Kohleria villosa var. anisophylla</taxonomicName>
|
|
. Based on limited material,
|
|
<bibRefCitation id="8EC387CE9E1138C898DAAEFE65887A04" author="Wiehler, H" journalOrPublisher="Selbyana" pageId="0" pageNumber="99" pagination="61 - 93" refId="B10" refString="Wiehler, H, 1978. Miscellaneous transfers and new species of neotropical Gesneriaceae. Selbyana 5 (1): 61 - 93" title="Miscellaneous transfers and new species of neotropical Gesneriaceae." volume="5" year="1978">Wiehler (1978)</bibRefCitation>
|
|
cited the type (
|
|
<emphasis id="88486E73A9D0A75D3FF6667BCA907943" italics="true" pageId="0" pageNumber="99">F.C. Lehmann 5843</emphasis>
|
|
) and a recently collected specimen from Ecuador (
|
|
<emphasis id="0EE12CEFDDD797881CF27CEDE8D87CCB" italics="true" pageId="0" pageNumber="99">C. Luer & A. Hirtz 2672</emphasis>
|
|
).
|
|
<bibRefCitation id="2BD6C78AEB965CADFE9774933B4694A0" DOI="https://doi.org/10.5479/si.0081024X.79" author="Kvist, L" journalOrPublisher="Smithsonian Contributions to Botany" pageId="0" pageNumber="99" pagination="1 - 83" refId="B4" refString="Kvist, L, Skog, LE, 1992. Revision of Kohleria (Gesneriaceae). Smithsonian Contributions to Botany 79 (79): 1 - 83, DOI: https://doi.org/10.5479/si.0081024X.79" title="Revision of Kohleria (Gesneriaceae)." url="https://doi.org/10.5479/si.0081024X.79" volume="79" year="1992">Kvist and Skog (1992)</bibRefCitation>
|
|
cited the same Ecuadorian collection and mentioned the study of eleven additional specimens. This species is common along the northwestern slopes of the Ecuadorian Andes, especially along the old road between Quito and Santo Domingo where many of the images were taken for Figure
|
|
<figureCitation id="AAABBF756D1CB60D1586BE532CF64F6E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Kohleria anisophylla (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A, B Clark et al. 10981 C Clark 10948 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure3" httpUri="https://binary.pensoft.net/fig/564819" pageId="0" pageNumber="99">3</figureCitation>
|
|
. Outlined here are characters to differentiate
|
|
<taxonomicName id="19FC8AA3D2545135FB13893DB85C6043" lsidName="K. anisophylla" pageId="0" pageNumber="99" rank="species" species="anisophylla">
|
|
<emphasis id="3284A371A345AE2F3AA0FAE7C1DDE60C" italics="true" pageId="0" pageNumber="99">K. anisophylla</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="04368A601E074E38E5AA8332770DBF8F" lsidName="K. villosa" pageId="0" pageNumber="99" rank="species" species="villosa">
|
|
<emphasis id="EA1EA3B9309F2E26CE084E1C976B023B" italics="true" pageId="0" pageNumber="99">K. villosa</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="374AB17D65A6A484018A19CB240F57E3" lsidName="K. hypertrichosa" pageId="0" pageNumber="99" rank="species" species="hypertrichosa">
|
|
<emphasis id="A6F0134B0AE6A0C532CD9E73FE66FE4C" italics="true" pageId="0" pageNumber="99">K. hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
(see Table
|
|
<tableCitation id="FD2E7803F63132802D5B722076368BE0" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological differences and general distribution of Kohleria anisophylla, K. villosa, and K. hypertrichosa." httpUri="http://table.plazi.org/id/27F1FFD5AEB783645E5ABB717EEAE8E8" pageId="0" pageNumber="99" tableUuid="27F1FFD5AEB783645E5ABB717EEAE8E8">2</tableCitation>
|
|
for a comparison of characters that are discussed below).
|
|
</paragraph>
|
|
<caption id="27F1FFD5AEB783645E5ABB717EEAE8E8" ID-Table-UUID="27F1FFD5AEB783645E5ABB717EEAE8E8" httpUri="http://table.plazi.org/id/27F1FFD5AEB783645E5ABB717EEAE8E8" pageId="0" pageNumber="99" start="Table 2" startId="T2">
|
|
<paragraph id="B3394BDC17F9E3345B88B13D694CCAB3" pageId="0" pageNumber="99">
|
|
<emphasis id="29DF01735295CA7F1E2B1710DB19B51B" bold="true" pageId="0" pageNumber="99">Table 2.</emphasis>
|
|
Morphological differences and general distribution of
|
|
<taxonomicName id="349A68D85975F283E1EB047A5F33434E" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="53716B43A303C575531BF97BA0001212" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="EBEA952E68E078C9FEDE53C05979F175" lsidName="K. villosa" pageId="0" pageNumber="99" rank="species" species="villosa">
|
|
<emphasis id="89D0E89AE8B1D926D0A75E4C7D2816D3" italics="true" pageId="0" pageNumber="99">K. villosa</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="45E3D1E6FCF6447053E1A01B3CDFD8F9" lsidName="K. hypertrichosa" pageId="0" pageNumber="99" rank="species" species="hypertrichosa">
|
|
<emphasis id="91F03973D8C5D73430F9801116D35D41" italics="true" pageId="0" pageNumber="99">K. hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="7814FDB92B5D4517A5A09D4C0E6C3908" pageId="0" pageNumber="99">
|
|
<table id="44C9483F1B073AF1163E73FADDF17651" pageId="0" pageNumber="99">
|
|
<tr id="D12F70A2742168A9856049C49D42BD6F" pageId="0" pageNumber="99">
|
|
<th id="5D32D186848D5ADA4FF169CB971F96B5" colspan="1" isEmpty="true" pageId="0" pageNumber="99" rowspan="1">-</th>
|
|
<th id="EABE3A95E69B4CD18C79B8AEE8CAB838" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<taxonomicName id="278F82F82528582DAAAA0D0076DA9763" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="3A4C21FC0901FDB9DEC1B9AB4D25716F" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
(Fritsch) Wiehler
|
|
</th>
|
|
<th id="801E49F96A76D8391DC5D99631FBE855" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<taxonomicName id="831431FD062A266937EBAB0AB97AAB82" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="F6DBC9906D1F59056604069C2F2805E8" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
(Fritsch) Wiehler
|
|
</th>
|
|
<th id="DC99C2EABC8EFA7F5166F7B5F92A3E2F" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<taxonomicName id="6F0B7941C746C0C8D047C8E1DD3B240E" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria hypertrichosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="hypertrichosa">
|
|
<emphasis id="65DB4CB661069BAF07DB907E15EB695A" italics="true" pageId="0" pageNumber="99">Kohleria hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
J.L. Clark & L.E. Skog
|
|
</th>
|
|
</tr>
|
|
<tr id="16C8296AA11E2843CB682E345B0FBA38" pageId="0" pageNumber="99">
|
|
<td id="052331B0E3B6F51C8E426210C6CFF0AF" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<emphasis id="594C78F44DE562C1570B55CEE4310427" bold="true" pageId="0" pageNumber="99">Habit</emphasis>
|
|
</td>
|
|
<td id="E1994AB8A11E98B416E31041C66BDF16" colspan="1" pageId="0" pageNumber="99" rowspan="1">facultative epiphyte</td>
|
|
<td id="B8567D0BC21824F0BFB2B7CF8E8EC362" colspan="1" pageId="0" pageNumber="99" rowspan="1">terrestrial</td>
|
|
<td id="7A2FC8CA9DBA560E8FAD4A193D821B70" colspan="1" pageId="0" pageNumber="99" rowspan="1">facultative epiphyte</td>
|
|
</tr>
|
|
<tr id="538AC7F31E74F6904C9BE9F44793F0DE" pageId="0" pageNumber="99">
|
|
<td id="417E304CDDE1A496C285526C31957DB1" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<emphasis id="231DBBB418DE6E72DDC94CF8BDE14E1D" bold="true" pageId="0" pageNumber="99">Shoots</emphasis>
|
|
</td>
|
|
<td id="403E32DF4E7231D904C4000CF5AD5239" colspan="1" pageId="0" pageNumber="99" rowspan="1">dorsiventral</td>
|
|
<td id="BB043A4BDC2178A3E9828AA7A9831043" colspan="1" pageId="0" pageNumber="99" rowspan="1">erect</td>
|
|
<td id="F9BAD4B2DCA1335FE820A89ED9CEE13A" colspan="1" pageId="0" pageNumber="99" rowspan="1">dorsiventral</td>
|
|
</tr>
|
|
<tr id="76E409538800BF2D2AE20AADD6D3C307" pageId="0" pageNumber="99">
|
|
<td id="8D7FBFC156A31995391AA4A2CD83D46C" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<emphasis id="C7CFAE2132C302684A687319E4F70071" bold="true" pageId="0" pageNumber="99">Relative leaf size</emphasis>
|
|
</td>
|
|
<td id="EF582A09164410C7B3EF21AB0691B818" colspan="1" pageId="0" pageNumber="99" rowspan="1">anisophyllous</td>
|
|
<td id="9CB6C7538D136DE509888989E907DA72" colspan="1" pageId="0" pageNumber="99" rowspan="1">isophyllous</td>
|
|
<td id="4E8AF3B3B65BFD34190D37CFFCD4147A" colspan="1" pageId="0" pageNumber="99" rowspan="1">anisophyllous</td>
|
|
</tr>
|
|
<tr id="1EAE28D1894836EBBA450A1E66DB1347" pageId="0" pageNumber="99">
|
|
<td id="0557E2335428DF378A0F77464C36DC70" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<emphasis id="39EF3F48A716117F0BCD490C507581BD" bold="true" pageId="0" pageNumber="99">Corolla vestiture</emphasis>
|
|
</td>
|
|
<td id="A9BC1D541D91CC7A583863464DD22E83" colspan="1" pageId="0" pageNumber="99" rowspan="1">villous</td>
|
|
<td id="AC9D8011C49A49127514B3F5FD163E3D" colspan="1" pageId="0" pageNumber="99" rowspan="1">villous</td>
|
|
<td id="188053422D45EB4BAD748F077937A542" colspan="1" pageId="0" pageNumber="99" rowspan="1">tomentose</td>
|
|
</tr>
|
|
<tr id="9451EA9595D489B1C67D87E648F175B1" pageId="0" pageNumber="99">
|
|
<td id="937F2883F5C61CF8B8E25206DED1B3ED" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
<emphasis id="B368E01C237DA6D6AB7F51134E87FF3F" bold="true" pageId="0" pageNumber="99">Distribution</emphasis>
|
|
</td>
|
|
<td id="4C39D8E1BC50444947D848754C72CC7E" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
Ecuador (
|
|
<normalizedToken id="5966A23FE89F3A4789962530136C5836" originalValue="Bolívar">Bolivar</normalizedToken>
|
|
, Carchi, Pichincha) and Colombia (
|
|
<normalizedToken id="396C20C2D526249E4321991CD1BCEA55" originalValue="Nariño">Narino</normalizedToken>
|
|
)
|
|
</td>
|
|
<td id="E717D719F244ECC5255109426AA711D5" colspan="1" pageId="0" pageNumber="99" rowspan="1">
|
|
Ecuador (
|
|
<normalizedToken id="0CBB946B4637F93B608E0F990D872499" originalValue="Bolívar">Bolivar</normalizedToken>
|
|
, Carchi, Cotopaxi, Esmeraldas, Imbabura, Pichincha)
|
|
</td>
|
|
<td id="F3A62E4FE4AC5507F47414E2ED064BFC" colspan="1" pageId="0" pageNumber="99" rowspan="1">northern Ecuador (Carchi and Esmeraldas)</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption id="285232BC0EDE999A52AD181174676A25" doi="10.3897/phytokeys.179.65990.figure3" httpUri="https://binary.pensoft.net/fig/564819" pageId="0" pageNumber="99" start="Figure 3" startId="F3">
|
|
<paragraph id="873DAEDFB373C9804798E9EC7C0BB689" pageId="0" pageNumber="99">
|
|
<emphasis id="A903A8D89FEDDA4EC01D47108FA3D4CE" bold="true" pageId="0" pageNumber="99">Figure 3.</emphasis>
|
|
<taxonomicName id="7D394C21DEA0AADCF28507BE6A7CF57E" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="DBB741DE47C1DBBEC4E9140C2850A970" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
(Fritsch) Wiehler
|
|
<emphasis id="4E3FFA5828F8CDF38F410D9F0FF2551B" bold="true" pageId="0" pageNumber="99">A</emphasis>
|
|
lateral view of flower
|
|
<emphasis id="8B002EAD92A85F79DB363A5935B54285" bold="true" pageId="0" pageNumber="99">B</emphasis>
|
|
front view of corolla
|
|
<emphasis id="58F877669CF2AB694AC8F1F9254CB48A" bold="true" pageId="0" pageNumber="99">C</emphasis>
|
|
mature fruit
|
|
<emphasis id="BA7ACFD1891D34DC6BB89A54D4F98675" bold="true" pageId="0" pageNumber="99">D</emphasis>
|
|
dorsiventral habit with anisophyllous leaves (
|
|
<emphasis id="172939D12C55B82C2EF0212AD85BCD80" bold="true" pageId="0" pageNumber="99">A, B</emphasis>
|
|
<emphasis id="174CDE7B809655E4BE164AED4C94D1E0" italics="true" pageId="0" pageNumber="99">Clark et al. 10981</emphasis>
|
|
<emphasis id="2DABFD56B82C76B6B8147784B0C96172" bold="true" pageId="0" pageNumber="99">C</emphasis>
|
|
<emphasis id="C5DC00C6DAEFFE041588F019C24DAF20" italics="true" pageId="0" pageNumber="99">Clark 10948</emphasis>
|
|
<emphasis id="3508B132F694D5A26254683859E77B68" bold="true" pageId="0" pageNumber="99">D</emphasis>
|
|
<emphasis id="7B804BDDF5B8BADBADBE9BD0B5FAB38A" italics="true" pageId="0" pageNumber="99">Clark et al. 14295</emphasis>
|
|
). Photos by J.L. Clark.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="9DAE359A439179E98EC5DAD093A10E4A" pageId="0" pageNumber="99">
|
|
The recent transfer (
|
|
<bibRefCitation id="DB3B1CC57C54725DDEB9E283F6E36E4C" DOI="https://doi.org/10.2307/25065368" author="Roalson, EH" journalOrPublisher="Taxon" pageId="0" pageNumber="99" pagination="389 - 410" refId="B6" refString="Roalson, EH, Boggan, JK, Skog, LE, Zimmer, EA, 2005b. Untangling Gloxinieae (Gesneriaceae). I. Phylogenetic patterns and generic boundaries inferred from nuclear, chloroplast, and morphological cladistic datasets. Taxon 54 (2): 389 - 410, DOI: https://doi.org/10.2307/25065368" title="Untangling Gloxinieae (Gesneriaceae). I. Phylogenetic patterns and generic boundaries inferred from nuclear, chloroplast, and morphological cladistic datasets." url="https://doi.org/10.2307/25065368" volume="54" year="2005 b">Roalson et al. 2005b</bibRefCitation>
|
|
) of
|
|
<taxonomicName id="B63903A6CD86828E1CA23548FED0F60D" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria affinis" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="affinis">
|
|
<emphasis id="5370A1248CC5543458269BDC889CFB7D" italics="true" pageId="0" pageNumber="99">Kohleria affinis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="CEE0C056FC8296D6A7CAFF5851063E5C" lsidName="K. tigridia" pageId="0" pageNumber="99" rank="species" species="tigridia">
|
|
<emphasis id="D64257F82CF61D66DDE94737F4F43CE9" italics="true" pageId="0" pageNumber="99">K. tigridia</emphasis>
|
|
</taxonomicName>
|
|
(Ohlend.) Roalson and Boggan represented an autapomorphic synapomorphy of epiphytism in traditionally recognized
|
|
<taxonomicName id="6A3DD8433419AA6F2281A889339754B2" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="F605EC971FCEA457AF610708D599F4E8" italics="true" pageId="0" pageNumber="99">Kohleria</emphasis>
|
|
</taxonomicName>
|
|
. What is noteworthy about
|
|
<taxonomicName id="6D30B664033138BE66FF56331A29E769" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="3E24204EEE4250C7E9AAB184AE12E9D4" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="3682022A0DE65EE28F59674806F784A3" lsidName="K. hypertrichosa" pageId="0" pageNumber="99" rank="species" species="hypertrichosa">
|
|
<emphasis id="006C6B189810794556AFB010282BBB7F" italics="true" pageId="0" pageNumber="99">K. hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
is their previously unreported epiphytic habits. Thus, the presence of an epiphytic habit in
|
|
<taxonomicName id="DC528407CC33AC9655988B14D9D05BB2" lsidName="K. anisophylla" pageId="0" pageNumber="99" rank="species" species="anisophylla">
|
|
<emphasis id="7E81E960D9260757EFA681B657A31C9F" italics="true" pageId="0" pageNumber="99">K. anisophylla</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="D8E10655D581AB305CEBA423F7BAE231" lsidName="K. hypertrichosa" pageId="0" pageNumber="99" rank="species" species="hypertrichosa">
|
|
<emphasis id="2C62E425D6B1096DAF4ABDF7B41A6644" italics="true" pageId="0" pageNumber="99">K. hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
could represent an additional independent origin of epiphytism in
|
|
<taxonomicName id="87ABD592FB003E8732FB275123B510D1" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="DE5FC6D8D1A1AF41279FF648E5A26E75" italics="true" pageId="0" pageNumber="99">Kohleria</emphasis>
|
|
</taxonomicName>
|
|
. Several populations of
|
|
<taxonomicName id="5B2B7D07C40F588EE319045A9643888E" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="04FDC88996536CD357B569397D6F2024" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
were observed and documented with dorsiventral shoots, a feature that is common in facultative epiphytes in other
|
|
<taxonomicName id="202382603567F7BFBB27029C2E3E3AE4" class="Magnoliopsida" family="Gesneriaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="family">Gesneriaceae</taxonomicName>
|
|
genera. Many members of
|
|
<taxonomicName id="1A5E2011D9D64F503F49BB3D4C069838" class="Magnoliopsida" family="Gesneriaceae" genus="Columnea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Columnea" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="6EFECA0D66FC841837F5F9300F615751" italics="true" pageId="0" pageNumber="99">Columnea</emphasis>
|
|
</taxonomicName>
|
|
have strongly anisophyllous leaves - especially species that are facultative epiphytes with dorsiventral shoots. Other species of
|
|
<taxonomicName id="8F480FA4B4D63FE475D44270EEC4A79E" class="Magnoliopsida" family="Gesneriaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="family">Gesneriaceae</taxonomicName>
|
|
that are facultative epiphytes with dorsiventral shoots include
|
|
<taxonomicName id="8CD2C5A0ABF2E538702A4E377C722321" class="Magnoliopsida" family="Gesneriaceae" genus="Cremosperma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cremosperma anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Magnoliophyta" rank="species" species="anisophylla">
|
|
<emphasis id="D810FC8371C06B358794AF94A43057F9" italics="true" pageId="0" pageNumber="99">Cremosperma anisophylla</emphasis>
|
|
</taxonomicName>
|
|
J.L. Clark & L.E. Skog,
|
|
<taxonomicName id="4132CCB74186BCEFA612F5B655D25631" authorityName="L.E.Skog & L.P.Kvist" authorityYear="1994" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Drymonia anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Magnoliophyta" rank="species" species="anisophylla">
|
|
<emphasis id="8E3875EFFC4DCE5F6F7C4E7EAEA42654" italics="true" pageId="0" pageNumber="99">Drymonia anisophylla</emphasis>
|
|
</taxonomicName>
|
|
L.E. Skog & L.P. Kvist, and the majority of species in
|
|
<taxonomicName id="84E24060152D20278BD39B3B99E633D5" class="Magnoliopsida" family="Gesneriaceae" genus="Monopyle" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Monopyle" order="Lamiales" pageId="0" pageNumber="99" phylum="Magnoliophyta" rank="genus">
|
|
<emphasis id="8EAC1010BDB5B1E1C825C92091367B05" italics="true" pageId="0" pageNumber="99">Monopyle</emphasis>
|
|
</taxonomicName>
|
|
Moritz ex Benth. and
|
|
<taxonomicName id="CCEAA04433CCB52FDB9B9BD3A0D5286E" class="Magnoliopsida" family="Gesneriaceae" genus="Trichodrymonia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Trichodrymonia" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="58DC1B45A342AB56BEF07E7B62666DB1" italics="true" pageId="0" pageNumber="99">Trichodrymonia</emphasis>
|
|
</taxonomicName>
|
|
Oerst. Likewise,
|
|
<taxonomicName id="73E3B34003146BEBBD7D98E144DA48EF" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="65703A4D2106F064D946D4F1098586C5" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="D1ED811EFE5250F2105E59BEC9C9FB72" lsidName="K. hypertrichosa" pageId="0" pageNumber="99" rank="species" species="hypertrichosa">
|
|
<emphasis id="7211B18BABCB45CA9F732D33CBD84B6A" italics="true" pageId="0" pageNumber="99">K. hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
are facultative epiphytes with dorsiventral shoots and anisophyllous leaves. In contrast,
|
|
<taxonomicName id="19FCADD41D97BFA8407FD3E5EF2969D7" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="B733A68688417604C67C85EF32F5D8C1" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
is a terrestrial herb with isophyllous leaves (Fig.
|
|
<figureCitation id="0321ADB63A1CC096CFC7C6F7FB5E22E6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Kohleria villosa (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D erect herbaceous habit with isophyllous leaves (A Clark et al. 14295 B Clark 7331 C Clark et al. 7400 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure4" httpUri="https://binary.pensoft.net/fig/564820" pageId="0" pageNumber="99">4D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="CACFB7C90683D5881130DE61D292E497" doi="10.3897/phytokeys.179.65990.figure4" httpUri="https://binary.pensoft.net/fig/564820" pageId="0" pageNumber="99" start="Figure 4" startId="F4">
|
|
<paragraph id="6884B69FF49A108EE4BAFCD62383DCD6" pageId="0" pageNumber="99">
|
|
<emphasis id="9235281C4A10587524938ACE7FD68DFE" bold="true" pageId="0" pageNumber="99">Figure 4.</emphasis>
|
|
<taxonomicName id="7BB9F7F283CCAD5F0145BC4FDFADC4DF" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="92383F55C309DB4738219A41A19F81D1" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
(Fritsch) Wiehler
|
|
<emphasis id="FC00D406931357E1E3F66929B6CF78A5" bold="true" pageId="0" pageNumber="99">A</emphasis>
|
|
lateral view of flower
|
|
<emphasis id="70E5F55C12FEB411082788D3499552B0" bold="true" pageId="0" pageNumber="99">B</emphasis>
|
|
front view of corolla
|
|
<emphasis id="679257BED440015DFDADA4D96B62D167" bold="true" pageId="0" pageNumber="99">C</emphasis>
|
|
mature fruit
|
|
<emphasis id="ABFABF0D68341EED3DE06BF08AC34180" bold="true" pageId="0" pageNumber="99">D</emphasis>
|
|
erect herbaceous habit with isophyllous leaves (
|
|
<emphasis id="11379E8252046EE68C7CC6C29E133217" bold="true" pageId="0" pageNumber="99">A</emphasis>
|
|
<emphasis id="92B6DE7304FE9EC7FAA911325FFD64F2" italics="true" pageId="0" pageNumber="99">Clark et al. 14295</emphasis>
|
|
<emphasis id="759325130B54D58C5E7CBC3EED0A1A71" bold="true" pageId="0" pageNumber="99">B</emphasis>
|
|
<emphasis id="B5C9FB51081E7DCC7196DA538144D917" italics="true" pageId="0" pageNumber="99">Clark 7331</emphasis>
|
|
<emphasis id="2AEA624101C7AF4DADEBC71E331F31BD" bold="true" pageId="0" pageNumber="99">C</emphasis>
|
|
<emphasis id="B893D0C0243D70D12BC973172F3E4AD4" italics="true" pageId="0" pageNumber="99">Clark et al. 7400</emphasis>
|
|
<emphasis id="8ECD1C33F490A80E38A6DA6126C92E3C" bold="true" pageId="0" pageNumber="99">D</emphasis>
|
|
<emphasis id="F70C8291C1E593031A57D5BB5857EEA3" italics="true" pageId="0" pageNumber="99">Clark et al. 14295</emphasis>
|
|
). Photos by J.L. Clark.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="6A81E726F9CB133C76AEC395D9B7BA6F" pageId="0" pageNumber="99">
|
|
The corollas of
|
|
<taxonomicName id="C72F12A3573A36F4B8B0B5EE8A0C7531" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="B6EE56A8E86FE8A423AB7E8F8F0DDBA0" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="B6FF6E31380DBFF65DFD55A4A4779BDE" lsidName="K. anisophylla" pageId="0" pageNumber="99" rank="species" species="anisophylla">
|
|
<emphasis id="6093A355352F36D3240A8694E9396359" italics="true" pageId="0" pageNumber="99">K. anisophylla</emphasis>
|
|
</taxonomicName>
|
|
are villous (Figs
|
|
<figureCitation id="98FE0BA1119EBA0ACDAD3E978971E455" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Kohleria anisophylla (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A, B Clark et al. 10981 C Clark 10948 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure3" httpUri="https://binary.pensoft.net/fig/564819" pageId="0" pageNumber="99">3</figureCitation>
|
|
,
|
|
<figureCitation id="C56E111F231B81AD1C7C5526E5E4ADD4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Kohleria villosa (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D erect herbaceous habit with isophyllous leaves (A Clark et al. 14295 B Clark 7331 C Clark et al. 7400 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure4" httpUri="https://binary.pensoft.net/fig/564820" pageId="0" pageNumber="99">4</figureCitation>
|
|
). The corollas of
|
|
<taxonomicName id="B5ADB69E5B62E0ADB90D3470D7348D52" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria hypertrichosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="hypertrichosa">
|
|
<emphasis id="EB4C06706E16D5ECFEEC2AD04113E742" italics="true" pageId="0" pageNumber="99">Kohleria hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
are densely tomentose (Fig.
|
|
<figureCitation id="7D5DCC843F796609C1ABD48A38DBECC0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Kohleria hypertrichosa J. L. Clark & L. E. Skog A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A Clark et al. 15900 B Clark 6539 C Clark et al. 10310 D Clark et al. 14942). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure5" httpUri="https://binary.pensoft.net/fig/564821" pageId="0" pageNumber="99">5</figureCitation>
|
|
). The specific epiphyte, "
|
|
<taxonomicName id="7C63AB6664834B9642B8FAEAD19C47A1" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria hypertrichosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="hypertrichosa">
|
|
<emphasis id="FCB18F5C635CE543C6F6B69B61D95DE2" italics="true" pageId="0" pageNumber="99">Kohleria hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
" refers to the abundance of trichomes, which is why it is commonly known in the horticultural community as
|
|
<normalizedToken id="27DE02C509588877ADABF90031F4E775" originalValue="“Chewbacca,”">"Chewbacca,"</normalizedToken>
|
|
a reference to the Wookie (fictional character) in the movie Star Wars.
|
|
</paragraph>
|
|
<caption id="D49BCB3BAD95DF8AC9E2C848C612B2A1" doi="10.3897/phytokeys.179.65990.figure5" httpUri="https://binary.pensoft.net/fig/564821" pageId="0" pageNumber="99" start="Figure 5" startId="F5">
|
|
<paragraph id="769611101FD3AB43CF02A6FEBEB0B6BB" pageId="0" pageNumber="99">
|
|
<emphasis id="4A1E52C754183FC72A8AF091EE0F6BF7" bold="true" pageId="0" pageNumber="99">Figure 5.</emphasis>
|
|
<taxonomicName id="F071F30300527B2903DFF0B7BF915392" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria hypertrichosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="hypertrichosa">
|
|
<emphasis id="3CCAD2F70EEF6492C8EE4A39B942043E" italics="true" pageId="0" pageNumber="99">Kohleria hypertrichosa</emphasis>
|
|
</taxonomicName>
|
|
J.L. Clark & L.E. Skog
|
|
<emphasis id="14C8454A3B51F3E2FCD5D53188849627" bold="true" pageId="0" pageNumber="99">A</emphasis>
|
|
lateral view of flower
|
|
<emphasis id="1C3046906597A5B9F0CC0E31A6F447D5" bold="true" pageId="0" pageNumber="99">B</emphasis>
|
|
front view of corolla
|
|
<emphasis id="B06D10661F7C90E880E1C36D841F3416" bold="true" pageId="0" pageNumber="99">C</emphasis>
|
|
mature fruit
|
|
<emphasis id="6EAFEB045BC35D0F6F83B1E3B2B8910E" bold="true" pageId="0" pageNumber="99">D</emphasis>
|
|
dorsiventral habit with anisophyllous leaves (
|
|
<emphasis id="4A5EC086D993CE6AE8A6EC6847BE88E5" bold="true" pageId="0" pageNumber="99">A</emphasis>
|
|
<emphasis id="58B1D2CB64EFC0B370C835737F710CFF" italics="true" pageId="0" pageNumber="99">Clark et al. 15900</emphasis>
|
|
<emphasis id="50FF01E433329CE306BC439A83E39565" bold="true" pageId="0" pageNumber="99">B</emphasis>
|
|
<emphasis id="DF11CB199644CCCBD8C96D3957EA44F6" italics="true" pageId="0" pageNumber="99">Clark 6539</emphasis>
|
|
<emphasis id="7D107B92336254B5C9B67F27BF6BA43A" bold="true" pageId="0" pageNumber="99">C</emphasis>
|
|
<emphasis id="EACF8D28E4A0C88E26C671645C43C5F5" italics="true" pageId="0" pageNumber="99">Clark et al. 10310</emphasis>
|
|
<emphasis id="80A4C6F56E8147595BCF44261D6031CF" bold="true" pageId="0" pageNumber="99">D</emphasis>
|
|
<emphasis id="8BFB86953D5204D326DD8EBC8BAC62B0" italics="true" pageId="0" pageNumber="99">Clark et al. 14942</emphasis>
|
|
). Photos by J.L. Clark.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="9C588A4070182E99C91A311B76EEFFA2" pageId="0" pageNumber="99">
|
|
<taxonomicName id="33CF5AD97F9026E4D53ABBA31A7ACFE3" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="2779AF2C9145A737F6E1281802CD3762" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="918553B8F1BFFD02452A93F289C8ABE9" lsidName="K. anisophylla" pageId="0" pageNumber="99" rank="species" species="anisophylla">
|
|
<emphasis id="0EC1E9122C349FC500888E197D16F50A" italics="true" pageId="0" pageNumber="99">K. anisophylla</emphasis>
|
|
</taxonomicName>
|
|
are easily recognized when sterile. The opposite leaves of
|
|
<taxonomicName id="2FAA3D901F4A240E6A4A89CFE84E8B83" authorityName="Wiehler" authorityYear="1978" baseAuthorityName="Fritsch" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria anisophylla" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="anisophylla">
|
|
<emphasis id="D7ED3176B2DC1F3499B686B11551442E" italics="true" pageId="0" pageNumber="99">Kohleria anisophylla</emphasis>
|
|
</taxonomicName>
|
|
are consistently unequal in size or anisophyllous (Fig.
|
|
<figureCitation id="EE8DB1FFDF2A7223CB2371373A5B32DC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Kohleria anisophylla (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D dorsiventral habit with anisophyllous leaves (A, B Clark et al. 10981 C Clark 10948 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure3" httpUri="https://binary.pensoft.net/fig/564819" pageId="0" pageNumber="99">3D</figureCitation>
|
|
). In contrast, the opposite leaves of
|
|
<taxonomicName id="2C254EE34AD014749F439CB581366399" class="Magnoliopsida" family="Gesneriaceae" genus="Kohleria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kohleria villosa" order="Lamiales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="villosa">
|
|
<emphasis id="E057B21B9BEC82370035995B5449A640" italics="true" pageId="0" pageNumber="99">Kohleria villosa</emphasis>
|
|
</taxonomicName>
|
|
are consistently equal in size or isophyllous (Fig.
|
|
<figureCitation id="902BADBC987D8181F0E877A848E8E33C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Kohleria villosa (Fritsch) Wiehler A lateral view of flower B front view of corolla C mature fruit D erect herbaceous habit with isophyllous leaves (A Clark et al. 14295 B Clark 7331 C Clark et al. 7400 D Clark et al. 14295). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.179.65990.figure4" httpUri="https://binary.pensoft.net/fig/564820" pageId="0" pageNumber="99">4D</figureCitation>
|
|
). In addition, the dorsiventral shoots distinguishes
|
|
<taxonomicName id="0793B246543BFB3592C803257E4C73AE" lsidName="K. anisophylla" pageId="0" pageNumber="99" rank="species" species="anisophylla">
|
|
<emphasis id="FD08383051C5287B5495CEDBA902940F" italics="true" pageId="0" pageNumber="99">K. anisophylla</emphasis>
|
|
</taxonomicName>
|
|
from the erect shoots of
|
|
<taxonomicName id="765606875F9DADF6143FC5C68A8206E8" lsidName="K. villosus" pageId="0" pageNumber="99" rank="species" species="villosus">
|
|
<emphasis id="9CE70D5FA8EDAC5DF471633578BF90C9" italics="true" pageId="0" pageNumber="99">K. villosus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |